1611 lines
151 KiB
XML
1611 lines
151 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1077.67081" ID-GBIF-Dataset="66b827c0-65ac-44a6-90e0-b78cd9ef81c2" ID-PMC="PMC8692307" ID-Pensoft-Pub="1313-2970-1077-1" ID-Pensoft-UUID="734088641608531C8E2CC69397B000ED" ID-PubMed="35027858" ID-ZooBank="C28AD65A76F242CFBED7DFB3702CABCE" ModsDocID="1313-2970-1077-1" checkinTime="1639570129388" checkinUser="pensoft" docAuthor="De Mattia, Willy, Reier, Susanne & Haring, Elisabeth" docDate="2021" docId="464D0C8C9C0B599590E50AE106263C30" docLanguage="en" docName="ZooKeys 1077: 1-175" docOrigin="ZooKeys 1077" docPubDate="2021-12-14" docSource="http://dx.doi.org/10.3897/zookeys.1077.67081" docTitle="Mauritanica perinni subsp. polygyra" docType="treatment" docVersion="5" id="734088641608531C8E2CC69397B000ED" lastPageNumber="1" masterDocId="734088641608531C8E2CC69397B000ED" masterDocTitle="Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)" masterLastPageNumber="175" masterPageNumber="1" pageNumber="1" updateTime="1668151235153" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae)</mods:title>
|
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>De Mattia, Willy</mods:namePart>
|
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0056-467X</mods:nameIdentifier>
|
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">willy.demattia@icgeb.org</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reier, Susanne</mods:namePart>
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<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Haring, Elisabeth</mods:namePart>
|
||
<mods:affiliation>Central Research Laboratories, of Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-12-14</mods:number>
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</mods:detail>
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||
<mods:detail type="volume">
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||
<mods:number>1077</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>175</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1077.67081</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1077.67081</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1077-1</mods:identifier>
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<mods:identifier type="ZooBank">C28AD65A76F242CFBED7DFB3702CABCE</mods:identifier>
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<mods:identifier type="Pensoft-UUID">734088641608531C8E2CC69397B000ED</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="191645927" LSID="urn:lsid:plazi:treatment:464D0C8C9C0B599590E50AE106263C30" httpUri="http://treatment.plazi.org/id/464D0C8C9C0B599590E50AE106263C30" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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||
<taxonomicName LSID="464D0C8C-9C0B-5995-90E5-0AE106263C30" authority="(O. Boettger, 1879)" baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">Mauritanica perinni polygyra (O. Boettger, 1879)</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Mauritanica scarificata (L. Pfeiffer, 1856), comb. nov., Marettimo 37.1 whole distal genital organs 37.2 internal distal part of genital organs 37.3 whole distal genital organs 37.4 internal distal part of genital organs 37.5 detail of the pseudopapilla. Mauritanica perinni polygyra (O. Boettger, 1879), Tunisia, Zaghouan 37.6 whole distal genital organs 37.7 internal distal part of genital organs 37.8 - 37.9 spermatophore 37.10 cross section of the spermatophore." figureDoi="10.3897/zookeys.1077.67081.figure37" httpUri="https://binary.pensoft.net/fig/622829" pageId="0" pageNumber="1">Figs 37.6-37.10, 38.5, 38.6</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Clausilia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Clausilia (Mauritanica) polygyra" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="polygyra" subGenus="Mauritanica">Clausilia (Mauritanica) polygyra</taxonomicName>
|
||
Boettger in Kobelt 1879: 153.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName family="Clausiliidae" genus="Charpenteria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Charpenteria (Mauritanica) perinni subsp. polygyra" order="Stylommatophora" pageId="0" pageNumber="1" rank="subSpecies" species="perinni" subGenus="Mauritanica" subSpecies="polygyra">Charpenteria (Mauritanica) perinni polygyra</taxonomicName>
|
||
-
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" pagination="27 - 39" refId="B64" refString="Nordsieck, H, 2002. Contributions to the knowldge of the Delimini (Gastropoda: Stylommatophora: Clausiliidae). Mitteilungen der deutschen malakozoologischen Gesellschaft 67: 27 - 39" title="Contributions to the knowldge of the Delimini (Gastropoda: Stylommatophora: Clausiliidae)." volume="67" year="2002">Nordsieck 2002</bibRefCitation>
|
||
: 27, 29, 35, 36.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName family="Clausiliidae" genus="Charpenteria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Charpenteria (Mauritanica) perinni subsp. polygyra" order="Stylommatophora" pageId="0" pageNumber="1" rank="subSpecies" species="perinni" subGenus="Mauritanica" subSpecies="polygyra">Charpenteria (Mauritanica) perinni polygyra</taxonomicName>
|
||
-
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B65" refString="Nordsieck, H, 2007. Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." title="Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." year="2007">Nordsieck 2007</bibRefCitation>
|
||
: 54.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">Mauritanica perinni polygyra</taxonomicName>
|
||
-
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. 2020</bibRefCitation>
|
||
: 318.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
was recently revised by
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. (2020</bibRefCitation>
|
||
: 318). Based on differences of the shell morphology alone,
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. (2020</bibRefCitation>
|
||
: 319) proposed species status for
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
, independent from
|
||
<taxonomicName authority="perinni" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. perinni" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="perinni">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni perinni</emphasis>
|
||
</taxonomicName>
|
||
(Bourguignat, 1876). A genital comparison with the nominate subspecies was not possible since the genital organs of this taxon were not provided by
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. (2020)</bibRefCitation>
|
||
. Moreover, a comparison between our anatomical results and those provided by
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. (2020</bibRefCitation>
|
||
: 337, figs 60-61) for
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
was rather difficult: Essential details, as the internal structure of the epiphallus and the presence/absence of the ER and ELP are not detectable. The fine structure of the internal penis is not clear and, contrary to our results that show a smooth internal wall, the vagina seems to have a smooth longitudinal pleat (2020: 337, fig. 61). The presence of the ET is not clear as well (2020: 337, fig. 60).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure37" httpUri="https://binary.pensoft.net/fig/622829" pageId="0" pageNumber="1" start="Figure 37" startId="F37">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 37.</emphasis>
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica scarificata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica scarificata</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer, 1856), comb. nov., Marettimo
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.1</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.2</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.3</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.4</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.5</emphasis>
|
||
detail of the pseudopapilla.
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
(O. Boettger, 1879), Tunisia, Zaghouan
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.6</emphasis>
|
||
whole distal genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.7</emphasis>
|
||
internal distal part of genital organs
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.8-37.9</emphasis>
|
||
spermatophore
|
||
<emphasis bold="true" pageId="0" pageNumber="1">37.10</emphasis>
|
||
cross section of the spermatophore.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<materialsCitation accessionNumber="MW758966, MW757044, MW757045" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Lab ID, Nordsieck" country="Tunisia" elevation="789" latitude="36.367344" location="Zaghouan" longLatPrecision="1" longitude="10.119947" specimenCount="1" stateProvince="Zaghouan">
|
||
<collectingCountry name="Tunisia">Tunisia</collectingCountry>
|
||
, above
|
||
<collectingRegion country="Tunisia" name="Zaghouan">Zaghouan</collectingRegion>
|
||
(road to summit),
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.89" unit="m" value="789.0">
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.89" unit="m" value="789.0">789 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<geoCoordinate degrees="36" direction="north" minutes="22" orientation="latitude" precision="1" seconds="2.45" value="36.367344">36°22'2.45"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="07" orientation="longitude" precision="1" seconds="11.81" value="10.119947">10°07'11.81"E</geoCoordinate>
|
||
, [
|
||
<collectorName>Lab ID</collectorName>
|
||
SMF335011_1, COI:
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/MW758966">MW758966</accessionNumber>
|
||
, ITS2:
|
||
<accessionNumber isEnumeration="true">MW757044, MW757045</accessionNumber>
|
||
], ex
|
||
<collectorName>Nordsieck</collectorName>
|
||
coll. (11639), currently SMF 335011/1, 1 dissected spm
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="shell">
|
||
<paragraph pageId="0" pageNumber="1">Shell</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
Figs
|
||
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Mauritanica scarificata (L. Pfeiffer, 1856), comb. nov., Marettimo 38.1 shell 38.2 shell 38.3 detail of the aperture 38.4 clausiliar plate double side. Mauritanica perinni polygyra (O. Boettger, 1879), Tunisia, Zaghouan 38.5 shell 38.6 shell (picture courtesy of Claude and Amandine Evanno)." figureDoi="10.3897/zookeys.1077.67081.figure38" httpUri="https://binary.pensoft.net/fig/622830" pageId="0" pageNumber="1">38</figureCitation>
|
||
.5, 38.6).
|
||
</emphasis>
|
||
Shell decollate or not decollate; whorls finely and irregularly striated, flat, with dense papillae along the sutures; dorsal keel indistinct; inferior lamella high; anterior upper palatal plicae absent; posterior upper palatal plica present; lunella dorso-lateral; little subclaustralis, clausilium plough-like with subrectangular plate (
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. 2020</bibRefCitation>
|
||
: 318).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Measurements</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">(n = 1, decollate).</emphasis>
|
||
shell height 22.4, whorl width 4.1, aperture height 3.3, aperture width 2.4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="external morphology of the genital organs">
|
||
<paragraph pageId="0" pageNumber="1">External morphology of the genital organs</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Mauritanica scarificata (L. Pfeiffer, 1856), comb. nov., Marettimo 37.1 whole distal genital organs 37.2 internal distal part of genital organs 37.3 whole distal genital organs 37.4 internal distal part of genital organs 37.5 detail of the pseudopapilla. Mauritanica perinni polygyra (O. Boettger, 1879), Tunisia, Zaghouan 37.6 whole distal genital organs 37.7 internal distal part of genital organs 37.8 - 37.9 spermatophore 37.10 cross section of the spermatophore." figureDoi="10.3897/zookeys.1077.67081.figure37" httpUri="https://binary.pensoft.net/fig/622829" pageId="0" pageNumber="1">37</figureCitation>
|
||
.6).
|
||
</emphasis>
|
||
The FO is longer than the V (FO/V = 2.1). The VD is thin along its whole course. The FDBC is slightly shorter than the BC+SDBC (FDBC/BC+SDBC = 0.9). The BC+SDBC is club-like and longer than the V (BC+SDBC/V = 1.4), with clear distinction between the SDBC and the BC. The SDBC is very short. The apex is big and round. The D is much longer than the V (D/V = 1.9) and longer that the BC+SDBC (D/BC+SDBC = 2.2), thinner than the BC+SDBC and with a small and round apex. The V is cylindrical but the presence of the spermatophore may modify its normal shape. The A is small. The PC is longer than the V (P+E/V = 2.8). The PR is long and robust. The ET is not clearly visible. The E is shorter than the P (E/P = 0.9), gradually shrinking and turning into the VD.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="internal morphology of the genital organs">
|
||
<paragraph pageId="0" pageNumber="1">Internal morphology of the genital organs</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Mauritanica scarificata (L. Pfeiffer, 1856), comb. nov., Marettimo 37.1 whole distal genital organs 37.2 internal distal part of genital organs 37.3 whole distal genital organs 37.4 internal distal part of genital organs 37.5 detail of the pseudopapilla. Mauritanica perinni polygyra (O. Boettger, 1879), Tunisia, Zaghouan 37.6 whole distal genital organs 37.7 internal distal part of genital organs 37.8 - 37.9 spermatophore 37.10 cross section of the spermatophore." figureDoi="10.3897/zookeys.1077.67081.figure37" httpUri="https://binary.pensoft.net/fig/622829" pageId="0" pageNumber="1">37</figureCitation>
|
||
.7).
|
||
</emphasis>
|
||
The A, V and the P show a very finely granulated surface. The PP is small, elongated. Its surface is very irregular, folded and its internal texture is spongy. The P-E transition is simple, with no ER. The origin of the PP is connected with one of the two ELP. The epiphallar formula is: PP(ELP). The E shows two smooth longitudinal pleats that fade before the VD. Between the ELP an irregular transverse small, interrupted pattern gives an overall coarse appearance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="spermatophore">
|
||
<paragraph pageId="0" pageNumber="1">Spermatophore</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(Figs
|
||
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Mauritanica scarificata (L. Pfeiffer, 1856), comb. nov., Marettimo 37.1 whole distal genital organs 37.2 internal distal part of genital organs 37.3 whole distal genital organs 37.4 internal distal part of genital organs 37.5 detail of the pseudopapilla. Mauritanica perinni polygyra (O. Boettger, 1879), Tunisia, Zaghouan 37.6 whole distal genital organs 37.7 internal distal part of genital organs 37.8 - 37.9 spermatophore 37.10 cross section of the spermatophore." figureDoi="10.3897/zookeys.1077.67081.figure37" httpUri="https://binary.pensoft.net/fig/622829" pageId="0" pageNumber="1">37</figureCitation>
|
||
.8-37.10).
|
||
</emphasis>
|
||
The spermatophore is thin, elongated. It is 4.7 mm long and 0.8 mm wide at its widest point. The tail is pointed and sharp. The head is missing, probably already digested. The upper keel is simple and goes from the tip of the tail as far as the broken head part. The lower keel is missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="ecology">
|
||
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
is a limestone dweller, on cliffs, crevices and under stones (
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. 2020</bibRefCitation>
|
||
: 319).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
is known only from Djebel Zaghouan area (
|
||
<bibRefCitation DOI="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" author="Sparacio, I" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="1" pagination="311 - 346" refId="B91" refString="Sparacio, I, Liberto, F, La Mantia, T, 2020. The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia. Biodiversity Journal 11 (1): 311 - 346, DOI: https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" title="The genus Mauritanica O. Boettger, 1879 (Gastropoda Stylommatophora Clausiliidae) in Tunisia." url="https://doi.org/10.31396/Biodiv.Jour.2020.11.1.311.346" volume="11" year="2020">Sparacio et al. 2020</bibRefCitation>
|
||
: 319).
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure38" httpUri="https://binary.pensoft.net/fig/622830" pageId="0" pageNumber="1" start="Figure 38" startId="F38">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 38.</emphasis>
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica scarificata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica scarificata</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer, 1856), comb. nov., Marettimo
|
||
<emphasis bold="true" pageId="0" pageNumber="1">38.1</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">38.2</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">38.3</emphasis>
|
||
detail of the aperture
|
||
<emphasis bold="true" pageId="0" pageNumber="1">38.4</emphasis>
|
||
clausiliar plate double side.
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica perinni subsp. polygyra" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="perinni" subSpecies="polygyra">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica perinni polygyra</emphasis>
|
||
</taxonomicName>
|
||
(O. Boettger, 1879), Tunisia, Zaghouan
|
||
<emphasis bold="true" pageId="0" pageNumber="1">38.5</emphasis>
|
||
shell
|
||
<emphasis bold="true" pageId="0" pageNumber="1">38.6</emphasis>
|
||
shell (picture courtesy of Claude and Amandine Evanno).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="an overview of siciliaria related genera charpentieria, gibbularia, and stigmatica">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
An overview of
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
related genera
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Gibbularia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbularia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gibbularia</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The separation of
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania</emphasis>
|
||
</taxonomicName>
|
||
(and provisionally
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
) from the other genera is well supported, but the relationships among
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Gibbularia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbularia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gibbularia</emphasis>
|
||
</taxonomicName>
|
||
remain unresolved and appear as polytomy in the trees.
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
(four species included) forms one (poorly supported) clade but, the relationships within this clade received no considerable support. The genera
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Gibbularia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbularia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gibbularia</emphasis>
|
||
</taxonomicName>
|
||
are not monophyletic and populations from distant areas show high genetic distances (Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Mean genetic distances (p-distances in %) for the COI dataset between the genera and / or lineages analysed. Mean genetic distances within the genera Siciliaria and Sicania are in bold. Gibbularia 1 = Gibbularia gibbula gibbula from Apricena; Gibbularia 2 = Gibbularia gibbula gibbula from Muggia." httpUri="http://table.plazi.org/id/F1793A68627A13C7D9E4BFFB44130BC6" pageId="0" pageNumber="1" tableUuid="F1793A68627A13C7D9E4BFFB44130BC6">2</tableCitation>
|
||
).
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
is based on
|
||
<taxonomicName authorityName="Rossmassler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stigmatica">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia stigmatica</emphasis>
|
||
</taxonomicName>
|
||
by tautonomy and, in our trees (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. BI tree of based on the COI data set of Siciliaria, Sicania and related genera. BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure4" httpUri="https://binary.pensoft.net/fig/622796" pageId="0" pageNumber="1">4</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. BI tree of based on the concatenated COI and ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Fig. 1 E, F). BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure6" httpUri="https://binary.pensoft.net/fig/622798" pageId="0" pageNumber="1">6</figureCitation>
|
||
), only this clade with the type species
|
||
<taxonomicName authority="stigmatica" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stigmatica">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica stigmatica</emphasis>
|
||
</taxonomicName>
|
||
can be deemed as a representative of this genus. Despite emerging from a polytomy, additional taxa included in our trees and currently considered as belonging to
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
(namely
|
||
<taxonomicName baseAuthorityName="Benoit" baseAuthorityYear="1860" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica vulcanica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="vulcanica">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica vulcanica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName baseAuthorityName="O. Boettger" baseAuthorityYear="1889" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica pantocratoris" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="pantocratoris">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica pantocratoris</emphasis>
|
||
</taxonomicName>
|
||
) are clearly separated from the type species. We thus only conventionally keep them as
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
species. A focused approach with more genetic markers and specimens is needed to identify the phylogenetic relationships among the species currently included into
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Stigmatica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stigmatica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stigmatica</emphasis>
|
||
</taxonomicName>
|
||
(following MolluscaBase 2021). A similar approach is used for
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Gibbularia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbularia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Gibbularia</emphasis>
|
||
</taxonomicName>
|
||
, which needs to be excluded from
|
||
<emphasis italics="true" pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Siciliaria</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Sicania</taxonomicName>
|
||
</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. BI tree of based on the COI data set of Siciliaria, Sicania and related genera. BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure4" httpUri="https://binary.pensoft.net/fig/622796" pageId="0" pageNumber="1">4</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. BI tree of based on the concatenated COI and ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Fig. 1 E, F). BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure6" httpUri="https://binary.pensoft.net/fig/622798" pageId="0" pageNumber="1">6</figureCitation>
|
||
). Its phylogenetic relationship with the other genera is far from being resolved.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1077.67081.figure39" httpUri="https://binary.pensoft.net/fig/622831" pageId="0" pageNumber="1" start="Figure 39" startId="F39">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 39.</emphasis>
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania crassicostata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="crassicostata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania crassicostata</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer, 1856), comb. nov., Monte Cofano
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.1</emphasis>
|
||
palatal plicae
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.2</emphasis>
|
||
parietal lamellae.
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="A. Schmidt" baseAuthorityYear="1868" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania eminens" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="eminens">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania eminens</emphasis>
|
||
</taxonomicName>
|
||
(A. Schmidt, 1868), comb. nov., Custonaci, Baglio Messina
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.3-39.4</emphasis>
|
||
palatal plicae.
|
||
<taxonomicName authority="nobilis" authorityName="nobilis (L. Pfeiffer" authorityYear="1848" baseAuthorityName="nobilis (L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania nobilis subsp. nobilis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="nobilis" subSpecies="nobilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania nobilis nobilis</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer, 1848), comb. nov., Castelluzzo
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.5-39.6</emphasis>
|
||
palatal plicae
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.7</emphasis>
|
||
parietal lamellae.
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1856" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica scarificata" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="scarificata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica scarificata</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer, 1856), comb. nov., Marettimo
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.8</emphasis>
|
||
palatal plicae
|
||
<emphasis bold="true" pageId="0" pageNumber="1">39.9</emphasis>
|
||
parietal lamellae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName authorityName="Hartmann" authorityYear="1842" class="Gastropoda" family="Clausiliidae" genus="Papillifera" kingdom="Animalia" lsidName="Papillifera" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Papillifera</emphasis>
|
||
</taxonomicName>
|
||
, which was used as outgroup, appeared monophyletic in our trees, even though the distances within this genus are in the same range as among different genera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="southern alpine charpentieria">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Southern alpine
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The taxa included in the alpine
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria ornata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ornata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria ornata</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
clade show a pan-Southern Alpine distribution, with
|
||
<taxonomicName lsidName="C. dyodon" pageId="0" pageNumber="1" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon</emphasis>
|
||
</taxonomicName>
|
||
isolated in Piemonte and Ticino,
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
along central and eastern Southern Alps whereas
|
||
<taxonomicName lsidName="C. ornata" pageId="0" pageNumber="1" rank="species" species="ornata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. ornata</emphasis>
|
||
</taxonomicName>
|
||
is present along the easternmost Southern Alps and northwestern Balkans. The taxa included in the
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
subclade occupy the eastern Southern Alps, from eastern Trentino-Alto Adige, Friuli-Venezia Giulia, W Slovenia and SE Austria.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Despite the low support values, the COI and the ITS2 phylogenetic trees (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. BI tree of based on the COI data set of Siciliaria, Sicania and related genera. BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure4" httpUri="https://binary.pensoft.net/fig/622796" pageId="0" pageNumber="1">4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. BI tree of based on the ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Figs 1 E, F). BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure5" httpUri="https://binary.pensoft.net/fig/622797" pageId="0" pageNumber="1">5</figureCitation>
|
||
) show a general picture where
|
||
<taxonomicName authority="dyodon" authorityName="dyodon (S. Studer" authorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon dyodon</emphasis>
|
||
</taxonomicName>
|
||
(S. Studer, 1820) appears as the sister species of
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
(with
|
||
<taxonomicName authority="itala" lsidName="C. itala subsp. itala" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala itala</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="C. itala subsp. baldensis" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="baldensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala baldensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. itala subsp. lorinae" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="lorinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala lorinae</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName baseAuthorityName="Rossmassler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii</emphasis>
|
||
</taxonomicName>
|
||
(with
|
||
<taxonomicName lsidName="C. stenzii subsp. cincta" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="cincta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii cincta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. stenzii subsp. letochana" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="letochana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii letochana</emphasis>
|
||
</taxonomicName>
|
||
) represents a separate clade.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The results presented by Scheel and Hausdorf (2012: 3799, 3801) and
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2020.106982" author="Xu, J" journalOrPublisher="Petersbourg, Classe physico-mathematique 11: [1, 2], [1 - 37]" pageId="0" pageNumber="1" refId="B106" refString="Xu, J, Hausdorf, B, 2021. Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982. https://doi.org/10.1016/j.ympev.2020.106982" title="Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982." url="https://doi.org/10.1016/j.ympev.2020.106982" year="2021">Xu and Hausdorf (2021</bibRefCitation>
|
||
: 8) showed that
|
||
<taxonomicName lsidName="C. dyodon" pageId="0" pageNumber="1" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon</emphasis>
|
||
</taxonomicName>
|
||
is the sister species of the
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
complex whereas
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
is sister to
|
||
<taxonomicName lsidName="C. ornata" pageId="0" pageNumber="1" rank="species" species="ornata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. ornata</emphasis>
|
||
</taxonomicName>
|
||
. These results are difficult to compare with our trees, but at least not in contrast.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The stenzioid subspecies of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
included in our genital anatomical investigations (
|
||
<taxonomicName baseAuthorityName="Rossmassler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria clavata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="clavata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria clavata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria variscoi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="variscoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria variscoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria balsamoi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="balsamoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria balsamoi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria lorinae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria lorinae</emphasis>
|
||
</taxonomicName>
|
||
) were first assigned to
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Käufel">Kaeufel</normalizedToken>
|
||
, 1928) but later transferred to
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="81 - 115" refId="B57" refString="Nordsieck, H, 1963a. Zur Anatomie und Systematik der Clausilien, I. Archiv fuer Molluskenkunde 92: 81 - 115" title="Zur Anatomie und Systematik der Clausilien, I." volume="92" year="1963 a">Nordsieck (1963a</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">b</bibRefCitation>
|
||
). In 2007,
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Mitteilungen der deutschen malakozoologischen Gesellschaft" pageId="0" pageNumber="1" refId="B65" refString="Nordsieck, H, 2007. Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." title="Worldwide Door Snails (Clausiliidae), recent and fossil. Conchbooks Hackenheim, 214 pp." year="2007">Nordsieck (2007)</bibRefCitation>
|
||
transferred these taxa to
|
||
<taxonomicName lsidName="C. clavata" pageId="0" pageNumber="1" rank="species" species="clavata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. clavata</emphasis>
|
||
</taxonomicName>
|
||
. Scheel and Hausdorf (2012), following their molecular genetic results considered the stenzioid
|
||
<taxonomicName lsidName="C. clavata" pageId="0" pageNumber="1" rank="species" species="clavata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. clavata</emphasis>
|
||
</taxonomicName>
|
||
as subspecies of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
, and thus, its subspecies obtained a new combination with
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
. Scheel and Hausdorf (2012: fig. 3) and
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2020.106982" author="Xu, J" journalOrPublisher="Petersbourg, Classe physico-mathematique 11: [1, 2], [1 - 37]" pageId="0" pageNumber="1" refId="B106" refString="Xu, J, Hausdorf, B, 2021. Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982. https://doi.org/10.1016/j.ympev.2020.106982" title="Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982." url="https://doi.org/10.1016/j.ympev.2020.106982" year="2021">Xu and Hausdorf (2021</bibRefCitation>
|
||
: 6, fig. 5, supplementary table S1) revealed gene flow between
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
and the stenzioid subspecies of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
across the contact zone and also highlighted several mixture events within the
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The list of the dissected material (at least two specimens per population) is found in the Table
|
||
<tableCitation captionStart="Table 8" captionStartId="T8" captionText="Table 8. Examined taxa of Charpentieria with information on shell type (stenzioid / nonstenzioid), availability of genetic data (DNA) and epiphallar formula." httpUri="http://table.plazi.org/id/66A5CD940D950B91226F96579AB448F1" pageId="0" pageNumber="1" tableUuid="66A5CD940D950B91226F96579AB448F1">8</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="66A5CD940D950B91226F96579AB448F1" httpUri="http://table.plazi.org/id/66A5CD940D950B91226F96579AB448F1" pageId="0" pageNumber="1" start="Table 8" startId="T8">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Table 8.</emphasis>
|
||
Examined taxa of
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
with information on shell type (stenzioid/nonstenzioid), availability of genetic data (DNA) and epiphallar formula.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<table pageId="0" pageNumber="1">
|
||
<tr pageId="0" pageNumber="1">
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Taxon</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Specimens examined</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Shell stenzioid/ nonstenzioid</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">DNA</th>
|
||
<th colspan="1" pageId="0" pageNumber="1" rowspan="1">Epiphallar formula</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authority="dyodon" authorityName="dyodon (S. Studer" authorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon dyodon</emphasis>
|
||
</taxonomicName>
|
||
(S. Studer, 1820)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Trasquera di Iselle, Verbano, Piedmont, Italy. 750 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="12" orientation="latitude" precision="1" seconds="29.53" value="46.208202">46°12'29.53"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="08" direction="east" minutes="12" orientation="longitude" precision="1" seconds="46.18" value="8.212828">08°12'46.18"E</geoCoordinate>
|
||
, I. Niero leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Y</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Stabile" baseAuthorityYear="1859" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. alpina" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="alpina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon alpina</emphasis>
|
||
</taxonomicName>
|
||
(Stabile, 1859)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Margone, Val di Lanzo, Usseglio, Torino, Piedmont, Italy. 1500 m asl,
|
||
<geoCoordinate degrees="45" direction="north" minutes="14" orientation="latitude" precision="1" seconds="04.89" value="45.23469">45°14'04.89"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="07" direction="east" minutes="12" orientation="longitude" precision="1" seconds="18.90" value="7.20525">07°12'18.90"E</geoCoordinate>
|
||
, leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Sentiero per
|
||
<normalizedToken originalValue="Prà">Pra</normalizedToken>
|
||
del Vecia, Piedicavallo, Biella, Piedmont, Italy. 1200 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="41" orientation="latitude" precision="1" seconds="32.37" value="45.692326">45°41'32.37"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="07" direction="east" minutes="58" orientation="longitude" precision="1" seconds="02.37" value="7.967325">07°58'02.37"E</geoCoordinate>
|
||
, leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon subsp. thomasiana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon thomasiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Santuario di Oropa, Biella, Piedmont, Italy. 1100 m asl
|
||
<geoCoordinate degrees="45" direction="north" minutes="37" orientation="latitude" precision="1" seconds="49.01" value="45.63028">45°37'49.01"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="07" direction="east" minutes="58" orientation="longitude" precision="1" seconds="42.07" value="7.978353">07°58'42.07"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria ornata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ornata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria ornata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
|
||
, 1836)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Ogulin city centre, Croatia. 320 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="16" orientation="latitude" precision="1" seconds="2.55" value="45.267376">45°16'2.55"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="15" direction="east" minutes="13" orientation="longitude" precision="1" seconds="33.34" value="15.225927">15°13'33.34"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP+ELP</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authority="itala" authorityName="itala (G. von Martens" authorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala itala</emphasis>
|
||
</taxonomicName>
|
||
(G. von Martens, 1824)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
San Donato, Barbarano Vicentino, Vicenza, Veneto, Italy. 300 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="24" orientation="latitude" precision="1" seconds="22.30" value="45.406197">45°24'22.30"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="11" direction="east" minutes="31" orientation="longitude" precision="1" seconds="11.93" value="11.51998">11°31'11.93"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Y</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="De Cristofori & Jan" baseAuthorityYear="1832" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. albopustolata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="albopustolata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala albopustolata</emphasis>
|
||
</taxonomicName>
|
||
(De Cristofori & Jan, 1832)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Brescia, city castle walls, Lombardia, Italy. 200 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="32" orientation="latitude" precision="1" seconds="29.87" value="45.54163">45°32'29.87"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="13" orientation="longitude" precision="1" seconds="28.37" value="10.224546">10°13'28.37"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER+ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Kaeufel" baseAuthorityYear="1928" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. allatollae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="allatollae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala allatollae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Käufel">Kaeufel</normalizedToken>
|
||
, 1928)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Albergo alla Tolla, Val Ampola, Storo, Trentino-Alto Adige, Italy. 720 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="51" orientation="latitude" precision="1" seconds="7.62" value="45.852116">45°51'7.62"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="38" orientation="longitude" precision="1" seconds="04.51" value="10.634586">10°38'04.51"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Strobel" baseAuthorityYear="1851" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. baldensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="baldensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala baldensis</emphasis>
|
||
</taxonomicName>
|
||
(Strobel, 1851)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Naole, San Zeno di Montagna, Verona, Veneto, Italy. 900 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="38" orientation="latitude" precision="1" seconds="43.67" value="45.645466">45°38'43.67"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="44" orientation="longitude" precision="1" seconds="15.86" value="10.73774">10°44'15.86"E</geoCoordinate>
|
||
, I. Niero leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Y</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ER+PP+ELP</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Strobel" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. balsamoi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="balsamoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala balsamoi</emphasis>
|
||
</taxonomicName>
|
||
(Strobel, 1850)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Loc. Galleria, Bracca, Val Serina, Bergamo, Lombardia, Italy.
|
||
<geoCoordinate degrees="45" direction="north" minutes="49" orientation="latitude" precision="1" seconds="24.49" value="45.823467">45°49'24.49"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="09" direction="east" minutes="43" orientation="longitude" precision="1" seconds="06.73" value="9.718535">09°43'06.73"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER(ELP))</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. clavata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="clavata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala clavata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
|
||
, 1836)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Ballabio, Lecco, Lombardia, Italy. 710 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="54" orientation="latitude" precision="1" seconds="25.96" value="45.90721">45°54'25.96"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="09" direction="east" minutes="25" orientation="longitude" precision="1" seconds="55.16" value="9.43199">09°25'55.16"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Kuester" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. latestriata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="latestriata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala latestriata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Bedulita, Bergamo, Lombardia, Italy. 625 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="47" orientation="latitude" precision="1" seconds="16.15" value="45.78782">45°47'16.15"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="09" direction="east" minutes="32" orientation="longitude" precision="1" seconds="42.26" value="9.545073">09°32'42.26"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER+ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Gredler" baseAuthorityYear="1869" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. lorinae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="lorinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala lorinae</emphasis>
|
||
</taxonomicName>
|
||
(Gredler, 1869)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Val di Lorina, Storo, Brescia, Lombardia, Italy. 600 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="50" orientation="latitude" precision="1" seconds="35.28" value="45.843132">45°50'35.28"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="37" orientation="longitude" precision="1" seconds="12.47" value="10.620131">10°37'12.47"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Y</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">ER+PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. zalloti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="zalloti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala zalloti</emphasis>
|
||
</taxonomicName>
|
||
nom. nov.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Udine, Friuli-Venezia Giulia, Italy. 140 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="03" orientation="latitude" precision="1" seconds="53.37" value="46.064823">46°03'53.37"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="13" direction="east" minutes="14" orientation="longitude" precision="1" seconds="12.35" value="13.236764">13°14'12.35"E</geoCoordinate>
|
||
, L. Anzil leg., W. De Mattia det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER(ELP))</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authorityName="De Mattia & Reier & Haring" authorityYear="2021" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. zalloti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="zalloti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala zalloti</emphasis>
|
||
</taxonomicName>
|
||
nom. nov.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Udine, Friuli-Venezia Giulia, Italy. 140 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="03" orientation="latitude" precision="1" seconds="53.37" value="46.064823">46°03'53.37"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="13" direction="east" minutes="14" orientation="longitude" precision="1" seconds="12.35" value="13.236764">13°14'12.35"E</geoCoordinate>
|
||
, L. Anzil leg., W. De Mattia det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER(ELP))</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Nordsieck" baseAuthorityYear="1963" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. serravalensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="serravalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala serravalensis</emphasis>
|
||
</taxonomicName>
|
||
(Nordsieck, 1963)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Serravalle near Vittorio Veneto, Veneto, Italy. 500 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="00" orientation="latitude" precision="1" seconds="04.46" value="46.00124">46°00'04.46"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12" direction="east" minutes="17" orientation="longitude" precision="1" seconds="11.13" value="12.286426">12°17'11.13"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">nonstenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER(ELP))</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Kaeufel" baseAuthorityYear="1928" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. trepida" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="trepida">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala trepida</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Käufel">Kaeufel</normalizedToken>
|
||
, 1928)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Forte Cima Ora, Anfo, Brescia, Lombardia, Italy. 1500 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="47" orientation="latitude" precision="1" seconds="50.57" value="45.79738">45°47'50.57"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="28" orientation="longitude" precision="1" seconds="03.60" value="10.467667">10°28'03.60"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authorityName="Nardi" authorityYear="2011" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. triumplinae" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="triumplinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala triumplinae</emphasis>
|
||
</taxonomicName>
|
||
Nardi, 2011
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Cima Caldoline, Lavenone, Brescia, Lombardia, Italy. 1700 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="48" orientation="latitude" precision="1" seconds="03.54" value="45.800983">45°48'03.54"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="10" direction="east" minutes="24" orientation="longitude" precision="1" seconds="20.12" value="10.405588">10°24'20.12"E</geoCoordinate>
|
||
, G. Nardi leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER(ELP))</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Pini" baseAuthorityYear="1883" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala subsp. variscoi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="itala" subSpecies="variscoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala variscoi</emphasis>
|
||
</taxonomicName>
|
||
(Pini, 1883)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Lavina, Val Taleggio, Bergamo, Lombardia, Italy. 650 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="53" orientation="latitude" precision="1" seconds="11.73" value="45.886593">45°53'11.73"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="09" direction="east" minutes="33" orientation="longitude" precision="1" seconds="55.46" value="9.565406">09°33'55.46"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">PP(ER+ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authority="stenzii" authorityName="stenzii (Rossmaessler" authorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. stenzii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii stenzii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
|
||
, 1836)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
San Romedio, Trento, Italy. 730 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="22" orientation="latitude" precision="1" seconds="03.83" value="46.36773">46°22'03.83"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="11" direction="east" minutes="06" orientation="longitude" precision="1" seconds="31.84" value="11.108845">11°06'31.84"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Bank" baseAuthorityYear="1987" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. butoti" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="butoti">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii butoti</emphasis>
|
||
</taxonomicName>
|
||
(Bank, 1987)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Fai della Paganella, Trento, Trentino-Alto Adige, Italy. 600 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="12" orientation="latitude" precision="1" seconds="57.04" value="46.215847">46°12'57.04"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="11" direction="east" minutes="04" orientation="longitude" precision="1" seconds="22.88" value="11.073022">11°04'22.88"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Brumati" baseAuthorityYear="1838" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. cincta" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="cincta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii cincta</emphasis>
|
||
</taxonomicName>
|
||
(Brumati, 1838)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Bosplans, Pordenone, Friuli-Venezia Giulia, Italy. 500 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="11" orientation="latitude" precision="1" seconds="58.11" value="46.199474">46°11'58.11"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12" direction="east" minutes="38" orientation="longitude" precision="1" seconds="5.24" value="12.6347885">12°38'5.24"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Y</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Brumati" baseAuthorityYear="1838" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. cincta" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="cincta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii cincta</emphasis>
|
||
</taxonomicName>
|
||
(Brumati, 1838)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Val Resia, sella di Carnizza, Tarvisio, Friuli-Venezia Giulia, Italy. 400 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="20" orientation="latitude" precision="1" seconds="15.55" value="46.33765">46°20'15.55"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="13" direction="east" minutes="19" orientation="longitude" precision="1" seconds="17.11" value="13.32142">13°19'17.11"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det. Mattia leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Bank" baseAuthorityYear="1987" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. faueri" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="faueri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii faueri</emphasis>
|
||
</taxonomicName>
|
||
(Bank, 1987)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Grotte di Oliero Valstagna, Vicenza, Veneto, Italy. 800 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="50" orientation="latitude" precision="1" seconds="47.79" value="45.846607">45°50'47.79"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="11" direction="east" minutes="40" orientation="longitude" precision="1" seconds="04.24" value="11.667845">11°40'04.24"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Gredler" baseAuthorityYear="1874" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. letochana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="letochana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii letochana</emphasis>
|
||
</taxonomicName>
|
||
(Gredler, 1874)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Val Fonda, Misurina, Belluno, Veneto, Italy. 1500 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="36" orientation="latitude" precision="1" seconds="27.42" value="46.607616">46°36'27.42"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12" direction="east" minutes="12" orientation="longitude" precision="1" seconds="20.01" value="12.205558">12°12'20.01"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">Y</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="Fauer" baseAuthorityYear="1991" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. nordsiecki" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="nordsiecki">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii nordsiecki</emphasis>
|
||
</taxonomicName>
|
||
Fauer, 1991
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Cison di Val Marino, Treviso, Veneto, Italy. 250 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="58" orientation="latitude" precision="1" seconds="05.53" value="45.968204">45°58'05.53"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12" direction="east" minutes="08" orientation="longitude" precision="1" seconds="52.60" value="12.147944">12°08'52.60"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">no pseudopapilla</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName baseAuthorityName="De Betta & Martinati" baseAuthorityYear="1855" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. paroliniana" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="paroliniana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii paroliniana</emphasis>
|
||
</taxonomicName>
|
||
(De Betta & Martinati, 1855)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Loc. La Goccia, Foza, Valstagna, Vicenza, Veneto, Italy. 800 m asl.
|
||
<geoCoordinate degrees="45" direction="north" minutes="52" orientation="latitude" precision="1" seconds="29.20" value="45.874775">45°52'29.20"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="11" direction="east" minutes="38" orientation="longitude" precision="1" seconds="59.73" value="11.649925">11°38'59.73"E</geoCoordinate>
|
||
, W. De Mattia and J. Macor leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="1">
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
<taxonomicName authorityName="Nordsieck" authorityYear="1993" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii subsp. westerlundi" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="stenzii" subSpecies="westerlundi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii westerlundi</emphasis>
|
||
</taxonomicName>
|
||
(Nordsieck, 1993)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
|
||
Val Fiscalina, Croda Rossa, Sesto, Bolzano, Trentino-Alto Adige, Italy. 1500 m asl.
|
||
<geoCoordinate degrees="46" direction="north" minutes="38" orientation="latitude" precision="1" seconds="02.83" value="46.63412">46°38'02.83"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="12" direction="east" minutes="21" orientation="longitude" precision="1" seconds="42.79" value="12.361886">12°21'42.79"E</geoCoordinate>
|
||
, I. Niero leg. and det.
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">stenzioid</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">N</td>
|
||
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">EP(ELP)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">The present anatomical investigation indeed generated consistent results that are congruent with the molecular genetic outcomes.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName baseAuthorityName="Rossmaessler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria ornata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="ornata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria ornata</emphasis>
|
||
</taxonomicName>
|
||
-
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
groups of taxa show the same external overall shape of the distal genital organs (Figs
|
||
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Charpentieria dyodon dyodon (S. Studer, 1820), Verbano 40.1 whole distal genital organs 40.2 internal distal part of genital organs. Charpentieria dyodon alpina (Stabile, 1859). Val di Lanzo, Usseglio 40.3 whole distal genital organs 40.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure40" httpUri="https://binary.pensoft.net/fig/622832" pageId="0" pageNumber="1">40</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Charpentieria ornata (Rossmaessler, 1836), Ogulin, HR 43.1 whole distal genital organs 43.2 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure43" httpUri="https://binary.pensoft.net/fig/622835" pageId="0" pageNumber="1">43</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 46" captionStartId="F46" captionText="Figure 46. Charpentieria itala clavata (Rossmaessler, 1836), Ballabio, Lecco 46.1 whole distal genital organs 46.2 internal distal part of genital organs 46.3 cross section of the pseudopapilla 46.3 spermatophore. Charpentieria itala latestriata (Kuester, 1850), Bedulita, Bergamo 46.5 whole distal genital organs 46.6 internal distal part of genital organs. Charpentieria itala lorinae (Gredler, 1869), Val Lorina, Storo 46.7 whole distal genital organs 46.8 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure46" httpUri="https://binary.pensoft.net/fig/622838" pageId="0" pageNumber="1">46</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 48" captionStartId="F48" captionText="Figure 48. Charpentieria itala zalloti nom. nov., Udine 48.1 whole distal genital organs 48.2 internal distal part of genital organs. Charpentieria itala serravalensis (Nordsieck, 1963), Serravalle, Vittorio Veneto 48.3 whole distal genital organs 48.4 internal distal part of genital organs. Charpentieria itala trepida (Kaeufel, 1928), Cima Ora, Anfo 48.5 whole distal genital organs 48.6 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure48" httpUri="https://binary.pensoft.net/fig/622840" pageId="0" pageNumber="1">48</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Charpentieria itala triumplinae Nardi, 2011, Lavenone, Brescia 49.1 whole distal genital organs 49.2 internal distal part of genital organs. Charpentieria itala variscoi (Pini, 1883), Val Taleggio, Bergamo 49.3 whole distal genital organs 49.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure49" httpUri="https://binary.pensoft.net/fig/622841" pageId="0" pageNumber="1">49</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Charpentieria stenzii stenzii (Rossmaessler, 1836), San Romedio, Trento 51.1 whole distal genital organs 51.2 internal distal part of genital organs 51.3 cross section of the epiphallus. Charpentieria stenzii butoti (Bank, 1987), Fai della Paganella 51.4 whole distal genital organs 51.5. internal distal part of genital organs 51.6 detail of the hemipapilla: uncut 51.7 detail of the hemipapilla: cut funnel 51.8 cross section of the epiphallus." figureDoi="10.3897/zookeys.1077.67081.figure51" httpUri="https://binary.pensoft.net/fig/622843" pageId="0" pageNumber="1">51</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 55" captionStartId="F55" captionText="Figure 55. Charpentieria stenzii nordsiecki Fauer, 1991, Cison di Valmarino, Treviso 55.1 whole distal genital organs 55.2 internal distal part of genital organs. Charpentieria stenzii paroliniana (De Betta & Martinati, 1855), Valstagna, Vicenza 55.3 whole distal genital organs 55.4 detail of the hemipapilla: uncut 55.5 internal distal part of genital organs with cut funnel. Charpentieria stenzii westerlundi Nordsieck, 1993, Val Fiscalina, Bolzano 55.6 whole distal genital organs 55.7 internal distal part of genital organs with cut funnel." figureDoi="10.3897/zookeys.1077.67081.figure55" httpUri="https://binary.pensoft.net/fig/622847" pageId="0" pageNumber="1">55</figureCitation>
|
||
). No major difference can be observed regarding the main genital parts as, e.g., the shape and relative dimensions of the OSD, FO, A and penial complex (P+E). Only the V is shorter in
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
, with an average P/V ratio of 1.4 for
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
and 2.3 for
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, this general external genital arrangement of
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
is not distinguishable by any means from
|
||
<taxonomicName authorityName="Vest" authorityYear="1867" class="Gastropoda" family="Clausiliidae" genus="Siciliaria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Siciliaria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Siciliaria</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Tomlin" authorityYear="1929" class="Gastropoda" family="Clausiliidae" genus="Sicania" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sicania" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Sicania</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authorityName="O. Boettger" authorityYear="1879" class="Gastropoda" family="Clausiliidae" genus="Mauritanica" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mauritanica" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Mauritanica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The genus
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
was introduced by
|
||
<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.12997" author="Stabile, J" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali" pageId="0" pageNumber="1" pagination="3 - 141" refId="B93" refString="Stabile, J, 1864. Mollusques terrestres vivants du Piemont. Atti della Societa Italiana di Scienze Naturali 7 (1): 3 - 141, DOI: https://doi.org/10.5962/bhl.title.12997" title="Mollusques terrestres vivants du Piemont." url="https://doi.org/10.5962/bhl.title.12997" volume="7" year="1864">Stabile (1864</bibRefCitation>
|
||
: 80) [type species:
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
(S. Studer, 1820)]. The type species was subsequently designated by
|
||
<bibRefCitation author="Kennard, AS" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="298 - 308" refId="B37" refString="Kennard, AS, Woodward, BB, 1923. Note on the nomenclature and systematic arrangement of the Clausiliidae. Proceedings of the Malacological Society of London 15 (6): 298 - 308" title="Note on the nomenclature and systematic arrangement of the Clausiliidae." volume="15" year="1923">Kennard and Woodward (1923</bibRefCitation>
|
||
: 306).
|
||
<bibRefCitation author="Boettger, O" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B8" refString="Boettger, O, 1877. Clausilienstudien. Palaeontographica (N. F.) Suppl.3: 1-122. [4 pls. Cassel]" title="Clausilienstudien. Palaeontographica (N. F.) Suppl. 3: 1 - 122. [4 pls. Cassel]" year="1877">Boettger (1877</bibRefCitation>
|
||
: 53) listed only
|
||
<taxonomicName lsidName="C. dyodon" pageId="0" pageNumber="1" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon</emphasis>
|
||
</taxonomicName>
|
||
and its subspecies for the genus
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation author="Boettger, O" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B8" refString="Boettger, O, 1877. Clausilienstudien. Palaeontographica (N. F.) Suppl.3: 1-122. [4 pls. Cassel]" title="Clausilienstudien. Palaeontographica (N. F.) Suppl. 3: 1 - 122. [4 pls. Cassel]" year="1877">Boettger (1877</bibRefCitation>
|
||
: 35) introduced the name
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Tirolica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tirolica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Tirolica</emphasis>
|
||
</taxonomicName>
|
||
as "Gruppe der
|
||
<taxonomicName genus="Stentzi" lsidName="Stentzi" pageId="0" pageNumber="1" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Stentzi</emphasis>
|
||
</taxonomicName>
|
||
" (type species:
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Delima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Delima stenzii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Delima stenzii</emphasis>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Charpentieria</taxonomicName>
|
||
<taxonomicName baseAuthorityName="Rossmassler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii</emphasis>
|
||
</taxonomicName>
|
||
), that included most of the stenzioid southern alpine taxa known in 1877. This genus included taxa currently belonging to both
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="C. i. subsp. balsamoi" pageId="0" pageNumber="1" rank="subSpecies" species="i." subSpecies="balsamoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. i. balsamoi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. i. subsp. lorinae" pageId="0" pageNumber="1" rank="subSpecies" species="i." subSpecies="lorinae">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. i. lorinae</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName lsidName="C. s. subsp. stenzii" pageId="0" pageNumber="1" rank="subSpecies" species="s." subSpecies="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. s. stenzii</emphasis>
|
||
</taxonomicName>
|
||
and its synonym
|
||
<taxonomicName authorityName="Kuster" authorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia saccata" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="saccata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia saccata</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1848,
|
||
<taxonomicName lsidName="C. stenzii subsp. cincta" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="cincta">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii cincta</emphasis>
|
||
</taxonomicName>
|
||
and its synonyms:
|
||
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1841" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia rossmaessleri" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="rossmaessleri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia rossmaessleri</emphasis>
|
||
</taxonomicName>
|
||
L. Pfeiffer, 1841,
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia cincta subsp. var. var. maior" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="cincta" subSpecies="var." variety="maior">Clausilia cincta var. maior</taxonomicName>
|
||
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
|
||
, 1842 and
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia cincta subsp. var. var. minor" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="variety" species="cincta" subSpecies="var." variety="minor">Clausilia cincta var. minor</taxonomicName>
|
||
Westerlund, 1878 and finally
|
||
<taxonomicName lsidName="C. s. subsp. letochana" pageId="0" pageNumber="1" rank="subSpecies" species="s." subSpecies="letochana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. s. letochana</emphasis>
|
||
</taxonomicName>
|
||
and its synonym
|
||
<taxonomicName authorityName="Gredler" authorityYear="1874" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia funki" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="funki">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia funki</emphasis>
|
||
</taxonomicName>
|
||
Gredler, 1874). Inexpicably,
|
||
<bibRefCitation author="Boettger, O" journalOrPublisher="Bullettino della Societa Malacologica Italiana" pageId="0" pageNumber="1" refId="B8" refString="Boettger, O, 1877. Clausilienstudien. Palaeontographica (N. F.) Suppl.3: 1-122. [4 pls. Cassel]" title="Clausilienstudien. Palaeontographica (N. F.) Suppl. 3: 1 - 122. [4 pls. Cassel]" year="1877">Boettger (1877</bibRefCitation>
|
||
:35) did not include the stenzioid
|
||
<taxonomicName lsidName="C. stenzii subsp. paroliniana" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="paroliniana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii paroliniana</emphasis>
|
||
</taxonomicName>
|
||
(De Betta & Martinati, 1855) and
|
||
<taxonomicName lsidName="C. itala subsp. clavata" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="clavata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala clavata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2020.106982" author="Xu, J" journalOrPublisher="Petersbourg, Classe physico-mathematique 11: [1, 2], [1 - 37]" pageId="0" pageNumber="1" refId="B106" refString="Xu, J, Hausdorf, B, 2021. Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982. https://doi.org/10.1016/j.ympev.2020.106982" title="Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982." url="https://doi.org/10.1016/j.ympev.2020.106982" year="2021">Xu and Hausdorf (2021</bibRefCitation>
|
||
: 6), after sequencing 843 loci, found that
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
is well embedded in
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
. Although our anatomical investigations revealed a peculiar internal structure of the P-E transition and the penial pseudopapilla is found exclusively in
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
(we named it hemipapilla, see point 6.6 for a detailed decription), we follow
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2020.106982" author="Xu, J" journalOrPublisher="Petersbourg, Classe physico-mathematique 11: [1, 2], [1 - 37]" pageId="0" pageNumber="1" refId="B106" refString="Xu, J, Hausdorf, B, 2021. Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982. https://doi.org/10.1016/j.ympev.2020.106982" title="Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982." url="https://doi.org/10.1016/j.ympev.2020.106982" year="2021">Xu and Hausdorf (2021)</bibRefCitation>
|
||
in not separating this group on generic level. The hemipapilla was not found in the stenzioid subspecies despite the notable gene flow with
|
||
<taxonomicName lsidName="C. stenzii" pageId="0" pageNumber="1" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii</emphasis>
|
||
</taxonomicName>
|
||
, as proven by
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2020.106982" author="Xu, J" journalOrPublisher="Petersbourg, Classe physico-mathematique 11: [1, 2], [1 - 37]" pageId="0" pageNumber="1" refId="B106" refString="Xu, J, Hausdorf, B, 2021. Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982. https://doi.org/10.1016/j.ympev.2020.106982" title="Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982." url="https://doi.org/10.1016/j.ympev.2020.106982" year="2021">Xu and Hausdorf (2021</bibRefCitation>
|
||
: 7).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Boettger, 1877, introduced the genus
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Itala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Itala</emphasis>
|
||
</taxonomicName>
|
||
as "Gruppe der
|
||
<taxonomicName authority="itala" class="Gastropoda" family="Clausiliidae" genus="Itala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itala itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Itala itala</emphasis>
|
||
</taxonomicName>
|
||
" (type species:
|
||
<taxonomicName baseAuthorityName="G. v. Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Delima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Delima itala" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Delima itala</emphasis>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Charpentieria</taxonomicName>
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
), including all the alpine non-stenzioid taxa, excluding
|
||
<taxonomicName lsidName="C. dyodon" pageId="0" pageNumber="1" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon</emphasis>
|
||
</taxonomicName>
|
||
but including two stenzioid taxa:
|
||
<taxonomicName lsidName="C. stenzii subsp. paroliniana" pageId="0" pageNumber="1" rank="subSpecies" species="stenzii" subSpecies="paroliniana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. stenzii paroliniana</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="C. itala subsp. clavata" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="clavata">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala clavata</emphasis>
|
||
</taxonomicName>
|
||
. He also included two taxa from the Balkans:
|
||
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Delima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Delima conspersa" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="conspersa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Delima conspersa</emphasis>
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName baseAuthorityName="L. Pfeiffer" baseAuthorityYear="1848" class="Gastropoda" family="Clausiliidae" genus="Strigilodelima" kingdom="Animalia" lsidName="Strigilodelima conspersa" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="conspersa">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Strigilodelima conspersa</emphasis>
|
||
</taxonomicName>
|
||
(L. Pfeiffer, 1848)] and its synonym
|
||
<taxonomicName baseAuthorityName="Kuster" baseAuthorityYear="1850" class="Gastropoda" family="Clausiliidae" genus="Delima" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Delima platystoma" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="platystoma">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Delima platystoma</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850. As for
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Tirolica" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Tirolica" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Tirolica</emphasis>
|
||
</taxonomicName>
|
||
, the recent results by
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2020.106982" author="Xu, J" journalOrPublisher="Petersbourg, Classe physico-mathematique 11: [1, 2], [1 - 37]" pageId="0" pageNumber="1" refId="B106" refString="Xu, J, Hausdorf, B, 2021. Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982. https://doi.org/10.1016/j.ympev.2020.106982" title="Repeated hybridization increased diversity in the door snail complex Charpentieria itala in the Southern Alps. Molecular Phylogenetics and Evolution 155: 106982." url="https://doi.org/10.1016/j.ympev.2020.106982" year="2021">Xu and Hausdorf (2021</bibRefCitation>
|
||
: 6-8) indicated that the
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
complex should not be separated on a (sub)generic level (i.e.,
|
||
<taxonomicName authorityName="O.Boettger" authorityYear="1877" class="Gastropoda" family="Clausiliidae" genus="Itala" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Itala</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation author="Wagner, AJ" journalOrPublisher="Annales Zoologici Musei Polonici Historiae Naturalis" pageId="0" pageNumber="1" pagination="1 - 73" refId="B100" refString="Wagner, AJ, 1925. Studien ueber die Systematik, Stammesgeschichte und geographische Verbreitung des Genus Delima (Hartmann) A. J. Wagner. Annales Zoologici Musei Polonici Historiae Naturalis 4 (1): 1 - 73" title="Studien ueber die Systematik, Stammesgeschichte und geographische Verbreitung des Genus Delima (Hartmann) A. J. Wagner." volume="4" year="1925">Wagner (1925</bibRefCitation>
|
||
: 17) introduced the genus subgenus
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia (Alpidelima)" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subGenus" subGenus="Alpidelima">Clausilia Alpidelima</taxonomicName>
|
||
[type species
|
||
<taxonomicName authorityName="G. von Martens" authorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia itala</emphasis>
|
||
</taxonomicName>
|
||
Martens, 1824, by subsequent designation by
|
||
<bibRefCitation author="Schileyko, AA" journalOrPublisher="Ruthenica, Supplement" pageId="0" pageNumber="1" pagination="563 - 729" refId="B88" refString="Schileyko, AA, 2000. Treatise on recent terrestrial pulmonate molluscs. 5. Clausiliidae. Ruthenica, Supplement 2: 563 - 729" title="Treatise on recent terrestrial pulmonate molluscs. 5. Clausiliidae." volume="2" year="2000">Schileyko (2000</bibRefCitation>
|
||
: 664)] that must be deemed as a junior synonym of
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="81 - 115" refId="B57" refString="Nordsieck, H, 1963a. Zur Anatomie und Systematik der Clausilien, I. Archiv fuer Molluskenkunde 92: 81 - 115" title="Zur Anatomie und Systematik der Clausilien, I." volume="92" year="1963 a">Nordsieck (1963a</bibRefCitation>
|
||
: 95, figs 9-10; 97, fig. 11; 1963b: 185, 189, 193, 197) depicted the genital organs of some taxa currently belonging to the genus
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
from the Alps. The sketchy drawings lacked important details, both concerning internal and external features. Except for fig. 11a of
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="81 - 115" refId="B57" refString="Nordsieck, H, 1963a. Zur Anatomie und Systematik der Clausilien, I. Archiv fuer Molluskenkunde 92: 81 - 115" title="Zur Anatomie und Systematik der Clausilien, I." volume="92" year="1963 a">Nordsieck (1963a)</bibRefCitation>
|
||
, the exact position of the insertion of the proximal vas deferens into the beginning of the free oviduct was not clear. The internal structure of the penis-epiphallus transition was not investigated. The pseudopapilla was shown, but its origin was missing as well as its connection to (and the presence/absence of) the epiphallar ring(s) and the epiphallar longitudinal pleats. Only in
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="81 - 115" refId="B57" refString="Nordsieck, H, 1963a. Zur Anatomie und Systematik der Clausilien, I. Archiv fuer Molluskenkunde 92: 81 - 115" title="Zur Anatomie und Systematik der Clausilien, I." volume="92" year="1963 a">Nordsieck (1963a</bibRefCitation>
|
||
: fig. 11b) the origin of the pseudopapilla and the epiphallar ring seem to be roughly depicted.
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="81 - 115" refId="B57" refString="Nordsieck, H, 1963a. Zur Anatomie und Systematik der Clausilien, I. Archiv fuer Molluskenkunde 92: 81 - 115" title="Zur Anatomie und Systematik der Clausilien, I." volume="92" year="1963 a">Nordsieck (1963a)</bibRefCitation>
|
||
reported the presence of the penial flagellum for all the four taxa, which is in contrast to our results, based on 63 specimens, where this particular anatomical structure was never detected despite careful check.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
He introduced a new subspecific taxon,
|
||
<taxonomicName lsidName="C. itala subsp. serravalensis" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="serravalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala serravalensis</emphasis>
|
||
</taxonomicName>
|
||
(1963a: 175), but depicted its external anatomy of the genital organs [Nordsieck, 1963b: 184-185, Abb. 47
|
||
<emphasis italics="true" pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">Charpentieria itala</taxonomicName>
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria serravalensis" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="serravalensis">Charpentieria serravalensis</taxonomicName>
|
||
</emphasis>
|
||
, Serravalle (
|
||
<normalizedToken originalValue="präp">praep</normalizedToken>
|
||
. 109)] with the same genital drawing provided for
|
||
<taxonomicName authority="itala" lsidName="C. itala subsp. itala" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala itala</emphasis>
|
||
</taxonomicName>
|
||
in Nordsieck, (1963a: 97, fig. 11a, b). The shell description, genital organs and measures provided for
|
||
<taxonomicName lsidName="C. itala subsp. serravalensis" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="serravalensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala serravalensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">Nordsieck 1963b</bibRefCitation>
|
||
: 175) match with the morphology of the specimens from Colli Euganei (San Donato, Barbarano Vicentino, Padova, Italy) (Fig.
|
||
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Charpentieria itala itala (G. von Martens, 1824), Barbarano Vicentino 44.1, 44.2 whole distal genital organs 44.3 internal distal part of genital organs 44.4 detail of the pseudopapilla. Charpentieria itala albopustolata (De Cristofori & Jan, 1832), Brescia 44.5 whole distal genital organs 44.6 internal distal part of genital organs. Charpentieria itala allatollae (Kaeufel, 1928), Val Ampola, Storo 44.7 whole distal genital organs 44.8 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure44" httpUri="https://binary.pensoft.net/fig/622836" pageId="0" pageNumber="1">44</figureCitation>
|
||
.1-44.4) studied in the present paper.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Nordsieck cites
|
||
<taxonomicName authority="itala" lsidName="C. itala subsp. itala" pageId="0" pageNumber="1" rank="subSpecies" species="itala" subSpecies="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala itala</emphasis>
|
||
</taxonomicName>
|
||
(1963a) but unexpectedly,
|
||
<bibRefCitation author="Nordsieck, H" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="169 - 203" refId="B58" refString="Nordsieck, H, 1963b. Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen. Archiv fuer Molluskenkunde 92: 169 - 203" title="Zur Anatomie und Systematik der Clausilien, II. Die Formenbildung des Genus Delima in den Suedalpen." volume="92" year="1963 b">Nordsieck (1963b)</bibRefCitation>
|
||
neither cited the nominate subspecies, nor did he provide information about the type locality of the species
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
was described by G. von
|
||
<bibRefCitation author="Martens, G von" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="1" refId="B47" refString="Martens, G von, 1824. Reise nach Venedig. Zweiter Theil. Venedig. Euganeen. Alpen von Belluno. Tirol Baiern Naturgeschichtlicher Angang. Stettin, Ulm, [I-VI,] 664 pp." title="Reise nach Venedig. Zweiter Theil. Venedig. Euganeen. Alpen von Belluno. Tirol Baiern Naturgeschichtlicher Angang. Stettin, Ulm, [I-VI,] 664 pp." year="1824">Martens (1824</bibRefCitation>
|
||
: 442), but he did not provide any indication about the collecting site. L.
|
||
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Paris, [1, I-IV = 1 - 4]" pageId="0" pageNumber="1" refId="B73" refString="Pfeiffer, L, 1853. Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen tertium. Lipsiae, [I-VIII,] 711 pp." title="Monographia heliceorum viventium. Sistens descriptiones systematicas et criticas omnium huius familiae generum et specierum hodie cognitarum. Volumen tertium. Lipsiae, [I-VIII,] 711 pp." year="1853">Pfeiffer (1853</bibRefCitation>
|
||
: 606) reports the description of the species given by Martens. In his "Reisebemerkungen
|
||
<normalizedToken originalValue="über">ueber</normalizedToken>
|
||
einige Binnenschnecken Italiens", E. von
|
||
<bibRefCitation author="Martens, E von" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="1" pagination="120 - 155" refId="B46" refString="Martens, E von, 1857. Reisebemerkungen ueber einige binnenschnecken Italiens. Malakozoologische Blaetter 4: 120 - 155" title="Reisebemerkungen ueber einige binnenschnecken Italiens." volume="4" year="1857">Martens (1857</bibRefCitation>
|
||
: 129), the son of G. von Martens, finally gives precise indications where his father collected the first series of specimens of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
(between Mira and Dolo): "Zwischen Mira und Dolo hatte mein Vater auch
|
||
<normalizedToken originalValue="früher">frueher</normalizedToken>
|
||
seine
|
||
<taxonomicName authorityName="G. von Martens" authorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Clausilia" kingdom="Animalia" lsidName="Clausilia itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Clausilia itala</emphasis>
|
||
</taxonomicName>
|
||
.". Dolo and Mira are located in the province of Venice, between Venice and Padova, along the river Brenta in the Pianura Padana. Being it a quaternary alluvial plain area, the specimens were probably collected on walls and houses. Eduard von
|
||
<bibRefCitation author="Martens, E von" journalOrPublisher="Malakozoologische Blaetter" pageId="0" pageNumber="1" pagination="120 - 155" refId="B46" refString="Martens, E von, 1857. Reisebemerkungen ueber einige binnenschnecken Italiens. Malakozoologische Blaetter 4: 120 - 155" title="Reisebemerkungen ueber einige binnenschnecken Italiens." volume="4" year="1857">Martens (1857</bibRefCitation>
|
||
: 129) collected more samples of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
from the walls of the Botanic Garden of Padova. These alluvial plain populations probably originate from downstream transport by creeks, deriving from the Colli Euganei and Colli Berici, which flow into the Brenta River.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="brief genital anatomical description of charpentieria dyodon subspecific taxa: the parapseudopapilla">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Brief genital anatomical description of
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
subspecific taxa: the parapseudopapilla
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
is the name-bearing species of the genus
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
. It comprises five subspecies [
|
||
<taxonomicName authority="dyodon" lsidName="C. dyodon subsp. dyodon" pageId="0" pageNumber="1" rank="subSpecies" species="dyodon" subSpecies="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon dyodon</emphasis>
|
||
</taxonomicName>
|
||
(Studer, 1820;
|
||
<taxonomicName lsidName="C. dyodon subsp. alpina" pageId="0" pageNumber="1" rank="subSpecies" species="dyodon" subSpecies="alpina">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon alpina</emphasis>
|
||
</taxonomicName>
|
||
(Stabile, 1859);
|
||
<taxonomicName lsidName="C. dyodon subsp. paulucciana" pageId="0" pageNumber="1" rank="subSpecies" species="dyodon" subSpecies="paulucciana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon paulucciana</emphasis>
|
||
</taxonomicName>
|
||
(Pollonera, 1885);
|
||
<taxonomicName lsidName="C. dyodon subsp. studeri" pageId="0" pageNumber="1" rank="subSpecies" species="dyodon" subSpecies="studeri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon studeri</emphasis>
|
||
</taxonomicName>
|
||
(Pini, 1885) and
|
||
<taxonomicName lsidName="C. dyodon subsp. thomasiana" pageId="0" pageNumber="1" rank="subSpecies" species="dyodon" subSpecies="thomasiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon thomasiana</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Küster">Kuester</normalizedToken>
|
||
, 1850)]. These taxa are distributed over an area that includes the western Southern Alps of Piemonte, Italy (from Dora Riparia River to Lago Maggiore) and the South Switzerland (Ticino and Valais). This species is not sympatric with any other
|
||
<taxonomicName authorityName="Stabile" authorityYear="1864" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
In our COI tree only the nominate subspecies
|
||
<taxonomicName class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria d. subsp. dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="subSpecies" species="d." subSpecies="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria d. dyodon</emphasis>
|
||
</taxonomicName>
|
||
was included (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. BI tree of based on the COI data set of Siciliaria, Sicania and related genera. BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure4" httpUri="https://binary.pensoft.net/fig/622796" pageId="0" pageNumber="1">4</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. BI tree of based on the concatenated COI and ITS 2 data set of Siciliaria, Sicania and related genera. Colours of Siciliaria and Sicania clades correspond to those in the distribution maps (Fig. 1 E, F). BI posterior probability values (> 0.90) of major nodes are indicated." figureDoi="10.3897/zookeys.1077.67081.figure6" httpUri="https://binary.pensoft.net/fig/622798" pageId="0" pageNumber="1">6</figureCitation>
|
||
). It emerged as the sister species of
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
, yet with only low node support.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Three out of five subspecies of
|
||
<taxonomicName lsidName="C. dyodon" pageId="0" pageNumber="1" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. dyodon</emphasis>
|
||
</taxonomicName>
|
||
were anatomically investigated (Table
|
||
<tableCitation captionStart="Table 8" captionStartId="T8" captionText="Table 8. Examined taxa of Charpentieria with information on shell type (stenzioid / nonstenzioid), availability of genetic data (DNA) and epiphallar formula." httpUri="http://table.plazi.org/id/66A5CD940D950B91226F96579AB448F1" pageId="0" pageNumber="1" tableUuid="66A5CD940D950B91226F96579AB448F1">8</tableCitation>
|
||
). They revealed a common overall genital arrangement (Figs
|
||
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Charpentieria dyodon dyodon (S. Studer, 1820), Verbano 40.1 whole distal genital organs 40.2 internal distal part of genital organs. Charpentieria dyodon alpina (Stabile, 1859). Val di Lanzo, Usseglio 40.3 whole distal genital organs 40.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure40" httpUri="https://binary.pensoft.net/fig/622832" pageId="0" pageNumber="1">40</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
|
||
), including a stable presence of the penial pseudopapilla. The pseudopapilla can be either simple or rooted. The internal wall of the penis is smooth, the epiphallus shows two fringed longitudinal pleats and the internal wall of V presents many smooth longitudinal pleats.
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
has a distribution detached from
|
||
<taxonomicName lsidName="C. itala" pageId="0" pageNumber="1" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">C. itala</emphasis>
|
||
</taxonomicName>
|
||
with no overlapping areas.
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
shows substantial differences of the shell characters (smooth or weakly striated surface, lunella mostly absent and no palatal folds, sutural papillae very scarce or absent) and epiphallar structure (see the following chapter).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The four populations of
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
included for dissection, revealed a very stable morphology of the genital organs (Figs
|
||
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Charpentieria dyodon dyodon (S. Studer, 1820), Verbano 40.1 whole distal genital organs 40.2 internal distal part of genital organs. Charpentieria dyodon alpina (Stabile, 1859). Val di Lanzo, Usseglio 40.3 whole distal genital organs 40.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure40" httpUri="https://binary.pensoft.net/fig/622832" pageId="0" pageNumber="1">40</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
|
||
). The internal surface of the P is smooth and the V presents many smooth longitudinal pleats that stop before entering the atrium. The external transition between P and E is always visible.
|
||
<taxonomicName baseAuthorityName="S. Studer" baseAuthorityYear="1820" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria dyodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="dyodon">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria dyodon</emphasis>
|
||
</taxonomicName>
|
||
revealed a peculiar morphology of the pseudopapilla and the internal transition area between penis and epiphallus (Figs
|
||
<figureCitation captionStart="Figure 40" captionStartId="F40" captionText="Figure 40. Charpentieria dyodon dyodon (S. Studer, 1820), Verbano 40.1 whole distal genital organs 40.2 internal distal part of genital organs. Charpentieria dyodon alpina (Stabile, 1859). Val di Lanzo, Usseglio 40.3 whole distal genital organs 40.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure40" httpUri="https://binary.pensoft.net/fig/622832" pageId="0" pageNumber="1">40</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. Charpentieria dyodon thomasiana (Kuester, 1850), Piedicavallo 41.1 whole distal genital organs 41.2 internal distal part of genital organs. Oropa, Biella 41.3 whole distal genital organs 41.4 internal distal part of genital organs." figureDoi="10.3897/zookeys.1077.67081.figure41" httpUri="https://binary.pensoft.net/fig/622833" pageId="0" pageNumber="1">41</figureCitation>
|
||
). The parapseudopapilla involves the pseudopapilla and the epiphallar longitudinal pleats: the pseudopapilla has its origin at the distal end of one epiphallar longitudinal pleat (
|
||
<normalizedToken originalValue="“occupied“">"occupied"</normalizedToken>
|
||
epiphallar longitudinal pleat) and the opposite wall is delimited and narrowed by the other epiphallar longitudinal pleat (
|
||
<normalizedToken originalValue="“free“">"free"</normalizedToken>
|
||
epiphallar longitudinal pleat) and connected by a ER, creating a funnel-like structure. The PPP, at a first glance, resembles a true penial papilla, but it remarkably lacks the seminal channel. The extremely large ER of the PPP completely obstructs the lumen of the transition area. The connection between the penis and epiphallus is represented by a short, narrow passage, named "epiphallar funnel" (EF), that is found at the base of the PPP and it is quite difficult to detect.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
This parapseudopapilla represents an intermediate form between the simple structure of
|
||
<taxonomicName baseAuthorityName="G. von Martens" baseAuthorityYear="1824" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria itala" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="itala">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria itala</emphasis>
|
||
</taxonomicName>
|
||
and the hemipapilla (HP) of
|
||
<taxonomicName baseAuthorityName="Rossmassler" baseAuthorityYear="1836" class="Gastropoda" family="Clausiliidae" genus="Charpentieria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Charpentieria stenzii" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="stenzii">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Charpentieria stenzii</emphasis>
|
||
</taxonomicName>
|
||
. In the normal PP the transition stretch E is relatively wide and the counter wall opposite of the PP is represented by the simple P-E transition wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |