treatments-xml/data/03/B0/87/03B087FC5243EB008C83EF9EE37B5FD7.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.4763693" ID-GBIF-Dataset="9ef44358-db5b-4d04-a08a-0d44711ce040" ID-GBIF-Taxon="182074635" ID-ISSN="1854-0392" ID-Zenodo-Dep="4763693" checkinTime="1620964150388" checkinUser="felipe" docAuthor="Baumann, Richard W. &amp; Call, Ronald G." docDate="2012" docId="03B087FC5243EB008C83EF9EE37B5FD7" docLanguage="en" docName="Illiesia.8.8.104-110.pdf" docOrigin="Illiesia 8 (8)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Lednia tetonica Baumann &amp; Call 2012, sp. n." docType="treatment" docVersion="6" lastPageId="6" lastPageNumber="110" masterDocId="FF89FF845243EB068F6CE967E157597E" masterDocTitle="Lednia Tetonica, A New Species Of Stonefly From Wyoming (Plecoptera: Nemouridae)" masterLastPageNumber="110" masterPageNumber="104" pageId="0" pageNumber="104" updateTime="1643540827215" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Lednia Tetonica, A New Species Of Stonefly From Wyoming (Plecoptera: Nemouridae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baumann, Richard W.</mods:namePart>
<mods:affiliation>Department of Biology and Monte L. Bean Life Science Museum, Brigham Young University, Provo, UT, U. S. A. 84602 E-mail: richard _ baumann @ byu. edu</mods:affiliation>
<mods:nameIdentifier type="email">richard_baumann@byu.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Call, Ronald G.</mods:namePart>
<mods:affiliation>Madison High School, 2300 University Blvd., Rexburg, ID, U. S. A. 83440 E-mail: callr @ msd 321. com</mods:affiliation>
<mods:nameIdentifier type="email">callr@msd</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Illiesia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>8</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>104</mods:start>
<mods:end>110</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760806</mods:identifier>
<mods:identifier type="GBIF-Dataset">9ef44358-db5b-4d04-a08a-0d44711ce040</mods:identifier>
<mods:identifier type="ISSN">1854-0392</mods:identifier>
<mods:identifier type="Zenodo-Dep">4760806</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4763693" ID-GBIF-Taxon="182074635" ID-Zenodo-Dep="4763693" LSID="urn:lsid:plazi:treatment:03B087FC5243EB008C83EF9EE37B5FD7" httpUri="http://treatment.plazi.org/id/03B087FC5243EB008C83EF9EE37B5FD7" lastPageId="6" lastPageNumber="110" pageId="0" pageNumber="104">
<subSubSection box="[1007,1271,1785,1810]" pageId="0" pageNumber="104" type="nomenclature">
<paragraph blockId="0.[1007,1271,1785,1810]" box="[1007,1271,1785,1810]" pageId="0" pageNumber="104">
<heading bold="true" box="[1007,1271,1785,1810]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="104" reason="2">
<emphasis bold="true" box="[1007,1271,1785,1810]" pageId="0" pageNumber="104">
<taxonomicName authority="Baumann &amp; Call, 2012" authorityName="Baumann &amp; Call" authorityYear="2012" box="[1007,1192,1785,1810]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="104" phylum="Arthropoda" rank="species" species="tetonica" status="sp. n.">
<emphasis bold="true" box="[1007,1192,1785,1810]" italics="true" pageId="0" pageNumber="104">Lednia tetonica</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1206,1271,1786,1810]" pageId="0" pageNumber="104" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[1081,1197,1821,1846]" pageId="0" pageNumber="104" type="description">
<paragraph blockId="0.[1081,1197,1821,1846]" box="[1081,1197,1821,1846]" pageId="0" pageNumber="104">
(
<figureCitation box="[1089,1189,1821,1846]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="1.[189,230,1468,1492]" captionStartId-1="2.[189,241,1788,1812]" captionTargetBox-0="[261,714,657,1422]" captionTargetBox-1="[268,1364,243,1740]" captionTargetId-0="figure-34@1.[226,755,618,1478]" captionTargetId-1="figure-34@2.[268,1364,243,1740]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="Fig. 1. Lednia tetonica male, epiproct, dorsal." captionText-1="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi-0="http://doi.org/10.5281/zenodo.4760808" figureDoi-1="http://doi.org/10.5281/zenodo.4760810" httpUri-0="https://zenodo.org/record/4760808/files/figure.png" httpUri-1="https://zenodo.org/record/4760810/files/figure.png" pageId="0" pageNumber="104">Figs. 1-9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="104" type="reference_group">
<paragraph blockId="0.[873,1433,1878,1933]" pageId="0" pageNumber="104">
<treatmentCitationGroup pageId="0" pageNumber="104">
<treatmentCitation ID-DOI="http://doi.org/10.5281/zenodo.4756959" ID-Zenodo-Dep="4756959" author="Baumann, R. W. &amp; B. C. Kondratieff" box="[873,1430,1878,1900]" httpUri="http://treatment.plazi.org/id/038E1501FFAF9611FC06F914FB234AE5" page="316" pageId="0" pageNumber="104" year="2010">
<taxonomicName authority=": Baumann and Kondratieff, 2010: 316" authorityName=": Baumann and Kondratieff" authorityPageNumber="316" authorityYear="2010" box="[873,1430,1878,1900]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="0" pageNumber="104" phylum="Arthropoda" rank="species" species="tumana">
<emphasis box="[873,1019,1878,1900]" italics="true" pageId="0" pageNumber="104">Lednia tumana</emphasis>
: 
<bibRefCitation author="Baumann, R. W. &amp; B. C. Kondratieff" box="[1035,1430,1878,1900]" pageId="0" pageNumber="104" pagination="315 - 327" refId="ref2643" refString="Baumann, R. W. and B. C. Kondratieff. 2010. The stonefly genus Lednia in North America (Plecoptera: Nemouridae). Illiesia, 6: 315 - 327." type="journal article" year="2010">Baumann and Kondratieff, 2010:316</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. (Distribution in part).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="105" type="materials_examined">
<paragraph blockId="1.[189,798,224,569]" pageId="1" pageNumber="105">
<emphasis bold="true" box="[189,429,225,249]" pageId="1" pageNumber="105">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111647302" collectingDate="2011-08-27" collectorName="R. G. Call" country="United States of America" county="Teton County" location="Darby Creek" municipality="South Fork" pageId="1" pageNumber="105" specimenCount="1" specimenCount-male="1" stateProvince="Wyoming" typeStatus="holotype">
<typeStatus box="[441,553,225,249]" pageId="1" pageNumber="105">Holotype</typeStatus>
<specimenCount box="[563,583,224,249]" pageId="1" pageNumber="105" type="male">♂</specimenCount>
, 
<collectingCountry box="[600,659,225,249]" name="United States of America" pageId="1" pageNumber="105">USA</collectingCountry>
: 
<collectingRegion box="[673,793,225,249]" country="United States of America" name="Wyoming" pageId="1" pageNumber="105">Wyoming</collectingRegion>
, 
<collectingCounty box="[189,353,261,285]" pageId="1" pageNumber="105">Teton County</collectingCounty>
, 
<collectingMunicipality box="[362,493,261,285]" pageId="1" pageNumber="105">South Fork</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:03B087FC5243EB008C83EF9EE37B5FD7:8EC660315242EB078E9BE862E3DA5863" box="[503,653,261,285]" country="United States of America" county="Teton County" municipality="South Fork" name="Darby Creek" pageId="1" pageNumber="105" stateProvince="Wyoming">Darby Creek</location>
, 
<location LSID="urn:lsid:plazi:treatment:03B087FC5243EB008C83EF9EE37B5FD7:8EC660315242EB078DFBE862E24F5863" box="[663,792,261,285]" country="United States of America" county="Teton County" municipality="South Fork" name="Wind Cave" pageId="1" pageNumber="105" stateProvince="Wyoming">Wind Cave</location>
, 
<locationDeviation box="[189,417,296,320]" pageId="1" pageNumber="105">southeast of Driggs</locationDeviation>
, Idaho, 
<date box="[504,681,296,320]" pageId="1" pageNumber="105" value="2011-08-27">
<collectingDate box="[504,681,296,320]" pageId="1" pageNumber="105" value="2011-08-27">27 August 2011</collectingDate>
</date>
, 
<collectorName box="[689,794,296,320]" pageId="1" pageNumber="105">R.G. Call</collectorName>
</materialsCitation>
. 
<materialsCitation ID-GBIF-Occurrence="3111647301" collectingDate="2008-08-14" collectingDateMax="2011-08-27" collectingDateMin="2008-08-14" collectionCode="BYUC, RCPC, CSUC" collectorName="R. G. Call" country="United States of America" location="Same" pageId="1" pageNumber="105" specimenCount="159" specimenCount-female="128" specimenCount-larva="9" specimenCount-male="22" stateProvince="Washington" typeStatus="paratype">
<typeStatus box="[189,308,332,356]" pageId="1" pageNumber="105">Paratypes</typeStatus>
: 
<location LSID="urn:lsid:plazi:treatment:03B087FC5243EB008C83EF9EE37B5FD7:8EC660315242EB078E2EE82BE0D6581A" box="[322,385,332,356]" country="United States of America" name="Same" pageId="1" pageNumber="105" stateProvince="Washington">Same</location>
locality and collector, 
<date pageId="1" pageNumber="105" value="2008-08-14">
<collectingDate pageId="1" pageNumber="105" value="2008-08-14">14 August 2008</collectingDate>
</date>
, 
<specimenCount box="[255,288,366,391]" pageId="1" pageNumber="105" type="female">1♀</specimenCount>
(
<collectionCode box="[301,380,367,391]" pageId="1" pageNumber="105">BYUC</collectionCode>
); 
<date box="[396,537,367,391]" pageId="1" pageNumber="105" value="2009-07-11">
<collectingDate box="[396,537,367,391]" pageId="1" pageNumber="105" value="2009-07-11">11 July 2009</collectingDate>
</date>
, 
<specimenCount box="[547,580,366,391]" pageId="1" pageNumber="105" type="female">2♀</specimenCount>
(
<collectionCode box="[593,671,367,391]" pageId="1" pageNumber="105">BYUC</collectionCode>
, 
<collectionCode box="[680,755,367,391]" pageId="1" pageNumber="105">RCPC</collectionCode>
); 
<date pageId="1" pageNumber="105" value="2011-08-27">
<collectingDate pageId="1" pageNumber="105" value="2011-08-27">27 August 2011</collectingDate>
</date>
, 
<specimenCount box="[368,415,402,427]" pageId="1" pageNumber="105" type="male">22♂</specimenCount>
, 
<specimenCount box="[436,497,402,427]" pageId="1" pageNumber="105" type="female">125♀</specimenCount>
(
<collectionCode box="[519,597,403,427]" pageId="1" pageNumber="105">BYUC</collectionCode>
, 
<collectionCode box="[616,691,403,427]" country="0" httpUri="http://grbio.org/cool/je3k-bvrg" name="California State University, Chico, Vertebrate Museum" pageId="1" pageNumber="105" type="University or college">CSUC</collectionCode>
, 
<collectionCode box="[710,786,403,427]" pageId="1" pageNumber="105">RCPC</collectionCode>
).
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3111647303" collectingDate="2008-08-14" collectingDateMax="2011-08-27" collectingDateMin="2008-08-14" collectionCode="BYUC, RCPC, CSUC" collectorName="R. G. Call" country="United States of America" location="Same" pageId="1" pageNumber="105" specimenCount="159" specimenCount-female="128" specimenCount-larva="9" specimenCount-male="22" stateProvince="Washington" typeStatus="paratype">
Larvae: Same locality and collector, 
<date box="[625,771,438,462]" pageId="1" pageNumber="105" value="2010-07-26">
<collectingDate box="[625,771,438,462]" pageId="1" pageNumber="105" value="2010-07-26">26 July 2010</collectingDate>
</date>
, 
<specimenCount pageId="1" pageNumber="105" type="larva">9 larvae</specimenCount>
(
<collectionCode box="[292,370,474,498]" pageId="1" pageNumber="105">BYUC</collectionCode>
, 
<collectionCode box="[396,472,474,498]" pageId="1" pageNumber="105">RCPC</collectionCode>
).
</materialsCitation>
<collectorName box="[506,550,474,498]" pageId="1" pageNumber="105">The</collectorName>
<typeStatus box="[573,678,474,498]" pageId="1" pageNumber="105">holotype</typeStatus>
will be deposited at the 
<collectingCountry box="[399,560,509,533]" name="United States of America" pageId="1" pageNumber="105">United States</collectingCountry>
<collectorName box="[571,792,509,533]" pageId="1" pageNumber="105">National Museum</collectorName>
, 
<collectorName box="[189,469,545,569]" pageId="1" pageNumber="105">Smithsonian Institution</collectorName>
, 
<collectingRegion box="[480,625,545,569]" country="United States of America" name="Washington" pageId="1" pageNumber="105">Washington</collectingRegion>
, D. C.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760808" ID-Zenodo-Dep="4760808" box="[189,708,1468,1493]" httpUri="https://zenodo.org/record/4760808/files/figure.png" pageId="1" pageNumber="105" startId="1.[189,230,1468,1492]" targetBox="[261,714,657,1422]" targetPageId="1">
<paragraph blockId="1.[189,708,1468,1493]" box="[189,708,1468,1493]" pageId="1" pageNumber="105">
Fig. 1. 
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" box="[267,431,1468,1492]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="105" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis box="[267,431,1468,1492]" italics="true" pageId="1" pageNumber="105">Lednia tetonica</emphasis>
</taxonomicName>
male, epiproct, dorsal.
</paragraph>
</caption>
<subSubSection pageId="1" pageNumber="105" type="description">
<paragraph blockId="1.[189,797,1575,1919]" lastBlockId="1.[835,1444,225,1919]" pageId="1" pageNumber="105">
<emphasis bold="true" box="[189,258,1575,1599]" pageId="1" pageNumber="105">Male.</emphasis>
Macropterous, wings hyaline, with darker veins near cord. Length of body 4.0-5.0 mm, color dark brown, anterior abdominal segments lightly sclerotized and lighter in color. Legs dark brown. Epiproct bilaterally symmetrical, with deep, median dorsal groove (
<figureCitation box="[370,435,1753,1777]" captionStart="Fig" captionStartId="1.[189,230,1468,1492]" captionTargetBox="[261,714,657,1422]" captionTargetId="figure-34@1.[226,755,618,1478]" captionTargetPageId="1" captionText="Fig. 1. Lednia tetonica male, epiproct, dorsal." figureDoi="http://doi.org/10.5281/zenodo.4760808" httpUri="https://zenodo.org/record/4760808/files/figure.png" pageId="1" pageNumber="105">Fig. 1</figureCitation>
); ventral sclerite thin, flat and darkly sclerotized, bearing scattered stout spines ventrally in apical third, basal-lateral margins shaped like small fans, apex extending beyond apical margin of dorsal sclerite, tip bluntly rounded with short triangular lateral points, appearing as a blunted arrowhead (
<figureCitation box="[980,1116,260,284]" captionStart="Figs" captionStartId="2.[189,241,1788,1812]" captionTargetBox="[268,1364,243,1740]" captionTargetId="figure-34@2.[268,1364,243,1740]" captionTargetPageId="2" captionText="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi="http://doi.org/10.5281/zenodo.4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="1" pageNumber="105">Figs. 3, 5, 7</figureCitation>
); dorsal sclerite with broad bare base, terminating in paired rounded lobes, sclerite narrowing gradually towards apex, anterior dorsal surface bearing paired lateral patches of large backward directed spines, containing from 50-60 spines per patch, spine patches elongate and covering more than a third of the length of the sclerotized lateral margins, spines variable in size with longest spines located medially, few tiny hairlike spines present at anterior base of large spines (
<figureCitation box="[843,996,615,640]" captionStart="Figs" captionStartId="2.[189,241,1788,1812]" captionTargetBox="[268,1364,243,1740]" captionTargetId="figure-34@2.[268,1364,243,1740]" captionTargetPageId="2" captionText="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi="http://doi.org/10.5281/zenodo.4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="1" pageNumber="105">Figs. 2, 4, 5, 7</figureCitation>
). Hypoproct sclerotized, broad at base, tapering towards narrow apex, vesicle absent (
<figureCitation box="[1366,1429,651,675]" captionStart="Figs" captionStartId="2.[189,241,1788,1812]" captionTargetBox="[268,1364,243,1740]" captionTargetId="figure-34@2.[268,1364,243,1740]" captionTargetPageId="2" captionText="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi="http://doi.org/10.5281/zenodo.4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="1" pageNumber="105">Fig. 8</figureCitation>
). Paraprocts composed of two lobes, outer lobe broad and short, covered with numerous short hairs, inner lobes long and thin, curving outward along base of epiproct, tip sharply pointed (
<figureCitation box="[1185,1287,793,817]" captionStart="Figs" captionStartId="2.[189,241,1788,1812]" captionTargetBox="[268,1364,243,1740]" captionTargetId="figure-34@2.[268,1364,243,1740]" captionTargetPageId="2" captionText="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi="http://doi.org/10.5281/zenodo.4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="1" pageNumber="105">Figs. 6, 8</figureCitation>
). Tergum ten with broad dished out dorsal area nearly naked of spines, bearing large paired lateral projections, covered with short stout spines and numerous sensory pits, tips appearing as large mace-like structures, which border the apex of the epiproct (
<figureCitation box="[843,949,1006,1030]" captionStart="Figs" captionStartId="2.[189,241,1788,1812]" captionTargetBox="[268,1364,243,1740]" captionTargetId="figure-34@2.[268,1364,243,1740]" captionTargetPageId="2" captionText="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi="http://doi.org/10.5281/zenodo.4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="1" pageNumber="105">Figs. 3, 7</figureCitation>
).
</paragraph>
<paragraph blockId="1.[835,1444,225,1919]" pageId="1" pageNumber="105">
<emphasis bold="true" box="[835,929,1042,1066]" pageId="1" pageNumber="105">Female.</emphasis>
General color of body and wings similar to male. Length 5.0-6.0 mm. Sternum seven with broadly rounded lightly sclerotized lobe posteriormedially. Subgenital plate covering most of sternum eight, darkly sclerotized, bearing numerous long hairs medially, base broad, tapering to narrow rounded apex, which is often divided medially by a thin groove, tips of apex covered with short stout spines (
<figureCitation box="[926,986,1326,1350]" captionStart="Figs" captionStartId="2.[189,241,1788,1812]" captionTargetBox="[268,1364,243,1740]" captionTargetId="figure-34@2.[268,1364,243,1740]" captionTargetPageId="2" captionText="Figs. 2-9. Lednia tetonica adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10th tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10th tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia." figureDoi="http://doi.org/10.5281/zenodo.4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="1" pageNumber="105">Fig.9</figureCitation>
). Lateral posterior margins of sternum eight formed into large nearly rounded sclerotized lobes, which border the apex of the subgenital plate.
</paragraph>
<paragraph blockId="1.[835,1444,225,1919]" pageId="1" pageNumber="105">
<emphasis bold="true" box="[835,910,1433,1457]" pageId="1" pageNumber="105">Larva.</emphasis>
Gills absent. Length: mature male 4.5-5.5 mm, mature female 5.0-6.0 mm. Dorsal surface of body nearly naked of spines except at anterior margins of thoracic segments, abdominal terga with few spines except along posterior margin of terminal segment. Legs with few dorsal spines, occasional larger spines present on anterior margin of segments. Female larvae exhibiting large truncate lobe on dorsomedial margin of sternum eight. Cerci with whorls of short stout spines at posterior margins of segments, few intercalary spines present, most numerous on basal segments.
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="107" pageId="1" pageNumber="105" type="etymology">
<paragraph blockId="1.[835,1444,225,1919]" lastBlockId="3.[189,798,225,1670]" lastPageId="3" lastPageNumber="107" pageId="1" pageNumber="105">
<emphasis bold="true" box="[835,973,1860,1884]" pageId="1" pageNumber="105">Etymology.</emphasis>
The species name 
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" box="[1201,1316,1860,1883]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="1" pageNumber="105" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis box="[1201,1316,1860,1883]" italics="true" pageId="1" pageNumber="105">L. tetonica</emphasis>
</taxonomicName>
is derived from the fact that the specimens were collected from a tributary of the Teton River, located in the Teton Mountains, Teton County, 
<collectingRegion box="[506,627,260,284]" country="United States of America" name="Wyoming" pageId="3" pageNumber="107">Wyoming</collectingRegion>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4760810" ID-Zenodo-Dep="4760810" httpUri="https://zenodo.org/record/4760810/files/figure.png" pageId="2" pageNumber="106" startId="2.[189,241,1788,1812]" targetBox="[268,1364,243,1740]" targetPageId="2">
<paragraph blockId="2.[189,1443,1787,1918]" pageId="2" pageNumber="106">
Figs. 2-9. 
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" box="[302,466,1788,1812]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="2" pageNumber="106" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis box="[302,466,1788,1812]" italics="true" pageId="2" pageNumber="106">Lednia tetonica</emphasis>
</taxonomicName>
adult terminalia: 2. Male, epiproct, dorsal. 3. Male, epiproct, lateral and 10 
<superScript attach="left" box="[1354,1369,1787,1801]" fontSize="6" pageId="2" pageNumber="106">th</superScript>
tergal prongs. 4. Male, epiproct, dorsal, close up. 5. Male, epiproct, anterior. 6. Male, terminalia, posterior, relaxed. 7. Male, epiproct, dorsolateral and 10 
<superScript attach="left" box="[617,632,1858,1872]" fontSize="6" pageId="2" pageNumber="106">th</superScript>
tergal prong. 8. Male, terminalia, posterior, contracted. 9. Female, subgenital plate and terminalia.
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="107" type="diagnosis">
<paragraph blockId="3.[189,798,225,1670]" pageId="3" pageNumber="107">
<emphasis bold="true" box="[189,319,296,320]" pageId="3" pageNumber="107">Diagnosis.</emphasis>
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" box="[324,486,296,320]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis box="[324,486,296,320]" italics="true" pageId="3" pageNumber="107">Lednia tetonica</emphasis>
</taxonomicName>
is most similar to 
<taxonomicName authorityName=": Baumann and Kondratieff" authorityYear="2010" box="[691,797,297,320]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tumana">
<emphasis box="[691,797,297,320]" italics="true" pageId="3" pageNumber="107">L. tumana</emphasis>
</taxonomicName>
and 
<taxonomicName authority="Baumann and Kondratieff. It" authorityName="Baumann and Kondratieff. It" box="[246,708,332,356]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sierra">
<emphasis box="[246,340,332,355]" italics="true" pageId="3" pageNumber="107">L. sierra</emphasis>
Baumann and Kondratieff. It
</taxonomicName>
differs mostly in the details of the epiproct. There are 70-80 spines in the spine patches on the dorsal surface of the epiproct in 
<taxonomicName authorityName=": Baumann and Kondratieff" authorityYear="2010" box="[366,477,439,462]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tumana">
<emphasis box="[366,477,439,462]" italics="true" pageId="3" pageNumber="107">L. tumana</emphasis>
</taxonomicName>
. The spine patch number is 50-60 spines in 
<emphasis box="[378,500,474,497]" italics="true" pageId="3" pageNumber="107">
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" box="[378,496,474,497]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tetonica">L. tetonica</taxonomicName>
.
</emphasis>
There are fewer than 40 spines in the patches of 
<emphasis box="[479,576,510,533]" italics="true" pageId="3" pageNumber="107">
<taxonomicName box="[479,572,510,533]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sierra">L. sierra</taxonomicName>
.
</emphasis>
The spine patches are also found on different areas of the epiproct and they are shaped differently. 
<taxonomicName authorityName=": Baumann and Kondratieff" authorityYear="2010" box="[535,700,580,604]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tumana">
<emphasis box="[535,700,580,604]" italics="true" pageId="3" pageNumber="107">Lednia tumana</emphasis>
</taxonomicName>
has the spines located only on the anterior half of the epiproct and they are confined to elongate oval patches, the spines are found further back toward the base of the epiproct in the other two species. 
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis italics="true" pageId="3" pageNumber="107">Lednia tetonica</emphasis>
</taxonomicName>
has most of the spines attached near the middle of the epiproct, with only a few located on the lateral margins, which extend toward the epiproct base. 
<taxonomicName box="[356,494,864,888]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sierra">
<emphasis box="[356,494,864,888]" italics="true" pageId="3" pageNumber="107">Lednia sierra</emphasis>
</taxonomicName>
has fewer spines medially while most of the spines are on the margins and extend well back toward the base. The spines are smaller and all about the same size in 
<taxonomicName authorityName=": Baumann and Kondratieff" authorityYear="2010" box="[675,791,972,995]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tumana">
<emphasis box="[675,791,972,995]" italics="true" pageId="3" pageNumber="107">L. tumana</emphasis>
</taxonomicName>
, while they vary in size in the other species. A patch of large spines is located close to the midline in 
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis italics="true" pageId="3" pageNumber="107">L. tetonica</emphasis>
</taxonomicName>
with fewer small spines covering the edges. In 
<taxonomicName box="[235,328,1114,1137]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sierra">
<emphasis box="[235,250,1114,1137]" italics="true" pageId="3" pageNumber="107">L</emphasis>
. 
<emphasis box="[268,328,1114,1137]" italics="true" pageId="3" pageNumber="107">sierra</emphasis>
</taxonomicName>
, there are a few large spines near the midline but many smaller spines extend along the midline from the middle to near the base of the epiproct. The shape of the epiproct also differs between species. Dorsally, the epiproct in 
<taxonomicName authorityName=": Baumann and Kondratieff" authorityYear="2010" box="[686,797,1256,1279]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tumana">
<emphasis box="[686,797,1256,1279]" italics="true" pageId="3" pageNumber="107">L. tumana</emphasis>
</taxonomicName>
is stouter in shape with a shorter apex. Both 
<taxonomicName authorityName="Baumann &amp; Call" authorityYear="2012" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="tetonica">
<emphasis italics="true" pageId="3" pageNumber="107">L. tetonica</emphasis>
</taxonomicName>
and 
<taxonomicName box="[333,422,1327,1350]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sierra">
<emphasis box="[333,422,1327,1350]" italics="true" pageId="3" pageNumber="107">L. sierra</emphasis>
</taxonomicName>
exhibit a long, narrow epiproct with an elongate tip which is more deeply grooved in 
<emphasis box="[189,285,1398,1421]" italics="true" pageId="3" pageNumber="107">
<taxonomicName box="[189,282,1398,1421]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sierra">L. sierra</taxonomicName>
.
</emphasis>
The females are very similar and cannot be separated at this time without associated males.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="107" type="discussion">
<paragraph blockId="3.[189,798,225,1670]" pageId="3" pageNumber="107">
<emphasis bold="true" box="[189,304,1469,1493]" pageId="3" pageNumber="107">Remarks.</emphasis>
The South Fork of Darby Creek is an excellent stonefly habitat and the following species were found between the cave and the confluence with Darby Creek. Collections were made during the summer months of June-August, while searching for the elusive 
<emphasis box="[323,403,1646,1670]" italics="true" pageId="3" pageNumber="107">
<taxonomicName box="[323,399,1646,1670]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="genus">Lednia</taxonomicName>
.
</emphasis>
</paragraph>
<paragraph blockId="3.[341,767,1718,1919]" box="[341,616,1718,1742]" pageId="3" pageNumber="107">
<taxonomicName authority="(Banks)" baseAuthorityName="Banks" baseAuthorityYear="1897" box="[341,616,1718,1742]" class="Insecta" family="Nemouridae" genus="Zapada" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis box="[341,521,1718,1742]" italics="true" pageId="3" pageNumber="107">Zapada cinctipes</emphasis>
(Banks)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[341,767,1718,1919]" box="[341,662,1753,1777]" pageId="3" pageNumber="107">
<taxonomicName authority="Claassen" authorityName="Claassen" box="[341,662,1753,1777]" class="Insecta" family="Nemouridae" genus="Zapada" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="columbiana">
<emphasis box="[341,551,1753,1777]" italics="true" pageId="3" pageNumber="107">Zapada columbiana</emphasis>
Claassen
</taxonomicName>
</paragraph>
<paragraph blockId="3.[341,767,1718,1919]" box="[341,582,1789,1813]" pageId="3" pageNumber="107">
<taxonomicName authority="(Ricker)" baseAuthorityName="Ricker" baseAuthorityYear="1952" box="[341,582,1789,1813]" class="Insecta" family="Nemouridae" genus="Zapada" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="haysi">
<emphasis box="[341,483,1789,1813]" italics="true" pageId="3" pageNumber="107">Zapada haysi</emphasis>
(Ricker)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[341,767,1718,1919]" box="[341,588,1824,1848]" pageId="3" pageNumber="107">
<taxonomicName authority="Claassen" authorityName="Claassen" authorityYear="1924" box="[341,588,1824,1848]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="nana">
<emphasis box="[341,477,1825,1848]" italics="true" pageId="3" pageNumber="107">Capnia nana</emphasis>
Claassen
</taxonomicName>
</paragraph>
<paragraph blockId="3.[341,767,1718,1919]" box="[341,564,1860,1884]" pageId="3" pageNumber="107">
<taxonomicName authority="Jewett" authorityName="Jewett" authorityYear="1954" box="[341,564,1860,1884]" class="Insecta" family="Capniidae" genus="Capnia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="petila">
<emphasis box="[341,483,1860,1884]" italics="true" pageId="3" pageNumber="107">Capnia petila</emphasis>
Jewett
</taxonomicName>
</paragraph>
<paragraph blockId="3.[341,767,1718,1919]" box="[341,767,1895,1919]" pageId="3" pageNumber="107">
<taxonomicName authority="Nebeker &amp; Gaufin" authorityName="Nebeker &amp; Gaufin" authorityYear="1966" box="[341,767,1895,1919]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="jewetti">
<emphasis box="[341,544,1895,1919]" italics="true" pageId="3" pageNumber="107">Paraleuctra jewetti</emphasis>
Nebeker &amp; Gaufin
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1301,225,249]" pageId="3" pageNumber="107">
<taxonomicName authority="(Frison)" baseAuthorityName="Frison" baseAuthorityYear="1942" box="[986,1301,225,249]" class="Insecta" family="Leuctridae" genus="Paraleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="projecta">
<emphasis box="[986,1203,225,249]" italics="true" pageId="3" pageNumber="107">Paraleuctra projecta</emphasis>
(Frison)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1257,260,284]" pageId="3" pageNumber="107">
<taxonomicName authority="Banks" authorityName="Banks" authorityYear="1906" box="[986,1257,260,284]" class="Insecta" family="Leuctridae" genus="Perlomyia" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="collaris">
<emphasis box="[986,1179,260,284]" italics="true" pageId="3" pageNumber="107">Perlomyia collaris</emphasis>
Banks
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1341,296,320]" pageId="3" pageNumber="107">
<taxonomicName authority="(Neave)" baseAuthorityName="Neave" baseAuthorityYear="1934" box="[986,1341,296,320]" class="Insecta" family="Leuctridae" genus="Pomoleuctra" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="purcellana">
<emphasis box="[986,1241,296,320]" italics="true" pageId="3" pageNumber="107">Pomoleuctra purcellana</emphasis>
(Neave)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1244,331,355]" pageId="3" pageNumber="107">
<taxonomicName authority="(Hagen)" baseAuthorityName="Hagen" baseAuthorityYear="1874" box="[986,1244,331,355]" class="Insecta" family="Perlodidae" genus="Isoperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="sobria">
<emphasis box="[986,1141,331,355]" italics="true" pageId="3" pageNumber="107">Isoperla sobria</emphasis>
(Hagen)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1279,367,391]" pageId="3" pageNumber="107">
<taxonomicName authority="(Hagen)" baseAuthorityName="Hagen" baseAuthorityYear="1874" box="[986,1279,367,391]" class="Insecta" family="Perlodidae" genus="Megarcys" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="signata">
<emphasis box="[986,1177,368,391]" italics="true" pageId="3" pageNumber="107">Megarcys signata</emphasis>
(Hagen)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" pageId="3" pageNumber="107">
<taxonomicName authority="Needham &amp; Claassen" authorityName="Needham &amp; Claassen" authorityYear="1925" box="[986,1426,402,426]" class="Insecta" family="Chloroperlidae" genus="Alloperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="serrata">
<emphasis box="[986,1166,402,426]" italics="true" pageId="3" pageNumber="107">Alloperla serrata</emphasis>
Needham &amp; Claassen
</taxonomicName>
<taxonomicName authority="(Frison)" baseAuthorityName="Frison" baseAuthorityYear="1935" box="[986,1288,438,462]" class="Insecta" family="Chloroperlidae" genus="Plumiperla" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="diversa">
<emphasis box="[986,1190,438,462]" italics="true" pageId="3" pageNumber="107">Plumiperla diversa</emphasis>
(Frison)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1252,473,497]" pageId="3" pageNumber="107">
<taxonomicName authority="(Banks)" baseAuthorityName="Banks" baseAuthorityYear="1895" box="[986,1252,473,497]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="borealis">
<emphasis box="[986,1157,473,497]" italics="true" pageId="3" pageNumber="107">Sweltsa borealis</emphasis>
(Banks)
</taxonomicName>
</paragraph>
<paragraph blockId="3.[986,1426,225,533]" box="[986,1235,509,533]" pageId="3" pageNumber="107">
<taxonomicName authority="(Banks)" baseAuthorityName="Banks" baseAuthorityYear="1920" box="[986,1235,509,533]" class="Insecta" family="Chloroperlidae" genus="Sweltsa" kingdom="Animalia" order="Plecoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="fidelis">
<emphasis box="[986,1139,509,533]" italics="true" pageId="3" pageNumber="107">Sweltsa fidelis</emphasis>
(Banks)
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="110" pageId="3" pageNumber="107" type="discussion">
<paragraph blockId="3.[835,1444,580,1919]" lastBlockId="6.[189,798,225,1705]" lastPageId="6" lastPageNumber="110" pageId="3" pageNumber="107">
<emphasis bold="true" box="[835,976,580,604]" pageId="3" pageNumber="107">Discussion.</emphasis>
The Teton Mountain Range lies along the border of 
<collectingRegion box="[1015,1132,616,640]" country="United States of America" name="Wyoming" pageId="3" pageNumber="107">Wyoming</collectingRegion>
and 
<collectingRegion box="[1204,1273,616,640]" country="United States of America" name="Idaho" pageId="3" pageNumber="107">Idaho</collectingRegion>
southwest of Grand Teton and Yellowstone National Parks. The Teton Mountains are geologically young, beginning to form about ten million years ago (
<bibRefCitation author="Crammerer, A. B." pageId="3" pageNumber="107" refId="ref2712" refString="Crammerer, A. B. 1937. Grand Teton [Wyoming] National Park. United States Government Printing Office, Washington, D. C., 33 pp." type="book" year="1937">Crammerer 1937</bibRefCitation>
). Varied geological forces including earthquakes, uplifts and volcanism from the Yellowstone Hotspot, and glacial advances and retreats have made the Tetons a unique area (
<bibRefCitation author="Smith, R. B. &amp; L. J. Siegel" box="[1110,1386,865,889]" pageId="3" pageNumber="107" refId="ref2805" refString="Smith, R. B. and L. J. Siegel. 2000. Windows into the Earth: The geologic story of Yellowstone and Grand Teton National Parks. Oxford University Press, 256 pp." type="book" year="2000">Smith and Siegel 2000</bibRefCitation>
). Six peaks reach an elevation greater than 
<quantity box="[1313,1442,900,924]" metricMagnitude="0" metricUnit="m" metricValue="3.6576" pageId="3" pageNumber="107" unit="ft" value="12.0">12,000 feet</quantity>
(
<quantity box="[842,936,936,960]" metricMagnitude="0" metricUnit="m" metricValue="3.657" pageId="3" pageNumber="107" unit="m" value="3.657">3,657 m</quantity>
), the highest of which is the Grand Teton at 
<quantity box="[835,959,971,996]" metricMagnitude="0" metricUnit="m" metricValue="4.197096" pageId="3" pageNumber="107" unit="ft" value="13.77">13,770 feet</quantity>
(
<quantity box="[974,1070,972,996]" metricMagnitude="0" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="107" unit="m" value="4.2">4,200 m</quantity>
). These mountains are popular for mountain climbing, back packing, hiking, horseback riding, camping, and skiing. Darby Canyon and the Wind Cave are within the Jedediah Smith Wilderness Area of the Caribou-Targhee National Forest, along the western edge of the Teton Range. Although Darby Canyon is located in 
<collectingRegion box="[835,955,1220,1244]" country="United States of America" name="Wyoming" pageId="3" pageNumber="107">Wyoming</collectingRegion>
, the only road to access the canyon is from 
<collectingRegion box="[835,903,1256,1280]" country="United States of America" name="Idaho" pageId="3" pageNumber="107">Idaho</collectingRegion>
on the western side of the mountains. Geologically, Darby Canyon cuts through the Lodgepole formation, part of the Madison Limestone formation. The rocks are from the early Mississippian period. Fossils of corals and other marine organisms occur throughout the canyon. The forest is primarily lodgepole pine with other conifers, some aspen, and mixed deciduous trees. Wildflowers of many species are abundant. Wind Cave is at the end of Darby Canyon at an elevation of about 
<quantity box="[1217,1328,1575,1599]" metricMagnitude="0" metricUnit="m" metricValue="2.7432" pageId="3" pageNumber="107" unit="ft" value="9.0">9,000 feet</quantity>
(
<quantity box="[1342,1430,1575,1599]" metricMagnitude="3" metricUnit="m" metricValue="2.743" pageId="3" pageNumber="107" unit="m" value="2743.0">2743 m</quantity>
). The opening to the Wind Cave is about 
<quantity box="[1309,1399,1611,1635]" metricMagnitude="1" metricUnit="m" metricValue="3.6576" pageId="3" pageNumber="107" unit="ft" value="120.0">120 feet</quantity>
(
<quantity metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="107" unit="m" value="36.0">36 m</quantity>
) high by about 
<quantity box="[1045,1122,1646,1671]" metricMagnitude="1" metricUnit="m" metricValue="1.2192" pageId="3" pageNumber="107" unit="ft" value="40.0">40 feet</quantity>
(12 meters) wide. The rock walls and floor are jagged chunks of limestone that have fallen from the ceiling. Water percolates down from the melting snowpack on the top of the mountain giving rise to a stream that flows from the mouth of the cave. Within 15 or 
<quantity box="[1233,1312,1824,1848]" metricMagnitude="0" metricUnit="m" metricValue="6.096" pageId="3" pageNumber="107" unit="ft" value="20.0">20 feet</quantity>
(5 or 
<quantity box="[1388,1435,1824,1848]" metricMagnitude="0" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="107" unit="m" value="6.0">6 m</quantity>
) inside the cave, the stream no longer flows over the tops of the rocks, but can be heard flowing beneath the rocks, as an underground stream. Deep inside the cave, the temperature is low enough to maintain year-round ice. The cave connects to another opening about one-half mile (0.8 km) away on the same side of the mountain called the Ice Cave. About 
<quantity box="[706,797,367,391]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="6" pageNumber="110" unit="ft" value="300.0">300 feet</quantity>
(
<quantity box="[197,261,402,426]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="110" unit="m" value="90.0">90 m</quantity>
) downstream from Wind Cave, the stream drops over a waterfall about 
<quantity box="[525,600,438,462]" metricMagnitude="1" metricUnit="m" metricValue="2.1336" pageId="6" pageNumber="110" unit="ft" value="70.0">70 feet</quantity>
(
<quantity box="[614,672,438,462]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="110" unit="m" value="20.0">20 m</quantity>
) high then continues to flow down Darby Canyon as part of the South Fork of Darby Creek.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4765817" ID-Zenodo-Dep="4765817" box="[189,534,971,995]" httpUri="https://zenodo.org/record/4765817/files/figure.png" pageId="4" pageNumber="108" startId="4.[189,230,971,995]" targetBox="[287,1345,227,924]" targetPageId="4">
<paragraph blockId="4.[189,534,971,995]" box="[189,534,971,995]" pageId="4" pageNumber="108">Fig. 10. Wind Cave in winter.</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4765819" ID-Zenodo-Dep="4765819" box="[189,666,1894,1919]" httpUri="https://zenodo.org/record/4765819/files/figure.png" pageId="4" pageNumber="108" startId="4.[189,230,1894,1918]" targetBox="[287,1345,1058,1848]" targetPageId="4">
<paragraph blockId="4.[189,666,1894,1919]" box="[189,666,1894,1919]" pageId="4" pageNumber="108">Fig. 11. Wind Cave Valley and waterfall.</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4765821" ID-Zenodo-Dep="4765821" box="[189,554,1006,1030]" httpUri="https://zenodo.org/record/4765821/files/figure.png" pageId="5" pageNumber="109" startId="5.[189,230,1006,1030]" targetBox="[306,1293,224,965]" targetPageId="5">
<paragraph blockId="5.[189,554,1006,1030]" box="[189,554,1006,1030]" pageId="5" pageNumber="109">Fig. 12. Wind Cave in summer.</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4765823" ID-Zenodo-Dep="4765823" box="[189,533,1894,1919]" httpUri="https://zenodo.org/record/4765823/files/figure.png" pageId="5" pageNumber="109" startId="5.[189,230,1894,1918]" targetBox="[305,1292,1102,1842]" targetPageId="5">
<paragraph blockId="5.[189,533,1894,1919]" box="[189,533,1894,1919]" pageId="5" pageNumber="109">Fig. 13. Wind Cave at mouth.</paragraph>
</caption>
<paragraph blockId="6.[189,798,225,1705]" pageId="6" pageNumber="110">
The first 
<taxonomicName box="[329,401,544,568]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis box="[329,401,544,568]" italics="true" pageId="6" pageNumber="110">Lednia</emphasis>
</taxonomicName>
specimen found in Darby Canyon, a female, was fortuitously collected by one of the authors (Ron Call), while hiking with his family. Three years after discovering the first female 
<taxonomicName box="[725,797,651,675]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis box="[725,797,651,675]" italics="true" pageId="6" pageNumber="110">Lednia</emphasis>
</taxonomicName>
specimen, and after many unsuccessful collecting trips, adults were found inside the cave. They were abundant inside the cave but were not collected outside. This species may be a troglophile because of its cavernicolous life style. Cave dwelling 
<taxonomicName box="[675,797,829,853]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="110" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
species have been found previously in Europe in studies such as 
<bibRefCitation author="Sivec, I." box="[373,513,900,924]" pageId="6" pageNumber="110" pagination="39 - 44" refId="ref2768" refString="Sivec, I. 1979. Notes on the drift of stoneflies (Plecoptera) in subterranean environment of the sinking-river Pivka (Slovenija, Yugoslavia). Gewasser und Abwasser 64: 39 - 44." type="journal article" year="1979">Sivec (1979)</bibRefCitation>
and 
<bibRefCitation author="Tierno de Figueroa, J. M. &amp; M. J. Lopez-Rodriguez" pageId="6" pageNumber="110" pagination="48 - 54" refId="ref2843" refString="Tierno de Figueroa, J. M. and M. J. Lopez-Rodriguez. 2010. Protonemoura gevi sp. n., a cavernicolous new species of stonefly (Insecta: Plecoptera). Zootaxa, 2365: 48 - 54." type="journal article" year="2010">Tierno de Figueroa and Lopez-Rodriguez (2010)</bibRefCitation>
. Finding 
<taxonomicName box="[644,716,935,959]" class="Insecta" family="Nemouridae" genus="Lednia" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis box="[644,716,935,959]" italics="true" pageId="6" pageNumber="110">Lednia</emphasis>
</taxonomicName>
at this location highlights several important points: First, that this genus is likely more widespread than recently believed. Second, it was finally only successfully collected after years of work, reminding us that it is important to remember that lack of data does not make for sound conclusions about species distribution. Species often exist in an area but have eluded collection because of the difficulty in getting to remote areas, or with problems gaining access to the habitat during the adult emergence period. So, if the new species emerged then, it would have been impossible to access the cave during the winter or early spring and it would probably still not have been discovered. Third, finding this species in a remote, backcountry, high elevation area illustrates the importance of collecting in special difficult to reach areas. A study of the greater Teton- Yellowstone region, particularly remote, backcountry habitats, would be useful and could provide a much clearer picture about the distribution of this and other rare stonefly species.
</paragraph>
</subSubSection>
</treatment>
</document>