270 lines
34 KiB
XML
270 lines
34 KiB
XML
<document id="7C052908A792ABBF43E3BCB0F724620D" ID-CLB-Dataset="8749" ID-DOI="10.11646/zootaxa.4838.1.10" ID-GBIF-Dataset="6ac7702d-58e9-4fc4-a74c-b45d965eb4b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="4403518" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609350870738" checkinUser="plazi" docAuthor="Skibińska, Kornelia, Szadziewski, Ryszard, Kania, Iwona & Krzemiński, Wiesław" docDate="2020" docId="03B087A09F1CA60653AC7C93FEC4FDDA" docLanguage="en" docName="zootaxa.4838.1.10.pdf" docOrigin="Zootaxa 4838 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoffeinsodes adamowiczi Skibińska & Szadziewski & Kania & Krzemiński 2020, sp. nov." docType="treatment" docUuid="91E406F1-1AE9-483D-B66D-67D93417EC6A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="150" masterDocId="FF89FFD89F1CA605533B7B13FFF0FFBA" masterDocTitle="A new species of Bruchomyiinae (Diptera: Psychodidae) from Baltic amber" masterLastPageNumber="150" masterPageNumber="147" pageNumber="147" updateTime="1698850103700" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="6613C6BDFC0073A74BF4821140E95CDF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8A3ED65B0B9C2AC791925792ECDF12D4">
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<mods:title id="FA023C34C3555A53573F16E2FEC076CC">A new species of Bruchomyiinae (Diptera: Psychodidae) from Baltic amber</mods:title>
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</mods:titleInfo>
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<mods:name id="A4C73A1699C753D717CE4C7A78FFC813" type="personal">
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<mods:roleTerm id="D0838B5977056A17F3992903FC290614">Author</mods:roleTerm>
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<mods:namePart id="94845C1BA42402B14692FE25F830E80B">Skibińska, Kornelia</mods:namePart>
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<mods:affiliation id="1AF8FF8B4246A04990FEA196D490B05C">Institute of Systematics and Evolution of Animals Polish Academy of Sciences, Poland.</mods:affiliation>
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</mods:name>
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<mods:name id="986BEF19ED00A8FD8D1B46844D756CD4" type="personal">
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<mods:role id="F091D492FA5FF09915CD76A0818A9C12">
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<mods:roleTerm id="531FACC14227C6A6B13BC05ED3688C6C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7D27A2FF512E8950D7050372D40FCB4E">Szadziewski, Ryszard</mods:namePart>
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<mods:nameIdentifier id="01F6CCDA0C006B593BC174EB1120E224" type="ORCID">0000-0002-2899-6861</mods:nameIdentifier>
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<mods:affiliation id="95CDD35F4D0E0B4B36AC7E7E7ED05264">Department of Invertebrate Zoology and Parasitology, University of Gdańsk, Wita Stwosza 59, 80 - 308 Gdańsk Poland. ryszard. szadziewski @ ug. edu. pl; https: // orcid. org / 0000 - 0002 - 2899 - 6861</mods:affiliation>
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<mods:nameIdentifier id="93EA05CE7C1A328E49ACAEFC7ADBBE45" type="email">ryszard.szadziewski@ug.edu.pl</mods:nameIdentifier>
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</mods:name>
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<mods:name id="E0C847AD75A49333EF1F0BFC12DC7039" type="personal">
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<mods:role id="D885CA8C5A82A37E141337314DD4FF42">
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<mods:roleTerm id="30C189EB08FAC68210D603F7B5DBA29E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="948E0E1311ECC96B85094C154424E379">Kania, Iwona</mods:namePart>
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<mods:nameIdentifier id="482110DDF5F85D3E379C0CB71EE2BFA4" type="ORCID">0000-0002-2325-4308</mods:nameIdentifier>
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<mods:affiliation id="51643DA357AE9D20FE67171E4DBEFC69">Department of Paleobiology and Evolutionary Biology, University of Rzeszów, Zelwerowicza 4, 35 - 601 Rzeszów, Poland. iwonakania @ onet. eu; https: // orcid. org / 0000 - 0002 - 2325 - 4308</mods:affiliation>
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<mods:nameIdentifier id="79AB6BED9924E184BC39EA92A6D35D05" type="email">iwonakania@onet.eu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="FC0F628A0C9318BAB7979F8CBB7CAC76" type="personal">
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<mods:roleTerm id="A9029C9E33810EB7574A7C368942D830">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0002786F7D47F3116C37E9E133634443">Krzemiński, Wiesław</mods:namePart>
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<mods:affiliation id="119766551AED1D42AE25C52FB435B41D">Institute of Biology, Pedagogical University of Kraków, Podbrzezie 3, 31 - 054 Kraków, Poland.</mods:affiliation>
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<mods:typeOfResource id="6E3706BB63088F74AB79B81D7C67D258">text</mods:typeOfResource>
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<mods:title id="203F3BC4639578E5447033DC1E3800BA">Zootaxa</mods:title>
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<mods:part id="D3CF56964B3E4EEADEEAAB42A6048E86">
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<mods:date id="668F016F1C6ECA52F867E16E52C62989">2020</mods:date>
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<mods:number id="87D866544BBEF97778973D9F2078E5F5">2020-08-25</mods:number>
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<mods:number id="71959F0288E593F2F6F591F18C8E2525">4838</mods:number>
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<mods:start id="C11C0DF5AA75A45E918DBAABB875B15B">147</mods:start>
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<mods:identifier id="5CF594C77BCAD4A4D35AA6003A9416C5" type="CLB-Dataset">8749</mods:identifier>
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<mods:identifier id="108B9193F79733A564735B4826B9474B" type="DOI">10.11646/zootaxa.4838.1.10</mods:identifier>
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<treatment id="03B087A09F1CA60653AC7C93FEC4FDDA" ID-DOI="http://doi.org/10.5281/zenodo.4453273" ID-GBIF-Taxon="177012376" ID-Zenodo-Dep="4453273" LSID="urn:lsid:zoobank.org:act:91E406F1-1AE9-483D-B66D-67D93417EC6A" httpUri="http://treatment.plazi.org/id/03B087A09F1CA60653AC7C93FEC4FDDA" lastPageId="3" lastPageNumber="150" pageId="0" pageNumber="147">
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<subSubSection id="C303653D9F1CA60553AC7C93FC51F804" pageId="0" pageNumber="147" type="nomenclature">
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<paragraph id="8BA636B69F1CA60553AC7C93FD4AF820" blockId="0.[151,929,1920,1982]" box="[151,698,1920,1947]" pageId="0" pageNumber="147">
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<heading id="D0EE81DA9F1CA60553AC7C93FD4AF820" bold="true" box="[151,698,1920,1947]" fontSize="11" level="1" pageId="0" pageNumber="147" reason="1">
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<emphasis id="B96DEAA49F1CA60553AC7C93FD4AF820" bold="true" box="[151,698,1920,1947]" pageId="0" pageNumber="147">
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<taxonomicName id="4C194D359F1CA60553AC7C93FE3CF821" ID-CoL="9HLZF" authority="Skibińska & Szadziewski & Kania & Krzemiński, 2020" authorityName="Skibińska & Szadziewski & Kania & Krzemiński" authorityYear="2020" box="[151,460,1920,1947]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="0" pageNumber="147" phylum="Arthropoda" rank="species" species="adamowiczi" status="sp. nov.">
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<emphasis id="B96DEAA49F1CA60553AC7C93FE3CF821" bold="true" box="[151,460,1920,1947]" italics="true" pageId="0" pageNumber="147">Hoffeinsodes adamowiczi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25E57DF9F1CA60552E87C92FDC0F821" box="[467,560,1921,1947]" pageId="0" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
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(
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<figureCitation id="13222A339F1CA605517B7C93FD42F820" box="[576,690,1920,1946]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,244,1840,1862]" captionStartId-1="2.[151,244,1928,1950]" captionTargetBox-0="[158,1427,883,1813]" captionTargetBox-1="[161,1427,197,1895]" captionTargetId-0="figure-414@1.[158,1428,883,1814]" captionTargetId-1="figure-22@2.[151,1436,182,1903]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Photomicrographs of Hoffeinsodes adamowiczi sp. nov., holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia." captionText-1="FIGURE 2. Line drawings of Hoffeinsodes adamowiczi sp. nov. holotype male.A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci.Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci." figureDoi-0="http://doi.org/10.5281/zenodo.4403520" figureDoi-1="http://doi.org/10.5281/zenodo.4403522" httpUri-0="https://zenodo.org/record/4403520/files/figure.png" httpUri-1="https://zenodo.org/record/4403522/files/figure.png" pageId="0" pageNumber="147">Figs. 1–2</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA636B69F1CA60553AC7CB5FC51F804" blockId="0.[151,929,1920,1982]" box="[151,929,1958,1982]" pageId="0" pageNumber="147">
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<uri id="FF883AB49F1CA60553AC7CB5FC51F804" box="[151,929,1958,1982]" pageId="0" pageNumber="147">
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urn:lsid:zoobank.org:act:
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<uuid id="FFBF0C639F1CA60552A57CB5FC51F804" box="[414,929,1958,1982]" pageId="0" pageNumber="147">91E406F1-1AE9-483D-B66D-67D93417EC6A</uuid>
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</uri>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C303653D9F1DA60453AC7B8BFA83FF6E" pageId="1" pageNumber="148" type="etymology">
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<paragraph id="8BA636B69F1DA60453AC7B8BFA83FF6E" blockId="1.[151,1437,151,859]" pageId="1" pageNumber="148">
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<emphasis id="B96DEAA49F1DA60453AC7B8BFEE8FF0A" bold="true" box="[151,280,152,176]" inLineHeading="true" pageId="1" pageNumber="148" reason="1">Etymology.</emphasis>
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The new species is dedicated to the late Paweł Adamowicz, ambassador of Baltic amber and promoter of related research, who served many years as the Mayor of the city Gdańsk until his assassination on
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<date id="FFA710769F1DA60457877BAFFA80FF6E" box="[1212,1392,188,212]" pageId="1" pageNumber="148" value="2019-01-13">13 January 2019</date>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C303653D9F1DA60453FC7BCCFE3BFE85" pageId="1" pageNumber="148" type="diagnosis">
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<paragraph id="8BA636B69F1DA60453FC7BCCFE3BFE85" blockId="1.[151,1437,151,859]" pageId="1" pageNumber="148">
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<emphasis id="B96DEAA49F1DA60453FC7BCCFECCFF4D" box="[199,316,223,247]" inLineHeading="true" pageId="1" pageNumber="148" reason="1">
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<emphasis id="B96DEAA49F1DA60453FC7BCCFEC5FF4D" bold="true" box="[199,309,223,247]" pageId="1" pageNumber="148">Diagnosis</emphasis>
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.
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</emphasis>
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The species is characterized by having cross-vein r-m close to the fork M
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<subScript id="179D34F39F1DA60457787BE3FBABFF44" attach="left" box="[1091,1115,240,254]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
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(about 5/6 length of M
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<subScript id="179D34F39F1DA60456757BE3FA97FF44" attach="left" box="[1358,1383,240,254]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
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); M
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<subScript id="179D34F39F1DA60456AF7BE3FA6CFF44" attach="left" box="[1428,1436,240,254]" fontSize="6" pageId="1" pageNumber="148">1</subScript>
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about 2 times longer than M
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<subScript id="179D34F39F1DA60452F27A07FE11FE98" attach="left" box="[457,481,276,290]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
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; epandrium with elongated epandrial lobes greatly extended at its ends; gonocoxite with two lobes. Female unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="C303653D9F1DA60653FC7A58FEC4FDDA" lastPageId="3" lastPageNumber="150" pageId="1" pageNumber="148" type="description">
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<paragraph id="8BA636B69F1DA60453FC7A58FDA3FC82" blockId="1.[151,1437,151,859]" pageId="1" pageNumber="148">
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<emphasis id="B96DEAA49F1DA60453FC7A58FEA2FED9" box="[199,338,331,355]" inLineHeading="true" pageId="1" pageNumber="148" reason="1">
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<emphasis id="B96DEAA49F1DA60453FC7A58FEBBFED9" bold="true" box="[199,331,331,355]" pageId="1" pageNumber="148">Description</emphasis>
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.
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</emphasis>
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Male. Head broad with oval eyes (
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<figureCitation id="13222A339F1DA60451E57A58FCC4FEDE" box="[734,820,331,356]" captionStart="FIGURE 1" captionStartId="1.[151,244,1840,1862]" captionTargetBox="[158,1427,883,1813]" captionTargetId="figure-414@1.[158,1428,883,1814]" captionTargetPageId="1" captionText="FIGURE 1. Photomicrographs of Hoffeinsodes adamowiczi sp. nov., holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia." figureDoi="http://doi.org/10.5281/zenodo.4403520" httpUri="https://zenodo.org/record/4403520/files/figure.png" pageId="1" pageNumber="148">Fig. 1B</figureCitation>
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); palpus four segmented (length in mm: 1/0.15; 2/0.23; 3/0.30; 4/0.63), last segment almost equal in length to the total length of the previous two (
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<figureCitation id="13222A339F1DA604574B7A7CFB3EFE32" box="[1136,1230,367,392]" captionStart="FIGURE 1" captionStartId="1.[151,244,1840,1862]" captionTargetBox="[158,1427,883,1813]" captionTargetId="figure-414@1.[158,1428,883,1814]" captionTargetPageId="1" captionText="FIGURE 1. Photomicrographs of Hoffeinsodes adamowiczi sp. nov., holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia." figureDoi="http://doi.org/10.5281/zenodo.4403520" httpUri="https://zenodo.org/record/4403520/files/figure.png" pageId="1" pageNumber="148">Figs. 1B</figureCitation>
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,
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<figureCitation id="13222A339F1DA60457E07A63FB0DFE32" box="[1243,1277,368,392]" captionStart="FIGURE 2" captionStartId="2.[151,244,1928,1950]" captionTargetBox="[161,1427,197,1895]" captionTargetId="figure-22@2.[151,1436,182,1903]" captionTargetPageId="2" captionText="FIGURE 2. Line drawings of Hoffeinsodes adamowiczi sp. nov. holotype male.A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci.Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci." figureDoi="http://doi.org/10.5281/zenodo.4403522" httpUri="https://zenodo.org/record/4403522/files/figure.png" pageId="1" pageNumber="148">2B</figureCitation>
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); antenna with scape (
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<quantity id="4CE19B539F1DA60453D97A87FEB4FE11" box="[226,324,404,428]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="148" unit="mm" value="0.04">0.04 mm</quantity>
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long), pedicel (
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<quantity id="4CE19B539F1DA60452CA7A87FDA3FE11" box="[497,595,404,428]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="148" unit="mm" value="0.1">0.10 mm</quantity>
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long) and 14 flagellomeres; flagellomeres long, tubular, decreasing in length (length in mm: 1/0.39; 14/0.12) (
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<figureCitation id="13222A339F1DA60452C67AA4FDABFE6A" box="[509,603,439,464]" captionStart="FIGURE 1" captionStartId="1.[151,244,1840,1862]" captionTargetBox="[158,1427,883,1813]" captionTargetId="figure-414@1.[158,1428,883,1814]" captionTargetPageId="1" captionText="FIGURE 1. Photomicrographs of Hoffeinsodes adamowiczi sp. nov., holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia." figureDoi="http://doi.org/10.5281/zenodo.4403520" httpUri="https://zenodo.org/record/4403520/files/figure.png" pageId="1" pageNumber="148">Figs. 1B</figureCitation>
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,
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<figureCitation id="13222A339F1DA60451537AABFD7DFE6A" box="[616,653,440,464]" captionStart="FIGURE 2" captionStartId="2.[151,244,1928,1950]" captionTargetBox="[161,1427,197,1895]" captionTargetId="figure-22@2.[151,1436,182,1903]" captionTargetPageId="2" captionText="FIGURE 2. Line drawings of Hoffeinsodes adamowiczi sp. nov. holotype male.A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci.Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci." figureDoi="http://doi.org/10.5281/zenodo.4403522" httpUri="https://zenodo.org/record/4403522/files/figure.png" pageId="1" pageNumber="148">2A</figureCitation>
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). Wing length
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<quantity id="4CE19B539F1DA60450167AABFC73FE75" box="[813,899,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="1" pageNumber="148" unit="mm" value="4.3">4.3 mm</quantity>
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; width
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<quantity id="4CE19B539F1DA60450EF7AABFBC8FE75" box="[980,1080,440,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.22" pageId="1" pageNumber="148" unit="mm" value="1.22">1.22 mm</quantity>
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; Sc ending before half of a wing length and opposite to vein r-m; sc-r about its length before end of Sc; R
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<subScript id="179D34F39F1DA60450AC7AFFFC6FFE40" attach="left" box="[919,927,492,506]" fontSize="6" pageId="1" pageNumber="148">1</subScript>
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long, ends opposite about 3/4 length of R
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<subScript id="179D34F39F1DA60456617AFFFA92FE40" attach="left" box="[1370,1378,492,506]" fontSize="6" pageId="1" pageNumber="148">2</subScript>
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; fork of Rb into R
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<subScript id="179D34F39F1DA60452227903FED1FDA4" attach="left" box="[281,289,528,542]" fontSize="6" pageId="1" pageNumber="148">1</subScript>
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and Rs before fork of Mb into M
|
||
<subScript id="179D34F39F1DA60451407903FD64FDA4" attach="left" box="[635,660,528,542]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
|
||
and M
|
||
<subScript id="179D34F39F1DA60451E67903FD05FDA4" attach="left" box="[733,757,528,542]" fontSize="6" pageId="1" pageNumber="148">3+4</subScript>
|
||
; R
|
||
<subScript id="179D34F39F1DA604502E7903FCDDFDA4" attach="left" box="[789,813,528,542]" fontSize="6" pageId="1" pageNumber="148">2+3</subScript>
|
||
more than 2 times longer than Rs; R
|
||
<subScript id="179D34F39F1DA60457907903FB43FDA4" attach="left" box="[1195,1203,528,542]" fontSize="6" pageId="1" pageNumber="148">2</subScript>
|
||
almost 3 times longer than Rs; r-m located between R
|
||
<subScript id="179D34F39F1DA60452DC7927FE1FFDF8" attach="left" box="[487,495,564,578]" fontSize="6" pageId="1" pageNumber="148">5</subScript>
|
||
and M
|
||
<subScript id="179D34F39F1DA60451027927FDA2FDF8" attach="left" box="[569,594,564,578]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
|
||
; M
|
||
<subScript id="179D34F39F1DA604514C7927FD60FDF8" attach="left" box="[631,656,564,578]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
|
||
two times longer than Mb; M
|
||
<subScript id="179D34F39F1DA60450F57927FC26FDF8" attach="left" box="[974,982,564,578]" fontSize="6" pageId="1" pageNumber="148">1</subScript>
|
||
about 2 times longer than M
|
||
<subScript id="179D34F39F1DA60456327927FAD2FDF8" attach="left" box="[1289,1314,564,578]" fontSize="6" pageId="1" pageNumber="148">1+2</subScript>
|
||
; M
|
||
<subScript id="179D34F39F1DA604567C7927FABFFDF8" attach="left" box="[1351,1359,564,578]" fontSize="6" pageId="1" pageNumber="148">3</subScript>
|
||
almost 11 times longer than M
|
||
<subScript id="179D34F39F1DA60452AA794BFE59FDDC" attach="left" box="[401,425,600,614]" fontSize="6" pageId="1" pageNumber="148">3+4</subScript>
|
||
; M
|
||
<subScript id="179D34F39F1DA60452EB794BFE28FDDC" attach="left" box="[464,472,600,614]" fontSize="6" pageId="1" pageNumber="148">4</subScript>
|
||
about 9 times longer than M
|
||
<subScript id="179D34F39F1DA6045035794BFCD6FDDC" attach="left" box="[782,806,600,614]" fontSize="6" pageId="1" pageNumber="148">3+4</subScript>
|
||
; opening of cells m
|
||
<subScript id="179D34F39F1DA60450C7794BFBF4FDDC" attach="left" box="[1020,1028,600,614]" fontSize="6" pageId="1" pageNumber="148">1</subScript>
|
||
, m
|
||
<subScript id="179D34F39F1DA604571C794BFBDFFDDC" attach="left" box="[1063,1071,600,614]" fontSize="6" pageId="1" pageNumber="148">2</subScript>
|
||
and m
|
||
<subScript id="179D34F39F1DA6045743794BFB70FDDC" attach="left" box="[1144,1152,600,614]" fontSize="6" pageId="1" pageNumber="148">3</subScript>
|
||
almost equal in width; Cu ends opposite fork of Rs; m-cu located in fork of M
|
||
<subScript id="179D34F39F1DA604518A796FFD39FD30" attach="left" box="[689,713,636,650]" fontSize="6" pageId="1" pageNumber="148">3+4</subScript>
|
||
; A
|
||
<subScript id="179D34F39F1DA60451D3796FFD00FD30" attach="left" box="[744,752,636,650]" fontSize="6" pageId="1" pageNumber="148">1</subScript>
|
||
quite long ends clearly after m-cu (
|
||
<figureCitation id="13222A339F1DA60457587978FB4FFD3E" box="[1123,1215,619,644]" captionStart="FIGURE 1" captionStartId="1.[151,244,1840,1862]" captionTargetBox="[158,1427,883,1813]" captionTargetId="figure-414@1.[158,1428,883,1814]" captionTargetPageId="1" captionText="FIGURE 1. Photomicrographs of Hoffeinsodes adamowiczi sp. nov., holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia." figureDoi="http://doi.org/10.5281/zenodo.4403520" httpUri="https://zenodo.org/record/4403520/files/figure.png" pageId="1" pageNumber="148">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="13222A339F1DA60457F1797FFB1CFD3E" box="[1226,1260,620,644]" captionStart="FIGURE 2" captionStartId="2.[151,244,1928,1950]" captionTargetBox="[161,1427,197,1895]" captionTargetId="figure-22@2.[151,1436,182,1903]" captionTargetPageId="2" captionText="FIGURE 2. Line drawings of Hoffeinsodes adamowiczi sp. nov. holotype male.A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci.Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci." figureDoi="http://doi.org/10.5281/zenodo.4403522" httpUri="https://zenodo.org/record/4403522/files/figure.png" pageId="1" pageNumber="148">2C</figureCitation>
|
||
). Male genitalia. Gonocoxites fused with hypandrium, each with two lobes tapering to a point; compared to each other, lower gonocoxal lobe broad, slightly arched, upper lobe narrow; aedeagus not discernable; gonostyli simple, about twice as long as wide, sub-rectangular in lateral view, slightly narrowing to blunt apices, covered with setae; epandrium with symmetrical, elongated epandrial lobes clearly broadening at apices; proctiger tubular, elongate, extending clearly beyond epandrium; cerci ovate in lateral view (
|
||
<figureCitation id="13222A339F1DA6045244780CFE2DFC82" box="[383,477,799,824]" captionStart="FIGURE 1" captionStartId="1.[151,244,1840,1862]" captionTargetBox="[158,1427,883,1813]" captionTargetId="figure-414@1.[158,1428,883,1814]" captionTargetPageId="1" captionText="FIGURE 1. Photomicrographs of Hoffeinsodes adamowiczi sp. nov., holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia." figureDoi="http://doi.org/10.5281/zenodo.4403520" httpUri="https://zenodo.org/record/4403520/files/figure.png" pageId="1" pageNumber="148">Figs. 1D</figureCitation>
|
||
,
|
||
<figureCitation id="13222A339F1DA60452D27833FDB6FC82" box="[489,582,800,824]" captionStart="FIGURE 2" captionStartId="2.[151,244,1928,1950]" captionTargetBox="[161,1427,197,1895]" captionTargetId="figure-22@2.[151,1436,182,1903]" captionTargetPageId="2" captionText="FIGURE 2. Line drawings of Hoffeinsodes adamowiczi sp. nov. holotype male.A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci.Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci." figureDoi="http://doi.org/10.5281/zenodo.4403522" httpUri="https://zenodo.org/record/4403522/files/figure.png" pageId="1" pageNumber="148">2D, E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636B69F1DA60453FC7850FE64FCE1" blockId="1.[151,1437,151,859]" box="[199,404,835,859]" pageId="1" pageNumber="148">Female. Unknown.</paragraph>
|
||
<caption id="DF66663E9F1DA60453AC7C23FEBBF8D0" ID-DOI="http://doi.org/10.5281/zenodo.4403520" ID-Zenodo-Dep="4403520" httpUri="https://zenodo.org/record/4403520/files/figure.png" pageId="1" pageNumber="148" startId="1.[151,244,1840,1862]" targetBox="[158,1427,883,1813]" targetPageId="1">
|
||
<paragraph id="8BA636B69F1DA60453AC7C23FEBBF8D0" blockId="1.[151,1437,1840,1898]" pageId="1" pageNumber="148">
|
||
<emphasis id="B96DEAA49F1DA60453AC7C23FEFEF8FF" bold="true" box="[151,270,1840,1862]" pageId="1" pageNumber="148">FIGURE 1.</emphasis>
|
||
Photomicrographs of
|
||
<taxonomicName id="4C194D359F1DA60452DE7C23FD2BF8FF" authority="Skibińska & Szadziewski & Kania & Krzemiński, 2020" authorityName="Skibińska & Szadziewski & Kania & Krzemiński" authorityYear="2020" box="[485,731,1840,1861]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="148" phylum="Arthropoda" rank="species" species="adamowiczi" status="sp. nov.">
|
||
<emphasis id="B96DEAA49F1DA60452DE7C23FD2BF8FF" box="[485,731,1840,1861]" italics="true" pageId="1" pageNumber="148">Hoffeinsodes adamowiczi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E57DF9F1DA60451D97C23FCDFF8FF" box="[738,815,1840,1861]" pageId="1" pageNumber="148" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype male no. MAIG 5945 A. whole specimen; B. head; C. wing; D. genitalia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF66663E9F1EA60753AC7C9BFB1CF85C" ID-DOI="http://doi.org/10.5281/zenodo.4403522" ID-Zenodo-Dep="4403522" httpUri="https://zenodo.org/record/4403522/files/figure.png" pageId="2" pageNumber="149" startId="2.[151,244,1928,1950]" targetBox="[161,1427,197,1895]" targetPageId="2">
|
||
<paragraph id="8BA636B69F1EA60753AC7C9BFB1CF85C" blockId="2.[151,1437,1928,2022]" pageId="2" pageNumber="149">
|
||
<emphasis id="B96DEAA49F1EA60753AC7C9BFEFDF827" bold="true" box="[151,269,1928,1950]" pageId="2" pageNumber="149">FIGURE 2.</emphasis>
|
||
Line drawings of
|
||
<taxonomicName id="4C194D359F1EA60752877C9BFD42F827" authority="Skibińska & Szadziewski & Kania & Krzemiński, 2020" authorityName="Skibińska & Szadziewski & Kania & Krzemiński" authorityYear="2020" box="[444,690,1928,1949]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="149" phylum="Arthropoda" rank="species" species="adamowiczi" status="sp. nov.">
|
||
<emphasis id="B96DEAA49F1EA60752877C9BFD42F827" box="[444,690,1928,1949]" italics="true" pageId="2" pageNumber="149">Hoffeinsodes adamowiczi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E57DF9F1EA60751837C9BFCF2F827" box="[696,770,1928,1949]" pageId="2" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
|
||
holotype male. A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci. Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA636B69F1FA60653FC7B8BFD7DFE16" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="150">
|
||
Two genera of
|
||
<taxonomicName id="4C194D359F1FA60652517B84FDFEFF15" authorityName="Alexander" authorityYear="1920" box="[362,526,151,175]" class="Insecta" family="Psychodidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="subFamily" subFamily="Bruchomyiinae">Bruchomyiinae</taxonomicName>
|
||
are known from Baltic amber:
|
||
<taxonomicName id="4C194D359F1FA60650597B84FB54FF15" authority="Meunier, 1905" authorityName="Meunier" authorityYear="1905" box="[866,1188,151,175]" class="Insecta" family="Psychodidae" genus="Palaeosycorax" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96DEAA49F1FA60650597B84FBF0FF15" box="[866,1024,151,175]" italics="true" pageId="3" pageNumber="150">Palaeosycorax</emphasis>
|
||
<bibRefCitation id="EF884B479F1FA60657337B84FB54FF15" author="Meunier, F." box="[1032,1188,151,175]" pageId="3" pageNumber="150" pagination="235 - 255" refId="ref1961" refString="Meunier, F. (1905) Monographie des Psychodidae de l'ambre de la Baltique. Annales Musei Nationalis Hungarici, 3, 235 - 255, tas. V & VI." type="journal article" year="1905">Meunier, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C194D359F1FA60657E17B84FA95FF15" authorityName="Wagner" authorityYear="2017" box="[1242,1381,151,175]" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96DEAA49F1FA60657E17B84FA95FF15" box="[1242,1381,151,175]" italics="true" pageId="3" pageNumber="150">Hoffeinsodes</emphasis>
|
||
</taxonomicName>
|
||
. The latter genus, now including seven species, is readily characterized by the fusion of gonocoxites and hypandrium but can also be recognized by an exceptionally large epandrium with elongated lobes. Regarding the wing venation,
|
||
<taxonomicName id="4C194D359F1FA60656127BF3FF3EFEA1" authority="Skibińska & Szadziewski & Kania & Krzemiński, 2020" authorityName="Skibińska & Szadziewski & Kania & Krzemiński" authorityYear="2020" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="adamowiczi" status="sp. nov.">
|
||
<emphasis id="B96DEAA49F1FA60656127BF3FF3EFEA1" italics="true" pageId="3" pageNumber="150">H. adamowiczi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25E57DF9F1FA60653ED7A17FEDBFEA1" box="[214,299,260,283]" pageId="3" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be easily distinguished by location of cross-vein r-m (
|
||
<figureCitation id="13222A339F1FA60650907A10FC0FFEA6" box="[939,1023,259,284]" captionStart="FIGURE 2" captionStartId="2.[151,244,1928,1950]" captionTargetBox="[161,1427,197,1895]" captionTargetId="figure-22@2.[151,1436,182,1903]" captionTargetPageId="2" captionText="FIGURE 2. Line drawings of Hoffeinsodes adamowiczi sp. nov. holotype male.A. antenna; B. palpus; C. wing venation; D. genitalia, lateral view; E. gonostylus and gonocoxites fused with hypandrium; F. epandrium with cerci.Abbreviations: ep, epandrium; epl, epandrial lobes; fhg, fused hypandrium and gonocoxites; gst, gonostylus; gxl, gonocoxal lobe; p+c, proctiger with cerci." figureDoi="http://doi.org/10.5281/zenodo.4403522" httpUri="https://zenodo.org/record/4403522/files/figure.png" pageId="3" pageNumber="150">Fig. 2C</figureCitation>
|
||
) and proportions of radial and medial veins. Also, external morphology of the male genitalia is unique among described species, especially the epandrium with elongated and apically expanded epandrial lobes. Male genitalia of the new species bear some similarity to those of
|
||
<taxonomicName id="4C194D359F1FA60656507A5FFE8AFE3D" authorityName="Wagner" authorityYear="2017" class="Insecta" family="Psychodidae" genus="Hoffeinsodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="150" phylum="Arthropoda" rank="species" species="longicauda">
|
||
<emphasis id="B96DEAA49F1FA60656507A5FFE8AFE3D" italics="true" pageId="3" pageNumber="150">Hoffeinsodes longicauda</emphasis>
|
||
</taxonomicName>
|
||
from Baltic amber. In the latter species, however, gonocoxites and gonostyli are lacking divisions and gonocoxites have no lobes (
|
||
<bibRefCitation id="EF884B479F1FA60652D77A87FD71FE16" author="Wagner, R." box="[492,641,404,428]" pageId="3" pageNumber="150" pagination="100 - 120" refId="ref2226" refString="Wagner, R. (2017) Synopsis of extinct Bruchomyiinae (Diptera, Psychodidae) from Burmese, Baltic and Dominican amber, with description of new genera and species. Zootaxa, 4320 (1), 100 - 120. https: // doi. org / 10.11646 / zootaxa. 4320.1.6" type="journal article" year="2017">Wagner 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA636B69F1FA60653FC7AABFEC4FDDA" blockId="3.[151,1437,151,608]" pageId="3" pageNumber="150">
|
||
<materialsCitation id="3B713CEB9F1FA60653FC7AABFBBBFD81" ID-GBIF-Occurrence="3020149302" country="Poland" county="National Science Centre" location="Museum of Amber" municipality="Funding" pageId="3" pageNumber="150" specimenCount="1" typeStatus="holotype">
|
||
We are much indebted to amber dealer Jonas Damzen for donation of the
|
||
<typeStatus id="54A288149F1FA60650DA7AA4FBCEFE75" box="[993,1086,439,463]" pageId="3" pageNumber="150" type="holotype">holotype</typeStatus>
|
||
to the
|
||
<location id="8EC6606D9F1FA60657B17AABFAA5FE75" LSID="urn:lsid:plazi:treatment:03B087A09F1CA60653AC7C93FEC4FDDA:8EC6606D9F1FA60657B17AABFAA5FE75" box="[1162,1365,439,463]" country="Poland" county="National Science Centre" municipality="Funding" name="Museum of Amber" pageId="3" pageNumber="150">Museum of Amber</location>
|
||
Inclusions at
|
||
<location id="8EC6606D9F1FA60653CA7AC8FD7DFE49" LSID="urn:lsid:plazi:treatment:03B087A09F1CA60653AC7C93FEC4FDDA:8EC6606D9F1FA60653CA7AC8FD7DFE49" box="[241,653,475,500]" country="Poland" county="National Science Centre" municipality="Funding" name="University of Gdansk. Gregory Curler" pageId="3" pageNumber="150">University of Gdańsk. Gregory Curler</location>
|
||
is greatly appreciated for suggestions to improve the original version of this paper and providing encouragement.
|
||
<location id="8EC6606D9F1FA60651687AECFD52FDAD" LSID="urn:lsid:plazi:treatment:03B087A09F1CA60653AC7C93FEC4FDDA:8EC6606D9F1FA60651687AECFD52FDAD" box="[595,674,511,535]" country="Poland" county="National Science Centre" municipality="Funding" name="Thanks" pageId="3" pageNumber="150">Thanks</location>
|
||
as well to two reviewers for their helpful suggestions.
|
||
<collectingMunicipality id="6BC2ACCC9F1FA60657CB7AECFAB9FDAD" box="[1264,1353,511,535]" pageId="3" pageNumber="150">Funding</collectingMunicipality>
|
||
for this research was partially provided by the grant from the
|
||
<collectingCounty id="62C74E3A9F1FA60651E37930FC2CFD86" box="[728,988,547,572]" pageId="3" pageNumber="150">National Science Centre</collectingCounty>
|
||
of
|
||
<collectingCountry id="F30E76269F1FA606573A7930FBBBFD81" box="[1025,1099,547,571]" name="Poland" pageId="3" pageNumber="150">Poland</collectingCountry>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B713CEB9F1FA60657697930FF29FDDA" ID-GBIF-Occurrence="3020149301" collectingDate="2018-12" country="New Zealand" location="NZ" pageId="3" pageNumber="150" specimenCount="1" typeStatus="holotype">
|
||
Miniatura 2 no.
|
||
<date id="FFA710769F1FA60657C47937FA82FD86" box="[1279,1394,548,572]" pageId="3" pageNumber="150" value="2018-12">
|
||
<collectingDate id="EFE3E99E9F1FA60657C47937FA82FD86" box="[1279,1394,548,572]" pageId="3" pageNumber="150" value="2018-12">DEC-2018</collectingDate>
|
||
</date>
|
||
/02/ X/
|
||
<collectingCountry id="F30E76269F1FA606538F795BFF29FDDA" box="[180,217,584,608]" name="New Zealand" pageId="3" pageNumber="150">NZ</collectingCountry>
|
||
</materialsCitation>
|
||
8/01001.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |