283 lines
38 KiB
XML
283 lines
38 KiB
XML
<document id="B49709B9EB684FD1A7B30A1B436CA274" ID-DOI="10.5281/zenodo.5303571" ID-ISSN="0374-1036" ID-Zenodo-Dep="5303571" ID-ZooBank="0360453D-E09E-4CC4-9BB0-981AEC7BED66S" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630104360660" checkinUser="marcus" docAuthor="Tsai, Jing-Fu & Rédei, Dávid" docDate="2015" docId="03B63F659F551C5BFE16FC73FE05FE9B" docLanguage="en" docName="ActaEntMusNatPra.2015.55.2.625-664.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Acanthosoma axicia Tsai & Rédei 2015, sp. nov." docType="treatment" docVersion="3" lastPageNumber="642" masterDocId="FF8F471D9F5A1C49FFADFFB2FFA0FFF4" masterDocTitle="The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)" masterLastPageNumber="664" masterPageNumber="625" pageNumber="640" updateTime="1699153447944" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="FB1C5C49D7F48F137E9B5F4BAF579A47">
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<mods:title id="CCA905603FBB5CD09CF75A1E2B04729D">The genus Acanthosoma in Taiwan (Hemiptera: Heteroptera: Acanthosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F8AE2981D8A30E2DFE4268F45A6DBE1C" type="personal">
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<mods:roleTerm id="DD72775F03F3A16D7AD856CEB1BCF7D1">Author</mods:roleTerm>
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<mods:namePart id="4023D3AE732C895CFF61DED14EF28E0D">Tsai, Jing-Fu</mods:namePart>
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<mods:affiliation id="6F88C5B6850CA65DC3F928503E6FA75A">Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589 Japan; e-mail: jingfu. tsai @ gmail. com</mods:affiliation>
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<mods:roleTerm id="AF3AAEB552408A97232A80160E92DD67">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="872A75C66D2FA1AC3805610BA0AB7A3E">Rédei, Dávid</mods:namePart>
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<mods:affiliation id="CF71F9EE8B4E88AFF365F675DD464648">Institute of Entomology, College of Life Sciences, Nankai University, Weijin Road 94, 300071 Tianjin, China; e-mail: david. redei @ gmail. com & Department of Zoology, Hungarian Natural History Museum, H- 1088 Budapest, Baross u. 13, Hungary</mods:affiliation>
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<mods:typeOfResource id="D3F949D7144819B8C0FEA05442020F5D">text</mods:typeOfResource>
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<mods:title id="90F1477D920B1FC03F5D5B08DF664DE9">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="521F63092FAE2278B534AA2ED5476C03">
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<mods:date id="7A6D2B2DF7F30BC0CF6676DE8DBBEAC2">2015</mods:date>
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<mods:number id="3A61FDC1D6B4C18A1D2BDA8869AF877C">2015-12-31</mods:number>
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<mods:number id="9494D333115940C2321F7108735C350B">55</mods:number>
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<mods:number id="770D7645191CA51A96814F5DDB57E509">2</mods:number>
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<mods:start id="737F5EA2E11F83BF95182ED34550E935">625</mods:start>
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<mods:classification id="A7E4B82D7677B8832D6E66C4203D9B00">journal article</mods:classification>
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<mods:identifier id="253F136523F73DDC6814DB6C8C514504" type="DOI">10.5281/zenodo.5303571</mods:identifier>
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<mods:identifier id="4B530C64F9921BD4397006ACA1153417" type="ISSN">0374-1036</mods:identifier>
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<treatment id="03B63F659F551C5BFE16FC73FE05FE9B" LSID="urn:lsid:plazi:treatment:03B63F659F551C5BFE16FC73FE05FE9B" httpUri="http://treatment.plazi.org/id/03B63F659F551C5BFE16FC73FE05FE9B" lastPageId="18" lastPageNumber="642" pageId="15" pageNumber="640">
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<subSubSection id="C305DDF89F551C46FE16FC73FD57FC2D" box="[443,759,961,986]" pageId="15" pageNumber="640" type="nomenclature">
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<paragraph id="8BA08E739F551C46FE16FC73FD57FC2D" blockId="15.[443,759,961,986]" box="[443,759,961,986]" pageId="15" pageNumber="640">
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<heading id="D0E8391F9F551C46FE16FC73FD57FC2D" bold="true" box="[443,759,961,986]" centered="true" fontSize="8" level="2" pageId="15" pageNumber="640" reason="3">
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<emphasis id="B96B52619F551C46FE16FC73FD57FC2D" bold="true" box="[443,759,961,986]" pageId="15" pageNumber="640">
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<taxonomicName id="4C1FF5F09F551C46FE16FC73FD3CFC2E" authority="Tsai & Rédei, 2015" authorityName="Tsai & Rédei" authorityYear="2015" box="[443,668,961,986]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="640" phylum="Arthropoda" rank="species" species="axicia" status="sp. nov.">
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<emphasis id="B96B52619F551C46FE16FC73FD3CFC2E" bold="true" box="[443,668,961,986]" italics="true" pageId="15" pageNumber="640">Acanthosoma axicia</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A258EF1A9F551C46FD0FFC70FD57FC2D" box="[674,759,962,985]" pageId="15" pageNumber="640" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C305DDF89F551C46FE7EFC52FD40FC07" box="[467,736,992,1011]" pageId="15" pageNumber="640" type="description">
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<paragraph id="8BA08E739F551C46FE7EFC52FD40FC07" blockId="15.[113,1090,992,1134]" box="[467,736,992,1011]" pageId="15" pageNumber="640">
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(
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||
<figureCitation id="132492F69F551C46FE75FC52FD9AFC07" box="[472,570,992,1011]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="15" pageNumber="640">Figs 11–14</figureCitation>
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||
,
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||
<figureCitation id="132492F69F551C46FDEEFC52FDDAFC07" box="[579,634,992,1011]" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="15" pageNumber="640">37–39</figureCitation>
|
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,
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<figureCitation id="132492F69F551C46FD2EFC52FD3AFC07" box="[643,666,992,1011]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="15" pageNumber="640">52</figureCitation>
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,
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<figureCitation id="132492F69F551C46FD0EFC52FD79FC07" box="[675,729,992,1011]" captionStart="Figs 76–79" captionStartId="27.[113,150,1553,1572]" captionTargetBox="[155,1044,235,1489]" captionTargetId="figure-12@27.[113,1088,220,1520]" captionTargetPageId="27" captionText="Figs 76–79. Acanthosoma axicia sp. nov. 76–77 – male; 78–79 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303593" httpUri="https://zenodo.org/record/5303593/files/figure.png" pageId="15" pageNumber="640">76–79</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C305DDF89F551C46FFDCFBB8FBE2FB99" pageId="15" pageNumber="640" type="materials_examined">
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<paragraph id="8BA08E739F551C46FFDCFBB8FBE2FB99" blockId="15.[113,1090,992,1134]" pageId="15" pageNumber="640">
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||
<emphasis id="B96B52619F551C46FFDCFBB8FF54FBE9" bold="true" box="[113,244,1034,1053]" pageId="15" pageNumber="640">Type material.</emphasis>
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<materialsCitation id="3B77842E9F551C46FF55FBB8FCA8FBCC" collectingDate="2010-11-21" collectionCode="NMNS" collectorName="C. T. Tang & Y. C. Li" country="Taiwan" location="Lishan" pageId="15" pageNumber="640" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
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<typeStatus id="54A430D19F551C46FF55FBB8FEEBFBEF" box="[248,331,1034,1053]" pageId="15" pageNumber="640" type="holotype">HOLOTYPE</typeStatus>
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||
: J,
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||
<emphasis id="B96B52619F551C46FEC3FBB8FD9FFBE9" bold="true" box="[366,575,1034,1053]" pageId="15" pageNumber="640">
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<collectingCountry id="F308CEE39F551C46FEC3FBB8FE67FBE9" box="[366,455,1034,1053]" name="Taiwan" pageId="15" pageNumber="640">TAIWAN</collectingCountry>
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:
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<collectingRegion id="49DB40919F551C46FE62FBB8FDB4FBEF" box="[463,532,1034,1053]" country="Taiwan" name="Nantou" pageId="15" pageNumber="640">NANTOU</collectingRegion>
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||
CO.:
|
||
</emphasis>
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||
<location id="8EC0D8A89F551C46FDEEFBB8FDDFFBE9" LSID="urn:lsid:plazi:treatment:03B63F659F551C5BFE16FC73FE05FE9B:8EC0D8A89F551C46FDEEFBB8FDDFFBE9" box="[579,639,1034,1053]" country="Taiwan" name="Lishan" pageId="15" pageNumber="640" stateProvince="Nantou">Lishan</location>
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,
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||
<date id="FFA1A8B39F551C46FD2BFBB8FD43FBE9" box="[646,739,1034,1053]" pageId="15" pageNumber="640" value="2010-11-21">
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||
<collectingDate id="EFE5515B9F551C46FD2BFBB8FD43FBE9" box="[646,739,1034,1053]" pageId="15" pageNumber="640" value="2010-11-21">21.xi.2010</collectingDate>
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||
</date>
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||
, ex
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||
<taxonomicName id="4C1FF5F09F551C46FCAEFBB8FBBAFBE9" authority="(Makino)" baseAuthorityName="Makino" box="[771,1050,1034,1053]" class="Magnoliopsida" family="Pittosporaceae" genus="Pittosporum" kingdom="Plantae" order="Apiales" pageId="15" pageNumber="640" phylum="Tracheophyta" rank="species" species="illicioides">
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||
<emphasis id="B96B52619F551C46FCAEFBB8FC64FBE9" box="[771,964,1034,1053]" italics="true" pageId="15" pageNumber="640">Pittosporum illicioides</emphasis>
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||
(Makino)
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</taxonomicName>
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||
, leg.
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<collectorName id="26EAEBA59F551C46FFDCFB97FF63FBCC" box="[113,195,1061,1080]" pageId="15" pageNumber="640">C.T.Tang</collectorName>
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||
&
|
||
<collectorName id="26EAEBA59F551C46FF76FB97FEB9FBCC" box="[219,281,1061,1080]" pageId="15" pageNumber="640">Y.C. Li</collectorName>
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||
; mounted on card, intact; deposited in
|
||
<collectionCode id="ED0E16B69F551C46FDF4FB97FD36FBCC" box="[601,662,1061,1080]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="15" pageNumber="640">NMNS</collectionCode>
|
||
(
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||
<figureCitation id="132492F69F551C46FD0DFB97FCA2FBCC" box="[672,770,1061,1080]" captionStart="Figs 76–79" captionStartId="27.[113,150,1553,1572]" captionTargetBox="[155,1044,235,1489]" captionTargetId="figure-12@27.[113,1088,220,1520]" captionTargetPageId="27" captionText="Figs 76–79. Acanthosoma axicia sp. nov. 76–77 – male; 78–79 – female. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303593" httpUri="https://zenodo.org/record/5303593/files/figure.png" pageId="15" pageNumber="640">Figs 76–77</figureCitation>
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||
)
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</materialsCitation>
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.
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||
<materialsCitation id="3B77842E9F551C46FCA2FB97FDF3FBA7" collectionCode="NMNS, NCHU" country="Taiwan" location="NANTOU" pageId="15" pageNumber="640" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="paratype">
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<typeStatus id="54A430D19F551C46FCA2FB97FCC6FBC2" box="[783,870,1061,1080]" pageId="15" pageNumber="640" type="paratype">PARATYPES</typeStatus>
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:
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||
<emphasis id="B96B52619F551C46FCDDFB97FBE0FBCC" bold="true" box="[880,1088,1061,1080]" pageId="15" pageNumber="640">
|
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<collectingCountry id="F308CEE39F551C46FCDDFB97FC69FBCC" box="[880,969,1061,1080]" name="Taiwan" pageId="15" pageNumber="640">TAIWAN</collectingCountry>
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:
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<collectingRegion id="49DB40919F551C46FC7CFB97FBB5FBC2" box="[977,1045,1061,1080]" country="Taiwan" name="Nantou" pageId="15" pageNumber="640">NANTOU</collectingRegion>
|
||
CO.:
|
||
</emphasis>
|
||
same as
|
||
<typeStatus id="54A430D19F551C46FF16FBF2FEA6FBA7" box="[187,262,1088,1107]" pageId="15" pageNumber="640" type="holotype">holotype</typeStatus>
|
||
(
|
||
<specimenCount id="9D1945FA9F551C46FEB8FBF2FE95FBA6" box="[277,309,1081,1107]" pageId="15" pageNumber="640" type="female">1 ♀</specimenCount>
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||
<collectionCode id="ED0E16B69F551C46FE97FBF2FEDAFBA7" box="[314,378,1088,1107]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="15" pageNumber="640">NMNS</collectionCode>
|
||
, 1 J [dissected]
|
||
<collectionCode id="ED0E16B69F551C46FDA1FBF2FDECFBA7" box="[524,588,1088,1107]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="15" pageNumber="640">NCHU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B77842E9F551C46FDF1FBF2FEB7FB99" collectingDate="1990-04-26" collectionCode="NSMT" collectorName="He" country="Taiwan" location="Shihtyutou" municipality="Renai" pageId="15" pageNumber="640" specimenCount="1" stateProvince="Nantou" typeStatus="paratype">
|
||
<collectingMunicipality id="6BC414099F551C46FDF1FBF2FD30FBA7" box="[604,656,1088,1107]" pageId="15" pageNumber="640">Renai</collectingMunicipality>
|
||
,
|
||
<location id="8EC0D8A89F551C46FD34FBF2FD57FBA7" LSID="urn:lsid:plazi:treatment:03B63F659F551C5BFE16FC73FE05FE9B:8EC0D8A89F551C46FD34FBF2FD57FBA7" box="[665,759,1088,1107]" country="Taiwan" municipality="Renai" name="Shihtyutou" pageId="15" pageNumber="640" stateProvince="Nantou">Shihtyutou</location>
|
||
[= Shizitou],
|
||
<date id="FFA1A8B39F551C46FCDDFBF2FC71FBA7" box="[880,977,1088,1107]" pageId="15" pageNumber="640" value="1990-04-26">
|
||
<collectingDate id="EFE5515B9F551C46FCDDFBF2FC71FBA7" box="[880,977,1088,1107]" pageId="15" pageNumber="640" value="1990-04-26">26.IV.1990</collectingDate>
|
||
</date>
|
||
, NSMT-I
|
||
<collectorName id="26EAEBA59F551C46FB85FBF2FBE1FBA7" box="[1064,1089,1088,1107]" pageId="15" pageNumber="640">He</collectorName>
|
||
36136 (1J
|
||
<collectionCode id="ED0E16B69F551C46FF7EFBE8FEB1FB99" box="[211,273,1114,1133]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="15" pageNumber="640">NSMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B77842E9F551C46FEB0FBE8FB9EFB99" box="[285,1086,1108,1133]" collectingDate="2010-07-06" collectionCode="NCHU" collectorName="Y. C. Liao" country="Taiwan" location="Dasyueshan" pageId="15" pageNumber="640" specimenCount="1" specimenCount-female="1" stateProvince="Taichung" typeStatus="paratype">
|
||
<emphasis id="B96B52619F551C46FEB0FBE8FE3DFB99" bold="true" box="[285,413,1114,1133]" pageId="15" pageNumber="640">
|
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<collectingRegion id="49DB40919F551C46FEB0FBE8FED2FB9F" box="[285,370,1114,1133]" country="Taiwan" name="Taichung" pageId="15" pageNumber="640">TAICHUNG</collectingRegion>
|
||
CO.:
|
||
</emphasis>
|
||
<location id="8EC0D8A89F551C46FE0CFBE8FDA7FB99" LSID="urn:lsid:plazi:treatment:03B63F659F551C5BFE16FC73FE05FE9B:8EC0D8A89F551C46FE0CFBE8FDA7FB99" box="[417,519,1114,1133]" country="Taiwan" name="Dasyueshan" pageId="15" pageNumber="640" stateProvince="Taichung">Dasyueshan</location>
|
||
logging Rd. 32K,
|
||
<date id="FFA1A8B39F551C46FD30FBE8FD5CFB99" box="[669,764,1114,1133]" pageId="15" pageNumber="640" value="2010-07-06">
|
||
<collectingDate id="EFE5515B9F551C46FD30FBE8FD5CFB99" box="[669,764,1114,1133]" pageId="15" pageNumber="640" value="2010-07-06">6.VII.2010</collectingDate>
|
||
</date>
|
||
, MV lamp,
|
||
<collectorName id="26EAEBA59F551C46FCF7FBE8FC0AFB99" box="[858,938,1114,1133]" pageId="15" pageNumber="640">Y.C. Liao</collectorName>
|
||
leg. (
|
||
<specimenCount id="9D1945FA9F551C46FC75FBE8FC53FB99" box="[984,1011,1108,1133]" pageId="15" pageNumber="640" type="female">1♀</specimenCount>
|
||
<collectionCode id="ED0E16B69F551C46FC55FBE8FB98FB99" box="[1016,1080,1114,1133]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="15" pageNumber="640">NCHU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305DDF89F551C46FFDCFB34FB91FAA9" pageId="15" pageNumber="640" type="diagnosis">
|
||
<paragraph id="8BA08E739F551C46FFDCFB34FB91FAA9" blockId="15.[113,1089,1158,1598]" pageId="15" pageNumber="640">
|
||
<emphasis id="B96B52619F551C46FFDCFB34FF49FB69" bold="true" box="[113,233,1158,1181]" pageId="15" pageNumber="640">Diagnosis.</emphasis>
|
||
Recognized by the following combination of characters: humeri spiniform, yellowish to pale orange, directed subhorizontally, slightly elevated, gradually narrowed, apically obtuse; hind tibia simple, not dilated (J,
|
||
<specimenCount id="9D1945FA9F551C46FD2EFB0DFD36FB2A" box="[643,662,1215,1246]" pageId="15" pageNumber="640" type="female">♀</specimenCount>
|
||
); pregenital abdomen with conspicuous black markings at intersegmental sutures dorsally and ventrally (
|
||
<figureCitation id="132492F69F551C46FCB5FB54FCC8FB09" box="[792,872,1254,1277]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="15" pageNumber="640">Figs 11</figureCitation>
|
||
,
|
||
<figureCitation id="132492F69F551C46FCDFFB54FC30FB09" box="[882,912,1254,1277]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="15" pageNumber="640">52</figureCitation>
|
||
); genital capsule of male with a pair of posteriorly directed, rather straight, apically obtuse lateral projections with inner margins rather straight and enclosing an angle of about 40°, ventral surface rather densely covered by long setae, provided with a distinct tuft of setae apically (
|
||
<figureCitation id="132492F69F551C46FC04FAF4FB84FAA9" box="[937,1060,1350,1373]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="15" pageNumber="640">Figs 11–14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305DDF89F551C59FFDCFAD4FBFBFA0D" lastPageId="16" lastPageNumber="641" pageId="15" pageNumber="640" type="description">
|
||
<paragraph id="8BA08E739F551C59FFDCFAD4FB91FE6D" blockId="15.[113,1089,1158,1598]" lastBlockId="16.[151,1127,226,1593]" lastPageId="16" lastPageNumber="641" pageId="15" pageNumber="640">
|
||
<emphasis id="B96B52619F551C46FFDCFAD4FEF6FA89" bold="true" box="[113,342,1381,1405]" pageId="15" pageNumber="640">
|
||
Description.
|
||
<emphasis id="B96B52619F551C46FEA9FAD7FEF6FA89" bold="true" box="[260,342,1381,1405]" italics="true" pageId="15" pageNumber="640">Colour.</emphasis>
|
||
</emphasis>
|
||
Dorsum and venter rather uniformly stramineous, humeral processes somewhat paler (yellowish to pale orange); abdomen pale yellow, with distinct red suffusion on exposed parts of segments VII (J,
|
||
<specimenCount id="9D1945FA9F551C46FE72FA2DFE52FA4A" box="[479,498,1439,1470]" pageId="15" pageNumber="640" type="female">♀</specimenCount>
|
||
), VIII (
|
||
<specimenCount id="9D1945FA9F551C46FDEEFA2DFDF6FA4A" box="[579,598,1439,1470]" pageId="15" pageNumber="640" type="female">♀</specimenCount>
|
||
), and dorsal surface of basal portion of genital capsule (J); intersegmental sutures between connexival plates of abdominal segments II–VII each enclosed by a black marking extending in whole width of connexivum, each continuous with a marginal marking on the respective ventrites, dorsal parts (mediotergite + connexival plates) of segment VII (J,
|
||
<specimenCount id="9D1945FA9F551C46FE3EF9ADFE06F9CA" box="[403,422,1567,1598]" pageId="15" pageNumber="640" type="female">♀</specimenCount>
|
||
) broadly margined with black posteriorly, lateral portions and median, excised portion of ventrite VII of female broadly margined with black at both sides; tergite VIII of female with a large median, posteriorly narrowing triangular marking confluent with a pair of transverse lateral fasciae along basal margin of tergite VIII; lateral and mesal margins of valvifers VIII of female suffused with black (
|
||
<figureCitation id="132492F69F4A1C59FD59FEF0FCE4FEAD" box="[756,836,322,345]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="16" pageNumber="641">Fig. 52</figureCitation>
|
||
); posterolateral projections of genital capsule of male pale yellowish, inner margins broadly suffused with dark brown; antennal segment I and legs as general body colour, antennal segments II–IV blackish.
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F4A1C59FF18FE10FBC7FDAD" blockId="16.[151,1127,226,1593]" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF18FE10FE73FE4D" bold="true" box="[181,467,417,442]" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF18FE10FE6CFE4E" bold="true" box="[181,460,417,442]" italics="true" pageId="16" pageNumber="641">Integument and vestiture</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body glabrous except of fine, scattered hairs on appendages and terminalia; mandibular plates and interocular area virtually unpunctured; maxillary plate with an obtuse tubercle anteriad and mesad of antennal insertion; pronotum, scutellum and sclerotized parts of fore wings with rather uniform black punctation except on calli, humeral processes and narrow costal margin of fore wing; punctation of exocorium and neighbouring part of endocorium not conspicuously different; ventral surface of body virtually unpunctured.
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F4A1C59FF18FDD0FDDDFC6D" blockId="16.[151,1127,226,1593]" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF18FDD0FEC2FD8D" box="[181,354,610,634]" italics="true" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF18FDD0FE84FD8E" bold="true" box="[181,292,610,634]" italics="true" pageId="16" pageNumber="641">Structure.</emphasis>
|
||
Body
|
||
</emphasis>
|
||
elongate oval with distinctly produced humeri, distance between tips of humeral processes about 1.5 times as long as greatest width of body posteriad of humeri.
|
||
<emphasis id="B96B52619F4A1C59FF3AFD10FF71FD4D" box="[151,209,674,697]" italics="true" pageId="16" pageNumber="641">Head</emphasis>
|
||
1.1–1.25 times as broad as its median length, about 1.5–1.6 times as broad as interocular distance, finely transversely rugose, anterior portion of clypeus depressed, with a broad median longitudinal groove.
|
||
<emphasis id="B96B52619F4A1C59FE7DFD50FD81FD0D" box="[464,545,738,761]" italics="true" pageId="16" pageNumber="641">Labium</emphasis>
|
||
projecting between or slightly surpassing mid coxae.
|
||
<emphasis id="B96B52619F4A1C59FF3AFCB0FEA1FCED" box="[151,257,770,793]" italics="true" pageId="16" pageNumber="641">Pronotum</emphasis>
|
||
with anterolateral margin nearly straight anteriorly, then gradually continued in humeral process, therefore deeply emarginate; humeri spiniform, directed subhorizontally, apically rather obtuse.
|
||
<emphasis id="B96B52619F4A1C59FE2CFCF0FD1CFCAD" box="[385,700,834,857]" italics="true" pageId="16" pageNumber="641">Thoracic pleuron and sternum</emphasis>
|
||
. Mesosternal carina distinctly surpassing base of head, highly elevated, rather broadly rounded anteriorly; metathoracic scent gland ostiole with a long, slightly arched peritreme.
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F4A1C59FF18FC10FD1DFAED" blockId="16.[151,1127,226,1593]" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF18FC10FD93FC4D" box="[181,563,930,953]" italics="true" pageId="16" pageNumber="641">Male and female terminalia. Male</emphasis>
|
||
. Genital capsule (
|
||
<figureCitation id="132492F69F4A1C59FD52FC10FCDCFC4D" box="[767,892,930,953]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="16" pageNumber="641">Figs 11–14</figureCitation>
|
||
) with a pair of long, robust, scissors-like lateral projections directed posteriad, surpassing apex of membrane by their apical halves, lateral margin nearly straight, apically rounded with a tuft of hairs, dorsal rim nearly V-shaped, ventral rim with a pair of rigid and pigmented denticles and a pair of setal tufts laterad of paramere sockets, ventral infolding densely pilose; dorsal wall of projection with pigmented thickening along basal half of dorsal rim; distal portion of lateral projection slightly curved dorsad in lateral view (
|
||
<figureCitation id="132492F69F4A1C59FD53FBD0FCEFFB8D" box="[766,847,1122,1145]" captionStart="Figs 11–19" captionStartId="11.[112,149,1491,1510]" captionTargetBox="[112,1087,503,1436]" captionTargetId="figure-12@11.[112,1087,503,1452]" captionTargetPageId="11" captionText="Figs 11–19. Acanthosoma spp., males. 11 – A. axicia sp. nov., posterior part of abdomen, ventral view; 12 – same, genital capsule, ventral view; 13 – same, lateral view; 14 – same, dorsal view; 15 – A. laevicorne Dallas, 1851, posterior part of abdomen, ventral view; 16 – A. expansum Horváth, 1905, posterior part of abdomen, ventral view; 17 – same, posteroventral view; 18 – same, genital capsule, posterior view; 19 – same, ventral view. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303575" httpUri="https://zenodo.org/record/5303575/files/figure.png" pageId="16" pageNumber="641">Fig. 13</figureCitation>
|
||
). Body of paramere (
|
||
<figureCitation id="132492F69F4A1C59FB9BFBD0FF7DFB6D" captionStart="Figs 20–44" captionStartId="14.[151,188,1196,1215]" captionTargetBox="[151,1126,231,1174]" captionTargetId="figure-84@14.[151,1126,231,1178]" captionTargetPageId="14" captionText="Figs 20–44. Acanthosoma spp., right parameres in different views. 20–21 – A. atayal sp. nov.; 22 – A. haemorrhoidale formosanum subsp. nov., holotype; 23 – A. haemorrhoidale ouchii Ishihara, 1950 (China: Sichuan, Mt. Emei, USNM); 24 – A. haemorrhoidale haemorrhoidale (Linnaeus, 1758) (Spain: Fuente Dé, HNHM); 25–28 – A. asahinai Ishihara, 1943; 29–32 – A. fallax sp. nov.; 33–36 – A. pugnax sp. nov.; 37–39 – A. axicia sp. nov.; 40–41 – A. laevicorne Dallas, 1851; 42–44 – A. expansum Horváth, 1905. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303577" httpUri="https://zenodo.org/record/5303577/files/figure.png" pageId="16" pageNumber="641">Figs 33–36</figureCitation>
|
||
) L-shaped, apex transversely bifurcate, anchor-shaped.
|
||
<emphasis id="B96B52619F4A1C59FC93FB30FC2FFB6D" box="[830,911,1154,1177]" italics="true" pageId="16" pageNumber="641">Female</emphasis>
|
||
(
|
||
<figureCitation id="132492F69F4A1C59FC32FB30FC51FB6D" box="[927,1009,1154,1177]" captionStart="Figs 52–55" captionStartId="18.[151,188,1522,1541]" captionTargetBox="[151,1124,840,1491]" captionTargetId="figure-10@18.[151,1126,839,1496]" captionTargetPageId="18" captionText="Figs 52–55. Acanthosoma spp., female terminalia, posteroventral view. 52 – A. axicia sp. nov.; 53 – A. pugnax sp. nov.; 54 – A. laevicorne Dallas, 1851; 55 – A. expansum Horváth, 1905. PO6, PO7 = Pendergrast’s organs of ventrites VI and VII, respectively. Scales in mm." figureDoi="http://doi.org/10.5281/zenodo.5303581" httpUri="https://zenodo.org/record/5303581/files/figure.png" pageId="16" pageNumber="641">Fig. 52</figureCitation>
|
||
). Posterior margin of ventrite VII with a deep, shield-shaped median incision margined with black, surrounding valvifers VIII; posterior margin of laterotergites VIII broadly arcuate, posteriorly far surpassing posterolateral angles of abdominal ventrite VII; Pendergrast’s organs of ventrites VI and VII subelliptical, of equal size.
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F4A1C59FF18FA90FBFBFA0D" blockId="16.[151,1127,226,1593]" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF18FA90FEF5FACD" bold="true" box="[181,341,1314,1337]" italics="true" pageId="16" pageNumber="641">Measurements</emphasis>
|
||
(3JJ
|
||
<specimenCount id="9D1945FA9F4A1C59FE0DFA90FE76FACE" box="[416,470,1307,1338]" pageId="16" pageNumber="641" type="female">2♀♀</specimenCount>
|
||
) (in mm). Body length from apex of head to apex of membrane 13.7–14.2 (J) / 15.3–15.6 (
|
||
<specimenCount id="9D1945FA9F4A1C59FE6FFA89FE75FAAE" box="[450,469,1339,1370]" pageId="16" pageNumber="641" type="female">♀</specimenCount>
|
||
), from apex of head to line connecting apices of projections of genital capsule 15.7–17.1 (J); greatest width of body posteriad of humeral processes 6.70–7.60; median length of head 1.90–2.36, width across eyes 2.45–2.60, interocular distance 1.55–1.75; length of antennal segments (I) 1.50–1.70: (IIa) 1.75–1.93: (IIb) 1.55–1.70: (III) 2.30–2.40: (IV) 1.95–2.00; median length of pronotum 2.90–3.45, greatest width (across tips of humeri) 10.3–11.6; median length of scutellum 4.25–4.75, width at base 3.85–4.50.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305DDF89F4A1C58FF3AF9B0FE86FEFB" lastPageId="17" lastPageNumber="642" pageId="16" pageNumber="641" type="etymology">
|
||
<paragraph id="8BA08E739F4A1C59FF3AF9B0FC26F9CD" blockId="16.[151,1127,226,1593]" pageId="16" pageNumber="641">
|
||
<emphasis id="B96B52619F4A1C59FF3AF9B0FEB8F9ED" bold="true" box="[151,280,1538,1561]" pageId="16" pageNumber="641">Etymology.</emphasis>
|
||
The specific epithet is the Latin noun
|
||
<taxonomicName id="4C1FF5F09F4A1C59FD1DF9B0FD51F9ED" authorityName="Tsai & Rédei" authorityYear="2015" box="[688,753,1538,1561]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="641" phylum="Arthropoda" rank="species" species="axicia">
|
||
<emphasis id="B96B52619F4A1C59FD1DF9B0FD51F9ED" box="[688,753,1538,1561]" italics="true" pageId="16" pageNumber="641">axicia</emphasis>
|
||
</taxonomicName>
|
||
(‘a pair of scissors’) in apposition, referring to the shape of the genital capsule; ending not to be changed.
|
||
</paragraph>
|
||
<paragraph id="8BA08E739F4B1C58FFDCFF6AFE86FEFB" blockId="17.[113,1089,216,463]" pageId="17" pageNumber="642">
|
||
<emphasis id="B96B52619F4B1C58FFDCFF6AFF4EFF1B" bold="true" box="[113,238,216,239]" pageId="17" pageNumber="642">Bionomics.</emphasis>
|
||
Some of the examined adults were collected from
|
||
<taxonomicName id="4C1FF5F09F4B1C58FD54FF6AFE82FEFB" authority="Makino" authorityName="Makino" class="Magnoliopsida" family="Pittosporaceae" genus="Pittosporum" kingdom="Plantae" order="Apiales" pageId="17" pageNumber="642" phylum="Tracheophyta" rank="species" species="illicioides">
|
||
<emphasis id="B96B52619F4B1C58FD54FF6AFC49FF1B" box="[761,1001,216,239]" italics="true" pageId="17" pageNumber="642">Pittosporum illicioides</emphasis>
|
||
Makino (Pittosporaceae)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305DDF89F4B1C58FFDCFEAAFE9DFE9B" pageId="17" pageNumber="642" type="distribution">
|
||
<paragraph id="8BA08E739F4B1C58FFDCFEAAFE9DFE9B" blockId="17.[113,1089,216,463]" pageId="17" pageNumber="642">
|
||
<emphasis id="B96B52619F4B1C58FFDCFEAAFEA4FEDB" bold="true" box="[113,260,280,303]" pageId="17" pageNumber="642">Distribution.</emphasis>
|
||
The species is apparently endemic to
|
||
<collectingCountry id="F308CEE39F4B1C58FD0EFEAAFD55FEDB" box="[675,757,280,303]" name="Taiwan" pageId="17" pageNumber="642">Taiwan</collectingCountry>
|
||
. The few specimens which we could access were collected in primary forests of high mountains at an altitude of about
|
||
<quantity id="4CE723969F4B1C58FFDCFEEAFEA5FE9B" box="[113,261,344,367]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.5" metricValueMin="1.8" pageId="17" pageNumber="642" unit="m" value="2150.0" valueMax="2500.0" valueMin="1800.0">1800–2500 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C305DDF89F4B1C5BFFDCFECAFE05FE9B" lastPageId="18" lastPageNumber="643" pageId="17" pageNumber="642" type="discussion">
|
||
<paragraph id="8BA08E739F4B1C5BFFDCFECAFE05FE9B" blockId="17.[113,1089,216,463]" lastBlockId="18.[151,1127,216,367]" lastPageId="18" lastPageNumber="643" pageId="17" pageNumber="642">
|
||
<emphasis id="B96B52619F4B1C58FFDCFECAFF40FE7B" bold="true" box="[113,224,376,399]" pageId="17" pageNumber="642">Remarks.</emphasis>
|
||
Although the relatively elongate posterolateral projections of the genital capsule superficially resemble the condition found in members of the
|
||
<taxonomicName id="4C1FF5F09F4B1C58FCBBFE2AFCD3FE5B" authorityName="Dallas" authorityYear="1851" box="[790,883,408,431]" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="642" phylum="Arthropoda" rank="species" species="forfex">
|
||
<emphasis id="B96B52619F4B1C58FCBBFE2AFCD3FE5B" box="[790,883,408,431]" italics="true" pageId="17" pageNumber="642">A. forfex</emphasis>
|
||
</taxonomicName>
|
||
species group, the similarity is only apparent. The narrow, apically rather pointed, dosally unpunctured head, the shape and direction of the humeral process, and particularly the shape of the terminalia of both sexes (posterolateral projections of genital capsule of male robust, with flattened, densely pilose ventral infolding, body of paramere L-shaped, median excision of the posterior margin of ventrite VII of female broad) suggest that the new species is related to
|
||
<taxonomicName id="4C1FF5F09F481C5BFC7BFE8AFE01FE9B" authority="(Walker, 1868)" baseAuthorityName="Walker" baseAuthorityYear="1868" class="Insecta" family="Acanthosomatidae" genus="Acanthosoma" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="643" phylum="Arthropoda" rank="species" species="firmatum">
|
||
<emphasis id="B96B52619F481C5BFC7BFE8AFF58FE9B" italics="true" pageId="18" pageNumber="643">Acanthosoma firmatum</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8EF3829F481C5BFEA8FEEAFE39FE9B" author="WALKER F." box="[261,409,344,367]" pageId="18" pageNumber="643" pagination="419 - 599" refId="ref21358" refString="WALKER F. 1868: Catalogue of the specimens of Hemiptera Heteroptera in the collection of the British Museum. Part III. Trustees of the British Museum, London, pp. 419 - 599." type="book chapter" year="1868">Walker, 1868</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |