285 lines
36 KiB
XML
285 lines
36 KiB
XML
<document id="199A8F56E88B261214F2161E873B1AFF" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629576139757" checkinUser="felipe" docAuthor="McAlpine, DK" docDate="2007" docId="03FA02406A6A651B379849A5BC9FD5E8" docLanguage="en" docName="RecAustMus.59.3.143-219.pdf" docOrigin="Records of the Australian Museum 59" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Borboroides menura McAlpine 2007, n.sp." docType="treatment" docVersion="2" lastPageNumber="193" masterDocId="FFC37A386A456529342A4C0DBD22D171" masterDocTitle="Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with Reconsideration of the Status of Families Heleomyzidae and Sphaeroceridae, and Descriptions of Femoral Gland-baskets" masterLastPageNumber="219" masterPageNumber="143" pageNumber="190" updateTime="1699078874016" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods id="0EB602FE78A81C6992F4E18AFC2519AE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="AFD85D0B4B5DCA615A8CCAF198719326">
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<mods:title id="B67CB92BADAA6BF623C722291CDC25F1">Review of the Borboroidini or Wombat Flies (Diptera: Heteromyzidae), with Reconsideration of the Status of Families Heleomyzidae and Sphaeroceridae, and Descriptions of Femoral Gland-baskets</mods:title>
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</mods:titleInfo>
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<mods:name id="A0F44CC601F6A1D03FC75C601EF8DB10" type="personal">
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<mods:role id="E29A23719468D1A1765B7C4F449E5343">
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<mods:roleTerm id="4477CE251B59472A5AE5AAC0E2798621">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EE5AB1AFFAC01795BEBA59A3887F4507">McAlpine, DK</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="175087A949BFC11BE6A4371E130DCAFD">text</mods:typeOfResource>
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<mods:relatedItem id="73C90B70A2A6F1617478C03B3497D705" type="host">
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<mods:titleInfo id="3708BADAB70B151368E3D51431364A96">
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<mods:title id="E48B08C07AC7538C3CDF4A9CB2850A1C">Records of the Australian Museum</mods:title>
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</mods:titleInfo>
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<mods:part id="3D915D837A0161A7D56D186A2F87E9C2">
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<mods:date id="4DEFC824978E26FBFF936B286192C4F7">2007</mods:date>
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<mods:detail id="FAD250A30252892D35B29688A4737B51" type="volume">
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<mods:number id="7D2C72B5A973D10DAAAB9BE4ED5971CF">59</mods:number>
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</mods:detail>
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<mods:extent id="E0DF65FBD87405A5FA7CBCDD392063AC" unit="page">
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<mods:start id="F8DE7F61EDA149866B395BB1F013C224">143</mods:start>
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<mods:end id="37EE253B4E53BEDE5C923D1AC2559EF7">219</mods:end>
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<mods:classification id="84AA864ED5F5B7365F4B19DBED662183">journal article</mods:classification>
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<mods:identifier id="6EF881DDFDDCFBE271535061B082D2B8" type="ISSN">2201-4349</mods:identifier>
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</mods:mods>
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<treatment id="03FA02406A6A651B379849A5BC9FD5E8" LSID="urn:lsid:plazi:treatment:03FA02406A6A651B379849A5BC9FD5E8" httpUri="http://treatment.plazi.org/id/03FA02406A6A651B379849A5BC9FD5E8" lastPageId="50" lastPageNumber="193" pageId="47" pageNumber="190">
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<subSubSection id="C349E0DD6A6A6506379849A5B9D7D4B3" box="[946,1269,1448,1474]" pageId="47" pageNumber="190" type="nomenclature">
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<paragraph id="8BECB3566A6A6506379849A5B9D7D4B3" blockId="47.[946,1269,1448,1474]" box="[946,1269,1448,1474]" pageId="47" pageNumber="190">
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<heading id="D0A4043A6A6A6506379849A5B9D7D4B3" box="[946,1269,1448,1474]" centered="true" fontSize="11" level="2" pageId="47" pageNumber="190" reason="2">
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<emphasis id="B9276F446A6A6506379849A5B9D7D4B3" bold="true" box="[946,1269,1448,1474]" pageId="47" pageNumber="190">
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<taxonomicName id="4C53C8D56A6A6506379849A5B993D4B3" ID-CoL="68PB4" authority="McAlpine, 2007" authorityName="McAlpine" authorityYear="2007" box="[946,1201,1448,1474]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="menura" status="sp. nov.">
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<emphasis id="B9276F446A6A6506379849A5B993D4B3" bold="true" box="[946,1201,1448,1474]" italics="true" pageId="47" pageNumber="190">Borboroides menura</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A214D23F6A6A6506309349A5B9D7D4B3" box="[1209,1269,1448,1474]" pageId="47" pageNumber="190" rank="species">n.sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E0DD6A6A650637C149EBB99ED771" box="[1003,1212,1510,1536]" pageId="47" pageNumber="190" type="description">
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<paragraph id="8BECB3566A6A650637C149EBB99ED771" blockId="47.[1003,1212,1510,1536]" box="[1003,1212,1510,1536]" pageId="47" pageNumber="190">
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<figureCitation id="1368AFD36A6A650637C149EBB977D771" box="[1003,1109,1510,1536]" captionStart="Figs 7–10" captionStartId="6.[166,207,1047,1068]" captionTargetBox="[166,1466,151,1034]" captionTargetId="figure-286@6.[166,1466,151,1039]" captionTargetPageId="6" captionText="Figs 7–10. (7) Borboroides menura, part of posteroventral surface of hind femur, showing many gland-baskets; scale = 3 µm. (8) Detail of same gland-baskets, showing many microtrichia cohering through secretion; scale = 3 µm. (9) B. gorodkovi, part of hind femur showing posteroventral tract of convergent microtrichia covering glandular groove; scale = 10 µm. (10) Detail of glandular groove of same showing macrotrichia (socket-based) and microtrichia of various types; scale = 3 µm." pageId="47" pageNumber="190">Figs 7, 8</figureCitation>
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,
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<figureCitation id="1368AFD36A6A6506304949EBB99ED771" box="[1123,1212,1510,1536]" captionStart-0="Figs 97–98" captionStart-1="Figs 99–101" captionStartId-0="48.[1027,1069,1815,1836]" captionStartId-1="49.[121,162,1521,1542]" captionTargetBox-0="[166,1468,1005,1996]" captionTargetBox-1="[165,1274,174,1498]" captionTargetId-0="figure-10@48.[166,1466,967,2003]" captionTargetId-1="figure-13@49.[121,1421,151,1515]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionText-0="Figs 97–98. (97) Borboroides menura, male postabdomen, right lateral view, setulae omitted from tergites and sternites; scale = 0.2 mm. (98) Wing of same. c, cercus; e, epandrium; g, gonite; ss, surstylus; s5–s8, sternites 5 to 8; t5, t6, tergites 5 and 6; vb, ventral bridge of sternite 8." captionText-1="Figs 99–101. (99) Borboroides menura, epandrial and some hypandrial structures, anteroventral view, distiphallus and macrotrichia of epandrium and surstyli omitted; scale = 0.1 mm. (100) Left surstylus of same, anterior view; scale = 0.1 mm. (101) Distiphallus of same, anterodistal view; scale = 0.05 mm. aa, aedeagal apodeme (seen through prehypandrial membrane); af, anterior foramen of epandrium; bc, fused basal body of cerci; bp, basiphallus; dc, free distal lobe of cercus; dp, distiphallus; g, gonite; pp, prehypandrial process; ss, surstylus." pageId="47" pageNumber="190">97–101</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C349E0DD6A6A650637304A2FBE56D655" pageId="47" pageNumber="190" type="materials_examined">
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<paragraph id="8BECB3566A6A650637304A2FBE56D655" blockId="47.[794,1422,1569,1828]" pageId="47" pageNumber="190">
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<emphasis id="B9276F446A6A650637304A2FBED8D74B" bold="true" box="[794,1018,1570,1594]" pageId="47" pageNumber="190">Material examined</emphasis>
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.
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<materialsCitation id="3B3BB90B6A6A650630264A2FB8A8D7E0" collectingDate="2005-09-22" collectionCode="AM" collectorName="DK McAlpine & Near & Mounted" elevation="1200" location="Kanangra-Boyd National Park" pageId="47" pageNumber="190" specimenCode="K219753" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
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<typeStatus id="54E80DF46A6A650630264A2FB9A1D748" box="[1036,1155,1570,1593]" pageId="47" pageNumber="190" type="holotype">HOLOTYPE</typeStatus>
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.?,
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<collectingRegion id="49977DB46A6A650630964A2FB8A8D748" box="[1212,1418,1570,1593]" country="Australia" name="New South Wales" pageId="47" pageNumber="190">New South Wales</collectingRegion>
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:
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<location id="8E8CE58D6A6A650637304A4DB954D726" LSID="urn:lsid:plazi:treatment:03FA02406A6A651B379849A5BC9FD5E8:8E8CE58D6A6A650637304A4DB954D726" box="[794,1142,1600,1623]" name="Kanangra-Boyd National Park" pageId="47" pageNumber="190" stateProvince="New South Wales">Kanangra-Boyd National Park</location>
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,
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<locationDeviation id="9A5EAA746A6A650630AE4A4DB8AFD726" box="[1156,1421,1600,1623]" location="Boyd River" pageId="47" pageNumber="190">
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<quantity id="4CAB1EB36A6A650630AE4A4DB9E2D726" box="[1156,1216,1600,1623]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="47" pageNumber="190" unit="km" value="1.0">1 km</quantity>
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N of Boyd River
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</locationDeviation>
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[or Morong Creek] bridge,
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<quantity id="4CAB1EB36A6A650630654A50B984D705" box="[1103,1190,1629,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="47" pageNumber="190" unit="m" value="1200.0">
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<elevation id="007E54656A6A650630654A50B984D705" box="[1103,1190,1629,1652]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="47" pageNumber="190" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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,
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<date id="FFED95966A6A6506309F4A50B809D705" box="[1205,1323,1629,1652]" pageId="47" pageNumber="190" value="2005-09-22">
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<collectingDate id="EFA96C7E6A6A6506309F4A50B809D705" box="[1205,1323,1629,1652]" pageId="47" pageNumber="190" value="2005-09-22">22.ix.2005</collectingDate>
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</date>
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,
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<collectorName id="26A6D6806A6A650631104A50B8ACD705" box="[1338,1422,1629,1652]" name="DK McAlpine" pageId="47" pageNumber="190">D.K.M.</collectorName>
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(
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<collectionCode id="ED422B936A6A6506370B4A77BE6CD7E0" box="[801,846,1658,1681]" country="Australia" name="Australian Museum" pageId="47" pageNumber="190" type="Museum">AM</collectionCode>
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<specimenCode id="DBF51B2D6A6A6506377F4A77BE9FD7E0" box="[853,957,1658,1681]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="47" pageNumber="190" type="Museum">K219753</specimenCode>
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).
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<collectorName id="26A6D6806A6A650637E44A77B920D7E0" box="[974,1026,1658,1681]" pageId="47" pageNumber="190">Near</collectorName>
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dead chook.
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<collectorName id="26A6D6806A6A650630B84A77B9D0D7E0" box="[1170,1266,1658,1681]" pageId="47" pageNumber="190">Mounted</collectorName>
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on card point
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</materialsCitation>
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.
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<materialsCitation id="3B3BB90B6A6A650637304A95B833D7BD" collectingDate="2005-10" collectionCode="AM, USNM" collectorName="DK McAlpine & Near & Mounted" elevation="1200" location="Kanangra-Boyd National Park" pageId="47" pageNumber="190" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
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<typeStatus id="54E80DF46A6A650637304A95BEB7D7DF" box="[794,917,1688,1711]" pageId="47" pageNumber="190" type="paratype">PARATYPES</typeStatus>
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.
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||
<collectingRegion id="49977DB46A6A650637894A95B945D7DE" box="[931,1127,1688,1711]" country="Australia" name="New South Wales" pageId="47" pageNumber="190">New South Wales</collectingRegion>
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: 9??, 2!!, same data as holotype, but some
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<date id="FFED95966A6A650637D34AB8B945D7BD" box="[1017,1127,1717,1740]" pageId="47" pageNumber="190" value="2005-10">
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<collectingDate id="EFA96C7E6A6A650637D34AB8B945D7BD" box="[1017,1127,1717,1740]" pageId="47" pageNumber="190" value="2005-10">Oct. 2005</collectingDate>
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</date>
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||
(
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<collectionCode id="ED422B936A6A650630524AB8B985D7BD" box="[1144,1191,1717,1740]" country="Australia" name="Australian Museum" pageId="47" pageNumber="190" type="Museum">AM</collectionCode>
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,
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<collectionCode id="ED422B936A6A6506309C4AB8B82BD7BD" box="[1206,1289,1717,1740]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="47" pageNumber="190" type="Museum">USNM</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B3BB90B6A6A650631364AB8BE52D655" collectingDate="1978-04" collectionCode="ANIC" elevation="800" latitude="-35.366665" location="Picadilly Circus" longLatPrecision="1193" longitude="148.81667" pageId="47" pageNumber="190" specimenCount="1" stateProvince="Australian Capital Territory" typeStatus="paratype">
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<collectingRegion id="49977DB46A6A650631364AB8BEEDD798" country="Australia" name="Australian Capital Territory" pageId="47" pageNumber="190">Australian Capital Territory</collectingRegion>
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: 1?,
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<location id="8E8CE58D6A6A6506302D4ADFB990D798" LSID="urn:lsid:plazi:treatment:03FA02406A6A651B379849A5BC9FD5E8:8E8CE58D6A6A6506302D4ADFB990D798" box="[1031,1202,1746,1769]" latitude="-35.366665" longLatPrecision="1193" longitude="148.81667" name="Picadilly Circus" pageId="47" pageNumber="190" stateProvince="Australian Capital Territory">Picadilly Circus</location>
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[Brindabella Range,
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<geoCoordinate id="EE67D5916A6A650637304AFDBE50D676" box="[794,882,1776,1799]" degrees="35" direction="south" minutes="22" orientation="latitude" pageId="47" pageNumber="190" precision="925" value="-35.366665">35°22'S</geoCoordinate>
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<geoCoordinate id="EE67D5916A6A650637574AFDBEC8D676" box="[893,1002,1776,1799]" degrees="148" direction="east" minutes="49" orientation="longitude" pageId="47" pageNumber="190" precision="925" value="148.81667">148°49'E</geoCoordinate>
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],
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<quantity id="4CAB1EB36A6A6506302A4AFDB96FD676" box="[1024,1101,1776,1799]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="190" unit="m" value="800.0">
|
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<elevation id="007E54656A6A6506302A4AFDB96FD676" box="[1024,1101,1776,1799]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="190" unit="m" value="800.0">800 m</elevation>
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||
</quantity>
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||
,
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<date id="FFED95966A6A650630714AFDB9FFD676" box="[1115,1245,1776,1799]" pageId="47" pageNumber="190" value="1978-04">
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||
<collectingDate id="EFA96C7E6A6A650630714AFDB9FFD676" box="[1115,1245,1776,1799]" pageId="47" pageNumber="190" value="1978-04">April 1978</collectingDate>
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||
</date>
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||
, S.B.P., J.K.-P. (
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<collectionCode id="ED422B936A6A6506370B4B00BE4AD655" box="[801,872,1805,1828]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="47" pageNumber="190">ANIC</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C349E0DD6A6A651837304B4AB9EAD62F" lastPageId="49" lastPageNumber="192" pageId="47" pageNumber="190" type="description">
|
||
<paragraph id="8BECB3566A6A650637304B4AB85AD60D" blockId="47.[794,1421,1863,2004]" pageId="47" pageNumber="190">
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||
<emphasis id="B9276F446A6A650637304B4ABEBED62E" bold="true" box="[794,924,1863,1887]" pageId="47" pageNumber="190">Description</emphasis>
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(male, female). Small, slightly elongate, largely black fly, slightly larger than other species of
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<taxonomicName id="4C53C8D56A6A6506312A4B68B80ED60D" box="[1280,1324,1893,1916]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="47" pageNumber="190" phylum="Arthropoda" rank="species" species="atra">
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||
<emphasis id="B9276F446A6A6506312A4B68B80ED60D" box="[1280,1324,1893,1916]" italics="true" pageId="47" pageNumber="190">atra</emphasis>
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</taxonomicName>
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group.
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||
</paragraph>
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||
<paragraph id="8BECB3566A6A651937124B8EBC1BD33D" blockId="47.[794,1421,1863,2004]" lastBlockId="48.[166,795,154,1614]" lastPageId="48" lastPageNumber="191" pageId="47" pageNumber="190">
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||
<emphasis id="B9276F446A6A650637124B8EBE8DD6EB" box="[824,943,1923,1946]" italics="true" pageId="47" pageNumber="190">Coloration</emphasis>
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. Head largely black; face, anterior part of cheek, and anterior part of postfrons yellow in male, tawnybrown in female; postfrons pruinescent on rather more than anterior half, with large bare, shining to subshining zone on each side of ocelli and quite small subtriangular glossy zone in front of anterior ocellus. Antenna tawny-yellow, with arista and dorsal suffusion on segment 3 brown. Prelabrum brown in male, black in female; palpus yellowish brown to dark brown. Thorax normally black (fading to brown in old specimens); mesoscutum and scutellum evenly pruinescent; mesopleuron pruinescent on c. posterior half and on dorsal margin; sternopleuron pruinescent on dorsal margin; hypopleuron pruinescent both anteriorly and posteriorly. Legs largely blackish brown; fore coxa partly tawny-brown. Wing unmarked. Halter tawny, with pale yellow capitellum. Abdominal tergites black with brown pruinescence; tergite 5 of female shining, with generally distributed sparse pruinescence.
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</paragraph>
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||
<paragraph id="8BECB3566A75651934EF4E5FBC40D225" blockId="48.[166,795,154,1614]" pageId="48" pageNumber="191">
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||
<emphasis id="B9276F446A75651934EF4E5FBDDDD318" box="[197,255,594,617]" italics="true" pageId="48" pageNumber="191">Head</emphasis>
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||
. Eye c. as long as high or slightly longer; postfrons with numerous short setulae anteriorly; height of cheek c. 0.33–0.47 of height of eye; ocellar bristle slightly longer than posterior fronto-orbital bristle; anterior fronto-orbital bristle c. as long as posterior one, directed outwards. Antenna: segment 3 slightly longer than deep; arista c. as long as greatest diameter of eye. Prelabrum small and shallow in male, larger and more prominent in female; palpus moderately short.
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</paragraph>
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<paragraph id="8BECB3566A75651934EF4F57B854D035" blockId="48.[166,795,154,1614]" lastBlockId="48.[839,1467,154,969]" pageId="48" pageNumber="191">
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||
<emphasis id="B9276F446A75651934EF4F57BC30D200" box="[197,274,858,881]" italics="true" pageId="48" pageNumber="191">Thorax</emphasis>
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. Mesoscutum with intradorsocentral setulae on anterior half in two pairs of rows, with few in outer row; mesopleuron with c. four setulae on or near posterior margin; dorsocentral bristles two, length of anterior one c. 0.8 of that of posterior one; posterior intra-alar bristle moderately curved, slightly diverging from vein 4 apically; apical section of vein 4 2.1–2.3 times as long as penultimate section; anterior crossvein meeting vein 4 well before mid-length of discal cell; basal crossvein oblique; anal crossvein oblique, almost straight, but curved at anterior end; distal section of vein 6 as long as anal crossvein or somewhat longer.
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</paragraph>
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<paragraph id="8BECB3566A756518374F4D46BE4ED677" blockId="48.[839,1467,154,969]" lastBlockId="49.[794,1421,1716,1887]" lastPageId="49" lastPageNumber="192" pageId="48" pageNumber="191">
|
||
<emphasis id="B9276F446A756519374F4D46BEEAD013" box="[869,968,331,354]" italics="true" pageId="48" pageNumber="191">Abdomen</emphasis>
|
||
. Sternite 1 well sclerotized and undivided. Male: tergite 5 nearly as large as tergite 4; sternites 4 and 5 short, very broad, medially attenuated. Male postabdomen: tergite 6 narrowly transverse, undivided; compound protandrial sclerite rather short for
|
||
<taxonomicName id="4C53C8D56A75651930744DCDB9AED0A6" box="[1118,1164,448,471]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="191" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9276F446A75651930744DCDB9AED0A6" box="[1118,1164,448,471]" italics="true" pageId="48" pageNumber="191">atra</emphasis>
|
||
</taxonomicName>
|
||
group, anteriorly strongly asymmetrical, with sternite 6 extensively developed on left side and part of ventral surface, but not on right side; sternite 7 sclerotized; sternite 8 bridged ventrally, with bridge contracted on right side; epandrium, including its anteroventral bridge, asymmetrical (
|
||
<figureCitation id="1368AFD36A75651930EC4E5FB836D318" box="[1222,1300,594,617]" captionStart="Figs 99–101" captionStartId="49.[121,162,1521,1542]" captionTargetBox="[165,1274,174,1498]" captionTargetId="figure-13@49.[121,1421,151,1515]" captionTargetPageId="49" captionText="Figs 99–101. (99) Borboroides menura, epandrial and some hypandrial structures, anteroventral view, distiphallus and macrotrichia of epandrium and surstyli omitted; scale = 0.1 mm. (100) Left surstylus of same, anterior view; scale = 0.1 mm. (101) Distiphallus of same, anterodistal view; scale = 0.05 mm. aa, aedeagal apodeme (seen through prehypandrial membrane); af, anterior foramen of epandrium; bc, fused basal body of cerci; bp, basiphallus; dc, free distal lobe of cercus; dp, distiphallus; g, gonite; pp, prehypandrial process; ss, surstylus." pageId="48" pageNumber="191">Fig. 99</figureCitation>
|
||
); surstylus with two lobes, one directed distally, one directed posteriorly, both lobes with few setulae, distal lobe, in addition, heavily spinulose on inner surface, some of larger spinules distally compressed and bearing two to four rounded apical teeth; basal section of surstylus densely pubescent on inner surface; prehypandrial membrane rugose, without microtrichia; hypandrium with pair of narrowly bifid posteriorly directed lobes; a pair of prominent elongate processes arising from prehypandrial membrane where it borders on hypandrium, each minutely bifid and bearing two fine hair-like apical projections; basiphallus very short, sclerotized at least on posterior surface; distiphallus more elongate, consisting of three main irregularly bulbous sections, not strongly sclerotized but its basal and mid sections with many subparallel denticulate sclerotized ridges; distal section of aedeagus largely smooth, transparent, bearing beaklike terminal process with apical gonopore; cerci basally articulated with epandrium, their bases fused to form a stout body bearing a diversity of setulae and, on its posterodistal surface, a large process, forked from just beyond its base to form pair of very long, compressed, distally slightly spatulate lobes, which remain exposed when genital complex is infolded. Female postabdomen: segments 6 and 7 not attenuated, with undivided tergites and sternites; cercus not clavate.
|
||
</paragraph>
|
||
<caption id="DF2CE3DE6A75651930294B1AB959D6BD" pageId="48" pageNumber="191" startId="48.[1027,1069,1815,1836]" targetBox="[166,1468,1005,1996]" targetPageId="48">
|
||
<paragraph id="8BECB3566A75651930294B1AB959D6BD" blockId="48.[1027,1468,1815,1996]" pageId="48" pageNumber="191">
|
||
Figs 97–98. (
|
||
<emphasis id="B9276F446A75651930A34B1AB983D65D" box="[1161,1185,1815,1836]" italics="true" pageId="48" pageNumber="191">97</emphasis>
|
||
)
|
||
<taxonomicName id="4C53C8D56A75651930984B1AB85ED65D" authorityName="McAlpine" authorityYear="2007" box="[1202,1404,1815,1836]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="48" pageNumber="191" phylum="Arthropoda" rank="species" species="menura">
|
||
<emphasis id="B9276F446A75651930984B1AB85ED65D" box="[1202,1404,1815,1836]" italics="true" pageId="48" pageNumber="191">Borboroides menura</emphasis>
|
||
</taxonomicName>
|
||
, male postabdomen, right lateral view, setulae omitted from tergites and sternites; scale = 0.2 mm. (
|
||
<emphasis id="B9276F446A75651930214B6AB901D60D" box="[1035,1059,1895,1916]" italics="true" pageId="48" pageNumber="191">98</emphasis>
|
||
) Wing of same.
|
||
<emphasis id="B9276F446A75651930E74B6AB9FAD60D" box="[1229,1240,1895,1916]" italics="true" pageId="48" pageNumber="191">c</emphasis>
|
||
, cercus;
|
||
<emphasis id="B9276F446A756519311E4B6AB81DD60D" box="[1332,1343,1895,1916]" italics="true" pageId="48" pageNumber="191">e</emphasis>
|
||
, epandrium;
|
||
<emphasis id="B9276F446A75651930294B8FB92DD6E6" box="[1027,1039,1922,1943]" italics="true" pageId="48" pageNumber="191">g</emphasis>
|
||
, gonite;
|
||
<emphasis id="B9276F446A75651930404B8FB95FD6E6" box="[1130,1149,1922,1943]" italics="true" pageId="48" pageNumber="191">ss</emphasis>
|
||
, surstylus;
|
||
<emphasis id="B9276F446A75651930D84B8FB808D6E6" box="[1266,1322,1922,1943]" italics="true" pageId="48" pageNumber="191">s5–s8</emphasis>
|
||
, sternites 5 to 8;
|
||
<emphasis id="B9276F446A75651930344B91B970D6C0" box="[1054,1106,1948,1969]" italics="true" pageId="48" pageNumber="191">t5, t6</emphasis>
|
||
, tergites 5 and 6;
|
||
<emphasis id="B9276F446A75651931264B91B801D6C0" box="[1292,1315,1948,1969]" italics="true" pageId="48" pageNumber="191">vb</emphasis>
|
||
, ventral bridge of sternite 8.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2CE3DE6A746518345349FCBDF7D700" pageId="49" pageNumber="192" startId="49.[121,162,1521,1542]" targetBox="[165,1274,174,1498]" targetPageId="49">
|
||
<paragraph id="8BECB3566A746518345349FCBDF7D700" blockId="49.[121,1422,1521,1649]" pageId="49" pageNumber="192">
|
||
Figs 99–101. (
|
||
<emphasis id="B9276F446A746518352C49FCBC3CD777" box="[262,286,1521,1542]" italics="true" pageId="49" pageNumber="192">99</emphasis>
|
||
)
|
||
<taxonomicName id="4C53C8D56A746518350749FCBCD6D777" authorityName="McAlpine" authorityYear="2007" box="[301,500,1521,1542]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="49" pageNumber="192" phylum="Arthropoda" rank="species" species="menura">
|
||
<emphasis id="B9276F446A746518350749FCBCD6D777" box="[301,500,1521,1542]" italics="true" pageId="49" pageNumber="192">Borboroides menura</emphasis>
|
||
</taxonomicName>
|
||
, epandrial and some hypandrial structures, anteroventral view, distiphallus and macrotrichia of epandrium and surstyli omitted; scale = 0.1 mm. (
|
||
<emphasis id="B9276F446A74651836794A01BF55D750" box="[595,631,1548,1569]" italics="true" pageId="49" pageNumber="192">100</emphasis>
|
||
) Left surstylus of same, anterior view; scale = 0.1 mm. (
|
||
<emphasis id="B9276F446A74651830B84A01B994D750" box="[1170,1206,1548,1569]" italics="true" pageId="49" pageNumber="192">101</emphasis>
|
||
) Distiphallus of same, anterodistal view; scale = 0.05 mm.
|
||
<emphasis id="B9276F446A74651835FC4A2ABCCCD74D" box="[470,494,1575,1596]" italics="true" pageId="49" pageNumber="192">aa</emphasis>
|
||
, aedeagal apodeme (seen through prehypandrial membrane);
|
||
<emphasis id="B9276F446A746518306A4A2AB971D74D" box="[1088,1107,1575,1596]" italics="true" pageId="49" pageNumber="192">af</emphasis>
|
||
, anterior foramen of epandrium;
|
||
<emphasis id="B9276F446A74651834534A4CBDB2D727" box="[121,144,1601,1622]" italics="true" pageId="49" pageNumber="192">bc</emphasis>
|
||
, fused basal body of cerci;
|
||
<emphasis id="B9276F446A74651835884A4CBC98D727" box="[418,442,1601,1622]" italics="true" pageId="49" pageNumber="192">bp</emphasis>
|
||
, basiphallus;
|
||
<emphasis id="B9276F446A746518366B4A4CBF7AD727" box="[577,600,1601,1622]" italics="true" pageId="49" pageNumber="192">dc</emphasis>
|
||
, free distal lobe of cercus;
|
||
<emphasis id="B9276F446A746518374C4A4CBE5CD727" box="[870,894,1601,1622]" italics="true" pageId="49" pageNumber="192">dp</emphasis>
|
||
, distiphallus;
|
||
<emphasis id="B9276F446A74651830224A4CB936D727" box="[1032,1044,1601,1622]" italics="true" pageId="49" pageNumber="192">g</emphasis>
|
||
, gonite;
|
||
<emphasis id="B9276F446A74651830464A4CB9A6D727" box="[1132,1156,1601,1622]" italics="true" pageId="49" pageNumber="192">pp</emphasis>
|
||
, prehypandrial process;
|
||
<emphasis id="B9276F446A746518315E4A4CB8A5D727" box="[1396,1415,1601,1622]" italics="true" pageId="49" pageNumber="192">ss</emphasis>
|
||
, surstylus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BECB3566A74651837124B01B9EAD62F" blockId="49.[794,1421,1716,1887]" pageId="49" pageNumber="192">
|
||
<emphasis id="B9276F446A74651837124B01BE97D652" box="[824,949,1804,1827]" italics="true" pageId="49" pageNumber="192">Dimensions</emphasis>
|
||
. Total length,?
|
||
<quantity id="4CAB1EB36A746518304E4B01B9C6D652" box="[1124,1252,1804,1827]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="49" pageNumber="192" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 mm</quantity>
|
||
,!
|
||
<quantity id="4CAB1EB36A74651831224B01B8ABD652" box="[1288,1417,1804,1827]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="49" pageNumber="192" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1–2.3 mm</quantity>
|
||
; length of thorax,?
|
||
<quantity id="4CAB1EB36A74651837CD4B27B9A0D630" box="[999,1154,1834,1857]" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="8.7" metricValueMin="7.4" pageId="49" pageNumber="192" unit="mm" value="0.8049999999999999" valueMax="0.87" valueMin="0.74">0.74–0.87 mm</quantity>
|
||
,!
|
||
<quantity id="4CAB1EB36A746518308C4B27B860D630" box="[1190,1346,1834,1857]" metricMagnitude="-4" metricUnit="m" metricValue="8.899999999999999" metricValueMax="9.5" metricValueMin="8.299999999999999" pageId="49" pageNumber="192" unit="mm" value="0.8899999999999999" valueMax="0.95" valueMin="0.83">0.83–0.95 mm</quantity>
|
||
; length of wing,?
|
||
<quantity id="4CAB1EB36A74651837BC4B4AB93BD62F" box="[918,1049,1863,1886]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.5" metricValueMin="2.3" pageId="49" pageNumber="192" unit="mm" value="2.4" valueMax="2.5" valueMin="2.3">2.3–2.5 mm</quantity>
|
||
,!
|
||
<quantity id="4CAB1EB36A746518306B4B4AB9E6D62F" box="[1089,1220,1863,1886]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="2.8" metricValueMin="2.7" pageId="49" pageNumber="192" unit="mm" value="2.75" valueMax="2.8" valueMin="2.7">2.7–2.8 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C349E0DD6A74651B37304B8CBC9FD5E8" lastPageId="50" lastPageNumber="193" pageId="49" pageNumber="192" type="distribution">
|
||
<paragraph id="8BECB3566A74651837304B8CBE9AD6A2" blockId="49.[794,1422,1921,2003]" pageId="49" pageNumber="192">
|
||
<emphasis id="B9276F446A74651837304B8CBE87D6E8" bold="true" box="[794,933,1921,1945]" pageId="49" pageNumber="192">Distribution</emphasis>
|
||
.
|
||
<collectingRegion id="49977DB46A746518379E4B8FB958D6E8" box="[948,1146,1922,1945]" country="Australia" name="New South Wales" pageId="49" pageNumber="192">New South Wales</collectingRegion>
|
||
: highlands—few records.
|
||
<collectingRegion id="49977DB46A74651837304B92B96FD6C7" box="[794,1101,1951,1974]" country="Australia" name="Australian Capital Territory" pageId="49" pageNumber="192">Australian Capital Territory</collectingRegion>
|
||
: only known from highlands near Canberra.
|
||
</paragraph>
|
||
<paragraph id="8BECB3566A77651B35964C97BF21D1C5" blockId="50.[444,515,154,180]" box="[444,515,154,180]" pageId="50" pageNumber="193">
|
||
<emphasis id="B9276F446A77651B35964C97BF21D1C5" bold="true" box="[444,515,154,180]" pageId="50" pageNumber="193">Notes</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BECB3566A77651B348C4CDABFBAD2BA" blockId="50.[166,794,215,1177]" pageId="50" pageNumber="193">
|
||
<taxonomicName id="4C53C8D56A77651B348C4CDABCA6D19F" authorityName="McAlpine" authorityYear="2007" box="[166,388,215,238]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="menura">
|
||
<emphasis id="B9276F446A77651B348C4CDABCA6D19F" box="[166,388,215,238]" italics="true" pageId="50" pageNumber="193">Borboroides menura</emphasis>
|
||
</taxonomicName>
|
||
agrees with most species of the
|
||
<taxonomicName id="4C53C8D56A77651B36C44CDABE38D19F" box="[750,794,215,238]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9276F446A77651B36C44CDABE38D19F" box="[750,794,215,238]" italics="true" pageId="50" pageNumber="193">atra</emphasis>
|
||
</taxonomicName>
|
||
group in the long, outwardly directed anterior fronto-orbital bristle, the distribution of pruinescence on the mesopleuron and hypopleuron, the setulose mesopleuron, the short distal section of vein 6, and the well sclerotized undivided abdominal sternite 1. Within this group it is distinguished by its slightly larger size, quite large additional dorsocentral bristle near mid-length of mesoscutum, the different gland complexes on the hind femur, the strong distal curvature of vein 1, the more elongate anal and discal cells so that the discal crossvein is located well beyond mid-length of wing, and, in the male, the development of sternite 6 on the left side only and the much larger, strangely shaped cercus. These conditions for sternite 6 and the femoral gland complexes and the proportions of the anal and discal cells are probably more plesiomorphic than in all other species of the
|
||
<taxonomicName id="4C53C8D56A77651B35634EA1BC57D3B2" box="[329,373,684,707]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9276F446A77651B35634EA1BC57D3B2" box="[329,373,684,707]" italics="true" pageId="50" pageNumber="193">atra</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
<taxonomicName id="4C53C8D56A77651B35E54EA1BF8CD3B2" authorityName="McAlpine" authorityYear="2007" box="[463,686,684,707]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="menura">
|
||
<emphasis id="B9276F446A77651B35E54EA1BF8CD3B2" box="[463,686,684,707]" italics="true" pageId="50" pageNumber="193">Borboroides menura</emphasis>
|
||
</taxonomicName>
|
||
therefore appears to be a phylogenetically basal taxon within this group. The more oblique anal crossvein is probably also plesiomorphic for the group, but the condition is approached in
|
||
<taxonomicName id="4C53C8D56A77651B34E84F2FBC68D248" authorityName="McAlpine" authorityYear="2007" box="[194,330,802,825]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="gorodkovi">
|
||
<emphasis id="B9276F446A77651B34E84F2FBC68D248" box="[194,330,802,825]" italics="true" pageId="50" pageNumber="193">B. gorodkovi</emphasis>
|
||
</taxonomicName>
|
||
, which, with a little variation, has venation intermediate between that of
|
||
<taxonomicName id="4C53C8D56A77651B35FD4F32BF66D227" authorityName="McAlpine" authorityYear="2007" box="[471,580,831,854]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="menura">
|
||
<emphasis id="B9276F446A77651B35FD4F32BF66D227" box="[471,580,831,854]" italics="true" pageId="50" pageNumber="193">B. menura</emphasis>
|
||
</taxonomicName>
|
||
and other species of the
|
||
<taxonomicName id="4C53C8D56A77651B34E64F51BDDAD202" box="[204,248,860,883]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="atra">
|
||
<emphasis id="B9276F446A77651B34E64F51BDDAD202" box="[204,248,860,883]" italics="true" pageId="50" pageNumber="193">atra</emphasis>
|
||
</taxonomicName>
|
||
group. Absence of the usual very short microtrichia on the prehypandrial membrane of the male is a probable autapomorphy for
|
||
<taxonomicName id="4C53C8D56A77651B355E4F9ABCC4D2DF" authorityName="McAlpine" authorityYear="2007" box="[372,486,919,942]" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="menura">
|
||
<emphasis id="B9276F446A77651B355E4F9ABCC4D2DF" box="[372,486,919,942]" italics="true" pageId="50" pageNumber="193">B. menura</emphasis>
|
||
</taxonomicName>
|
||
, though it is repeated in
|
||
<taxonomicName id="4C53C8D56A77651B37294F9ABDC6D2BA" authorityName="McAlpine" authorityYear="2007" class="Insecta" family="Heleomyzidae" genus="Borboroides" kingdom="Animalia" order="Diptera" pageId="50" pageNumber="193" phylum="Arthropoda" rank="species" species="parva">
|
||
<emphasis id="B9276F446A77651B37294F9ABDC6D2BA" italics="true" pageId="50" pageNumber="193">B. parva</emphasis>
|
||
</taxonomicName>
|
||
, which appears not to be closely related.
|
||
</paragraph>
|
||
<paragraph id="8BECB3566A77651B34EF4FDFBCEED552" blockId="50.[166,794,215,1177]" pageId="50" pageNumber="193">The specimen from Picadilly Circus is labelled “carrion trap.” The other specimens were collected on or near old carcasses of domestic fowl.</paragraph>
|
||
<paragraph id="8BECB3566A77651B34EF4827BC9FD5E8" blockId="50.[166,794,215,1177]" pageId="50" pageNumber="193">The specific epithet (Greek, mighty tail) is the generic name of the lyre-bird, in reference to the similarity of the male cercal lobes to the major male tail feathers of that sympatric Australian bird.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |