424 lines
51 KiB
XML
424 lines
51 KiB
XML
<document id="37FFF12B235D29D62E31D386CFD411A3" ID-DOI="10.5281/zenodo.13154942" ID-ISSN="0068-547X" ID-Zenodo-Dep="13154942" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722537956198" checkinUser="felipe" docAuthor="Daniel, Thomas F." docDate="2015" docId="03ECC23AC45BB1067C8DFAAFFD0B851A" docLanguage="en" docName="ProcCalifAcadSci.62.19.451-455.pdf" docOrigin="Proceedings of the California Academy of Sciences 62 (19)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Anisotes tablensis T. F. Daniel 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="454" masterDocId="FFD5BA42C45BB1057D3FFF97FFD58321" masterDocTitle="Anisotes tablensis (Acanthaceae), a New Species from Southwestern Madagascar" masterLastPageNumber="455" masterPageNumber="451" pageNumber="451" updateTime="1722543696938" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="127F199BF154FC4350B6E9AE54D9D02E">
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<mods:title id="38798DE486AD9C9AAF6B46B887C8BD17">Anisotes tablensis (Acanthaceae), a New Species from Southwestern Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="D06EEAC096FACAC7DE1303C0A355C2DE" type="personal">
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<mods:roleTerm id="444DB9475B4DA5136AA685919A5CE118">Author</mods:roleTerm>
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<mods:namePart id="ED9AA2B64F590D67134D45D0929BA47F">Daniel, Thomas F.</mods:namePart>
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<mods:typeOfResource id="4814258945C574A624001A716FF60D7F">text</mods:typeOfResource>
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<mods:title id="2FFB0BF67175D62F99AF051822B495B1">Proceedings of the California Academy of Sciences</mods:title>
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</mods:titleInfo>
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<mods:part id="A47F612E154C04CD31882EF7F0D2DACE">
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<mods:date id="95366988D995A88746EF0BEFF61D1141">2015</mods:date>
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<mods:detail id="4C07DC0F93D990B96DD44F7DB5CBC645" type="pubDate">
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<mods:number id="619589F002B3299DDBDE528C12BD268C">2015-12-31</mods:number>
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<mods:number id="F9A73C8D65810493AF180ADD662013B5">62</mods:number>
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<mods:detail id="9870CA59F307ECEEAF0330068B931A4F" type="issue">
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<mods:number id="0F24F793C0EFED4C589C53A030ACFCA7">19</mods:number>
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<mods:start id="965DD40E54A40A0DFFFA3124DDECB383">451</mods:start>
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<mods:classification id="41FACCD7D5485B42D107844AD105CC72">journal article</mods:classification>
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<mods:identifier id="20AFCA90501AFB9E155C67C3C04DE285" type="DOI">10.5281/zenodo.13154942</mods:identifier>
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<treatment id="03ECC23AC45BB1067C8DFAAFFD0B851A" LSID="urn:lsid:plazi:treatment:03ECC23AC45BB1067C8DFAAFFD0B851A" httpUri="http://treatment.plazi.org/id/03ECC23AC45BB1067C8DFAAFFD0B851A" lastPageId="3" lastPageNumber="454" pageId="0" pageNumber="451">
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<subSubSection id="C35F20A7C45BB1057C8DFAAFFC8B8671" box="[434,862,1335,1360]" pageId="0" pageNumber="451" type="nomenclature">
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<heading id="D0B2C440C45BB1057C8DFAAFFC8B8671" box="[434,862,1335,1360]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="451" reason="4">
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<emphasis id="B931AF3EC45BB1057C8DFAAFFC8B8671" bold="true" box="[434,862,1335,1360]" pageId="0" pageNumber="451">
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<taxonomicName id="4C4508AFC45BB1057C8DFAAFFD2F8671" ID-CoL="6756F" authority="T.F. Daniel" authorityName="T. F. Daniel" authorityYear="2015" box="[434,762,1335,1360]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="451" phylum="Tracheophyta" rank="species" species="tablensis" status="sp. nov.">
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<emphasis id="B931AF3EC45BB1057C8DFAAFFDA2866E" bold="true" box="[434,631,1335,1360]" italics="true" pageId="0" pageNumber="451">Anisotes tablensis</emphasis>
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T.F. Daniel
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2021245C45BB1057E37FAAFFC8B8671" box="[776,862,1336,1360]" pageId="0" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F20A7C45BB1077D94FAFBFDD081B2" lastPageId="2" lastPageNumber="453" pageId="0" pageNumber="451" type="materials_examined">
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<paragraph id="8BFA732CC45BB1057D94FAFBFE5C86E4" blockId="0.[123,1174,1388,1478]" pageId="0" pageNumber="451">
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<materialsCitation id="3B2D7971C45BB1057D94FAFBFD0486A5" box="[171,721,1388,1412]" collectionCode="TYPE" collectorName="La Table" country="Madagascar" location="Toliara" pageId="0" pageNumber="451" specimenCount="1" stateProvince="Toliara">
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<emphasis id="B931AF3EC45BB1057D94FAFBFF3186A2" bold="true" box="[171,228,1388,1412]" pageId="0" pageNumber="451">
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<collectionCode id="ED54EBE9C45BB1057D94FAFBFF3186A2" box="[171,228,1388,1412]" pageId="0" pageNumber="451">TYPE</collectionCode>
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</emphasis>
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</smallCapsWord>
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.
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<collectingCountry id="F35233BCC45BB1057DCCFAFAFE5286A2" box="[243,391,1389,1412]" name="Madagascar" pageId="0" pageNumber="451">MADAGASCAR</collectingCountry>
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</smallCapsWord>
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:
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<emphasis id="B931AF3EC45BB1057CA8FAFBFE3C86A5" bold="true" box="[407,489,1388,1412]" pageId="0" pageNumber="451">
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<collectingRegion id="4981BDCEC45BB1057CA8FAFBFE3C86A5" box="[407,489,1388,1412]" country="Madagascar" name="Toliara" pageId="0" pageNumber="451">Toliara</collectingRegion>
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</emphasis>
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:
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<collectorName id="26B016FAC45BB1057CC6FAFAFD8D86A5" box="[505,600,1388,1412]" pageId="0" pageNumber="451">La Table</collectorName>
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, ca.
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</materialsCitation>
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<materialsCitation id="3B2D7971C45BB1057FE6FAFAFD0F8684" collectingDate="2004-05-19" collectionCode="N" country="Madagascar" elevation="85" latitude="-23.42389" longLatPrecision="21" longitude="43.767498" pageId="0" pageNumber="451" specimenCount="1" stateProvince="Toliara">
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<collectionCode id="ED54EBE9C45BB1057FE6FAFAFD3986A5" box="[729,748,1389,1412]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="0" pageNumber="451" type="Herbarium">N</collectionCode>
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[
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<emphasis id="B931AF3EC45BB1057FC1FAFAFCC986A5" box="[766,796,1389,1412]" italics="true" pageId="0" pageNumber="451">sic</emphasis>
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] of Toliara, slope and along ridge,
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,
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,
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<elevation id="0068941FC45BB1057CA5FA1AFDD38685" box="[410,518,1421,1445]" metricMagnitude="1" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="0" pageNumber="451" unit="m" value="85.0" valueMax="120.0" valueMin="50.0">50–120 m</elevation>
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</quantity>
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,
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<date id="FFFB55ECC45BB1057F2BFA1AFD738684" box="[532,678,1421,1445]" pageId="0" pageNumber="451" value="2004-05-19">
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</date>
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(flr)
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</materialsCitation>
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,
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<materialsCitation id="3B2D7971C45BB1057FD6FA19FF2186E7" collectionCode="Z, K, MO" country="Madagascar" pageId="0" pageNumber="451" specimenCount="2" stateProvince="Toliara" typeStatus="holotype">
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<emphasis id="B931AF3EC45BB1057FD6FA19FC168684" box="[745,963,1421,1445]" italics="true" pageId="0" pageNumber="451">
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<collectionCode id="ED54EBE9C45BB1057FD6FA19FD2D8684" box="[745,760,1422,1445]" country="Switzerland" name="Universität Zürich" pageId="0" pageNumber="451" type="Herbarium">Z</collectionCode>
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. Rogers et al. 483
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</emphasis>
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||
(
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<typeStatus id="54FECD8EC45BB1057EEBFA1AFBE48684" box="[980,1073,1421,1445]" pageId="0" pageNumber="451" type="holotype">holotype</typeStatus>
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||
:
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||
<collectionCode id="ED54EBE9C45BB105797EFA1AFB818685" box="[1089,1108,1421,1444]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="0" pageNumber="451" type="Herbarium">K</collectionCode>
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||
!;
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<typeStatus id="54FECD8EC45BB1057952FA1AFF7C86E7" pageId="0" pageNumber="451" type="isotype">isotype</typeStatus>
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||
:
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||
<collectionCode id="ED54EBE9C45BB1057D88FA39FF3786E7" box="[183,226,1454,1478]" country="USA" lsid="urn:lsid:biocol.org:col:15859" name="Missouri Botanical Garden" pageId="0" pageNumber="451" type="Herbarium">MO</collectionCode>
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!)
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||
</materialsCitation>
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||
.
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||
<figureCitation id="137E6FA9C45BB1057C3DFA3AFEB386E7" box="[258,358,1453,1478]" captionStart="FIGURE" captionStartId="1.[155,167,1582,1601]" captionTargetBox="[153,1136,248,1524]" captionTargetId="figure-58@1.[120,1178,216,1563]" captionTargetPageId="1" captionText="FIGURE 1. Anisotes tablensis (Rogers et al. 483). A. Habit. B. Leaf. C. Node with axillary inflorescence bearing a flower. D. Distal portion of stamen with anther. Drawn by Tom Davis." figureDoi="http://doi.org/10.5281/zenodo.13154944" httpUri="https://zenodo.org/record/13154944/files/figure.png" pageId="0" pageNumber="451">Figures 1</figureCitation>
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,
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<figureCitation id="137E6FA9C45BB1057C4AFA39FE5786E7" box="[373,386,1454,1478]" captionStart="FIGURE" captionStartId="4.[155,167,936,955]" captionTargetBox="[120,1178,216,927]" captionTargetId="figure-441@4.[120,1178,216,927]" captionTargetPageId="4" captionText="FIGURE 3. Pollen of Anisotes tablensis (Rogers et al. 483). A. Apertural view. B. Interapertural view." figureDoi="http://doi.org/10.5281/zenodo.13154948" httpUri="https://zenodo.org/record/13154948/files/figure.png" pageId="0" pageNumber="451">3</figureCitation>
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||
.
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</paragraph>
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<paragraph id="8BFA732CC45BB1077D94FA75FC3C8153" blockId="0.[123,1174,1505,1726]" lastBlockId="2.[122,1174,219,1075]" lastPageId="2" lastPageNumber="453" pageId="0" pageNumber="451">
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Divaricately branched shrubs to 4.5 dm tall. Young stems hexagonal, evenly and densely pubescent with a whitish, felt-like covering of antrorsely appressed eglandular trichomes to
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<quantity id="4CBDDEC9C45BB105794BF995FF71851B" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="0" pageNumber="451" unit="mm" value="0.1">0.1 mm</quantity>
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long, epidermis not visible, trichomes soon ± restricted to troughs of internodes. Leaves sessile to subsessile, petioles (if present) to
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<quantity id="4CBDDEC9C45BB1057F00F9D3FDA9857A" box="[575,636,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="0" pageNumber="451" unit="mm" value="2.0">2 mm</quantity>
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long, coriaceous, blades oblong to oblanceolate,
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<quantity id="4CBDDEC9C45BB1057D44F9F3FF35855D" box="[123,224,1636,1660]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="2.3" metricValueMin="0.4" pageId="0" pageNumber="451" unit="mm" value="13.5" valueMax="23.0" valueMin="4.0">4–23 mm</quantity>
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long, 1.4–
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<quantity id="4CBDDEC9C45BB1057C6CF9F3FE45855D" box="[339,400,1636,1660]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="0" pageNumber="451" unit="mm" value="3.0">3 mm</quantity>
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wide, 2.9–10.5 × longer than wide, gradually attenuate at base, rounded at apex, surfaces pubescent like young stems, margin flat. Inflorescence of axillary or terminal sessile short spikes (usually only 2 fertile bracts per spike) to
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<quantity id="4CBDDEC9C45BB1057FC6F931FCE3859C" box="[761,822,1702,1726]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="0" pageNumber="451" unit="mm" value="9.0">9 mm</quantity>
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long (excluding corollas), distal bracts fertile, elliptic,
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<quantity id="4CBDDEC9C459B1077C52FF4CFE0D83D3" box="[365,472,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="2" pageNumber="453" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
|
||
long,
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||
<quantity id="4CBDDEC9C459B1077F1EFF4CFD5883D3" box="[545,653,219,243]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.0" metricValueMin="3.6" pageId="2" pageNumber="453" unit="mm" value="3.8" valueMax="4.0" valueMin="3.6">3.6–4 mm</quantity>
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||
wide, apex rounded to obtuse-truncate, abaxial surface puberulent with antrorsely appressed eglandular trichomes to
|
||
<quantity id="4CBDDEC9C459B1077E4AFF6CFC138233" box="[885,966,251,275]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="453" unit="mm" value="0.1">0.1 mm</quantity>
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||
long and erect trichomes <
|
||
<quantity id="4CBDDEC9C459B1077DD4FE8CFE9F8213" box="[235,330,283,307]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="453" unit="mm" value="0.05">0.05 mm</quantity>
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||
long, margin ± hyaline, proximal bracts sterile and smaller than distal ones. Bracteoles absent. Calyx 5-lobed,
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||
<quantity id="4CBDDEC9C459B1077CD7FEACFDEA8273" box="[488,575,315,339]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="2" pageNumber="453" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
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||
long, lobes oblong-lanceolate,
|
||
<quantity id="4CBDDEC9C459B1077EB1FEACFC338273" box="[910,998,315,339]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="2" pageNumber="453" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">5–6 mm</quantity>
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||
long,
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||
<quantity id="4CBDDEC9C459B1077916FEACFB408273" box="[1065,1173,315,339]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="453" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
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||
wide, abaxial surface pubescent like bracts (or with antrorsely-appressed eglandular trichomes few or absent). Corolla pinkish red,
|
||
<quantity id="4CBDDEC9C459B1077CF7FEECFDEE82B3" box="[456,571,379,403]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="3.5" metricValueMin="2.7" pageId="2" pageNumber="453" unit="mm" value="31.0" valueMax="35.0" valueMin="27.0">27–35 mm</quantity>
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||
long, externally pubescent with erect glandular and flexuose to retrorse eglandular trichomes
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||
<quantity id="4CBDDEC9C459B1077F2EFE0CFD448293" box="[529,657,411,435]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="453" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
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long, tube
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||
<quantity id="4CBDDEC9C459B1077E33FE0CFCAB8293" box="[780,894,411,435]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="2" pageNumber="453" unit="mm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 mm</quantity>
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||
long, corolla tube length: corolla length = 0.44–0.49, upper lip
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||
<quantity id="4CBDDEC9C459B1077F38FE2CFDAC82F3" box="[519,633,443,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="2" pageNumber="453" unit="mm" value="19.0" valueMax="21.0" valueMin="17.0">17–21 mm</quantity>
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long, lower lip
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||
<quantity id="4CBDDEC9C459B1077E1DFE2CFC4182F3" box="[802,916,443,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="2" pageNumber="453" unit="mm" value="18.5" valueMax="19.0" valueMin="18.0">18–19 mm</quantity>
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long with lobes
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||
<quantity id="4CBDDEC9C459B107797AFE2CFF7182D3" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="453" unit="mm" value="3.0" valueMax="3.5" valueMin="2.5">2.5–3.5 mm</quantity>
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long. Stamens
|
||
<quantity id="4CBDDEC9C459B1077C78FE4CFE6C82D3" box="[327,441,475,499]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="2" pageNumber="453" unit="mm" value="21.5" valueMax="23.0" valueMin="20.0">20–23 mm</quantity>
|
||
long, filaments pubescent proximally with erect glandular trichomes
|
||
<quantity id="4CBDDEC9C459B1077D44FE6CFF2F8133" box="[123,250,507,531]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="2" pageNumber="453" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
|
||
long, glabrous distally, thecae
|
||
<quantity id="4CBDDEC9C459B1077F7BFE6CFD7A8133" box="[580,687,507,531]" metricMagnitude="-3" metricUnit="m" metricValue="2.15" metricValueMax="2.3" metricValueMin="2.0" pageId="2" pageNumber="453" unit="mm" value="2.15" valueMax="2.3" valueMin="2.0">2–2.3 mm</quantity>
|
||
long, glabrous, pollen 3-colporate, 6-pseudocolpate (the two pseudocolpi in one or more mesocolpia sometimes fused near one or both poles), 39–41 µm polar diameter (
|
||
<collectionCode id="ED54EBE9C459B1077C9DFDACFE678172" box="[418,434,571,595]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="453" type="Herbarium">P</collectionCode>
|
||
), 26–30 µm equatorial diameter (
|
||
<collectionCode id="ED54EBE9C459B1077E17FDACFCEE8172" box="[808,827,571,595]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="2" pageNumber="453" type="Herbarium">E</collectionCode>
|
||
),
|
||
<collectionCode id="ED54EBE9C459B1077E6CFDACFCB18173" box="[851,868,571,594]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="2" pageNumber="453" type="Herbarium">P</collectionCode>
|
||
:
|
||
<collectionCode id="ED54EBE9C459B1077E58FDACFCAC8173" box="[871,889,571,594]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="2" pageNumber="453" type="Herbarium">E</collectionCode>
|
||
= 1.37–1.50. Style
|
||
<quantity id="4CBDDEC9C459B107796DFDACFF768153" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.2" metricValueMin="2.8" pageId="2" pageNumber="453" unit="mm" value="30.0" valueMax="32.0" valueMin="28.0">28–32 mm</quantity>
|
||
long, glabrous, stigma equally 2-lobed, lobes
|
||
<quantity id="4CBDDEC9C459B1077FA9FDCCFD328153" box="[662,743,603,627]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="453" unit="mm" value="0.2">0.2 mm</quantity>
|
||
long. Capsule not seen.
|
||
</paragraph>
|
||
<caption id="DF3A23A4C45AB1047DA4F9B9FD43857B" ID-DOI="http://doi.org/10.5281/zenodo.13154944" ID-Zenodo-Dep="13154944" httpUri="https://zenodo.org/record/13154944/files/figure.png" pageId="1" pageNumber="452" startId="1.[155,167,1582,1601]" targetBox="[153,1136,248,1524]" targetPageId="1" targetType="figure">
|
||
<paragraph id="8BFA732CC45AB1047DA4F9B9FB428560" blockId="1.[123,1175,1582,1627]" box="[155,1175,1582,1601]" pageId="1" pageNumber="452">
|
||
<smallCapsWord id="8D1CE5F0C45AB1047DA4F9B9FF0D851E" baselines="1596,1595" box="[155,216,1582,1601]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="1" pageNumber="452">FIGURE</smallCapsWord>
|
||
1.
|
||
<taxonomicName id="4C4508AFC45AB1047DCDF9B9FE5E8560" authorityName="T. F. Daniel" authorityYear="2015" box="[242,395,1582,1601]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="452" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC45AB1047DCDF9B9FE5E8560" box="[242,395,1582,1601]" italics="true" pageId="1" pageNumber="452">Anisotes tablensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B931AF3EC45AB1047CA8F9B9FDF28560" box="[407,551,1582,1601]" italics="true" pageId="1" pageNumber="452">Rogers et al. 483</emphasis>
|
||
). A. Habit. B. Leaf. C. Node with axillary inflorescence bearing a flower.
|
||
</paragraph>
|
||
<paragraph id="8BFA732CC45AB1047D44F9DFFD43857B" blockId="1.[123,1175,1582,1627]" box="[123,662,1607,1627]" pageId="1" pageNumber="452">D. Distal portion of stamen with anther. Drawn by Tom Davis.</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFA732CC459B1077D95FDECFDD081B2" blockId="2.[122,1174,219,1075]" box="[170,517,635,659]" pageId="2" pageNumber="453">
|
||
<smallCapsWord id="8D1CE5F0C459B1077D95FDECFEE481B0" baselines="653,652" box="[170,305,635,659]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Phenology" pageId="2" pageNumber="453">
|
||
<emphasis id="B931AF3EC459B1077D95FDECFEE481B0" bold="true" box="[170,305,635,659]" pageId="2" pageNumber="453">PHENOLOGY</emphasis>
|
||
</smallCapsWord>
|
||
.—Flowering: May.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F20A7C459B1067D94FD0CFC1D8136" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453" type="distribution">
|
||
<paragraph id="8BFA732CC459B1077D94FD0CFCB981D2" blockId="2.[122,1174,219,1075]" pageId="2" pageNumber="453">
|
||
<emphasis id="B931AF3EC459B1077D94FD0CFE0F8190" bold="true" box="[171,474,667,691]" pageId="2" pageNumber="453">
|
||
<smallCapsWord id="8D1CE5F0C459B1077D94FD0CFE968190" baselines="685,684" box="[171,323,667,691]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Distribution" pageId="2" pageNumber="453">DISTRIBUTION</smallCapsWord>
|
||
<smallCapsWord id="8D1CE5F0C459B1077C73FD09FEA28190" baselines="684" box="[332,375,670,689]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="2" pageNumber="453">AND</smallCapsWord>
|
||
<smallCapsWord id="8D1CE5F0C459B1077CBEFD09FE0F8190" baselines="684" box="[385,474,670,689]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="habitat" pageId="2" pageNumber="453">HABITAT</smallCapsWord>
|
||
</emphasis>
|
||
.—Endemic to southwestern
|
||
<collectingCountry id="F35233BCC459B1077E26FD0CFC4F8192" box="[793,922,667,691]" name="Madagascar" pageId="2" pageNumber="453">Madagascar</collectingCountry>
|
||
(
|
||
<collectingRegion id="4981BDCEC459B1077E94FD0CFC2D8192" box="[939,1016,667,691]" country="Madagascar" name="Toliara" pageId="2" pageNumber="453">Toliara</collectingRegion>
|
||
) where plants occur in the arid thornscrub on soils derived from decomposed coral (sometimes referred to as “coral rag scrub”) at an elevation between 50 and 120 meters (
|
||
<figureCitation id="137E6FA9C459B1077E22FD4CFC8A81D2" box="[797,863,731,755]" captionStart="FIGURE" captionStartId="2.[153,165,1677,1696]" captionTargetBox="[118,1176,1097,1663]" captionTargetId="figure-559@2.[118,1176,1082,1663]" captionTargetPageId="2" captionText="FIGURE 2. La Table in southwestern Madagascar (looking westward toward Mozambique Channel), habitat of Anisotes tablensis. Photo by the author." figureDoi="http://doi.org/10.5281/zenodo.13154946" httpUri="https://zenodo.org/record/13154946/files/figure.png" pageId="2" pageNumber="453">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFA732CC459B1077D94FD6CFECA80D2" blockId="2.[122,1174,219,1075]" pageId="2" pageNumber="453">
|
||
<emphasis id="B931AF3EC459B1077D94FD6CFE0C8030" bold="true" box="[171,473,763,787]" pageId="2" pageNumber="453">
|
||
<smallCapsWord id="8D1CE5F0C459B1077D94FD6CFE9B8030" baselines="781,780" box="[171,334,763,787]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Conservation" pageId="2" pageNumber="453">CONSERVATION</smallCapsWord>
|
||
<smallCapsWord id="8D1CE5F0C459B1077C6AFD69FE0C8030" baselines="780" box="[341,473,766,785]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="assessment" pageId="2" pageNumber="453">ASSESSMENT</smallCapsWord>
|
||
</emphasis>
|
||
.—Because
|
||
<taxonomicName id="4C4508AFC459B1077F64FD6CFD0C8032" authorityName="T. F. Daniel" authorityYear="2015" box="[603,729,763,787]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="453" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC459B1077F64FD6CFD0C8032" box="[603,729,763,787]" italics="true" pageId="2" pageNumber="453">A. tablensis</emphasis>
|
||
</taxonomicName>
|
||
is known from a single, relatively recent collection from a population that was not observed during this study, it is difficult to assess its conservation status according to
|
||
<bibRefCitation id="EFD40EDDC459B1077C87FCACFD9D8072" author="IUCN" box="[440,584,827,851]" pageId="2" pageNumber="453" refId="ref2408" refString="IUCN. 2014. Guidelines for Using the IUCN Red List Categories and Criteria, Version 11. Standards and Petitions Subcommittee of the IUCN Species Survival Commission. <http: // www. iucnredlist. org / documents / RedListGuidelines. pdf> [accessed 16 November 2015]." type="url" year="2014">IUCN (2014)</bibRefCitation>
|
||
guidelines. Based on IUCN criteria for the threatened categories,
|
||
<taxonomicName id="4C4508AFC459B1077DCAFCCCFEA78052" authorityName="T. F. Daniel" authorityYear="2015" box="[245,370,859,883]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="453" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC459B1077DCAFCCCFEA78052" box="[245,370,859,883]" italics="true" pageId="2" pageNumber="453">A. tablensis</emphasis>
|
||
</taxonomicName>
|
||
likely has both EOO and AOO (criteria B1 and B2) sufficient for Critically Endangered status, but only one (i.e., one known location; i.e., criterion Ba) of two conditions required for assessment in that category. The only known collection does not occur in a locality that is protected. Lacking further data, this apparently rare species must be provisionally assessed as Data Deficient.
|
||
</paragraph>
|
||
<paragraph id="8BFA732CC459B1067D94FC6CFC1D8136" blockId="2.[122,1174,219,1075]" lastBlockId="3.[123,1174,219,535]" lastPageId="3" lastPageNumber="454" pageId="2" pageNumber="453">
|
||
<taxonomicName id="4C4508AFC459B1077D94FC6CFEB98732" authorityName="T. F. Daniel" authorityYear="2015" box="[171,364,1019,1043]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="453" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC459B1077D94FC6CFEB98732" box="[171,364,1019,1043]" italics="true" pageId="2" pageNumber="453">Anisotes tablensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54FECD8EC459B1077F46FC6CFD728732" box="[633,679,1019,1043]" pageId="2" pageNumber="453">type</typeStatus>
|
||
collection. It is the third species of the genus known from the dry regions of southwestern
|
||
<collectingCountry id="F35233BCC459B1077F66FB8CFD098712" box="[601,732,1051,1075]" name="Madagascar" pageId="2" pageNumber="453">Madagascar</collectingCountry>
|
||
. The geographic coordinates given on the label of the
|
||
<typeStatus id="54FECD8EC458B1067DC4FF4CFEFC83D2" box="[251,297,219,243]" pageId="3" pageNumber="454">type</typeStatus>
|
||
are about
|
||
<quantity id="4CBDDEC9C458B1067CA5FF4CFE0583D2" box="[410,464,219,243]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="454" unit="km" value="2.0">2 km</quantity>
|
||
southwest of the prominent local mesa-like physiographic feature known as La Table (
|
||
<figureCitation id="137E6FA9C458B1067C4BFF6CFE6F8235" box="[372,442,251,276]" captionStart="FIGURE" captionStartId="2.[153,165,1677,1696]" captionTargetBox="[118,1176,1097,1663]" captionTargetId="figure-559@2.[118,1176,1082,1663]" captionTargetPageId="2" captionText="FIGURE 2. La Table in southwestern Madagascar (looking westward toward Mozambique Channel), habitat of Anisotes tablensis. Photo by the author." figureDoi="http://doi.org/10.5281/zenodo.13154946" httpUri="https://zenodo.org/record/13154946/files/figure.png" pageId="3" pageNumber="454">Fig. 2</figureCitation>
|
||
), from which the epithet is derived. La Table (
|
||
<geoCoordinate id="EE7115EBC458B1067ECBFF6BFB5A8232" box="[1012,1167,250,276]" degrees="23" direction="south" minutes="24" orientation="latitude" pageId="3" pageNumber="454" precision="1" seconds="32. 40" value="-23.409">23°24ʹ32.40ʺS</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7115EBC458B1067D44FE8BFEC28215" box="[123,279,283,308]" degrees="43" direction="east" minutes="46" orientation="longitude" pageId="3" pageNumber="454" precision="1" seconds="51. 73" value="43.781036">43°46ʹ51.73ʺE</geoCoordinate>
|
||
) is a well-known collecting locale in a region of Tertiary limestone about
|
||
<quantity id="4CBDDEC9C458B106796DFE8BFB408215" box="[1106,1173,284,308]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="454" unit="km" value="12.0">12 km</quantity>
|
||
southeast of the city of
|
||
<collectingRegion id="4981BDCEC458B1067C41FEABFE1C8275" box="[382,457,316,340]" country="Madagascar" name="Toliara" pageId="3" pageNumber="454">Toliara</collectingRegion>
|
||
(Tulear) and home to a large number of
|
||
<taxonomicName id="4C4508AFC458B1067EB4FEABFBC68275" box="[907,1043,316,340]" class="Magnoliopsida" family="Acanthaceae" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="family">Acanthaceae</taxonomicName>
|
||
. It is also a popular birdwatching site where several rare birds, including the Red-shouldered Vanga (
|
||
<taxonomicName id="4C4508AFC458B1067908FECAFEFC82B4" authorityName="Goodman, Hawkins & Domergue" authorityYear="1997" class="Aves" family="Vangidae" genus="Calicalicus" kingdom="Animalia" order="Passeriformes" pageId="3" pageNumber="454" phylum="Chordata" rank="species" species="rufocarpalis">
|
||
<emphasis id="B931AF3EC458B1067908FECAFEFC82B4" italics="true" pageId="3" pageNumber="454">Calicalicus rufocarpalis</emphasis>
|
||
</taxonomicName>
|
||
) and Verreaux's Coua
|
||
<emphasis id="B931AF3EC458B1067F23FEE9FD1E82B4" box="[540,715,382,405]" italics="true" pageId="3" pageNumber="454">(Coua verreauxi</emphasis>
|
||
), are often observed. Given its accessibility, it is surprising that undescribed species are still being found on its slopes and ridge. Two other species of
|
||
<taxonomicName id="4C4508AFC458B1067DCDFE28FE9982F7" box="[242,332,447,470]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B931AF3EC458B1067DCDFE28FE9982F7" box="[242,332,447,470]" italics="true" pageId="3" pageNumber="454">Anisotes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4508AFC458B1067C62FE28FC0482F7" authority="T. F. Daniel, Mbola, Almeda & Phillipson" authorityName="T. F. Daniel, Mbola, Almeda & Phillipson" box="[349,977,446,471]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="divaricatus">
|
||
<emphasis id="B931AF3EC458B1067C62FE28FE2082F7" box="[349,501,446,470]" italics="true" pageId="3" pageNumber="454">A. divaricatus</emphasis>
|
||
T. F. Daniel, Mbola, Almeda & Phillipson
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4508AFC458B1067934FE28FEE782D7" authority="R. Ben." authorityName="R. Ben." class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||
<emphasis id="B931AF3EC458B1067934FE28FF0E82D6" italics="true" pageId="3" pageNumber="454">A. madagascariensis</emphasis>
|
||
R. Ben.
|
||
</taxonomicName>
|
||
, occur in nearby regions (
|
||
<bibRefCitation id="EFD40EDDC458B1067F79FE48FCD782D6" author="DANIEL, T. F. & B. A. V. MBOLA & F. ALMEDA & P. B. PHILLIPSON" box="[582,770,479,503]" pageId="3" pageNumber="454" pagination="121 - 131" refId="ref2355" refString="DANIEL, T. F., B. A. V. MBOLA, F. ALMEDA, AND P. B. PHILLIPSON. 2007. Anisotes (Acanthaceae) in Madagascar. Proceedings of the California Academy of Sciences, ser. 4, 58: 121 - 131." type="journal article" year="2007">Daniel et al. 2007</bibRefCitation>
|
||
), but neither is known from the vicinity of La Table. These three species can be distinguished by the following key:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3A23A4C459B1077DA6F91AFEAA8598" ID-DOI="http://doi.org/10.5281/zenodo.13154946" ID-Zenodo-Dep="13154946" httpUri="https://zenodo.org/record/13154946/files/figure.png" pageId="2" pageNumber="453" startId="2.[153,165,1677,1696]" targetBox="[118,1176,1097,1663]" targetPageId="2" targetType="figure">
|
||
<paragraph id="8BFA732CC459B1077DA6F91AFEAA8598" blockId="2.[120,1171,1677,1722]" pageId="2" pageNumber="453">
|
||
<smallCapsWord id="8D1CE5F0C459B1077DA6F91AFF0385BF" baselines="1691,1690" box="[153,214,1677,1696]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="2" pageNumber="453">FIGURE</smallCapsWord>
|
||
2. La Table in southwestern Madagascar (looking westward toward Mozambique Channel), habitat of
|
||
<taxonomicName id="4C4508AFC459B1077974F91AFF118598" authorityName="T. F. Daniel" authorityYear="2015" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="2" pageNumber="453" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC459B1077974F91AFF118598" italics="true" pageId="2" pageNumber="453">Anisotes tablensis</emphasis>
|
||
</taxonomicName>
|
||
. Photo by the author.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35F20A7C458B1067D44FDA4FD0B851A" pageId="3" pageNumber="454" type="key">
|
||
<key id="FFD4241DC458B1067D44FDA4FB4487A3" pageId="3" pageNumber="454">
|
||
<keyStep id="30B16B89C458B1067D44FDA4FB4481CF" pageId="3" pageNumber="454">
|
||
<paragraph id="8BFA732CC458B1067D44FDA4FB4481AD" blockId="3.[123,1174,563,750]" pageId="3" pageNumber="454">
|
||
<keyLead id="30B4D019C458B1067D44FDA4FB4481AD" pageId="3" pageNumber="454">
|
||
1a. Leaves sessile to subsessile, petioles (if present) to
|
||
<quantity id="4CBDDEC9C458B1067E3EFDA4FCEB816A" box="[769,830,563,587]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="454" unit="mm" value="2.0">2 mm</quantity>
|
||
long, blades 1.4–
|
||
<quantity id="4CBDDEC9C458B106792FFDA4FB98816A" box="[1040,1101,563,587]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="454" unit="mm" value="3.0">3 mm</quantity>
|
||
wide, length:width = 2.9–10.5; calyx
|
||
<quantity id="4CBDDEC9C458B1067CD2FDC3FD90814A" box="[493,581,596,620]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="3" pageNumber="454" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6–7 mm</quantity>
|
||
long; corolla externally pubescent with glandular and eglandular trichomes throughout, lobes of lower lip 2.5–
|
||
<quantity id="4CBDDEC9C458B1067FC9FDE3FC9281AD" box="[758,839,628,652]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="454" unit="mm" value="3.5">3.5 mm</quantity>
|
||
long...........
|
||
<taxonomicName id="4C4508AFC458B106792BFDE2FB4481AD" ID-CoL="6756F" authorityName="T. F. Daniel" authorityYear="2015" box="[1044,1169,628,652]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC458B106792BFDE2FB4481AD" box="[1044,1169,628,652]" italics="true" pageId="3" pageNumber="454">A. tablensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BFA732CC458B1067D44FD02FB4481CF" blockId="3.[123,1174,563,750]" pageId="3" pageNumber="454">
|
||
<keyLead id="30B4D019C458B1067D44FD02FB4481CF" pageId="3" pageNumber="454">
|
||
1b. Leaves petiolate, petioles to
|
||
<quantity id="4CBDDEC9C458B1067CD7FD02FDE7818D" box="[488,562,661,685]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="454" unit="mm" value="12.0">12 mm</quantity>
|
||
long, blades 4.
|
||
<quantity id="4CBDDEC9C458B1067FDEFD02FC93818D" box="[737,838,661,685]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="3.8" metricValueMin="0.5" pageId="3" pageNumber="454" unit="mm" value="21.5" valueMax="38.0" valueMin="5.0">5–38 mm</quantity>
|
||
wide, length:width = 0.7–2.7; calyx 1.3–
|
||
<quantity id="4CBDDEC9C458B1067C32FD22FE8B81EC" box="[269,350,693,717]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="454" unit="mm" value="3.5">3.5 mm</quantity>
|
||
long; corolla externally glabrous or occasionally with a few eglandular trichomes proximally, lobes of lower lip
|
||
<quantity id="4CBDDEC9C458B1067F71FD41FD6681CC" box="[590,691,726,750]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="3" pageNumber="454" unit="mm" value="11.0" valueMax="14.0" valueMin="8.0">8–14 mm</quantity>
|
||
long.............................. 2
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30B16B89C458B1067D44FC93FB4487A3" pageId="3" pageNumber="454">
|
||
<paragraph id="8BFA732CC458B1067D44FC93FB44809E" blockId="3.[123,1174,772,1154]" pageId="3" pageNumber="454">
|
||
<keyLead id="30B4D019C458B1067D44FC93FB44809E" pageId="3" pageNumber="454">
|
||
2a. Leaf blades broadly ovate to ovate-elliptic to elliptic to broadly elliptic,
|
||
<quantity id="4CBDDEC9C458B1067E94FC93FBC8803D" box="[939,1053,772,796]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="4.5" metricValueMin="1.4" pageId="3" pageNumber="454" unit="mm" value="29.5" valueMax="45.0" valueMin="14.0">14–45 mm</quantity>
|
||
long; margin of calyx lobes ± densely ciliate; corolla with internal surface conspicuously lighter in color than external surface, corolla tube
|
||
<quantity id="4CBDDEC9C458B1067F28FCD2FDA9807C" box="[535,636,837,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="1.5" metricValueMin="0.9" pageId="3" pageNumber="454" unit="mm" value="12.0" valueMax="15.0" valueMin="9.0">9–15 mm</quantity>
|
||
long, corolla tube:corolla = 0.23–0.38, upper lip
|
||
<quantity id="4CBDDEC9C458B1067DA8FCF1FEDF805C" box="[151,266,870,894]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="3" pageNumber="454" unit="mm" value="27.5" valueMax="35.0" valueMin="20.0">20–35 mm</quantity>
|
||
long and distally whitish to pinkish along margin, lower lip spirally coiled,
|
||
<quantity id="4CBDDEC9C458B106796CFCF1FF1580BF" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.8" metricValueMin="1.8" pageId="3" pageNumber="454" unit="mm" value="23.0" valueMax="28.0" valueMin="18.0">18–28 mm</quantity>
|
||
long, lobes
|
||
<quantity id="4CBDDEC9C458B1067C77FC11FE6E80BF" box="[328,443,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="3" pageNumber="454" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
|
||
long; stamens
|
||
<quantity id="4CBDDEC9C458B1067F5EFC11FD0680BF" box="[609,723,902,926]" metricMagnitude="-2" metricUnit="m" metricValue="2.95" metricValueMax="3.3" metricValueMin="2.6" pageId="3" pageNumber="454" unit="mm" value="29.5" valueMax="33.0" valueMin="26.0">26–33 mm</quantity>
|
||
long; capsule pubescent with flexuose to antrorse eglandular trichomes 0.05–
|
||
<quantity id="4CBDDEC9C458B1067F26FC30FDBF809F" box="[537,618,935,959]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="454" unit="mm" value="0.1">0.1 mm</quantity>
|
||
long....................
|
||
<taxonomicName id="4C4508AFC458B1067E8DFC30FB44809E" ID-CoL="66T6D" box="[946,1169,935,959]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||
<emphasis id="B931AF3EC458B1067E8DFC30FB44809E" box="[946,1169,935,959]" italics="true" pageId="3" pageNumber="454">A. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BFA732CC458B1067D44FC50FB4487A3" blockId="3.[123,1174,772,1154]" pageId="3" pageNumber="454">
|
||
<keyLead id="30B4D019C458B1067D44FC50FB4487A3" pageId="3" pageNumber="454">
|
||
2b. Leaf blades broadly obovate to subcircular to obcordate to obdeltate to oblate,
|
||
<quantity id="4CBDDEC9C458B1067ECDFC50FB8280FE" box="[1010,1111,967,991]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="14.0" metricValueMin="5.0" pageId="3" pageNumber="454" unit="mm" value="9.5" valueMax="14.0" valueMin="5.0">5–14 mm</quantity>
|
||
long; margin of calyx lobes eciliate to sparsely ciliate; corolla with internal surface not conspicuously lighter in color than external surface, corolla tube
|
||
<quantity id="4CBDDEC9C458B1067FD5FB9FFC888701" box="[746,861,1032,1056]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.0" metricValueMin="1.4" pageId="3" pageNumber="454" unit="mm" value="17.0" valueMax="20.0" valueMin="14.0">14–20 mm</quantity>
|
||
long, corolla tube:corolla = 0.46–0.56, upper lip (11–)
|
||
<quantity id="4CBDDEC9C458B1067C8BFBBEFDF38761" box="[436,550,1065,1089]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.8" metricValueMin="1.4" pageId="3" pageNumber="454" unit="mm" value="16.0" valueMax="18.0" valueMin="14.0">14–18 mm</quantity>
|
||
long and lacking a pale margin distally, lower lip recurved to reflexed (not spirally coiled),
|
||
<quantity id="4CBDDEC9C458B1067F3EFBDEFDA78740" box="[513,626,1097,1121]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.9" metricValueMin="1.1" pageId="3" pageNumber="454" unit="mm" value="15.0" valueMax="19.0" valueMin="11.0">11–19 mm</quantity>
|
||
long, lobes
|
||
<quantity id="4CBDDEC9C458B1067FC0FBDEFCB18740" box="[767,868,1097,1121]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="3" pageNumber="454" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
|
||
long; stamens 15–18.
|
||
<quantity id="4CBDDEC9C458B1067967FBDEFB408740" box="[1112,1173,1097,1121]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="454" unit="mm" value="5.0">5 mm</quantity>
|
||
long; capsule ± rugose-granulate but lacking noticeable eglandular trichomes..
|
||
<taxonomicName id="4C4508AFC458B1067EC4FBFDFB4487A3" ID-CoL="6756W" box="[1019,1169,1130,1154]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="divaricatus">
|
||
<emphasis id="B931AF3EC458B1067EC4FBFDFB4487A3" box="[1019,1169,1130,1154]" italics="true" pageId="3" pageNumber="454">A. divaricatus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
<paragraph id="8BFA732CC458B1067D94FB09FD0B851A" blockId="3.[123,1174,1182,1595]" pageId="3" pageNumber="454">
|
||
All three of the species of
|
||
<taxonomicName id="4C4508AFC458B1067CF5FB09FDF18794" ID-CoL="WH5" box="[458,548,1182,1205]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B931AF3EC458B1067CF5FB09FDF18794" box="[458,548,1182,1205]" italics="true" pageId="3" pageNumber="454">Anisotes</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F35233BCC458B1067FF4FB09FC998797" box="[715,844,1182,1206]" name="Madagascar" pageId="3" pageNumber="454">Madagascar</collectingCountry>
|
||
lack bracteoles and have coriaceous to subsucculent leaves, bracts lacking reticulate venation (or with venation not evident), and similar 3-aperturate pollen. They would appear to pertain to Baden’s section
|
||
<taxonomicName id="4C4508AFC458B1067EE5FB48FBE887D6" box="[986,1085,1247,1271]" pageId="3" pageNumber="454" rank="section" section="Spiciflori">
|
||
<emphasis id="B931AF3EC458B1067EE5FB48FBE887D6" box="[986,1085,1247,1271]" italics="true" pageId="3" pageNumber="454">Spiciflori</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFD40EDDC458B106796EFB48FF658636" author="BADEN, C." pageId="3" pageNumber="454" pagination="623 - 664" refId="ref2254" refString="BADEN, C. 1981. The genus Anisotes (Acanthaceae), a taxonomic revision. Nordic Journal of Botany 1: 623 - 664." type="journal article" year="1981">Baden 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFD40EDDC458B1067DFFFB68FE578636" author="DANIEL, T. F. & B. A. V. MBOLA & F. ALMEDA & P. B. PHILLIPSON" box="[192,386,1279,1303]" pageId="3" pageNumber="454" pagination="121 - 131" refId="ref2355" refString="DANIEL, T. F., B. A. V. MBOLA, F. ALMEDA, AND P. B. PHILLIPSON. 2007. Anisotes (Acanthaceae) in Madagascar. Proceedings of the California Academy of Sciences, ser. 4, 58: 121 - 131." type="journal article" year="2007">Daniel et al. 2007</bibRefCitation>
|
||
). Pollen of
|
||
<taxonomicName id="4C4508AFC458B1067F3AFB68FD518636" ID-CoL="6756F" authorityName="T. F. Daniel" authorityYear="2015" box="[517,644,1279,1303]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC458B1067F3AFB68FD518636" box="[517,644,1279,1303]" italics="true" pageId="3" pageNumber="454">A. tablensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="137E6FA9C458B1067FAAFB68FD018636" box="[661,724,1279,1303]" captionStart="FIGURE" captionStartId="4.[155,167,936,955]" captionTargetBox="[120,1178,216,927]" captionTargetId="figure-441@4.[120,1178,216,927]" captionTargetPageId="4" captionText="FIGURE 3. Pollen of Anisotes tablensis (Rogers et al. 483). A. Apertural view. B. Interapertural view." figureDoi="http://doi.org/10.5281/zenodo.13154948" httpUri="https://zenodo.org/record/13154948/files/figure.png" pageId="3" pageNumber="454">Fig. 3</figureCitation>
|
||
) greatly resembles that of both
|
||
<taxonomicName id="4C4508AFC458B106790DFB68FED58616" ID-CoL="66T6D" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="madagascariensis">
|
||
<emphasis id="B931AF3EC458B106790DFB68FED58616" italics="true" pageId="3" pageNumber="454">A. madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4508AFC458B1067C09FAB7FE198616" ID-CoL="6756W" box="[310,460,1311,1335]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="divaricatus">
|
||
<emphasis id="B931AF3EC458B1067C09FAB7FE198616" box="[310,460,1311,1335]" italics="true" pageId="3" pageNumber="454">A. divaricatus</emphasis>
|
||
</taxonomicName>
|
||
from southern
|
||
<collectingCountry id="F35233BCC458B1067F4DFA88FD278616" box="[626,754,1311,1335]" name="Madagascar" pageId="3" pageNumber="454">Madagascar</collectingCountry>
|
||
, except for its somewhat smaller polar diameter (39–41 vs. 62.5–7 µm). The four species from northern
|
||
<collectingCountry id="F35233BCC458B1067E76FAD7FC1F8679" box="[841,970,1344,1368]" name="Madagascar" pageId="3" pageNumber="454">Madagascar</collectingCountry>
|
||
(
|
||
<taxonomicName id="4C4508AFC458B1067EE3FAD7FDE08658" ID-CoL="66T7B" authority="T. F. Daniel, R. Letsara & Martin-Bravo" authorityName="T. F. Daniel, R. Letsara & Martin-Bravo" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="hygroscopicus">
|
||
<emphasis id="B931AF3EC458B1067EE3FAD7FB408679" box="[988,1173,1344,1368]" italics="true" pageId="3" pageNumber="454">A. hygroscopicus</emphasis>
|
||
T. F. Daniel, R. Letsara & Martín-Bravo
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4508AFC458B1067F7BFAF6FB5A8658" ID-CoL="5V56C" authority="T. F. Daniel, R. Letsara & Martin-Bravo" authorityName="T. F. Daniel, R. Letsara & Martin-Bravo" box="[580,1167,1376,1401]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="perplexus">
|
||
<emphasis id="B931AF3EC458B1067F7BFAF6FD198659" box="[580,716,1376,1400]" italics="true" pageId="3" pageNumber="454">A. perplexus</emphasis>
|
||
T. F. Daniel, R. Letsara & Martín-Bravo
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4508AFC458B1067D44FA16FD3786B8" ID-CoL="66T6F" authority="T. F. Daniel, R. Letsara & Martin-Bravo" authorityName="T. F. Daniel, R. Letsara & Martin-Bravo" box="[123,738,1409,1433]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="subcoriaceus">
|
||
<emphasis id="B931AF3EC458B1067D44FA16FEF386B8" box="[123,294,1409,1433]" italics="true" pageId="3" pageNumber="454">A. subcoriaceus</emphasis>
|
||
T. F. Daniel, R. Letsara & Martín-Bravo
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C4508AFC458B1067E21FA16FEFF869B" ID-CoL="5V56M" authority="T. F. Daniel, R. Letsara & Martin-Bravo" authorityName="T. F. Daniel, R. Letsara & Martin-Bravo" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="venosus">
|
||
<emphasis id="B931AF3EC458B1067E21FA16FC4486B8" box="[798,913,1409,1433]" italics="true" pageId="3" pageNumber="454">A. venosus</emphasis>
|
||
T. F. Daniel, R. Letsara & Martín-Bravo
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFD40EDDC458B1067C07FA36FE22869B" author="DANIEL, T. F. & R. LETSARA & S. MARTIN-BRAVO" box="[312,503,1441,1466]" pageId="3" pageNumber="454" pagination="396 - 408" refId="ref2319" refString="DANIEL, T. F., R. LETSARA, AND S. MARTIN-BRAVO. 2013. Four new species of Anisotes (Acanthaceae) in Madagascar. Novon 22: 396 - 408." type="journal article" year="2013">Daniel et al. 2013</bibRefCitation>
|
||
) show greater diversity in pollen shape and sculpturing (varying from 2- to 3-aperturate) and have membranous to subcoriaceous leaves, bracteoles, and bracts with reticulate venation evident.
|
||
<taxonomicName id="4C4508AFC458B1067CE5FA74FD4E86DB" ID-CoL="6756F" authorityName="T. F. Daniel" authorityYear="2015" box="[474,667,1506,1530]" class="Magnoliopsida" family="Acanthaceae" genus="Anisotes" kingdom="Plantae" order="Lamiales" pageId="3" pageNumber="454" phylum="Tracheophyta" rank="species" species="tablensis">
|
||
<emphasis id="B931AF3EC458B1067CE5FA74FD4E86DB" box="[474,667,1506,1530]" italics="true" pageId="3" pageNumber="454">Anisotes tablensis</emphasis>
|
||
</taxonomicName>
|
||
shows greater similarities to those four species than to the other two southern ones in the following characteristics: calyx length, external pubescence of the corolla, and length of lobes of the lower lip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |