171 lines
24 KiB
XML
171 lines
24 KiB
XML
<document id="E683C28784B9A3E8432CBFE0B571290D" ID-CLB-Dataset="53499" ID-DOI="10.11646/zootaxa.5296.3.6" ID-GBIF-Dataset="2825cf24-234a-4b85-b997-673ee08019bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984131" ID-ZooBank="59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1685427549966" checkinUser="plazi" docAuthor="Bartel, Christian, Dunlop, Jason A. & Giribet, Gonzalo" docDate="2023" docId="03D878433948FD2E8BB5F89CFD01FB0A" docLanguage="en" docName="zootaxa.5296.3.6.pdf" docOrigin="Zootaxa 5296 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Foveacorpus Bartel & Dunlop & Giribet 2023, gen. nov." docType="treatment" docUuid="695BD267-DC04-4076-932D-5D0A81E2CDB5" docUuidSource="ZooBank" docVersion="6" lastPageNumber="435" masterDocId="FFE1003B3945FD208B22FFC5FFC5FFCF" masterDocTitle="An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber" masterLastPageNumber="445" masterPageNumber="421" pageNumber="434" updateTime="1699510734804" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="28C9DFA9C564F41DB7BD8C9960DA4AD8">An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber</mods:title>
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<mods:namePart id="7174D1D60B8DF5D742DD4CF1C4C9EBB5">Bartel, Christian</mods:namePart>
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<mods:namePart id="626E1DC02C722B396B6D0F177FC20340">Dunlop, Jason A.</mods:namePart>
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<mods:namePart id="3A427F73549546C74E14AF731946883C">Giribet, Gonzalo</mods:namePart>
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<mods:title id="05BDE7E35C652A2C5D745F1FB6A8B03F">Zootaxa</mods:title>
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<mods:date id="0656242CEBFBEF981080A2117AB149CC">2023</mods:date>
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<mods:number id="FAB06D8195D49964670FC424EFC16B82">2023-05-29</mods:number>
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<treatment id="03D878433948FD2E8BB5F89CFD01FB0A" ID-DOI="http://doi.org/10.5281/zenodo.7984085" ID-GBIF-Taxon="209663740" ID-Zenodo-Dep="7984085" LSID="urn:lsid:zoobank.org:act:695BD267-DC04-4076-932D-5D0A81E2CDB5" httpUri="http://treatment.plazi.org/id/03D878433948FD2E8BB5F89CFD01FB0A" lastPageId="14" lastPageNumber="435" pageId="13" pageNumber="434">
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<subSubSection id="C36B9ADE3948FD2D8BB5F89CFC44F87F" pageId="13" pageNumber="434" type="nomenclature">
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<paragraph id="8BCEC9553948FD2D8BB5F89CFDC5F8BC" blockId="13.[151,512,1881,1907]" box="[151,512,1881,1907]" pageId="13" pageNumber="434">
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<heading id="D0867E393948FD2D8BB5F89CFDC5F8BC" bold="true" box="[151,512,1881,1907]" fontSize="11" level="1" pageId="13" pageNumber="434" reason="1">
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<emphasis id="B90515473948FD2D8BB5F89CFDC5F8BC" bold="true" box="[151,512,1881,1907]" pageId="13" pageNumber="434">
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Genus
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<taxonomicName id="4C71B2D63948FD2D8BCCF89CFE4FF8BC" ID-CoL="BHZMG" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[238,394,1881,1907]" class="Arachnida" family="Stylocellidae" genus="Foveacorpus" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="434" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90515473948FD2D8BCCF89CFE4FF8BC" bold="true" box="[238,394,1881,1907]" italics="true" pageId="13" pageNumber="434">Foveacorpus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A236A83C3948FD2D8AB0F89CFDC5F8BC" box="[402,512,1881,1907]" pageId="13" pageNumber="434" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCEC9553948FD2D8BB5F85DFC44F87F" blockId="13.[151,897,1944,1968]" box="[151,897,1944,1968]" pageId="13" pageNumber="434">
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<uri id="FFE0C5573948FD2D8BB5F85DFC44F87F" box="[151,897,1944,1968]" pageId="13" pageNumber="434">
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urn:lsid:zoobank.org:act:
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<uuid id="FFD7F3803948FD2D8AB2F85DFC44F87F" box="[400,897,1944,1968]" pageId="13" pageNumber="434">695BD267-DC04-4076-932D-5D0A81E2CDB5</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE3948FD2D8BB5F81CFD29F83B" box="[151,748,2009,2036]" pageId="13" pageNumber="434" type="type_taxon">
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<paragraph id="8BCEC9553948FD2D8BB5F81CFD29F83B" blockId="13.[151,748,2009,2036]" box="[151,748,2009,2036]" pageId="13" pageNumber="434">
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<emphasis id="B90515473948FD2D8BB5F81CFEF7F83C" bold="true" box="[151,306,2009,2035]" pageId="13" pageNumber="434">
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<typeStatus id="54CA77F73948FD2D8BB5F81CFF17F83C" box="[151,210,2009,2035]" pageId="13" pageNumber="434">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C71B2D63948FD2D8A1BF81FFDF9F83C" ID-CoL="BHZMJ" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[313,572,2010,2035]" class="Arachnida" family="Stylocellidae" genus="Foveacorpus" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="434" phylum="Arthropoda" rank="species" species="cretaceus" status="gen. et sp. nov.">
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<emphasis id="B90515473948FD2D8A1BF81FFDF9F83C" box="[313,572,2010,2035]" italics="true" pageId="13" pageNumber="434">Foveacorpus cretaceus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A236A83C3948FD2D8966F81FFD29F83B" box="[580,748,2010,2036]" pageId="13" pageNumber="434" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C36B9ADE394BFD2E8BE5FF52FB2AFF7E" box="[199,1263,151,177]" pageId="14" pageNumber="435" type="etymology">
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<paragraph id="8BCEC955394BFD2E8BE5FF52FB2AFF7E" blockId="14.[151,1437,151,1221]" box="[199,1263,151,177]" pageId="14" pageNumber="435">
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<emphasis id="B9051547394BFD2E8BE5FF52FE8BFF7E" bold="true" box="[199,334,151,177]" pageId="14" pageNumber="435">Etymology.</emphasis>
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From the Latin words “fovea” (= pit) and “corpus” (= body). Masculine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE394BFD2E8BE5FF7FFC0AFE8E" pageId="14" pageNumber="435" type="diagnosis">
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<paragraph id="8BCEC955394BFD2E8BE5FF7FFC0AFE8E" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="435">
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<emphasis id="B9051547394BFD2E8BE5FF7FFE87FF1B" bold="true" box="[199,322,186,212]" pageId="14" pageNumber="435">Diagnosis.</emphasis>
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<materialsCitation id="3B19C308394BFD2E8A69FF7EFC0EFE8E" ID-GBIF-Occurrence="4120750306" county="Anal region" location="Chelicerae" municipality="Typical" pageId="14" pageNumber="435" specimenCount="1">
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Body oval, small and completely covered with numerous small pits. Rounded ozophores in
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<typeStatus id="54CA77F7394BFD2E8E4EFF7EFA59FF1A" box="[1388,1436,187,213]" pageId="14" pageNumber="435">type</typeStatus>
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1 position.
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<location id="8EAE9F8E394BFD2E8A34FF1AFE4BFF36" LSID="urn:lsid:plazi:treatment:03D878433948FD2E8BB5F89CFD01FB0A:8EAE9F8E394BFD2E8A34FF1AFE4BFF36" box="[278,398,223,249]" county="Anal region" municipality="Typical" name="Chelicerae" pageId="14" pageNumber="435">Chelicerae</location>
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moderately long and thin.
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<location id="8EAE9F8E394BFD2E89E3FF1AFCEBFF36" LSID="urn:lsid:plazi:treatment:03D878433948FD2E8BB5F89CFD01FB0A:8EAE9F8E394BFD2E89E3FF1AFCEBFF36" box="[705,814,223,249]" county="Anal region" municipality="Typical" name="Pedipalps" pageId="14" pageNumber="435">Pedipalps</location>
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rather small and unmodified.
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<location id="8EAE9F8E394BFD2E8FA6FF1AFB7FFF36" LSID="urn:lsid:plazi:treatment:03D878433948FD2E8BB5F89CFD01FB0A:8EAE9F8E394BFD2E8FA6FF1AFB7FFF36" box="[1156,1210,223,249]" county="Anal region" municipality="Typical" name="Legs" pageId="14" pageNumber="435">Legs</location>
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relatively short and robust, with legs I and II directed forwards.
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<collectingMunicipality id="6BAA532F394BFD2E895DFEC6FD16FED2" box="[639,723,259,285]" pageId="14" pageNumber="435">Typical</collectingMunicipality>
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spiracles absent.
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<collectingCounty id="62AFB1D9394BFD2E88B7FEC6FBDFFED2" box="[917,1050,259,285]" pageId="14" pageNumber="435">Anal region</collectingCounty>
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with a corona analis (sternites 8, 9 and tergite IX fused and forming an unbroken ring around the anal plate)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE394BFD2E8BE5FE8EFD01FB0A" pageId="14" pageNumber="435" type="discussion">
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<paragraph id="8BCEC955394BFD2E8BE5FE8EFA86FD02" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="435">
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<emphasis id="B9051547394BFD2E8BE5FE8EFEFEFEAA" bold="true" box="[199,315,331,357]" pageId="14" pageNumber="435">Remarks.</emphasis>
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The
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<typeStatus id="54CA77F7394BFD2E8A51FE8EFE10FEAA" box="[371,469,331,357]" pageId="14" pageNumber="435" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="4C71B2D6394BFD2E8ADAFE8EFD3EFEAB" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[504,763,331,356]" class="Arachnida" family="Stylocellidae" genus="Foveacorpus" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="435" phylum="Arthropoda" rank="species" species="cretaceus" status="sp. nov.">
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<emphasis id="B9051547394BFD2E8ADAFE8EFD3EFEAB" box="[504,763,331,356]" italics="true" pageId="14" pageNumber="435">Foveacorpus cretaceus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A236A83C394BFD2E8823FE8EFC92FEAA" box="[769,855,331,357]" pageId="14" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
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(GPIH05129;
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<figureCitation id="134AD5D0394BFD2E88DEFE8EFBBAFEAA" box="[1020,1151,331,357]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="15.[151,250,1943,1967]" captionStartId-1="16.[151,250,1107,1131]" captionTargetBox-0="[172,1416,181,1910]" captionTargetBox-1="[151,1436,417,1084]" captionTargetId-0="figure-20@15.[172,1416,181,1920]" captionTargetId-1="figure-157@16.[151,1436,417,1084]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 11. Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ca, corona analis; ch, chelicerae; oz, ozophore; pp, pedipalp; legs numbered from I–IV. Scale bars equal 0.5 mm." captionText-1="FIGURE 12. Details of Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Close-up of chelicerae, pedipalps and gonostome (indicated by arrows), ventral view. B. Close-up of opisthosoma, ventral view. Abbreviations: ca, corona analis; ch, chelicerae; go, gonostome; pp, pedipalp; oz, ozophore; legs II, III and IV numbered. Scale bars equal 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7984159" figureDoi-1="http://doi.org/10.5281/zenodo.7984161" httpUri-0="https://zenodo.org/record/7984159/files/figure.png" httpUri-1="https://zenodo.org/record/7984161/files/figure.png" pageId="14" pageNumber="435">Figs. 11–12</figureCitation>
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) is interpreted as an adult female due to the open gonostome (
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<figureCitation id="134AD5D0394BFD2E8916FEAAFD78FE46" box="[564,701,367,393]" captionStart="FIGURE 11" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[172,1416,181,1910]" captionTargetId="figure-20@15.[172,1416,181,1920]" captionTargetPageId="15" captionText="FIGURE 11. Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ca, corona analis; ch, chelicerae; oz, ozophore; pp, pedipalp; legs numbered from I–IV. Scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7984159" httpUri="https://zenodo.org/record/7984159/files/figure.png" pageId="14" pageNumber="435">Fig. 11B, D</figureCitation>
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) and the absence of an adenostyle on tarsus IV. This very small specimen shows an unusual and potentially unique combination of characters. First, the whole body is completely covered with small pits (
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<figureCitation id="134AD5D0394BFD2E8A91FE72FDC3FE1E" box="[435,518,439,465]" captionStart="FIGURE 11" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[172,1416,181,1910]" captionTargetId="figure-20@15.[172,1416,181,1920]" captionTargetPageId="15" captionText="FIGURE 11. Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ca, corona analis; ch, chelicerae; oz, ozophore; pp, pedipalp; legs numbered from I–IV. Scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7984159" httpUri="https://zenodo.org/record/7984159/files/figure.png" pageId="14" pageNumber="435">Fig. 11</figureCitation>
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), a feature which has not been observed in any modern species, and the dorsum shows no sign of segmentation or of a longitudinal opisthosomal sulcus. There are extant stylocellid species, such as
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<taxonomicName id="4C71B2D6394BFD2E8BD0FE3AFC8DFDD6" authority="Clouse & Schwendinger, 2012" authorityName="Clouse & Schwendinger" authorityYear="2012" box="[242,840,511,537]" class="Arachnida" family="Stylocellidae" genus="Leptopsalis" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="435" phylum="Arthropoda" rank="species" species="foveolata">
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<emphasis id="B9051547394BFD2E8BD0FE3AFE26FDD6" box="[242,483,511,537]" italics="true" pageId="14" pageNumber="435">Leptopsalis foveolata</emphasis>
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<bibRefCitation id="EFE0B4A4394BFD2E8AC9FE3AFC8DFDD6" author="Clouse, R. M. & Schwendinger, P. J." box="[491,840,511,537]" pageId="14" pageNumber="435" pagination="529 - 546" refId="ref12602" refString="Clouse, R. M. & Schwendinger, P. J. (2012) Leptopsalis foveolata sp. nov., a new species of Stylocellidae from Thailand that displays a novel morphological feature in the suborder Cyphophthalmi (Arachnida, Opiliones). Revue suisse de Zoologie, 119, 529 - 546. https: // doi. org / 10.5962 / bhl. part. 150204" type="journal article" year="2012">Clouse & Schwendinger, 2012</bibRefCitation>
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</taxonomicName>
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, which feature a few much larger and deeper pits on the opisthosoma. It is possible that these small pits in the cuticle of this fossil are an artefact resulting from the fossilization process where, e.g., a substance coats different parts of the specimen, as seen in many Burmese amber fossils. However, the ventral region around the gonostome (
|
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<figureCitation id="134AD5D0394BFD2E881DFDAEFC63FD4A" box="[831,934,619,645]" captionStart="FIGURE 12" captionStartId="16.[151,250,1107,1131]" captionTargetBox="[151,1436,417,1084]" captionTargetId="figure-157@16.[151,1436,417,1084]" captionTargetPageId="16" captionText="FIGURE 12. Details of Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Close-up of chelicerae, pedipalps and gonostome (indicated by arrows), ventral view. B. Close-up of opisthosoma, ventral view. Abbreviations: ca, corona analis; ch, chelicerae; go, gonostome; pp, pedipalp; oz, ozophore; legs II, III and IV numbered. Scale bars equal 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7984161" httpUri="https://zenodo.org/record/7984161/files/figure.png" pageId="14" pageNumber="435">Fig. 12A</figureCitation>
|
||
) shows no signs of a coating as many small setae are clearly visible and the pits appear to be open. We conclude that most parts of the fossil are originally preserved, and that the pits on the prosoma and opisthosoma were presumably covered by some substance.
|
||
</paragraph>
|
||
<paragraph id="8BCEC955394BFD2E8BE5FD12FD01FB0A" blockId="14.[151,1437,151,1221]" pageId="14" pageNumber="435">
|
||
Placement of this fossil in the family
|
||
<taxonomicName id="4C71B2D6394BFD2E894CFD12FCC5FD3E" authorityName="Hansen & Sorensen" authorityYear="1904" box="[622,768,727,753]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="435" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
|
||
can be excluded with some confidence due to the presence of a corona analis in the fossil were sternites 8, 9 and tergite IX are fused (
|
||
<figureCitation id="134AD5D0394BFD2E88CDFD3EFB92FCDA" box="[1007,1111,763,789]" captionStart="FIGURE 12" captionStartId="16.[151,250,1107,1131]" captionTargetBox="[151,1436,417,1084]" captionTargetId="figure-157@16.[151,1436,417,1084]" captionTargetPageId="16" captionText="FIGURE 12. Details of Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Close-up of chelicerae, pedipalps and gonostome (indicated by arrows), ventral view. B. Close-up of opisthosoma, ventral view. Abbreviations: ca, corona analis; ch, chelicerae; go, gonostome; pp, pedipalp; oz, ozophore; legs II, III and IV numbered. Scale bars equal 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7984161" httpUri="https://zenodo.org/record/7984161/files/figure.png" pageId="14" pageNumber="435">Fig. 12B</figureCitation>
|
||
). In stylocellids, all sternites are fused except sternites 8, 9 and tergite IX that surround the anal plate. Margins between the fused tergites and sternites are completely absent in the amber specimen, whereas most modern
|
||
<taxonomicName id="4C71B2D6394BFD2E88DCFC86FB69FC92" authorityName="Simon" authorityYear="1879" box="[1022,1196,835,861]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="444" phylum="Arthropoda" rank="subOrder" subOrder="Cyphophthalmi">Cyphophthalmi</taxonomicName>
|
||
show at least a slight change in cuticle texture marking these borders. Besides this, there is an unusual setose region between the anterior coxae (
|
||
<figureCitation id="134AD5D0394BFD2E8BCBFC4EFE96FC6A" box="[233,339,907,933]" captionStart="FIGURE 12" captionStartId="16.[151,250,1107,1131]" captionTargetBox="[151,1436,417,1084]" captionTargetId="figure-157@16.[151,1436,417,1084]" captionTargetPageId="16" captionText="FIGURE 12. Details of Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Close-up of chelicerae, pedipalps and gonostome (indicated by arrows), ventral view. B. Close-up of opisthosoma, ventral view. Abbreviations: ca, corona analis; ch, chelicerae; go, gonostome; pp, pedipalp; oz, ozophore; legs II, III and IV numbered. Scale bars equal 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7984161" httpUri="https://zenodo.org/record/7984161/files/figure.png" pageId="14" pageNumber="435">Fig. 12A</figureCitation>
|
||
). The legs of the fossil are also untypical when compared to most extant
|
||
<taxonomicName id="4C71B2D6394BFD2E8FB4FC4EFA8DFC6A" authorityName="Simon" authorityYear="1879" box="[1174,1352,907,933]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="444" phylum="Arthropoda" rank="subOrder" subOrder="Cyphophthalmi">Cyphophthalmi</taxonomicName>
|
||
. Legs I and II seem to be directed forwards and the right leg III appears to be modified, as it looks more robust than the other legs. The latter is also covered by numerous thick setae, while the left leg III looks thinner, with normally sized setae (
|
||
<figureCitation id="134AD5D0394BFD2E8BFFFC32FEA6FBDE" box="[221,355,1015,1041]" captionStart="FIGURE 11" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[172,1416,181,1910]" captionTargetId="figure-20@15.[172,1416,181,1920]" captionTargetPageId="15" captionText="FIGURE 11. Foveacorpus cretaceus gen. et sp. nov., female holotype (GPIH05129). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ca, corona analis; ch, chelicerae; oz, ozophore; pp, pedipalp; legs numbered from I–IV. Scale bars equal 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7984159" httpUri="https://zenodo.org/record/7984159/files/figure.png" pageId="14" pageNumber="435">Fig. 11B, D</figureCitation>
|
||
). This limb condition (except for the modified leg III) is similar to a peculiar modern species from
|
||
<collectingCountry id="F36689C5394BFD2E8BB5FBDEFF26FBFA" box="[151,227,1051,1077]" name="Kenya" pageId="14" pageNumber="435">Kenya</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C71B2D6394BFD2E8BCCFBDEFDFEFBFA" authority="Shear, 1985" authorityName="Shear" authorityYear="1985" box="[238,571,1050,1077]" class="Arachnida" genus="Marwe" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="435" phylum="Arthropoda" rank="species" species="coarctata">
|
||
<emphasis id="B9051547394BFD2E8BCCFBDEFE68FBFB" box="[238,429,1051,1076]" italics="true" pageId="14" pageNumber="435">Marwe coarctata</emphasis>
|
||
<bibRefCitation id="EFE0B4A4394BFD2E8A96FBDEFDFEFBFA" author="Shear, W. A." box="[436,571,1050,1077]" pageId="14" pageNumber="435" pagination="1 - 6" refId="ref14255" refString="Shear, W. A. (1985) Marwe coarctata, a remarkable new cyphophthalmid from a limestone cave in Kenya (Arachnida, Opiliones). American Museum Novitates, 2830, 1 - 6." type="journal article" year="1985">Shear, 1985</bibRefCitation>
|
||
</taxonomicName>
|
||
, whose family assignment (like that of the fossil) is still unclear. Additionally, both species possess a corona analis,
|
||
<typeStatus id="54CA77F7394BFD2E891EFBFAFDA9FB96" box="[572,620,1087,1113]" pageId="14" pageNumber="435">type</typeStatus>
|
||
1 ozophores and dorsal areas without any visible sutures between them. On the other hand,
|
||
<taxonomicName id="4C71B2D6394BFD2E8A5AFBA6FDFEFBB3" authorityName="Shear" authorityYear="1985" box="[376,571,1123,1148]" class="Arachnida" genus="Marwe" kingdom="Animalia" order="Opiliones" pageId="14" pageNumber="435" phylum="Arthropoda" rank="species" species="coarctata">
|
||
<emphasis id="B9051547394BFD2E8A5AFBA6FDFEFBB3" box="[376,571,1123,1148]" italics="true" pageId="14" pageNumber="435">Marwe coarctata</emphasis>
|
||
</taxonomicName>
|
||
has a different habitus with an expanded, disc-like posterior portion of the prosoma, it is larger and has stouter chelicerae. In summary, the new fossil has a unique set of characters and cannot be easily placed within any of the extant families.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |