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<document id="614CD641BB219C299741393179191382" ID-CLB-Dataset="30502" ID-DOI="10.11646/zootaxa.4394.4.11" ID-GBIF-Dataset="b195c7f2-0a1a-4d59-ba88-7f1f95ac8486" ID-ISSN="1175-5326" ID-Zenodo-Dep="1199902" ID-ZooBank="F5A9FCF3-719D-4E1E-83DF-8D652EE2386B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1521181969001" checkinUser="plazi" docAuthor="Nadolny, Anton A." docDate="2018" docId="03D1BE0E95296611FF2A5F98FEF5E063" docLanguage="en" docName="zootaxa.4394.4.11.pdf" docOrigin="Zootaxa 4394 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alopecosa krynickii" docType="treatment" docVersion="8" lastPageNumber="599" masterDocId="FFE8C67695296614FFBD5B76FFABE322" masterDocTitle="Lost and found: Alopecosa krynickii (Araneae: Lycosidae) in the Crimea" masterLastPageNumber="599" masterPageNumber="594" pageNumber="594" updateTime="1698531946784" updateUser="ExternalLinkService">
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<mods:title id="7524C0BC0CA9299CD90E78A21F6966D0">Lost and found: Alopecosa krynickii (Araneae: Lycosidae) in the Crimea</mods:title>
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<mods:namePart id="E4FE3DA95636FD0FF59C0D4306968B53">Nadolny, Anton A.</mods:namePart>
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<mods:date id="704006B3FE2A50B9F6A5BF358B92BD6D">2018</mods:date>
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<mods:identifier id="1213FCD9E765CD61D44685E12D8DF6FA" type="DOI">10.11646/zootaxa.4394.4.11</mods:identifier>
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<treatment id="03D1BE0E95296611FF2A5F98FEF5E063" ID-DOI="http://doi.org/10.5281/zenodo.5978647" ID-GBIF-Taxon="142289703" ID-Zenodo-Dep="5978647" LSID="urn:lsid:plazi:treatment:03D1BE0E95296611FF2A5F98FEF5E063" httpUri="http://treatment.plazi.org/id/03D1BE0E95296611FF2A5F98FEF5E063" lastPageId="5" lastPageNumber="599" pageId="0" pageNumber="594">
<subSubSection id="C3625C9395296614FF2A5F98FEAAE60A" pageId="0" pageNumber="594" type="nomenclature">
<paragraph id="8BC70F1895296614FF2A5F98FDEEE62A" blockId="0.[151,885,1193,1424]" box="[151,581,1262,1288]" pageId="0" pageNumber="594">
<heading id="D08FB87495296614FF2A5F98FDEEE62A" bold="true" box="[151,581,1262,1288]" fontSize="11" level="1" pageId="0" pageNumber="594" reason="1">
<taxonomicName id="4C78749B95296614FF2A5F98FDEEE62A" ID-CoL="C4W8" authority="(Thorell, 1875)" baseAuthorityName="Thorell" baseAuthorityYear="1875" box="[151,581,1262,1288]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FF2A5F98FDEEE62A" bold="true" box="[151,581,1262,1288]" pageId="0" pageNumber="594">
<emphasis id="B90CD30A95296614FF2A5F98FE2DE62A" bold="true" box="[151,390,1262,1288]" italics="true" pageId="0" pageNumber="594">Alopecosa krynickii</emphasis>
(Thorell, 1875)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC70F1895296614FF2A5E67FEAAE60A" blockId="0.[151,885,1193,1424]" box="[151,257,1297,1320]" pageId="0" pageNumber="594">
<figureCitation id="1343139D95296614FF2A5E67FEAAE60A" box="[151,257,1297,1320]" captionStart-0="FIGURES1–7" captionStart-1="FIGURES 8–18" captionStart-2="FIGURES 19–21" captionStartId-0="2.[151,258,1941,1963]" captionStartId-1="3.[151,258,1668,1690]" captionStartId-2="4.[151,258,1725,1747]" captionTargetBox-0="[155,1431,201,1919]" captionTargetBox-1="[155,1432,198,1643]" captionTargetBox-2="[158,1429,573,1705]" captionTargetId-0="figure@2.[155,1432,191,1920]" captionTargetId-1="figure@3.[151,1436,191,1647]" captionTargetId-2="figure@4.[158,1429,573,1705]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." captionText-1="FIGURES 8–18. Copulatory organs of Alopecosa krynickii: 8 male palp, ventral; 9 bulbus, ventral-prolateral; 10 part of bulbus, retrolateral; 11–12 median apophysis, caudal and ventral; 13–14 embolic division, ventral and caudal-prolateral; 15 embolus, dorsal; 16–18 epigyne, ventral, dorsal, and dorsal-caudal. Abbreviations: AP anterior pocket of epigyne; D fertilization duct; FC functional conductor; MA median apophysis; P palea; S septum; ST spermatheca. Question mark indicates semi-transparent apophysis, explanationin the text.Scale bars 0.1 mm." captionText-2="FIGURES 19–21. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of Alopecosa krynickii. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of A.krynickii, where Villa Steven was situated.Map after Biodiversity Support Program (1999)." httpUri-0="https://zenodo.org/record/1199904/files/figure.png" httpUri-1="https://zenodo.org/record/1199908/files/figure.png" httpUri-2="https://zenodo.org/record/1199910/files/figure.png" pageId="0" pageNumber="594">Figs 1–21</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3625C9395296614FF2A5E42FC9CE6B2" pageId="0" pageNumber="594" type="reference_group">
<paragraph id="8BC70F1895296614FF2A5E42FC9CE6B2" blockId="0.[151,885,1193,1424]" pageId="0" pageNumber="594">
<treatmentCitationGroup id="AB68283695296614FF2A5E42FC9CE6B2" pageId="0" pageNumber="594">
<taxonomicName id="4C78749B95296614FF2A5E42FD91E669" authority="Thorell, 1875 a: 104" authorityName="Thorell" authorityPageNumber="104" authorityYear="1875" box="[151,570,1332,1355]" class="Arachnida" family="Lycosidae" genus="Tarentula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FF2A5E42FECBE669" box="[151,352,1332,1355]" italics="true" pageId="0" pageNumber="594">Tarentula krynickii</emphasis>
<treatmentCitation id="0AD9290995296614FEDA5E42FD91E669" author="" box="[359,570,1332,1355]" page="104" pageId="0" pageNumber="594">
<bibRefCitation id="EFE972E995296614FEDA5E42FDA3E669" box="[359,520,1332,1355]" pageId="0" pageNumber="594" refId="ref2707" refString="Thorell, T. (1875 a) Verzeichniss sudrussischer Spinnen. Horae Societatis entomologicae Rossicae, 11, 39 - 122." type="journal article">Thorell, 1875a</bibRefCitation>
: 104
</treatmentCitation>
</taxonomicName>
(♂);
<treatmentCitation id="0AD9290995296614FDC95E42FCE9E669" author="" box="[628,834,1332,1355]" page="157" pageId="0" pageNumber="594">
<bibRefCitation id="EFE972E995296614FDC95E42FCBBE669" box="[628,784,1332,1355]" pageId="0" pageNumber="594" refId="ref2730" refString="Thorell, T. (1875 b) Descriptions of several European and North African spiders. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar, 13 (5), 1 - 203." type="journal article">Thorell 1875b</bibRefCitation>
: 157
</treatmentCitation>
(♂).
<taxonomicName id="4C78749B95296614FF2A5E20FDFDE64F" ID-CoL="C4W8" authority=": Logunov 2013 a: 353" authorityName=": Logunov" authorityPageNumber="353" authorityYear="2013" box="[151,598,1366,1389]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FF2A5E20FEC2E64F" box="[151,361,1366,1389]" italics="true" pageId="0" pageNumber="594">Alopecosa krynickii</emphasis>
: Logunov 2013a: 353
</taxonomicName>
(♂). The complete reference list can be obtained from WSC (2017).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3625C9395296614FF2A5EC8FDD5E5EA" pageId="0" pageNumber="594" type="materials_examined">
<paragraph id="8BC70F1895296614FF2A5EC8FDD5E5EA" blockId="0.[151,1436,1470,2013]" pageId="0" pageNumber="594">
<emphasis id="B90CD30A95296614FF2A5EC8FEDAE6F7" bold="true" box="[151,369,1470,1493]" pageId="0" pageNumber="594">Material examined</emphasis>
. Crimea, Chernomorskoe Distr., Tarkhankut Peninsula: 1 ♀ (TNU-10059/2), Kipchak Valley, 8–
<date id="FFC629D895296614FF2A5E97FEABE6DA" box="[151,256,1505,1528]" pageId="0" pageNumber="594" value="2006-11-06">11.6.2006</date>
(M.M. Kovblyuk); 1 ♀ (TNU-10075/9), steppe between Medvedevo and Okunevka Villages,
<date id="FFC629D895296614FB4C5E97FAF0E6DA" box="[1265,1371,1505,1528]" pageId="0" pageNumber="594" value="2011-05-29">29.5.2011</date>
(O.V. Kukushkin); 1 juv (TNU-10010), steppe near Dzhangul coast, under a stone,
<date id="FFC629D895296614FC605D72FBECE539" box="[989,1095,1540,1563]" pageId="0" pageNumber="594" value="2014-07-19">19.7.2014</date>
(A.A. Nadolny); 1 ♀ 1 cocoon (TNU-10116/1), watershed of Malyi and Bolshoi Kastel Valleys, 45°26′58″N 32°32′50″E, steppe with
<emphasis id="B90CD30A95296614FA9A5D50FAC1E51F" box="[1319,1386,1574,1597]" italics="true" pageId="0" pageNumber="594">Stippa</emphasis>
sp.,
<taxonomicName id="4C78749B95296614FF2A5D3FFED9E542" box="[151,370,1609,1632]" class="Magnoliopsida" family="Asphodelaceae" genus="Asphodeline" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="0" pageNumber="594" phylum="Magnoliophyta" rank="species" species="taurica">
<emphasis id="B90CD30A95296614FF2A5D3FFED9E542" box="[151,370,1609,1632]" italics="true" pageId="0" pageNumber="594">Asphodeline taurica</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C78749B95296614FE085D3FFDE2E542" box="[437,585,1609,1632]" class="Magnoliopsida" family="Lamiaceae" genus="Salvia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="594" phylum="Tracheophyta" rank="species" species="nutans">
<emphasis id="B90CD30A95296614FE085D3FFDE2E542" box="[437,585,1609,1632]" italics="true" pageId="0" pageNumber="594">Salvia nutans</emphasis>
</taxonomicName>
, pitfalls, 13–
<date id="FFC629D895296614FD665D3FFCE3E542" box="[731,840,1609,1632]" pageId="0" pageNumber="594" value="2015-06-27">27.6.2015</date>
(A.A. Nadolny); 3 ♂♂ 2 ♀♀ 1 juv (TNU-10004/2), Bolshoi Kastel Valley, 45°27′11.9″N 32°33′29.1″E, at night,
<date id="FFC629D895296614FCA25D1AFC22E5A1" box="[799,905,1644,1667]" pageId="0" pageNumber="594" value="2015-09-19">19.9.2015</date>
(A.A. Nadolny); 5 ♂♂ 5 ♀♀
<docIssue id="CF1B70F795296614FB715D1AFB72E5A1" box="[1228,1241,1644,1667]" pageId="0" pageNumber="594">4</docIssue>
juv (TNU-10003/ 1) &amp; 1 ♂ 1 ♀ (MMUE-G7589.1), same locality, at night, 1–
<date id="FFC629D895296614FC815DF8FC00E587" box="[828,939,1678,1701]" pageId="0" pageNumber="594" value="2015-02-10">2.10.2015</date>
(A.A. Nadolny); 1 ♂ (TNU-10019/1), same locality, at night,
<date id="FFC629D895296614FEED5DC7FE63E5EA" box="[336,456,1713,1736]" pageId="0" pageNumber="594" value="2015-10-17">17.10.2015</date>
(A.A. Nadolny).
</paragraph>
</subSubSection>
<subSubSection id="C3625C9395296615FF7A5DA2FA3DE21B" lastPageId="1" lastPageNumber="595" pageId="0" pageNumber="594" type="diagnosis">
<paragraph id="8BC70F1895296614FF7A5DA2FDDDE4FF" blockId="0.[151,1436,1470,2013]" pageId="0" pageNumber="594">
<emphasis id="B90CD30A95296614FF7A5DA2FE9FE5C9" bold="true" box="[199,308,1748,1771]" pageId="0" pageNumber="594">Diagnosis</emphasis>
.
<taxonomicName id="4C78749B95296614FEFE5DA2FDBDE5C9" authorityName=": Logunov" authorityYear="2013" box="[323,534,1748,1771]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FEFE5DA2FDBDE5C9" box="[323,534,1748,1771]" italics="true" pageId="0" pageNumber="594">Alopecosa krynickii</emphasis>
</taxonomicName>
belongs to the
<emphasis id="B90CD30A95296614FD015DA2FCB4E5C9" box="[700,799,1748,1771]" italics="true" pageId="0" pageNumber="594">striatipes</emphasis>
species group (Logunov 2013a). The embolic division and median (=tegular) apophysis of
<taxonomicName id="4C78749B95296614FE435D80FDD6E42F" authorityName=": Logunov" authorityYear="2013" box="[510,637,1782,1805]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FE435D80FDD6E42F" box="[510,637,1782,1805]" italics="true" pageId="0" pageNumber="594">A. krynickii</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C78749B95296614FC325D80FB74E42F" authority="Logunov. Also" authorityName="Logunov. Also" box="[911,1247,1782,1805]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="marikovskii">
<emphasis id="B90CD30A95296614FC325D80FB85E42F" box="[911,1070,1782,1805]" italics="true" pageId="0" pageNumber="594">A. marikovskii</emphasis>
Logunov. Also
</taxonomicName>
, the males of
<taxonomicName id="4C78749B95296614FA385D80FF5FE412" authorityName=": Logunov" authorityYear="2013" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FA385D80FF5FE412" italics="true" pageId="0" pageNumber="594">A. krynickii</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="4C78749B95296614FE515C6FFD14E412" authority="(Thorell)" authorityName="Thorell" baseAuthorityName="Thorell" baseAuthorityYear="1875" box="[492,703,1817,1840]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B90CD30A95296614FE515C6FFDFCE412" box="[492,599,1817,1840]" italics="true" pageId="0" pageNumber="594">A. beckeri</emphasis>
(Thorell)
</taxonomicName>
and
<taxonomicName id="4C78749B95296614FD4E5C6FFBA5E412" authority="(Kulczynski)" authorityName="Kulczynski" baseAuthorityName="Kulczynski" baseAuthorityYear="1895" box="[755,1038,1817,1840]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="taeniopus">
<emphasis id="B90CD30A95296614FD4E5C6FFCD3E412" box="[755,888,1817,1840]" italics="true" pageId="0" pageNumber="594">A. taeniopus</emphasis>
(Kulczynski)
</taxonomicName>
: their embolic tips look like a singlebarbed harpoon. The embolus of
<taxonomicName id="4C78749B95296614FE415C4AFD3DE471" box="[508,662,1852,1875]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="marikovskii">
<emphasis id="B90CD30A95296614FE415C4AFD3DE471" box="[508,662,1852,1875]" italics="true" pageId="0" pageNumber="594">A. marikovskii</emphasis>
</taxonomicName>
is without harpoon. The epigyne of
<taxonomicName id="4C78749B95296614FBA25C4AFB31E471" authorityName=": Logunov" authorityYear="2013" box="[1055,1178,1852,1875]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FBA25C4AFB31E471" box="[1055,1178,1852,1875]" italics="true" pageId="0" pageNumber="594">A. krynickii</emphasis>
</taxonomicName>
is similar to those of
<taxonomicName id="4C78749B95296614FA385C4AFEE0E457" authority="Marusik" authorityName="Marusik" authorityYear="1995" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="akkolka">
<emphasis id="B90CD30A95296614FA385C4AFF41E457" italics="true" pageId="0" pageNumber="594">A. akkolka</emphasis>
Marusik
</taxonomicName>
,
<taxonomicName id="4C78749B95296614FEEB5C28FDB9E457" authority="Cordes" authorityName="Cordes" authorityYear="1996" box="[342,530,1886,1909]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="dryada">
<emphasis id="B90CD30A95296614FEEB5C28FE15E457" box="[342,446,1886,1909]" italics="true" pageId="0" pageNumber="594">A. dryada</emphasis>
Cordes
</taxonomicName>
,
<taxonomicName id="4C78749B95296614FDA05C28FC35E457" authority="Nadolny &amp; Ponomarev" authorityName="Nadolny &amp; Ponomarev" authorityYear="2012" box="[541,926,1886,1909]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="kovblyuki">
<emphasis id="B90CD30A95296614FDA05C28FD0BE457" box="[541,672,1886,1909]" italics="true" pageId="0" pageNumber="594">A. kovblyuki</emphasis>
Nadolny &amp; Ponomarev
</taxonomicName>
,
<taxonomicName id="4C78749B95296614FC155C28FBDFE457" authority="(Hahn)" authorityName="Hahn" baseAuthorityName="Hahn" baseAuthorityYear="1835" box="[936,1140,1886,1909]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="schmidti">
<emphasis id="B90CD30A95296614FC155C28FBB4E457" box="[936,1055,1886,1909]" italics="true" pageId="0" pageNumber="594">A. schmidti</emphasis>
(Hahn)
</taxonomicName>
, and
<taxonomicName id="4C78749B95296614FB165C28FADAE457" authority="(Pavesi)" authorityName="Pavesi" baseAuthorityName="Pavesi" baseAuthorityYear="1873" box="[1195,1393,1886,1909]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="sulzeri">
<emphasis id="B90CD30A95296614FB165C28FAA4E457" box="[1195,1295,1886,1909]" italics="true" pageId="0" pageNumber="594">A. sulzeri</emphasis>
(Pavesi)
</taxonomicName>
. All the listed species reliably differ from
<taxonomicName id="4C78749B95296614FD975CF7FD0EE4BA" authorityName=": Logunov" authorityYear="2013" box="[554,677,1921,1944]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="594" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95296614FD975CF7FD0EE4BA" box="[554,677,1921,1944]" italics="true" pageId="0" pageNumber="594">A. krynickii</emphasis>
</taxonomicName>
in the shape of median apophysis, the embolic division, the septum of epigyne, and spermathecae (cf. Lugetti &amp; Tongiorgi 1969; Eskov &amp; Marusik 1995; Cordes 1996;
<bibRefCitation id="EFE972E995296614FB0F5CD2FF7BE4FF" pageId="0" pageNumber="594" refId="ref2598" refString="Nadolny, A. A. &amp; Kovblyuk, M. M. (2010) On two closely related wolf spider species Alopecosa beckeri (Thorell, 1875) and A. taeniopus (Kulczynski, 1895) (Aranei: Lycosidae). Arthropoda Selecta, 19 (4), 237 - 247." type="journal article">Nadolny &amp; Kovblyuk 2010</bibRefCitation>
;
<bibRefCitation id="EFE972E995296614FF675CB0FE19E4FF" box="[218,434,1990,2013]" pageId="0" pageNumber="594" refId="ref2655" refString="Nadolny, A. A., Ponomarev, A. V. &amp; Dvadnenko, K. V. (2012) A new wolf spider species in the genus Alopecosa Simon, 1885 (Araneae: Lycosidae) from Eastern Europe. Zootaxa, 3484, 83 - 88." type="journal article">
Nadolny
<emphasis id="B90CD30A95296614FE815CB0FEC0E4FF" box="[316,363,1990,2013]" italics="true" pageId="0" pageNumber="594">et al</emphasis>
. 2012
</bibRefCitation>
; Logunov 2013b).
</paragraph>
<paragraph id="8BC70F1895286615FF7A5BEEFA3DE21B" blockId="1.[151,1436,152,348]" pageId="1" pageNumber="595">
<emphasis id="B90CD30A95286615FF7A5BEEFF57E38D" bold="true" box="[199,252,152,175]" pageId="1" pageNumber="595">Note</emphasis>
. The venter colour pattern in
<taxonomicName id="4C78749B95286615FDEA5BEEFD6DE38D" authorityName="Simon" authorityYear="1885" box="[599,710,152,175]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B90CD30A95286615FDEA5BEEFD6DE38D" box="[599,710,152,175]" italics="true" pageId="1" pageNumber="595">Alopecosa</emphasis>
</taxonomicName>
species can be black—in
<taxonomicName id="4C78749B95286615FC485BEEFBDAE38D" baseAuthorityName="Hahn" baseAuthorityYear="1835" box="[1013,1137,152,175]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="schmidti">
<emphasis id="B90CD30A95286615FC485BEEFBDAE38D" box="[1013,1137,152,175]" italics="true" pageId="1" pageNumber="595">A. schmidti</emphasis>
</taxonomicName>
, yellowish—
<taxonomicName id="4C78749B95286615FABB5BEEFF52E3F3" authority="(Thorell)" authorityName="Thorell" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="cronebergi">
<emphasis id="B90CD30A95286615FABB5BEEFA37E38D" box="[1286,1436,152,175]" italics="true" pageId="1" pageNumber="595">A. cronebergi</emphasis>
(Thorell)
</taxonomicName>
, with a large black spot behind the epigastric furrow—
<taxonomicName id="4C78749B95286615FCE15BCCFC52E3F3" box="[860,1017,186,209]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="marikovskii">
<emphasis id="B90CD30A95286615FCE15BCCFC52E3F3" box="[860,1017,186,209]" italics="true" pageId="1" pageNumber="595">A. marikovskii</emphasis>
</taxonomicName>
, and with a black area in front of the epigastric furrow—
<taxonomicName id="4C78749B95286615FED25BABFDE2E3D6" authority="Esyunin." authorityName="Esyunin." box="[367,585,221,244]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="uiensis">
<emphasis id="B90CD30A95286615FED25BABFE76E3D6" box="[367,477,221,244]" italics="true" pageId="1" pageNumber="595">A. uiensis</emphasis>
Esyunin.
</taxonomicName>
Juveniles and adults of
<taxonomicName id="4C78749B95286615FCD75BABFC40E3D6" authorityName=": Logunov" authorityYear="2013" box="[874,1003,221,244]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95286615FCD75BABFC40E3D6" box="[874,1003,221,244]" italics="true" pageId="1" pageNumber="595">A. krynickii</emphasis>
</taxonomicName>
have the uniquely bicolour venter of abdomen: black areas in front and behind the epigastric furrow, but the rear end of venter is light (
<figureCitation id="1343139D95286615FB4D5A76FAFEE235" box="[1264,1365,256,279]" captionStart="FIGURES1–7" captionStartId="2.[151,258,1941,1963]" captionTargetBox="[155,1431,201,1919]" captionTargetId="figure@2.[155,1432,191,1920]" captionTargetPageId="2" captionText="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="1" pageNumber="595">Figs 5–7</figureCitation>
). This character distinguishes
<taxonomicName id="4C78749B95286615FE2C5A54FDA7E21B" authorityName=": Logunov" authorityYear="2013" box="[401,524,290,313]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95286615FE2C5A54FDA7E21B" box="[401,524,290,313]" italics="true" pageId="1" pageNumber="595">A. krynickii</emphasis>
</taxonomicName>
from all other species of
<taxonomicName id="4C78749B95286615FCA15A54FC20E21B" authorityName="Simon" authorityYear="1885" box="[796,907,290,313]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="genus">
<emphasis id="B90CD30A95286615FCA15A54FC20E21B" box="[796,907,290,313]" italics="true" pageId="1" pageNumber="595">Alopecosa</emphasis>
</taxonomicName>
in the
<collectingRegion id="49BCC1FA95286615FC695A54FB8EE21B" box="[980,1061,290,313]" country="Ukraine" name="Crimea" pageId="1" pageNumber="595">Crimea</collectingRegion>
, and perhaps in the whole Europe.
</paragraph>
</subSubSection>
<subSubSection id="C3625C9395286610FF7A5A33FF5AE2C5" lastPageId="4" lastPageNumber="598" pageId="1" pageNumber="595" type="description">
<paragraph id="8BC70F1895286615FF7A5A33FAF8E27E" blockId="1.[151,1436,152,348]" box="[199,1363,325,348]" pageId="1" pageNumber="595">
<emphasis id="B90CD30A95286615FF7A5A33FEE0E27E" bold="true" box="[199,331,325,348]" pageId="1" pageNumber="595">Description</emphasis>
. Measurements of female / male: total length 16 / 13; carapace: 7.5 / 7.2 long and 5.0 / 4.8 wide.
</paragraph>
<caption id="DF075F9095286615FF2A5AFCFC5CE283" box="[151,1015,394,417]" pageId="1" pageNumber="595">
<paragraph id="8BC70F1895286615FF2A5AFCFC5CE283" blockId="1.[151,1015,394,417]" box="[151,1015,394,417]" pageId="1" pageNumber="595">
<emphasis id="B90CD30A95286615FF2A5AFCFEA1E283" bold="true" box="[151,266,394,417]" pageId="1" pageNumber="595">TABLE 1.</emphasis>
Female / male palp and leg segment measures of
<taxonomicName id="4C78749B95286615FCA05AFCFC5BE283" authorityName=": Logunov" authorityYear="2013" box="[797,1008,394,417]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95286615FCA05AFCFC5BE283" box="[797,1008,394,417]" italics="true" pageId="1" pageNumber="595">Alopecosa krynickii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC70F1895286615FEC85ACBFAB5E1B8" pageId="1" pageNumber="595">
<table id="F978FDB8952899EBFF225ACBFAB5E1B8" box="[159,1310,445,666]" gridcols="6" gridrows="6" pageId="1" pageNumber="595">
<tr id="35480D5A952899EBFF225ACBFAB5E2F0" box="[159,1310,445,466]" gridrow="0" pageId="1" pageNumber="595" rowspan-0="1">
<th id="76996426952899EBFEC85ACBFE6FE2F0" box="[373,452,445,466]" gridcol="1" gridrow="0" pageId="1" pageNumber="595">Femur</th>
<th id="76996426952899EBFDF15ACBFD30E2F0" box="[588,667,445,466]" gridcol="2" gridrow="0" pageId="1" pageNumber="595">Patella</th>
<th id="76996426952899EBFC9E5ACBFCDAE2F0" box="[803,881,445,466]" gridcol="3" gridrow="0" pageId="1" pageNumber="595">Tibia</th>
<th id="76996426952899EBFC445ACBFBC9E2F0" box="[1017,1122,445,466]" gridcol="4" gridrow="0" pageId="1" pageNumber="595">Metatarsus</th>
<th id="76996426952899EBFB6D5ACBFAB5E2F0" box="[1232,1310,445,466]" gridcol="5" gridrow="0" pageId="1" pageNumber="595">Tarsus</th>
</tr>
<tr id="35480D5A952899EBFF225A93FAB5E2D8" box="[159,1310,485,506]" gridrow="1" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225A93FF4FE2D8" box="[159,228,485,506]" gridcol="0" gridrow="1" pageId="1" pageNumber="595">Palp</th>
<td id="76996426952899EBFEC85A93FE6FE2D8" box="[373,452,485,506]" gridcol="1" gridrow="1" pageId="1" pageNumber="595">2.7 / 3.0</td>
<td id="76996426952899EBFDF15A93FD30E2D8" box="[588,667,485,506]" gridcol="2" gridrow="1" pageId="1" pageNumber="595">1.3 / 1.5</td>
<td id="76996426952899EBFC9E5A93FCDAE2D8" box="[803,881,485,506]" gridcol="3" gridrow="1" pageId="1" pageNumber="595">1.5 / 1.6</td>
<td id="76996426952899EBFC445A93FBC9E2D8" box="[1017,1122,485,506]" gridcol="4" gridrow="1" pageId="1" pageNumber="595">–</td>
<td id="76996426952899EBFB6D5A93FAB5E2D8" box="[1232,1310,485,506]" gridcol="5" gridrow="1" pageId="1" pageNumber="595">2.1 / 2.4</td>
</tr>
<tr id="35480D5A952899EBFF22597BFAB5E100" box="[159,1310,525,546]" gridrow="2" pageId="1" pageNumber="595">
<th id="76996426952899EBFF22597BFF4FE100" box="[159,228,525,546]" gridcol="0" gridrow="2" pageId="1" pageNumber="595">Leg I</th>
<td id="76996426952899EBFEC8597BFE6FE100" box="[373,452,525,546]" gridcol="1" gridrow="2" pageId="1" pageNumber="595">5.2 / 5.5</td>
<td id="76996426952899EBFDF1597BFD30E100" box="[588,667,525,546]" gridcol="2" gridrow="2" pageId="1" pageNumber="595">2.6 / 2.8</td>
<td id="76996426952899EBFC9E597BFCDAE100" box="[803,881,525,546]" gridcol="3" gridrow="2" pageId="1" pageNumber="595">3.8 / 4.2</td>
<td id="76996426952899EBFC44597BFBC9E100" box="[1017,1122,525,546]" gridcol="4" gridrow="2" pageId="1" pageNumber="595">3.8 / 4.7</td>
<td id="76996426952899EBFB6D597BFAB5E100" box="[1232,1310,525,546]" gridcol="5" gridrow="2" pageId="1" pageNumber="595">2.2 / 2.6</td>
</tr>
<tr id="35480D5A952899EBFF225943FAB5E168" box="[159,1310,565,586]" gridrow="3" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225943FF4FE168" box="[159,228,565,586]" gridcol="0" gridrow="3" pageId="1" pageNumber="595">Leg II</th>
<td id="76996426952899EBFEC85943FE6FE168" box="[373,452,565,586]" gridcol="1" gridrow="3" pageId="1" pageNumber="595">4.9 / 5.2</td>
<td id="76996426952899EBFDF15943FD30E168" box="[588,667,565,586]" gridcol="2" gridrow="3" pageId="1" pageNumber="595">2.6 / 2.6</td>
<td id="76996426952899EBFC9E5943FCDAE168" box="[803,881,565,586]" gridcol="3" gridrow="3" pageId="1" pageNumber="595">3.4 / 3.9</td>
<td id="76996426952899EBFC445943FBC9E168" box="[1017,1122,565,586]" gridcol="4" gridrow="3" pageId="1" pageNumber="595">3.6 / 4.6</td>
<td id="76996426952899EBFB6D5943FAB5E168" box="[1232,1310,565,586]" gridcol="5" gridrow="3" pageId="1" pageNumber="595">2.2 / 2.5</td>
</tr>
<tr id="35480D5A952899EBFF22592BFAB5E150" box="[159,1310,605,626]" gridrow="4" pageId="1" pageNumber="595">
<th id="76996426952899EBFF22592BFF4FE150" box="[159,228,605,626]" gridcol="0" gridrow="4" pageId="1" pageNumber="595">Leg III</th>
<td id="76996426952899EBFEC8592BFE6FE150" box="[373,452,605,626]" gridcol="1" gridrow="4" pageId="1" pageNumber="595">4.3 / 4.8</td>
<td id="76996426952899EBFDF1592BFD30E150" box="[588,667,605,626]" gridcol="2" gridrow="4" pageId="1" pageNumber="595">2.2 / 2.4</td>
<td id="76996426952899EBFC9E592BFCDAE150" box="[803,881,605,626]" gridcol="3" gridrow="4" pageId="1" pageNumber="595">3.0 / 3.4</td>
<td id="76996426952899EBFC44592BFBC9E150" box="[1017,1122,605,626]" gridcol="4" gridrow="4" pageId="1" pageNumber="595">4.0 / 4.6</td>
<td id="76996426952899EBFB6D592BFAB5E150" box="[1232,1310,605,626]" gridcol="5" gridrow="4" pageId="1" pageNumber="595">2.0 / 2.2</td>
</tr>
<tr id="35480D5A952899EBFF2259F3FAB5E1B8" box="[159,1310,645,666]" gridrow="5" pageId="1" pageNumber="595">
<th id="76996426952899EBFF2259F3FF4FE1B8" box="[159,228,645,666]" gridcol="0" gridrow="5" pageId="1" pageNumber="595">Leg IV</th>
<td id="76996426952899EBFEC859F3FE6FE1B8" box="[373,452,645,666]" gridcol="1" gridrow="5" pageId="1" pageNumber="595">5.6 / 6.0</td>
<td id="76996426952899EBFDF159F3FD30E1B8" box="[588,667,645,666]" gridcol="2" gridrow="5" pageId="1" pageNumber="595">2.5 / 2.6</td>
<td id="76996426952899EBFC9E59F3FCDAE1B8" box="[803,881,645,666]" gridcol="3" gridrow="5" pageId="1" pageNumber="595">4.2 / 4.6</td>
<td id="76996426952899EBFC4459F3FBC9E1B8" box="[1017,1122,645,666]" gridcol="4" gridrow="5" pageId="1" pageNumber="595">6.0 / 6.6</td>
<td id="76996426952899EBFB6D59F3FAB5E1B8" box="[1232,1310,645,666]" gridcol="5" gridrow="5" pageId="1" pageNumber="595">2.6 / 2.8</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC70F1895286615FF7A5996FBB7E1D5" blockId="1.[199,1052,736,759]" box="[199,1052,736,759]" pageId="1" pageNumber="595">Female palp spination: femur d1-1-1, p1, r1; patella p1; tibia p2; tarsus p2-1, r1.</paragraph>
<caption id="DF075F9095286615FF2A5853FCE0E01E" box="[151,843,805,828]" pageId="1" pageNumber="595">
<paragraph id="8BC70F1895286615FF2A5853FCE0E01E" blockId="1.[151,843,805,828]" box="[151,843,805,828]" pageId="1" pageNumber="595">
<emphasis id="B90CD30A95286615FF2A5853FEA2E01E" bold="true" box="[151,265,805,828]" pageId="1" pageNumber="595">TABLE 2.</emphasis>
Female leg segment spination of
<taxonomicName id="4C78749B95286615FDCF5853FCEFE01E" authorityName=": Logunov" authorityYear="2013" box="[626,836,805,828]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A95286615FDCF5853FCEFE01E" box="[626,836,805,828]" italics="true" pageId="1" pageNumber="595">Alopecosa krynickii</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BC70F1895286615FEF55821FAF6E72E" pageId="1" pageNumber="595">
<table id="F978FDB8952899EBFF225821FAF6E72E" box="[159,1373,855,1036]" gridcols="5" gridrows="5" pageId="1" pageNumber="595" rowsContinueIn="1.[159,1354,1226,1407]">
<tr id="35480D5A952899EBFF225821FAF6E04E" box="[159,1373,855,876]" gridrow="0" pageId="1" pageNumber="595" rowspan-0="1">
<th id="76996426952899EBFEF55821FDA5E04E" box="[328,526,855,876]" gridcol="1" gridrow="0" pageId="1" pageNumber="595">Femur</th>
<th id="76996426952899EBFDD25821FD1BE04E" box="[623,688,855,876]" gridcol="2" gridrow="0" pageId="1" pageNumber="595">Patella</th>
<th id="76996426952899EBFC9D5821FBA0E04E" box="[800,1035,855,876]" gridcol="3" gridrow="0" pageId="1" pageNumber="595">Tibia</th>
<th id="76996426952899EBFBDE5821FAF6E04E" box="[1123,1373,855,876]" gridcol="4" gridrow="0" pageId="1" pageNumber="595">Metatarsus</th>
</tr>
<tr id="35480D5A952899EBFF225809FAF6E0B6" box="[159,1373,895,916]" gridrow="1" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225809FF4FE0B6" box="[159,228,895,916]" gridcol="0" gridrow="1" pageId="1" pageNumber="595">Leg I</th>
<td id="76996426952899EBFEF55809FDA5E0B6" box="[328,526,895,916]" gridcol="1" gridrow="1" pageId="1" pageNumber="595">d1-1-1, p1, r1-1-1.</td>
<td id="76996426952899EBFDD25809FD1BE0B6" box="[623,688,895,916]" gridcol="2" gridrow="1" pageId="1" pageNumber="595">p1.</td>
<td id="76996426952899EBFC9D5809FBA0E0B6" box="[800,1035,895,916]" gridcol="3" gridrow="1" pageId="1" pageNumber="595">p1-1, v2-2-2.</td>
<td id="76996426952899EBFBDE5809FAF6E0B6" box="[1123,1373,895,916]" gridcol="4" gridrow="1" pageId="1" pageNumber="595">p1a, v2-2-3a.</td>
</tr>
<tr id="35480D5A952899EBFF2258D1FAF6E09E" box="[159,1373,935,956]" gridrow="2" pageId="1" pageNumber="595">
<th id="76996426952899EBFF2258D1FF4FE09E" box="[159,228,935,956]" gridcol="0" gridrow="2" pageId="1" pageNumber="595">Leg II</th>
<td id="76996426952899EBFEF558D1FDA5E09E" box="[328,526,935,956]" gridcol="1" gridrow="2" pageId="1" pageNumber="595">d1-1-1, p1, r1-1-1.</td>
<td id="76996426952899EBFDD258D1FD1BE09E" box="[623,688,935,956]" gridcol="2" gridrow="2" pageId="1" pageNumber="595">p1.</td>
<td id="76996426952899EBFC9D58D1FBA0E09E" box="[800,1035,935,956]" gridcol="3" gridrow="2" pageId="1" pageNumber="595">p1-1, v2-2-2.</td>
<td id="76996426952899EBFBDE58D1FAF6E09E" box="[1123,1373,935,956]" gridcol="4" gridrow="2" pageId="1" pageNumber="595">p1-1a, v2-2-3a.</td>
</tr>
<tr id="35480D5A952899EBFF2258B9FAF6E0C6" box="[159,1373,975,996]" gridrow="3" pageId="1" pageNumber="595">
<th id="76996426952899EBFF2258B9FF4FE0C6" box="[159,228,975,996]" gridcol="0" gridrow="3" pageId="1" pageNumber="595">Leg III</th>
<td id="76996426952899EBFEF558B9FDA5E0C6" box="[328,526,975,996]" gridcol="1" gridrow="3" pageId="1" pageNumber="595">d1-1-1, p1-1, r1-2-1.</td>
<td id="76996426952899EBFDD258B9FD1BE0C6" box="[623,688,975,996]" gridcol="2" gridrow="3" pageId="1" pageNumber="595">p1, r1.</td>
<td id="76996426952899EBFC9D58B9FBA0E0C6" box="[800,1035,975,996]" gridcol="3" gridrow="3" pageId="1" pageNumber="595">d1-1, p1-1, r1-1, v2-2-2.</td>
<td id="76996426952899EBFBDE58B9FAF6E0C6" box="[1123,1373,975,996]" gridcol="4" gridrow="3" pageId="1" pageNumber="595">p1-1-1a, r1-1-1a, v2-2-3a.</td>
</tr>
<tr id="35480D5A952899EBFF225881FAF6E72E" box="[159,1373,1015,1036]" gridrow="4" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225881FF4FE72E" box="[159,228,1015,1036]" gridcol="0" gridrow="4" pageId="1" pageNumber="595">Leg IV</th>
<td id="76996426952899EBFEF55881FDA5E72E" box="[328,526,1015,1036]" gridcol="1" gridrow="4" pageId="1" pageNumber="595">d1-1-1, p1-1, r1-2-1.</td>
<td id="76996426952899EBFDD25881FD1BE72E" box="[623,688,1015,1036]" gridcol="2" gridrow="4" pageId="1" pageNumber="595">p1, r1.</td>
<td id="76996426952899EBFC9D5881FBA0E72E" box="[800,1035,1015,1036]" gridcol="3" gridrow="4" pageId="1" pageNumber="595">d1-1, p1-1, r1-1, v2-2-2.</td>
<td id="76996426952899EBFBDE5881FAF6E72E" box="[1123,1373,1015,1036]" gridcol="4" gridrow="4" pageId="1" pageNumber="595">p1-1-1a, r1-1-1a, v2-2-3a.</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC70F1895286615FF7A5F24FCDFE74B" blockId="1.[199,884,1106,1129]" box="[199,884,1106,1129]" pageId="1" pageNumber="595">Male palp spination: femur d1-1-1-2, p1, r1; patella p1; tibia p2.</paragraph>
<caption id="DF075F9095286615FF2A5FEEFC86E78D" box="[151,813,1176,1199]" pageId="1" pageNumber="595">
<paragraph id="8BC70F1895286615FF2A5FEEFC86E78D" blockId="1.[151,813,1176,1199]" box="[151,813,1176,1199]" pageId="1" pageNumber="595">
<emphasis id="B90CD30A95286615FF2A5FEEFEA2E78D" bold="true" box="[151,265,1176,1199]" pageId="1" pageNumber="595">TABLE 3.</emphasis>
Male leg segment spination of
<taxonomicName id="4C78749B95286615FDE75FEEFC86E78D" authorityName=": Logunov" authorityYear="2013" box="[602,813,1176,1199]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="595" phylum="Arthropoda" rank="species" species="krynickii">Alopecosa krynickii</taxonomicName>
</paragraph>
</caption>
<paragraph id="8BC70F1895286615FE825FBCFAE1E65D" pageId="1" pageNumber="595">
<table id="F978FDB8952899EBFF225FBCFAE1E65D" box="[159,1354,1226,1407]" gridcols="5" gridrows="5" pageId="1" pageNumber="595" rowsContinueFrom="1.[159,1373,855,1036]">
<tr id="35480D5A952899EBFF225FBCFAE1E7FD" box="[159,1354,1226,1247]" gridrow="0" pageId="1" pageNumber="595" rowspan-0="1">
<th id="76996426952899EBFE825FBCFDAFE7FD" box="[319,516,1226,1247]" gridcol="1" gridrow="0" pageId="1" pageNumber="595">Femur</th>
<th id="76996426952899EBFDD25FBCFD1BE7FD" box="[623,688,1226,1247]" gridcol="2" gridrow="0" pageId="1" pageNumber="595">Patella</th>
<th id="76996426952899EBFD475FBCFC44E7FD" box="[762,1007,1226,1247]" gridcol="3" gridrow="0" pageId="1" pageNumber="595">Tibia</th>
<th id="76996426952899EBFBED5FBCFAE1E7FD" box="[1104,1354,1226,1247]" gridcol="4" gridrow="0" pageId="1" pageNumber="595">Metatarsus</th>
</tr>
<tr id="35480D5A952899EBFF225F84FAE1E625" box="[159,1354,1266,1287]" gridrow="1" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225F84FF4FE625" box="[159,228,1266,1287]" gridcol="0" gridrow="1" pageId="1" pageNumber="595">Leg I</th>
<td id="76996426952899EBFE825F84FDAFE625" box="[319,516,1266,1287]" gridcol="1" gridrow="1" pageId="1" pageNumber="595">d1-1-1, p1, r1-1-1.</td>
<td id="76996426952899EBFDD25F84FD1BE625" box="[623,688,1266,1287]" gridcol="2" gridrow="1" pageId="1" pageNumber="595">p1, r1.</td>
<td id="76996426952899EBFD475F84FC44E625" box="[762,1007,1266,1287]" gridcol="3" gridrow="1" pageId="1" pageNumber="595">d1, p1-1, r1-1, v2-2-2a.</td>
<td id="76996426952899EBFBED5F84FAE1E625" box="[1104,1354,1266,1287]" gridcol="4" gridrow="1" pageId="1" pageNumber="595">p1-1-1a, r1-1-1a, v2-2-3a.</td>
</tr>
<tr id="35480D5A952899EBFF225E6CFAE1E60D" box="[159,1354,1306,1327]" gridrow="2" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225E6CFF4FE60D" box="[159,228,1306,1327]" gridcol="0" gridrow="2" pageId="1" pageNumber="595">Leg II</th>
<td id="76996426952899EBFE825E6CFDAFE60D" box="[319,516,1306,1327]" gridcol="1" gridrow="2" pageId="1" pageNumber="595">d1-1-1, p1-1, r1-1-1.</td>
<td id="76996426952899EBFDD25E6CFD1BE60D" box="[623,688,1306,1327]" gridcol="2" gridrow="2" pageId="1" pageNumber="595">p1, r1.</td>
<td id="76996426952899EBFD475E6CFC44E60D" box="[762,1007,1306,1327]" gridcol="3" gridrow="2" pageId="1" pageNumber="595">d1, p1-1, r1-1, v2-2-2a.</td>
<td id="76996426952899EBFBED5E6CFAE1E60D" box="[1104,1354,1306,1327]" gridcol="4" gridrow="2" pageId="1" pageNumber="595">p1-1-1a, r1-1-1a, v2-2-3a.</td>
</tr>
<tr id="35480D5A952899EBFF225E34FAE1E675" box="[159,1354,1346,1367]" gridrow="3" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225E34FF4FE675" box="[159,228,1346,1367]" gridcol="0" gridrow="3" pageId="1" pageNumber="595">Leg III</th>
<td id="76996426952899EBFE825E34FDAFE675" box="[319,516,1346,1367]" gridcol="1" gridrow="3" pageId="1" pageNumber="595">d1-1-1, p1-1, r1-1-1.</td>
<td id="76996426952899EBFDD25E34FD1BE675" box="[623,688,1346,1367]" gridcol="2" gridrow="3" pageId="1" pageNumber="595">p1, r1.</td>
<td id="76996426952899EBFD475E34FC44E675" box="[762,1007,1346,1367]" gridcol="3" gridrow="3" pageId="1" pageNumber="595">d1-1, p1-1, r1-1, v2-2-2a.</td>
<td id="76996426952899EBFBED5E34FAE1E675" box="[1104,1354,1346,1367]" gridcol="4" gridrow="3" pageId="1" pageNumber="595">p1-1-1a, r1-1-1a, v2-2-3a.</td>
</tr>
<tr id="35480D5A952899EBFF225E1CFAE1E65D" box="[159,1354,1386,1407]" gridrow="4" pageId="1" pageNumber="595">
<th id="76996426952899EBFF225E1CFF4FE65D" box="[159,228,1386,1407]" gridcol="0" gridrow="4" pageId="1" pageNumber="595">Leg IV</th>
<td id="76996426952899EBFE825E1CFDAFE65D" box="[319,516,1386,1407]" gridcol="1" gridrow="4" pageId="1" pageNumber="595">d1-1-1, p1-1, r1.</td>
<td id="76996426952899EBFDD25E1CFD1BE65D" box="[623,688,1386,1407]" gridcol="2" gridrow="4" pageId="1" pageNumber="595">p1, r1.</td>
<td id="76996426952899EBFD475E1CFC44E65D" box="[762,1007,1386,1407]" gridcol="3" gridrow="4" pageId="1" pageNumber="595">d1-1, p1-1, r1-1, v2-2-2a.</td>
<td id="76996426952899EBFBED5E1CFAE1E65D" box="[1104,1354,1386,1407]" gridcol="4" gridrow="4" pageId="1" pageNumber="595">p1-1-1a, r1-1-1a, v2-2-3a.</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC70F1895286615FF7A5EB3FB70E4E3" blockId="1.[151,1436,1477,1985]" pageId="1" pageNumber="595">
FEMALE. Habitus is shown in
<figureCitation id="1343139D95286615FD9A5EB3FD26E6FE" box="[551,653,1477,1500]" captionStart="FIGURES1–7" captionStartId="2.[151,258,1941,1963]" captionTargetBox="[155,1431,201,1919]" captionTargetId="figure@2.[155,1432,191,1920]" captionTargetPageId="2" captionText="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="1" pageNumber="595">Figs 1, 6</figureCitation>
. Prosoma: Carapace with a poorly marked gradual descend of thoracic region, eye field not elevated (
<figureCitation id="1343139D95286615FE5D5E9EFD89E6DD" box="[480,546,1512,1535]" captionStart="FIGURES1–7" captionStartId="2.[151,258,1941,1963]" captionTargetBox="[155,1431,201,1919]" captionTargetId="figure@2.[155,1432,191,1920]" captionTargetPageId="2" captionText="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="1" pageNumber="595">Fig. 3</figureCitation>
). Carapace brown, with light lines radiating from median oval spot of white setae, lateral wide stripes of black setae, submarginal narrow stripes of white setae, and marginal broken stripes of black setae. Eye area is covered with light setae. Behind eyes there are four black spots (better visible in the males). Clypeus covered with white setae. Sternum and labium brown, covered with black setae. Chelicerae brown, their anterior side (except the distal part) covered with white setae, other sides covered with black setae. Palps light brown, with white and black setae. Coxae brown, covered with black setae. Other leg segments light brown, mainly covered with white setae; black setae forms spots on femora, also all femora with narrow retrolateral longitudinal stripes of black setae. Tarsi I–IV and metatarsi I–II with scopula. Scopulae of tarsi I–II consist of grey setae, III–IV consist of grey and black setae. Spinules are absent from all legs. Ventral sides of tibiae I–II with grey setae (similar to those of scopula). Opisthosoma: Dorsum dark grey, mainly covered with black setae; white setae forms a pattern of lanceolate cardiac mark, two longitudinal parallel lines of rounded spots and a line of oval spots in between them. Venter light brown or yellowish, the anterior half dark, covered with black setae, the posterior half light, mainly with white setae. In adults, this black spot occupies not less than a half of the venter (in juveniles less than a half,
<figureCitation id="1343139D95286615FCAF5CFEFCFFE4BD" box="[786,852,1928,1951]" captionStart="FIGURES1–7" captionStartId="2.[151,258,1941,1963]" captionTargetBox="[155,1431,201,1919]" captionTargetId="figure@2.[155,1432,191,1920]" captionTargetPageId="2" captionText="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="1" pageNumber="595">Fig. 5</figureCitation>
). Copulatory organs as in
<figureCitation id="1343139D95286615FBCE5CFEFB44E4BD" box="[1139,1263,1928,1951]" captionStart="FIGURES 8–18" captionStartId="3.[151,258,1668,1690]" captionTargetBox="[155,1432,198,1643]" captionTargetId="figure@3.[151,1436,191,1647]" captionTargetPageId="3" captionText="FIGURES 8–18. Copulatory organs of Alopecosa krynickii: 8 male palp, ventral; 9 bulbus, ventral-prolateral; 10 part of bulbus, retrolateral; 11–12 median apophysis, caudal and ventral; 13–14 embolic division, ventral and caudal-prolateral; 15 embolus, dorsal; 16–18 epigyne, ventral, dorsal, and dorsal-caudal. Abbreviations: AP anterior pocket of epigyne; D fertilization duct; FC functional conductor; MA median apophysis; P palea; S septum; ST spermatheca. Question mark indicates semi-transparent apophysis, explanationin the text.Scale bars 0.1 mm." httpUri="https://zenodo.org/record/1199908/files/figure.png" pageId="1" pageNumber="595">Figs 16–18</figureCitation>
: epigyne with 2 small anterior pockets, septum vase-shaped, with a median longitudinal ridge, spermathecae spherical.
</paragraph>
<caption id="DF075F90952B6616FF2A5CE3FCDAE4EA" httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="2" pageNumber="596" startId="2.[151,258,1941,1963]" targetBox="[155,1431,201,1919]" targetPageId="2">
<paragraph id="8BC70F18952B6616FF2A5CE3FCDAE4EA" blockId="2.[151,1436,1941,1992]" pageId="2" pageNumber="596">
<emphasis id="B90CD30A952B6616FF2A5CE3FE80E489" bold="true" box="[151,299,1941,1963]" pageId="2" pageNumber="596">FIGURES 1–7</emphasis>
. Habitus of
<taxonomicName id="4C78749B952B6616FE1C5CE3FDF4E488" authorityName=": Logunov" authorityYear="2013" box="[417,607,1941,1962]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="596" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952B6616FE1C5CE3FDF4E488" box="[417,607,1941,1962]" italics="true" pageId="2" pageNumber="596">Alopecosa krynickii</emphasis>
</taxonomicName>
: 1–2 dorsum of female and male; 3–4 carapace profile of female and male; 5–7 venter of juvenile, female, and male. Arrows indicate position of epigastric furrow.
</paragraph>
</caption>
<caption id="DF075F90952A6617FF2A5DF2FD9AE42D" httpUri="https://zenodo.org/record/1199908/files/figure.png" pageId="3" pageNumber="597" startId="3.[151,258,1668,1690]" targetBox="[155,1432,198,1643]" targetPageId="3">
<paragraph id="8BC70F18952A6617FF2A5DF2FD9AE42D" blockId="3.[151,1436,1668,1807]" pageId="3" pageNumber="597">
<emphasis id="B90CD30A952A6617FF2A5DF2FE90E5B8" bold="true" box="[151,315,1668,1690]" pageId="3" pageNumber="597">FIGURES 8–18</emphasis>
. Copulatory organs of
<taxonomicName id="4C78749B952A6617FD985DF2FD4DE5BB" authorityName=": Logunov" authorityYear="2013" box="[549,742,1668,1689]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="597" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952A6617FD985DF2FD4DE5BB" box="[549,742,1668,1689]" italics="true" pageId="3" pageNumber="597">Alopecosa krynickii</emphasis>
</taxonomicName>
: 8 male palp, ventral; 9 bulbus, ventral-prolateral; 10 part of bulbus, retrolateral; 11–12 median apophysis, caudal and ventral; 13–14 embolic division, ventral and caudal-prolateral; 15 embolus, dorsal; 16–18 epigyne, ventral, dorsal, and dorsal-caudal. Abbreviations:
<emphasis id="B90CD30A952A6617FCA75DC9FC9CE5F6" box="[794,823,1727,1748]" italics="true" pageId="3" pageNumber="597">AP</emphasis>
anterior pocket of epigyne;
<emphasis id="B90CD30A952A6617FBF05DC9FBF5E5F6" box="[1101,1118,1727,1748]" italics="true" pageId="3" pageNumber="597">D</emphasis>
fertilization duct;
<emphasis id="B90CD30A952A6617FAA95DC9FA98E5F6" box="[1300,1331,1727,1748]" italics="true" pageId="3" pageNumber="597">FC</emphasis>
functional conductor;
<emphasis id="B90CD30A952A6617FEB75DAAFE86E5D3" box="[266,301,1756,1777]" italics="true" pageId="3" pageNumber="597">MA</emphasis>
median apophysis;
<emphasis id="B90CD30A952A6617FDBC5DAAFDBBE5D3" box="[513,528,1756,1777]" italics="true" pageId="3" pageNumber="597">P</emphasis>
palea;
<emphasis id="B90CD30A952A6617FDDE5DAAFDC4E5D3" box="[611,623,1756,1777]" italics="true" pageId="3" pageNumber="597">S</emphasis>
septum;
<emphasis id="B90CD30A952A6617FD695DAAFD46E5D3" box="[724,749,1756,1777]" italics="true" pageId="3" pageNumber="597">ST</emphasis>
spermatheca. Question mark indicates semi-transparent apophysis, explanation in the text. Scale bars 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC70F18952A6610FF7A5C43FF5AE2C5" blockId="3.[151,1436,1845,2006]" lastBlockId="4.[151,1436,152,521]" lastPageId="4" lastPageNumber="598" pageId="3" pageNumber="597">
MALE. Habitus is shown in
<figureCitation id="1343139D952A6617FE475C43FDF1E46E" box="[506,602,1845,1868]" captionStart="FIGURES1–7" captionStartId="2.[151,258,1941,1963]" captionTargetBox="[155,1431,201,1919]" captionTargetId="figure@2.[155,1432,191,1920]" captionTargetPageId="2" captionText="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="3" pageNumber="597">Figs 2, 7</figureCitation>
. The colour pattern of males and females is similar, but dorsal side in females is darker. Prosoma: Carapace with a posterior pitch, gradual descend of thoracic region is poorly marked, eye field not elevated (
<figureCitation id="1343139D952A6617FEBD5C0CFEE8E4B3" box="[256,323,1914,1937]" captionStart="FIGURES1–7" captionStartId="2.[151,258,1941,1963]" captionTargetBox="[155,1431,201,1919]" captionTargetId="figure@2.[155,1432,191,1920]" captionTargetPageId="2" captionText="FIGURES1–7. Habitus ofAlopecosa krynickii: 1–2dorsum of female and male; 3–4 carapaceprofile of female and male; 5–7 venter of juvenile, female,and male.Arrows indicate position of epigastric furrow." httpUri="https://zenodo.org/record/1199904/files/figure.png" pageId="3" pageNumber="597">Fig. 4</figureCitation>
). Males have well-visible four black spots behind eyes and, compared to the females, a better-marked median oval spot of white setae on carapace. Cymbium brown, with short black setae, other segments of palp are light brown, covered with white and black setae. Tarsi I–IV with scopula and spinules: tarsi I–II have well developed scopula and poorly developed spinules, tarsi III–IV—on the contrary. Metatarsi I–VI with scopula: metatarsi I–II with welldeveloped scopula and few spinules on distal part, metatarsi III–IV with poorly developed scopula, spinules are absent. Venter of tibia I with grey setae (similar to those of scopula). Opistosoma: Dorsum with a lanceolate cardiac mark of black and white setae and two longitudinal parallel lines of white and black spots. Copulatory organs as in
<figureCitation id="1343139D952D6610FA955A76FA3CE235" box="[1320,1431,256,279]" captionStart="FIGURES 8–18" captionStartId="3.[151,258,1668,1690]" captionTargetBox="[155,1432,198,1643]" captionTargetId="figure@3.[151,1436,191,1647]" captionTargetPageId="3" captionText="FIGURES 8–18. Copulatory organs of Alopecosa krynickii: 8 male palp, ventral; 9 bulbus, ventral-prolateral; 10 part of bulbus, retrolateral; 11–12 median apophysis, caudal and ventral; 13–14 embolic division, ventral and caudal-prolateral; 15 embolus, dorsal; 16–18 epigyne, ventral, dorsal, and dorsal-caudal. Abbreviations: AP anterior pocket of epigyne; D fertilization duct; FC functional conductor; MA median apophysis; P palea; S septum; ST spermatheca. Question mark indicates semi-transparent apophysis, explanationin the text.Scale bars 0.1 mm." httpUri="https://zenodo.org/record/1199908/files/figure.png" pageId="4" pageNumber="598">Figs 8–15</figureCitation>
: retrolateral side of median apophysis curved ventrally (=spoon-shaped lateral extension, after Logunov 2013a), tip of this curve with semi-transparent apophysis on retrolateral side (see
<figureCitation id="1343139D952D6610FCF75A33FC33E27E" box="[842,920,325,348]" captionStart="FIGURES 8–18" captionStartId="3.[151,258,1668,1690]" captionTargetBox="[155,1432,198,1643]" captionTargetId="figure@3.[151,1436,191,1647]" captionTargetPageId="3" captionText="FIGURES 8–18. Copulatory organs of Alopecosa krynickii: 8 male palp, ventral; 9 bulbus, ventral-prolateral; 10 part of bulbus, retrolateral; 11–12 median apophysis, caudal and ventral; 13–14 embolic division, ventral and caudal-prolateral; 15 embolus, dorsal; 16–18 epigyne, ventral, dorsal, and dorsal-caudal. Abbreviations: AP anterior pocket of epigyne; D fertilization duct; FC functional conductor; MA median apophysis; P palea; S septum; ST spermatheca. Question mark indicates semi-transparent apophysis, explanationin the text.Scale bars 0.1 mm." httpUri="https://zenodo.org/record/1199908/files/figure.png" pageId="4" pageNumber="598">Fig. 11</figureCitation>
, indicated by question mark), this apophysis is found only in one male, perhaps it can be easily broken or seems to be an aberration; retrolateral part of median apophysis is variable (
<figureCitation id="1343139D952D6610FE3B5AFCFE5FE283" box="[390,500,394,417]" captionStart="FIGURES 8–18" captionStartId="3.[151,258,1668,1690]" captionTargetBox="[155,1432,198,1643]" captionTargetId="figure@3.[151,1436,191,1647]" captionTargetPageId="3" captionText="FIGURES 8–18. Copulatory organs of Alopecosa krynickii: 8 male palp, ventral; 9 bulbus, ventral-prolateral; 10 part of bulbus, retrolateral; 11–12 median apophysis, caudal and ventral; 13–14 embolic division, ventral and caudal-prolateral; 15 embolus, dorsal; 16–18 epigyne, ventral, dorsal, and dorsal-caudal. Abbreviations: AP anterior pocket of epigyne; D fertilization duct; FC functional conductor; MA median apophysis; P palea; S septum; ST spermatheca. Question mark indicates semi-transparent apophysis, explanationin the text.Scale bars 0.1 mm." httpUri="https://zenodo.org/record/1199908/files/figure.png" pageId="4" pageNumber="598">Figs 8, 12</figureCitation>
); embolus flat and wide, embolic tip looks like a single-barbed harpoon; the palea with a poorly marked pointed prominence. Also, see the description of the
<typeStatus id="54C3B1BA952D6610FC2D5ADBFC46E2E6" box="[912,1005,429,452]" pageId="4" pageNumber="598" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C78749B952D6610FBED5ADBFB66E2E6" authorityName=": Logunov" authorityYear="2013" box="[1104,1229,429,452]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="598" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952D6610FBED5ADBFB66E2E6" box="[1104,1229,429,452]" italics="true" pageId="4" pageNumber="598">A. krynickii</emphasis>
</taxonomicName>
given by Logunov (2013a).
</paragraph>
</subSubSection>
<subSubSection id="C3625C93952D6610FF7A5A84FF42E450" pageId="4" pageNumber="598" type="distribution">
<paragraph id="8BC70F18952D6610FF7A5A84FC43E12B" blockId="4.[151,1436,152,521]" box="[199,1000,498,521]" pageId="4" pageNumber="598">
<emphasis id="B90CD30A952D6610FF7A5A84FEF9E12B" bold="true" box="[199,338,498,521]" pageId="4" pageNumber="598">Distribution</emphasis>
. The
<collectingRegion id="49BCC1FA952D6610FE325A84FE49E12B" box="[399,482,498,521]" country="Ukraine" name="Crimea" pageId="4" pageNumber="598">Crimea</collectingRegion>
: Tarkhankut peninsula and Simferopol (
<figureCitation id="1343139D952D6610FC315A84FC70E12B" box="[908,987,498,521]" captionStart="FIGURES 19–21" captionStartId="4.[151,258,1725,1747]" captionTargetBox="[158,1429,573,1705]" captionTargetId="figure@4.[158,1429,573,1705]" captionTargetPageId="4" captionText="FIGURES 19–21. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of Alopecosa krynickii. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of A.krynickii, where Villa Steven was situated.Map after Biodiversity Support Program (1999)." httpUri="https://zenodo.org/record/1199910/files/figure.png" pageId="4" pageNumber="598">Fig. 19</figureCitation>
).
</paragraph>
<caption id="DF075F90952D6610FF2A5DCBFB8BE42F" httpUri="https://zenodo.org/record/1199910/files/figure.png" pageId="4" pageNumber="598" startId="4.[151,258,1725,1747]" targetBox="[158,1429,573,1705]" targetPageId="4">
<paragraph id="8BC70F18952D6610FF2A5DCBFB8BE42F" blockId="4.[151,1436,1725,1805]" pageId="4" pageNumber="598">
<emphasis id="B90CD30A952D6610FF2A5DCBFEEDE5F1" bold="true" box="[151,326,1725,1747]" pageId="4" pageNumber="598">FIGURES 19–21</emphasis>
. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of
<taxonomicName id="4C78749B952D6610FF2A5DADFEFDE5D2" authorityName=": Logunov" authorityYear="2013" box="[151,342,1755,1776]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="598" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952D6610FF2A5DADFEFDE5D2" box="[151,342,1755,1776]" italics="true" pageId="4" pageNumber="598">Alopecosa krynickii</emphasis>
</taxonomicName>
. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of
<taxonomicName id="4C78749B952D6610FF2A5D8EFEAEE42F" authorityName=": Logunov" authorityYear="2013" box="[151,261,1784,1805]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="598" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952D6610FF2A5D8EFEAEE42F" box="[151,261,1784,1805]" italics="true" pageId="4" pageNumber="598">A. krynickii</emphasis>
</taxonomicName>
, where Villa Steven was situated. Map after Biodiversity Support Program (1999).
</paragraph>
</caption>
<paragraph id="8BC70F18952D6610FF7A5C4EFF42E450" blockId="4.[151,1436,1848,2010]" pageId="4" pageNumber="598">
<emphasis id="B90CD30A952D6610FF7A5C4EFE82E46D" bold="true" box="[199,297,1848,1871]" pageId="4" pageNumber="598">Habitats</emphasis>
. The stony steppe in two major habitat zones of the
<collectingRegion id="49BCC1FA952D6610FCEC5C4EFC0FE46D" box="[849,932,1848,1871]" country="Ukraine" name="Crimea" pageId="4" pageNumber="598">Crimea</collectingRegion>
: true steppe and premontane forest steppe (
<figureCitation id="1343139D952D6610FAD15C4EFF75E450" captionStart="FIGURES 19–21" captionStartId="4.[151,258,1725,1747]" captionTargetBox="[158,1429,573,1705]" captionTargetId="figure@4.[158,1429,573,1705]" captionTargetPageId="4" captionText="FIGURES 19–21. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of Alopecosa krynickii. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of A.krynickii, where Villa Steven was situated.Map after Biodiversity Support Program (1999)." httpUri="https://zenodo.org/record/1199910/files/figure.png" pageId="4" pageNumber="598">Figs 19–21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3625C93952D6610FF7A5C08FCA7E4F8" pageId="4" pageNumber="598" type="description">
<paragraph id="8BC70F18952D6610FF7A5C08FCA7E4F8" blockId="4.[151,1436,1848,2010]" pageId="4" pageNumber="598">
<emphasis id="B90CD30A952D6610FF7A5C08FE97E4B7" bold="true" box="[199,316,1918,1941]" pageId="4" pageNumber="598">Phenology</emphasis>
. In the
<collectingRegion id="49BCC1FA952D6610FE2C5C08FE49E4B7" box="[401,482,1918,1941]" country="Ukraine" name="Crimea" pageId="4" pageNumber="598">Crimea</collectingRegion>
, Tarkhankut peninsula: juveniles can be collected in July and September–October, adult females in May–June and September–October, adult males in September–October; females with egg sacs in June; the peak of adult activity was observed in September–October.
</paragraph>
</subSubSection>
<subSubSection id="C3625C93952C6611FF7A5BEEFEF5E063" pageId="5" pageNumber="599" type="discussion">
<paragraph id="8BC70F18952C6611FF7A5BEEFC52E153" blockId="5.[151,1436,152,833]" pageId="5" pageNumber="599">
<emphasis id="B90CD30A952C6611FF7A5BEEFE94E38D" bold="true" box="[199,319,152,175]" pageId="5" pageNumber="599">Discussion</emphasis>
.
<taxonomicName id="4C78749B952C6611FEEC5BEEFD83E38D" authorityName=": Logunov" authorityYear="2013" box="[337,552,152,175]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FEEC5BEEFD83E38D" box="[337,552,152,175]" italics="true" pageId="5" pageNumber="599">Alopecosa krynickii</emphasis>
</taxonomicName>
is still known as endemic of the
<collectingRegion id="49BCC1FA952C6611FC165BEEFC56E38D" box="[939,1021,152,175]" country="Ukraine" name="Crimea" pageId="5" pageNumber="599">Crimea</collectingRegion>
. It was assumed that endemic spider species of the
<collectingRegion id="49BCC1FA952C6611FE905BCCFED0E3F3" box="[301,379,186,209]" country="Ukraine" name="Crimea" pageId="5" pageNumber="599">Crimea</collectingRegion>
predominantly occur in mountains and/or the southern coast (
<bibRefCitation id="EFE972E9952C6611FBB05BCCFB56E3F3" box="[1037,1277,186,209]" pageId="5" pageNumber="599" refId="ref2554" refString="Kovblyuk, M. M. (2014) Spiders (Arachnida, Aranei) of the Crimea: faunogenesis and hypothesis of Pontida. Ukrainska Entomofaunistyka, 5 (2), 29 - 53. [in Russian with English summary]" type="journal article">Kovblyuk 2014: fig. 6</bibRefCitation>
), in the habitat zones 3–7 (
<figureCitation id="1343139D952C6611FEA55BABFEC0E3D6" box="[280,363,221,244]" captionStart="FIGURES 19–21" captionStartId="4.[151,258,1725,1747]" captionTargetBox="[158,1429,573,1705]" captionTargetId="figure@4.[158,1429,573,1705]" captionTargetPageId="4" captionText="FIGURES 19–21. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of Alopecosa krynickii. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of A.krynickii, where Villa Steven was situated.Map after Biodiversity Support Program (1999)." httpUri="https://zenodo.org/record/1199910/files/figure.png" pageId="5" pageNumber="599">Fig. 19</figureCitation>
). As this species has been found from outside the mountain part of the
<collectingRegion id="49BCC1FA952C6611FB2A5BABFB43E3D6" box="[1175,1256,221,244]" country="Ukraine" name="Crimea" pageId="5" pageNumber="599">Crimea</collectingRegion>
, its status of an endemic species is likely to be temporary.
<taxonomicName id="4C78749B952C6611FDD95A76FD4BE235" authorityName=": Logunov" authorityYear="2013" box="[612,736,256,279]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FDD95A76FD4BE235" box="[612,736,256,279]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
has been found in Tarkhankut peninsula, in stony steppes which has never been ploughed (
<figureCitation id="1343139D952C6611FE115A54FE50E21B" box="[428,507,290,313]" captionStart="FIGURES 19–21" captionStartId="4.[151,258,1725,1747]" captionTargetBox="[158,1429,573,1705]" captionTargetId="figure@4.[158,1429,573,1705]" captionTargetPageId="4" captionText="FIGURES 19–21. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of Alopecosa krynickii. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of A.krynickii, where Villa Steven was situated.Map after Biodiversity Support Program (1999)." httpUri="https://zenodo.org/record/1199910/files/figure.png" pageId="5" pageNumber="599">Fig. 20</figureCitation>
). Until the 20th century, virgin steppes had been widespread in the plains of the
<collectingRegion id="49BCC1FA952C6611FAF05A54FA30E21B" box="[1357,1435,290,313]" country="Ukraine" name="Crimea" pageId="5" pageNumber="599">Crimea</collectingRegion>
(the habitat zone 2 and partly the zones 1 &amp; 3). For example, until 1950s, a stony steppe had stretched in the vicinity of Simferopol, in the
<typeStatus id="54C3B1BA952C6611FEE05A1EFE20E25D" box="[349,395,360,383]" pageId="5" pageNumber="599">type</typeStatus>
locality of
<taxonomicName id="4C78749B952C6611FDB95A1EFDD5E25D" authorityName=": Logunov" authorityYear="2013" box="[516,638,360,383]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FDB95A1EFDD5E25D" box="[516,638,360,383]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
. At present this area is house-built and forested (
<figureCitation id="1343139D952C6611FB395A1EFB78E25D" box="[1156,1235,360,383]" captionStart="FIGURES 19–21" captionStartId="4.[151,258,1725,1747]" captionTargetBox="[158,1429,573,1705]" captionTargetId="figure@4.[158,1429,573,1705]" captionTargetPageId="4" captionText="FIGURES 19–21. Distribution records (19) and habitats (20, Bolshoi Kastel Valley in October; 21, Simferopol in September) of Alopecosa krynickii. Black circles indicate collecting localities of specimens examined in this study. Arrows indicate type locality of A.krynickii, where Villa Steven was situated.Map after Biodiversity Support Program (1999)." httpUri="https://zenodo.org/record/1199910/files/figure.png" pageId="5" pageNumber="599">Fig. 21</figureCitation>
). It is likely that
<taxonomicName id="4C78749B952C6611FA385A1EFF5FE283" authorityName=": Logunov" authorityYear="2013" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FA385A1EFF5FE283" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
can be found in other sites of virgin steppes of the Palaearctic, especially in
<collectingCountry id="F36F4F88952C6611FB8E5AFCFB22E283" box="[1075,1161,394,417]" name="Ukraine" pageId="5" pageNumber="599">Ukraine</collectingCountry>
and the European part of
<collectingCountry id="F36F4F88952C6611FF2A5ADBFF49E2E6" box="[151,226,429,452]" name="Russia" pageId="5" pageNumber="599">Russia</collectingCountry>
. It is difficult to collect
<taxonomicName id="4C78749B952C6611FE445ADBFDD3E2E6" authorityName=": Logunov" authorityYear="2013" box="[505,632,429,452]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FE445ADBFDD3E2E6" box="[505,632,429,452]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
by pitfalls. Among the 26 collected specimens of
<taxonomicName id="4C78749B952C6611FB085ADBFA9FE2E6" authorityName=": Logunov" authorityYear="2013" box="[1205,1332,429,452]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FB085ADBFA9FE2E6" box="[1205,1332,429,452]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
, a single specimen was caught by pitfall traps; in order to collect spiders, a total of 60 pitfalls were set in Tarkhankut and exposed for seven months, April–October (the catch was taken twice a month). Thus, hand picking in autumn at night, with the aid of head-torch, seems to be the best method of collecting
<taxonomicName id="4C78749B952C6611FC975963FC0DE10E" authorityName=": Logunov" authorityYear="2013" box="[810,934,533,556]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FC975963FC0DE10E" box="[810,934,533,556]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
, as well as other burrowing wolf spiders. The somatic characters, such as carapace profile, scopula and spinules on legs of
<taxonomicName id="4C78749B952C6611FC79594EFB95E16D" authorityName=": Logunov" authorityYear="2013" box="[964,1086,568,591]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FC79594EFB95E16D" box="[964,1086,568,591]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
well correspond to the structural and functional features of burrowing wolf spiders as described by
<bibRefCitation id="EFE972E9952C6611FCE4592CFC5EE153" box="[857,1013,602,625]" pageId="5" pageNumber="599" refId="ref2798" refString="Zyuzin, A. A. (1990) Studies on burrowing spiders of the family Lycosidae (Araneae). I. Preliminary data on structural and functional features. Acta Zoologica Fennica, 190, 419 - 422." type="journal article">Zyuzin (1990)</bibRefCitation>
.
</paragraph>
<paragraph id="8BC70F18952C6611FF7A590BFEF5E063" blockId="5.[151,1436,152,833]" pageId="5" pageNumber="599">
I am grateful to Mykola Kovblyuk and Oleg Kukushkin who collected some specimens of
<taxonomicName id="4C78749B952C6611FB34590BFAA8E1B6" authorityName=": Logunov" authorityYear="2013" box="[1161,1283,637,660]" class="Arachnida" family="Lycosidae" genus="Alopecosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="599" phylum="Arthropoda" rank="species" species="krynickii">
<emphasis id="B90CD30A952C6611FB34590BFAA8E1B6" box="[1161,1283,637,660]" italics="true" pageId="5" pageNumber="599">A. krynickii</emphasis>
</taxonomicName>
. Logistic help with the 2015 expeditions was provided by the Natural Park &quot;Tarkhankutskii&quot;, Chernomorskoe, the
<collectingRegion id="49BCC1FA952C6611FB7F59D6FABFE195" box="[1218,1300,672,695]" country="Ukraine" name="Crimea" pageId="5" pageNumber="599">Crimea</collectingRegion>
. I also thank reviewers (Dmitri Logunov and Galina Azarkina) and editor (Adalberto Santos) for their valuable comments and remarks to the manuscript. English of the manuscript was kindly checked and edited by Dmitri Logunov and Aatemmaa Sennedokkao. This study was supported by Ministry of Education and Science of the
<collectingCountry id="F36F4F88952C6611FB3E587EFAFCE03D" box="[1155,1367,776,799]" name="Russia" pageId="5" pageNumber="599">Russian Federation</collectingCountry>
grant #14.W03.31.0015.
</paragraph>
</subSubSection>
</treatment>
</document>
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