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<document id="6D460EEC6A84983DC583FC3493107912" ID-CLB-Dataset="291999" ID-DOI="10.25674/so95iss1id311" ID-GBIF-Dataset="06b03cc4-e211-4f65-be07-742740b0053c" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFE4FFD1BF70FF080C0903BA" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus sulawesianus Frisch 2022, spec. nov." docType="treatment" docVersion="4" lastPageNumber="60" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="59" updateTime="1711467745701" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BDC0DE2FFE1110302FD4EB96B1DDBF9A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3FC3E69E14494AD3124D632545DDEC71">
<mods:title id="07E358E3A2BA8C2782EB48E3519389EA">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
</mods:titleInfo>
<mods:name id="03607FB737503D6BB4AFF333195B7BFC" type="personal">
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<mods:roleTerm id="A4FC643015C18B084296D136BB928218">Author</mods:roleTerm>
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<mods:namePart id="E2A6F0BCD6BAFB7C5ED4F23EE0A90657">Frisch, Johannes</mods:namePart>
</mods:name>
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<mods:role id="8C5D384DFA453768B008D9976C6DB36E">
<mods:roleTerm id="97AFFCC11731BF32FA810AF88C141CD7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="58EC5418749AA7E0B023E28EB59D60D4">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="BE52C66805A75EE6BA670B986D03A12C">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F75538A8D53AA8C99F2D15BFC66367F7">text</mods:typeOfResource>
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<mods:title id="240AC98614C91A2FE96D85430E22F1A3">Soil Organisms</mods:title>
</mods:titleInfo>
<mods:part id="5840F68907FE3AACD0FB4758A13E4ADF">
<mods:date id="6E96EA24D630256D2226E8A972494C5B">2022</mods:date>
<mods:detail id="0EA3C8F304D3C071DAFAA1ACF591B27F" type="pubDate">
<mods:number id="204DAB2D84E609796181AACB3F12F296">2022-04-01</mods:number>
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<mods:number id="C38D163EA688705E41A4B302E265B944">95</mods:number>
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<mods:number id="3FEA8497B7D7C4A25B2D1424E4B46321">1</mods:number>
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<mods:start id="8268B6FB6BEED4388D725E3A25E84E04">23</mods:start>
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<mods:url id="DFA5CF5C506A1A71B775A207A10A6714">http://dx.doi.org/10.25674/so95iss1id311</mods:url>
</mods:location>
<mods:classification id="CE00A152D71B8D419E1315E67E46EBBA">journal article</mods:classification>
<mods:identifier id="BF8E3583BC1276A77E667D4145581874" type="CLB-Dataset">291999</mods:identifier>
<mods:identifier id="01930E3547508CD7EDCAE15CE067D9B2" type="DOI">10.25674/so95iss1id311</mods:identifier>
<mods:identifier id="782E4BFE2097B9679FACBD73584C0F02" type="GBIF-Dataset">06b03cc4-e211-4f65-be07-742740b0053c</mods:identifier>
<mods:identifier id="DC486239D5FECF3CA2D329254FF1F7A5" type="ISSN">2509-9523</mods:identifier>
<mods:identifier id="0593B62BB74A17D1E737FD4155E04CFE" type="Zenodo-Dep">10724236</mods:identifier>
</mods:mods>
<treatment id="03B7C77AFFE4FFD1BF70FF080C0903BA" ID-DOI="http://doi.org/10.5281/zenodo.10880272" ID-Zenodo-Dep="10880272" LSID="urn:lsid:plazi:treatment:03B7C77AFFE4FFD1BF70FF080C0903BA" httpUri="http://treatment.plazi.org/id/03B7C77AFFE4FFD1BF70FF080C0903BA" lastPageId="37" lastPageNumber="60" pageId="36" pageNumber="59">
<subSubSection id="C30425E7FFE4FFD0BF70FF08083E0507" box="[811,1315,239,266]" pageId="36" pageNumber="59" type="nomenclature">
<paragraph id="8BA1766CFFE4FFD0BF70FF08083E0507" blockId="36.[811,1315,239,299]" box="[811,1315,239,266]" pageId="36" pageNumber="59">
<heading id="D0E9C100FFE4FFD0BF70FF08083E0507" box="[811,1315,239,266]" fontSize="11" level="2" pageId="36" pageNumber="59" reason="5">
<taxonomicName id="4C1E0DEFFFE4FFD0BF70FF0809BC0504" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[811,1185,239,266]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="species" species="sulawesianus" status="sp. nov.">
<emphasis id="B96AAA7EFFE4FFD0BF70FF0809BC0504" bold="true" box="[811,1185,239,266]" pageId="36" pageNumber="59">
<emphasis id="B96AAA7EFFE4FFD0BF70FF0809550507" bold="true" box="[811,1096,239,266]" italics="true" pageId="36" pageNumber="59">Scopaeus sulawesianus</emphasis>
Frisch
</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2591705FFE4FFD0B8F4FF08083E0507" box="[1199,1315,240,266]" pageId="36" pageNumber="59" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE4FFD0BF70FEE809820526" box="[811,1183,272,299]" pageId="36" pageNumber="59" type="description">
<paragraph id="8BA1766CFFE4FFD0BF70FEE809820526" blockId="36.[811,1315,239,299]" box="[811,1183,272,299]" pageId="36" pageNumber="59">
(
<figureCitation id="13256AE9FFE4FFD0BF68FEE80E900526" box="[819,909,272,299]" captionStart="Figures 23–26" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 23–26. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="36" pageNumber="59">Figs 24</figureCitation>
,
<figureCitation id="13256AE9FFE4FFD0BFC2FEE90EAA0526" box="[921,951,273,299]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="36" pageNumber="59">49</figureCitation>
,
<figureCitation id="13256AE9FFE4FFD0BF98FEE809370526" box="[963,1066,272,299]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="36" pageNumber="59">116–118</figureCitation>
, 145, 178)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE4FFD0BF70FEAC090E063F" pageId="36" pageNumber="59" type="materials_examined">
<paragraph id="8BA1766CFFE4FFD0BF70FEAC090E063F" blockId="36.[788,1399,340,923]" pageId="36" pageNumber="59">
<emphasis id="B96AAA7EFFE4FFD0BF70FEAC0EF20561" bold="true" box="[811,1007,340,364]" pageId="36" pageNumber="59">Type specimens:</emphasis>
<materialsCitation id="3B767C31FFE4FFD0BFA5FEAC09CE05A3" collectingDate="2015-07-27" collectionCode="MZB" collectorName="Puchner" country="Indonesia" elevation="46" latitude="-3.5666666" location="Sadang River" longLatPrecision="1307" longitude="119.76667" municipality="Engrekang" pageId="36" pageNumber="59" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Selatan" typeStatus="holotype">
<collectingCountry id="F30936FCFFE4FFD0BFA5FEAC09760561" box="[1022,1131,340,364]" name="Indonesia" pageId="36" pageNumber="59">Indonesia</collectingCountry>
:
<typeStatus id="54A5C8CEFFE4FFD0B827FEAC09FD0561" box="[1148,1248,340,364]" pageId="36" pageNumber="59" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFE4FFD0B8B4FEAD081E0561" box="[1263,1283,341,364]" count="1" pageId="36" pageNumber="59" type="male">♂</specimenCount>
,
<collectingRegion id="49DAB88EFFE4FFD0B94DFEAC0E7B0580" country="Indonesia" name="Sulawesi Selatan" pageId="36" pageNumber="59">Sulawesi Selatan</collectingRegion>
,
<collectingMunicipality id="6BC5EC16FFE4FFD0BF2AFE8D0EFB0580" box="[881,998,373,397]" pageId="36" pageNumber="59">Engrekang</collectingMunicipality>
(
<location id="8EC120B7FFE4FFD0BFAFFE8D09970580" LSID="urn:lsid:plazi:treatment:03B7C77AFFE4FFD1BF70FF080C0903BA:8EC120B7FFE4FFD0BFAFFE8D09970580" box="[1012,1162,373,397]" country="Indonesia" latitude="-3.5666666" longLatPrecision="1307" longitude="119.76667" municipality="Engrekang" name="Sadang River" pageId="36" pageNumber="59" stateProvince="Sulawesi Selatan">Sadang River</location>
), (
<geoCoordinate id="EE2A10ABFFE4FFD0B8FFFE8D09E60580" box="[1188,1275,373,397]" degrees="03" direction="south" minutes="34" orientation="latitude" pageId="36" pageNumber="59" precision="925" value="-3.5666666">03°34’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFE4FFD0B95DFE8D08760580" box="[1286,1387,373,397]" degrees="119" direction="east" minutes="46" orientation="longitude" pageId="36" pageNumber="59" precision="925" value="119.76667">119°46’E</geoCoordinate>
),
<quantity id="4CE6DB89FFE4FFD0BF4FFE6E0E4B05A0" box="[788,854,406,430]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="36" pageNumber="59" unit="m" value="46.0">
<elevation id="0033915FFFE4FFD0BF4FFE6E0E4B05A0" box="[788,854,406,430]" metricMagnitude="1" metricUnit="m" metricValue="4.6" pageId="36" pageNumber="59" unit="m" value="46.0">46 m</elevation>
</quantity>
,
<date id="FFA050ACFFE4FFD0BF30FE6E0ECD05A3" box="[875,976,406,430]" pageId="36" pageNumber="59" value="2015-07-27">
<collectingDate id="EFE4A944FFE4FFD0BF30FE6E0ECD05A3" box="[875,976,406,430]" pageId="36" pageNumber="59" value="2015-07-27">27.7.2015</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFE4FFD0B840FE6E096905A3" box="[1051,1140,406,430]" pageId="36" pageNumber="59">Puchner</collectorName>
(
<collectionCode id="ED0FEEA9FFE4FFD0B8D7FE6E09D605A3" box="[1164,1227,406,430]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="36" pageNumber="59" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54A5C8CEFFE4FFD0B8BDFE6E084C05A3" box="[1254,1361,406,430]" pageId="36" pageNumber="59" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D18BDE5FFE4FFD0B932FE6E0E9405C2" count="9" pageId="36" pageNumber="59" type="generic">9 specimens</specimenCount>
):
<materialsCitation id="3B767C31FFE4FFD0BFC5FE4F0E6F05FD" collectingDate="2015-07-27" collectionCode="MFNB, MZMB" collectorName="Puchner" country="Indonesia" elevation="46" latitude="-3.5666666" location="Sadang River" longLatPrecision="1307" longitude="119.76667" municipality="Engrekang" pageId="36" pageNumber="59" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sulawesi Selatan" typeStatus="paratype">
<specimenCount id="9D18BDE5FFE4FFD0BFC5FE4F0ED705C3" box="[926,970,439,463]" count="2" pageId="36" pageNumber="59" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFE4FFD0BF82FE4F091905C3" box="[985,1028,439,463]" count="1" pageId="36" pageNumber="59" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED0FEEA9FFE4FFD0B97BFE4F086F05C2" box="[1312,1394,439,463]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="36" pageNumber="59">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFE4FFD0BF4FFE200E7705FD" box="[788,874,472,496]" pageId="36" pageNumber="59">MZMB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFE4FFD0BF27FE20090D063F" collectingDate="1999-11-07" collectionCode="MFNB, SMNS" collectorName="Bolm" country="Indonesia" elevation="50" latitude="-1.1875001" location="6 km E Tambarana" longLatPrecision="21" longitude="120.46833" pageId="36" pageNumber="59" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<specimenCount id="9D18BDE5FFE4FFD0BF27FE200EB805E2" box="[892,933,472,496]" count="3" pageId="36" pageNumber="59" type="male">3 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFE4FFD0BFEBFE200EC505E2" box="[944,984,472,496]" count="3" pageId="36" pageNumber="59" type="female">3 ♀</specimenCount>
,
<collectingRegion id="49DAB88EFFE4FFD0BFBEFE20098305FD" box="[997,1182,472,496]" country="Indonesia" name="Sulawesi Tengah" pageId="36" pageNumber="59">Sulawesi Tengah</collectingRegion>
,
<location id="8EC120B7FFE4FFD0B8F3FE20086B05FD" LSID="urn:lsid:plazi:treatment:03B7C77AFFE4FFD1BF70FF080C0903BA:8EC120B7FFE4FFD0B8F3FE20086B05FD" box="[1192,1398,472,496]" country="Indonesia" latitude="-1.1875001" longLatPrecision="21" longitude="120.46833" name="6 km E Tambarana" pageId="36" pageNumber="59" stateProvince="Sulawesi Tengah">
<quantity id="4CE6DB89FFE4FFD0B8F3FE2009FD05FD" box="[1192,1248,472,496]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="59" unit="km" value="6.0">6 km</quantity>
E Tambarana
</location>
(
<geoCoordinate id="EE2A10ABFFE4FFD0BF47FE000E88061D" box="[796,917,504,528]" degrees="01" direction="south" minutes="11" orientation="latitude" pageId="36" pageNumber="59" precision="15" seconds="15" value="-1.1875001">01°11’15’’S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFE4FFD0BFF1FE000921061D" box="[938,1084,504,528]" degrees="120" direction="east" minutes="28" orientation="longitude" pageId="36" pageNumber="59" precision="15" seconds="06" value="120.46833">120°28’06’’E</geoCoordinate>
),
<quantity id="4CE6DB89FFE4FFD0B802FE000981061D" box="[1113,1180,504,528]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="59" unit="m" value="50.0">
<elevation id="0033915FFFE4FFD0B802FE000981061D" box="[1113,1180,504,528]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="59" unit="m" value="50.0">50 m</elevation>
</quantity>
, 9. –
<date id="FFA050ACFFE4FFD0B883FE010826061C" box="[1240,1339,505,529]" pageId="36" pageNumber="59" value="1999-11-07">
<collectingDate id="EFE4A944FFE4FFD0B883FE010826061C" box="[1240,1339,505,529]" pageId="36" pageNumber="59" value="1999-11-07">11.7.1999</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFE4FFD0BF4FFDE10E52063C" box="[788,847,537,561]" pageId="36" pageNumber="59">Bolm</collectorName>
(
<collectionCode id="ED0FEEA9FFE4FFD0BF05FDE20EAD063F" box="[862,944,538,562]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="36" pageNumber="59">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFE4FFD0BFE0FDE20915063F" box="[955,1032,538,562]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="36" pageNumber="59">SMNS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE4FFD1BF70FDC20FA3016A" lastPageId="37" lastPageNumber="60" pageId="36" pageNumber="59" type="description">
<paragraph id="8BA1766CFFE4FFD0BF70FDC2094F0754" blockId="36.[788,1399,340,923]" pageId="36" pageNumber="59">
<emphasis id="B96AAA7EFFE4FFD0BF70FDC20EA4065F" bold="true" box="[811,953,570,594]" pageId="36" pageNumber="59">Description:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFE4FFD0B947FDC2086F065F" box="[1308,1394,570,594]" captionStart="Figures 23–26" captionStartId="14.[151,224,1930,1952]" captionTargetBox="[263,1352,435,1899]" captionTargetId="figure-131@14.[151,1398,422,1911]" captionTargetPageId="14" captionText="Figures 23–26. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26)." figureDoi="http://doi.org/10.5281/zenodo.10724252" httpUri="https://zenodo.org/record/10724252/files/figure.png" pageId="36" pageNumber="59">Fig. 24</figureCitation>
. Head subquadrate. Penultimate antennal segment elongate. Mesotibia moderately thickened. Body surface dull with extremely fine, dense, setose punctation; microreticulation absent. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Body including appendages light brown; disc of abdominal segments darker brown. Total body length
<quantity id="4CE6DB89FFE4FFD0B8B1FCD9086E0735" box="[1258,1395,801,825]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="36" pageNumber="59" unit="mm" value="3.2" valueMax="3.3" valueMin="3.1">3.1 – 3.3 mm</quantity>
; forebody length
<quantity id="4CE6DB89FFE4FFD0BF9DFCB909530754" box="[966,1102,833,857]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="1.9" metricValueMin="1.8" pageId="36" pageNumber="59" unit="mm" value="1.85" valueMax="1.9" valueMin="1.8">1.8 – 1.9 mm</quantity>
.
</paragraph>
<paragraph id="8BA1766CFFE4FFD1BF70FC9A0FCD0108" blockId="36.[788,1399,340,923]" lastBlockId="37.[189,802,241,1975]" lastPageId="37" lastPageNumber="60" pageId="36" pageNumber="59">
Male: Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII in almost posterior half with deep, triangular emargination with somewhat concave margins and acute anterior end; disc of sternite VIII with large depression wider than half sternite width stretching from emarginate posterior margin close to subbasal ridge; depression of sternite VIII surrounded laterally by long, medioposteriorly pointing macrosetae neighboured by sublateral, lateroposteriorly pointing macrosetae situated within depression (Fig. 145). Aedeagus (
<figureCitation id="13256AE9FFE5FFD1BD23FE000CD1061C" box="[376,460,504,529]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="37" pageNumber="60">Figs 49</figureCitation>
,
<figureCitation id="13256AE9FFE5FFD1BD80FE000F25061C" box="[475,568,504,529]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">116 – 118</figureCitation>
) about
<quantity id="4CE6DB89FFE5FFD1BED4FE000FFD061D" box="[655,736,504,528]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="60" unit="mm" value="0.5">0.5 mm</quantity>
long; phallobase stout, strongly sclerotized, thus dark reddish brown, shorter than distal lobes; distal portion of aedeagus seemingly attached to phallobase ventromedially owing to remarkable, dorsodistal extension of phallobase evenly narrowed towards wide, membranous, dorsodistal window (
<figureCitation id="13256AE9FFE5FFD1BD70FD460C9806DB" box="[299,389,702,726]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="37" pageNumber="60">Figs 49</figureCitation>
,
<figureCitation id="13256AE9FFE5FFD1BDC2FD460CDC06DB" box="[409,449,702,726]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">116</figureCitation>
); apical lobes in lateral view very narrow basally but convexly widened distad (
<figureCitation id="13256AE9FFE5FFD1BEA9FD270DC50715" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="37" pageNumber="60">Figs 49</figureCitation>
,
<figureCitation id="13256AE9FFE5FFD1BCB2FD070C0C071A" box="[233,273,767,791]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">116</figureCitation>
), in ventral and dorsal view with pronounced proximal waist, widened distolaterally and extended in remarkable, wing-like lobes with triangularly enlarged, mediodistad curved ends (
<figureCitation id="13256AE9FFE5FFD1BDB0FC9A0F990777" box="[491,644,866,890]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">Figs 117, 118</figureCitation>
); apical lobes ventrally separated in lateral halves with straight, parallel inner margins; ends of apical lobes membranous, in lateral view narrow and curved dorsad (
<figureCitation id="13256AE9FFE5FFD1BE79FC3D0F6E07D0" box="[546,627,965,989]" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="37" pageNumber="60">Figs 49</figureCitation>
,
<figureCitation id="13256AE9FFE5FFD1BE24FC3D0FBA07D0" box="[639,679,965,989]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">116</figureCitation>
), in ventral and dorsal view widened laterally, subflabellate apically (
<figureCitation id="13256AE9FFE5FFD1BC9FFBFF0C410012" box="[196,348,1031,1055]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">Figs 117, 118</figureCitation>
); dorsal lobe curved apicoventrad (
<figureCitation id="13256AE9FFE5FFD1BEA9FBFF0DC5004D" captionStart="Figures 42–49" captionStartId="17.[190,263,1886,1908]" captionTargetBox="[263,1341,889,1832]" captionTargetId="figure-379@17.[189,1436,851,1867]" captionTargetPageId="17" captionText="Figures 42–49. Aedeagus in lateral view of Scopaeus velifer, holotype, Banten, Lebak: Citorek Kidul (42); S. elegantulus, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (43); S. uncinatus, holotype,Banten, Lebak:Citorek Kidul (44); S. bipenicillatus, holotype,Sulawesi Tengah, Sigi: Pagana (45); S. bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso (46); S. cuspidatus, holotype, Bali, Jembrana: Cekik (47); S. crassipunctatus, holotype, Jawa Barat, Sukabumi, Sirnarasa:Cisarua (48); S. sulawesianus, holotype, Sulawesi Selatan: Engrekang (49)." figureDoi="http://doi.org/10.5281/zenodo.10724262" httpUri="https://zenodo.org/record/10724262/files/figure.png" pageId="37" pageNumber="60">Figs 49</figureCitation>
,
<figureCitation id="13256AE9FFE5FFD1BCBEFBDF0C100032" box="[229,269,1063,1087]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">116</figureCitation>
), with membranous apex triangularly projecting between membranous ends of apical lobes (
<figureCitation id="13256AE9FFE5FFD1BED0FBB00E08006D" box="[651,789,1096,1120]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">Figs 117, 118</figureCitation>
); flagellum inconspicuous (
<figureCitation id="13256AE9FFE5FFD1BD80FB910F25008C" box="[475,568,1129,1153]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">Fig. 117</figureCitation>
); small convex lobe, presumably ventral lobe, projecting from distal portion of aedeagus ventroproximally; median foramen oblong, with ends of lateral ridges curved proximolaterad; transverse ridge slender, curved distad (
<figureCitation id="13256AE9FFE5FFD1BE30FB150FD90108" box="[619,708,1261,1285]" captionStart="Figures 116–118" captionStartId="36.[154,227,1907,1929]" captionTargetBox="[224,1268,1020,1861]" captionTargetId="figure-354@36.[152,1399,977,1888]" captionTargetPageId="36" captionText="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">Fig. 117</figureCitation>
).
</paragraph>
<caption id="DF6126E4FFE4FFD0BCC1F88B0C9103BB" ID-DOI="http://doi.org/10.5281/zenodo.10724302" ID-Zenodo-Dep="10724302" httpUri="https://zenodo.org/record/10724302/files/figure.png" pageId="36" pageNumber="59" startId="36.[154,227,1907,1929]" targetBox="[224,1268,1020,1861]" targetPageId="36" targetType="figure">
<paragraph id="8BA1766CFFE4FFD0BCC1F88B0C9103BB" blockId="36.[154,1402,1907,1974]" pageId="36" pageNumber="59">
<emphasis id="B96AAA7EFFE4FFD0BCC1F88B0C260385" bold="true" box="[154,315,1907,1929]" pageId="36" pageNumber="59">Figures 116–118.</emphasis>
Aedeagus of
<taxonomicName id="4C1E0DEFFFE4FFD0BDE3F88B0F910385" authorityName="Frisch" authorityYear="2022" box="[440,652,1907,1928]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="59" phylum="Arthropoda" rank="species" species="sulawesianus">
<emphasis id="B96AAA7EFFE4FFD0BDE3F88B0F910385" box="[440,652,1907,1928]" italics="true" pageId="36" pageNumber="59">Scopaeus sulawesianus</emphasis>
</taxonomicName>
, holotype, Sulawesi Selatan: Engrekang in lateral (
<emphasis id="B96AAA7EFFE4FFD0B806F88B09630385" bold="true" box="[1117,1150,1907,1928]" pageId="36" pageNumber="59">116</emphasis>
), ventral (
<emphasis id="B96AAA7EFFE4FFD0B886F88B09E30385" bold="true" box="[1245,1278,1907,1928]" pageId="36" pageNumber="59">117</emphasis>
), dorsal view (
<emphasis id="B96AAA7EFFE4FFD0BCF9F8710DDE0393" bold="true" box="[162,195,1929,1950]" pageId="36" pageNumber="59">118</emphasis>
).
<emphasis id="B96AAA7EFFE4FFD0BC8DF8710C420392" bold="true" box="[214,351,1929,1951]" pageId="36" pageNumber="59">Abbreviations</emphasis>
:
<emphasis id="B96AAA7EFFE4FFD0BD37F8720C630392" bold="true" box="[364,382,1930,1951]" pageId="36" pageNumber="59">al</emphasis>
– apical lobes,
<emphasis id="B96AAA7EFFE4FFD0BE4BF8720F300392" bold="true" box="[528,557,1930,1951]" pageId="36" pageNumber="59">dw</emphasis>
– dorsodistal window of phallobase,
<emphasis id="B96AAA7EFFE4FFD0BFDDF8710EBD0392" bold="true" box="[902,928,1929,1951]" pageId="36" pageNumber="59">mf</emphasis>
– median foramen,
<emphasis id="B96AAA7EFFE4FFD0B800F87209700392" bold="true" box="[1115,1133,1930,1951]" pageId="36" pageNumber="59">tr</emphasis>
– transverse ridge of median foramen,
<emphasis id="B96AAA7EFFE4FFD0BCAAF8580C1E03B8" bold="true" box="[241,259,1952,1973]" pageId="36" pageNumber="59">vl</emphasis>
– ventral lobe.
</paragraph>
</caption>
<paragraph id="8BA1766CFFE5FFD1BC8FFAF60FA3016A" blockId="37.[189,802,241,1975]" pageId="37" pageNumber="60">
Female: Sperm pump with long, slender process segment; chamber without apophysis; bursal duct long, spirally tortuous (
<figureCitation id="13256AE9FFE5FFD1BD26FAB70CCA0165" box="[381,471,1359,1384]" captionStart="Figures 173–178" captionStartId="43.[189,262,1885,1907]" captionTargetId="figure-111@43.[189,1436,294,1840]" captionTargetPageId="43" captionText="Figures 173–178. Sperm pump (173–178) and bursa (173) of ♀ of Scopaeus bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (173, with lateral gonocoxal plates IX; 174); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (175); S. cuspidatus, paratype, Jawa Barat: Mt Salak (176); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (177); S. sulawesianus, paratype, Central Sulawesi, Tambarana (178, with bursal duct). Long scale bar: 174–178; short scale bar: 173." figureDoi="http://doi.org/10.5281/zenodo.10880230" httpUri="https://zenodo.org/record/10880230/files/figure.png" pageId="37" pageNumber="60">Fig. 178</figureCitation>
); bursa membranous.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BC8FFA880C4401A4" pageId="37" pageNumber="60" type="distribution">
<paragraph id="8BA1766CFFE5FFD1BC8FFA880C4401A4" blockId="37.[189,802,241,1975]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BC8FFA880C770185" bold="true" box="[212,362,1392,1416]" pageId="37" pageNumber="60">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFE5FFD1BD28FA890F680185" authorityName="Frisch" authorityYear="2022" box="[371,629,1392,1416]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="sulawesianus">
<emphasis id="B96AAA7EFFE5FFD1BD28FA890F680185" box="[371,629,1392,1416]" italics="true" pageId="37" pageNumber="60">Scopaeus sulawesianus</emphasis>
</taxonomicName>
is known from
<collectingRegion id="49DAB88EFFE5FFD1BCE6FA690C0001A4" box="[189,285,1425,1449]" country="Indonesia" name="Sulawesi Utara" pageId="37" pageNumber="60">Sulawesi</collectingRegion>
only.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BC8FFA4A0F0D0243" pageId="37" pageNumber="60" type="etymology">
<paragraph id="8BA1766CFFE5FFD1BC8FFA4A0F0D0243" blockId="37.[189,802,241,1975]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BC8FFA4A0C4701C7" bold="true" box="[212,346,1458,1482]" pageId="37" pageNumber="60">Etymology:</emphasis>
With the epithet
<taxonomicName id="4C1E0DEFFFE5FFD1BE41FA4A0FB601C7" authorityName="Frisch" authorityYear="2022" box="[538,683,1458,1482]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="sulawesianus">
<emphasis id="B96AAA7EFFE5FFD1BE41FA4A0FB601C7" box="[538,683,1458,1482]" italics="true" pageId="37" pageNumber="60">sulawesianus</emphasis>
</taxonomicName>
(adjective, Latin, composed of the geographic name
<collectingRegion id="49DAB88EFFE5FFD1BE9AFA2B0E3C01E6" box="[705,801,1491,1515]" country="Indonesia" name="Sulawesi Utara" pageId="37" pageNumber="60">Sulawesi</collectingRegion>
and the suffix –
<emphasis id="B96AAA7EFFE5FFD1BD2BFA0D0CBE0201" box="[368,419,1525,1548]" italics="true" pageId="37" pageNumber="60">anus</emphasis>
, which indicates the geographical affiliation), reference is made to the distribution of this presumed Sulawesian endemic.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFE5FFD1BC8FF9AE0C0903BA" pageId="37" pageNumber="60" type="discussion">
<paragraph id="8BA1766CFFE5FFD1BC8FF9AE0C0903BA" blockId="37.[189,802,241,1975]" pageId="37" pageNumber="60">
<emphasis id="B96AAA7EFFE5FFD1BC8FF9AE0C520263" bold="true" box="[212,335,1622,1646]" pageId="37" pageNumber="60">Comment:</emphasis>
<taxonomicName id="4C1E0DEFFFE5FFD1BD03F9AF0F4B0263" authorityName="Frisch" authorityYear="2022" box="[344,598,1622,1646]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="bipectenatus">
<emphasis id="B96AAA7EFFE5FFD1BD03F9AF0F4B0263" box="[344,598,1622,1646]" italics="true" pageId="37" pageNumber="60">Scopaeus bipectenatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E0DEFFFE5FFD1BE3DF9AF0E070263" authorityName="Frisch" authorityYear="2022" box="[614,794,1622,1646]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="bipenicillatus">
<emphasis id="B96AAA7EFFE5FFD1BE3DF9AF0E070263" box="[614,794,1622,1646]" italics="true" pageId="37" pageNumber="60">S. bipenicillatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E0DEFFFE5FFD1BCE6F9800C440282" authorityName="Frisch" authorityYear="2022" box="[189,345,1655,1679]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="cuspidatus">
<emphasis id="B96AAA7EFFE5FFD1BCE6F9800C440282" box="[189,345,1655,1679]" italics="true" pageId="37" pageNumber="60">S. cuspidatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E0DEFFFE5FFD1BDF9F9800F410282" authorityName="Frisch" authorityYear="2022" box="[418,604,1655,1679]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="species" species="sulawesianus">
<emphasis id="B96AAA7EFFE5FFD1BDF9F9800F410282" box="[418,604,1655,1679]" italics="true" pageId="37" pageNumber="60">S. sulawesianus</emphasis>
</taxonomicName>
share the dark, strongly sclerotized aedeagus with small phallobase, which has a dorsodistal, membranous window of unknown function (
<figureCitation id="13256AE9FFE5FFD1BDB7F9220F8702FF" box="[492,666,1754,1778]" captionStart-0="Figures 107–109" captionStart-1="Figures 110–112" captionStart-2="Figures 113–115" captionStart-3="Figures 116–118" captionStartId-0="33.[189,262,1243,1265]" captionStartId-1="34.[151,224,1885,1907]" captionStartId-2="35.[189,262,1908,1930]" captionStartId-3="36.[154,227,1907,1929]" captionTargetBox-0="[261,1344,258,1187]" captionTargetBox-1="[223,1310,947,1822]" captionTargetBox-2="[261,1322,1006,1857]" captionTargetBox-3="[224,1268,1020,1861]" captionTargetId-0="figure-309@33.[189,1436,235,1224]" captionTargetId-1="figure-411@34.[151,1398,912,1866]" captionTargetId-2="figure-356@35.[189,1436,978,1889]" captionTargetId-3="figure-354@36.[152,1399,977,1888]" captionTargetPageId-0="33" captionTargetPageId-1="34" captionTargetPageId-2="35" captionTargetPageId-3="36" captionText-0="Figures 107–109. Aedeagus of Scopaeus bipenicillatus, holotype, Sulawesi Tengah, Sigi: Pagana in lateral (107), ventral (108), dorsal view (109). Abbreviations: al – apical lobes, dl – dorsal lobe, dw – dorsodistal window of phallobase, f – flagellum, mf – median foramen, tr– transverse ridge. Arrow Fig. 108: ventral window of phallobase." captionText-1="Figures 110–112. Aedeagus of Scopaeus bipectenatus, holotype, Sulawesi Tengah, Poso: Lake Poso in lateral (110), ventral (111), dorsal view (112). Abbreviations: al – apical lobes, dl – dorsal lobe, dw – dorsodistal window of phallobase, f – flagellum, mf – median foramen, tr – transverse ridge of median foramen. Arrows: dorsal lobe projecting from apical lobes dorsally and ventrally (I, II); ventral, circular windows of apical lobes (III, IV); ventral, bipartite window of phallobase (V)." captionText-2="Figures 113–115. Aedeagus of Scopaeus cuspidatus, holotype, Bali, Jembrana: Cekik in lateral (113), ventral (114), dorsal view (115). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." captionText-3="Figures 116–118. Aedeagus of Scopaeus sulawesianus, holotype, Sulawesi Selatan: Engrekang in lateral (116), ventral (117), dorsal view (118). Abbreviations: al – apical lobes, dw – dorsodistal window of phallobase, mf – median foramen, tr – transverse ridge of median foramen, vl – ventral lobe." figureDoi-0="http://doi.org/10.5281/zenodo.10724292" figureDoi-1="http://doi.org/10.5281/zenodo.10724294" figureDoi-2="http://doi.org/10.5281/zenodo.10724298" figureDoi-3="http://doi.org/10.5281/zenodo.10724302" httpUri-0="https://zenodo.org/record/10724292/files/figure.png" httpUri-1="https://zenodo.org/record/10724294/files/figure.png" httpUri-2="https://zenodo.org/record/10724298/files/figure.png" httpUri-3="https://zenodo.org/record/10724302/files/figure.png" pageId="37" pageNumber="60">Figs 107 – 118</figureCitation>
), and the missing apophysis of the chamber segment of the sperm pump (
<figureCitation id="13256AE9FFE5FFD1BD31F8E40F530339" box="[362,590,1820,1844]" captionStart="Figures 173–178" captionStartId="43.[189,262,1885,1907]" captionTargetId="figure-111@43.[189,1436,294,1840]" captionTargetPageId="43" captionText="Figures 173–178. Sperm pump (173–178) and bursa (173) of ♀ of Scopaeus bipenicillatus, paratype, Sulawesi Tengah, Sigi: Pagana (173, with lateral gonocoxal plates IX; 174); S. bipectenatus, paratype, Sulawesi Tengah: Lake Poso (175); S. cuspidatus, paratype, Jawa Barat: Mt Salak (176); S. crassipunctatus, paratype, Jawa Barat, Sukabumi: Sirnarasa (177); S. sulawesianus, paratype, Central Sulawesi, Tambarana (178, with bursal duct). Long scale bar: 174–178; short scale bar: 173." figureDoi="http://doi.org/10.5281/zenodo.10880230" httpUri="https://zenodo.org/record/10880230/files/figure.png" pageId="37" pageNumber="60">Figs 173 – 175, 178</figureCitation>
). They constitute a distinctive, speciose phylogenetic lineage of
<taxonomicName id="4C1E0DEFFFE5FFD1BCE6F8A60C3B0378" box="[189,294,1886,1909]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="60" phylum="Arthropoda" rank="genus">
<emphasis id="B96AAA7EFFE5FFD1BCE6F8A60C3B0378" box="[189,294,1886,1909]" italics="true" pageId="37" pageNumber="60">Scopaeus</emphasis>
</taxonomicName>
in the Oriental, Australasian, Nearctic and Neotropical regions with a large number of undescribed species.
</paragraph>
</subSubSection>
</treatment>
</document>
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