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<document id="A9D6A2EBBD050300776F26F0230AB16F" ID-CLB-Dataset="291999" ID-DOI="10.25674/so95iss1id311" ID-GBIF-Dataset="06b03cc4-e211-4f65-be07-742740b0053c" ID-ISSN="2509-9523" ID-Zenodo-Dep="10724236" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1709147239332" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Narakusumo, Raden Pramesa" docDate="2022" docId="03B7C77AFFCCFFF9BF72FAB5082D0546" docLanguage="en" docName="SoilOrg.95.1.23-73.pdf" docOrigin="Soil Organisms 95 (1)" docSource="http://dx.doi.org/10.25674/so95iss1id311" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus spiraliflagellatus Frisch 2022, spec. nov." docType="treatment" docVersion="4" lastPageNumber="36" masterDocId="FF8EBF02FFC0FFF4BC5BFFF80D1D040D" masterDocTitle="Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species" masterLastPageNumber="73" masterPageNumber="23" pageNumber="35" updateTime="1711467745701" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="33CCA7082475C80C7AD240E31D6DEB72">Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species</mods:title>
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<mods:namePart id="EFB6824D229F327242233B5F2A2E4A4E">Frisch, Johannes</mods:namePart>
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<mods:namePart id="D7EB4CF513226C704510A722395BE368">Narakusumo, Raden Pramesa</mods:namePart>
<mods:affiliation id="61B2AAD230147BBDBF1CFEFF2E0C4CE7">MUseUm ZoologicUm Bogoriense, Research Center for Biosystematics and EvolUtion, National Research and Innovation Agency (BRIN), Jl. Raya Jakarta-Bogor KM 46, Cibinong, West Java, 16911, Indonesia</mods:affiliation>
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<mods:title id="1E87F6B1478C744AE266BD830A2C1C58">Soil Organisms</mods:title>
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<mods:date id="073D531446AF1AF44848801E3285491D">2022</mods:date>
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<treatment id="03B7C77AFFCCFFF9BF72FAB5082D0546" ID-DOI="http://doi.org/10.5281/zenodo.10880251" ID-Zenodo-Dep="10880251" LSID="urn:lsid:plazi:treatment:03B7C77AFFCCFFF9BF72FAB5082D0546" httpUri="http://treatment.plazi.org/id/03B7C77AFFCCFFF9BF72FAB5082D0546" lastPageId="13" lastPageNumber="36" pageId="12" pageNumber="35">
<subSubSection id="C30425E7FFCCFFF8BF72FAB508560165" box="[809,1355,1357,1384]" pageId="12" pageNumber="35" type="nomenclature">
<paragraph id="8BA1766CFFCCFFF8BF72FAB508560165" blockId="12.[809,1355,1357,1417]" box="[809,1355,1357,1384]" pageId="12" pageNumber="35">
<heading id="D0E9C100FFCCFFF8BF72FAB508560165" box="[809,1355,1357,1384]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="35" reason="5">
<taxonomicName id="4C1E0DEFFFCCFFF8BF72FAB509D4016A" authority="Frisch" authorityName="Frisch" authorityYear="2022" box="[809,1225,1357,1384]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="35" phylum="Arthropoda" rank="species" species="spiraliflagellatus" status="sp. nov.">
<emphasis id="B96AAA7EFFCCFFF8BF72FAB509D4016A" bold="true" box="[809,1225,1357,1384]" pageId="12" pageNumber="35">
<emphasis id="B96AAA7EFFCCFFF8BF72FAB5096C0165" bold="true" box="[809,1137,1357,1384]" italics="true" pageId="12" pageNumber="35">Scopaeus spiraliflagellatus</emphasis>
Frisch
</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2591705FFCCFFF8B88CFAB608560165" box="[1239,1355,1358,1384]" pageId="12" pageNumber="35" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCCFFF8BF72FA9609A30184" box="[809,1214,1390,1417]" pageId="12" pageNumber="35" type="description">
<paragraph id="8BA1766CFFCCFFF8BF72FA9609A30184" blockId="12.[809,1355,1357,1417]" box="[809,1214,1390,1417]" pageId="12" pageNumber="35">
(
<figureCitation id="13256AE9FFCCFFF8BF6AFA960E970184" box="[817,906,1390,1417]" captionStart="Figures 810" captionStartId="5.[189,262,1930,1952]" captionTargetBox="[237,1406,902,1874]" captionTargetId="figure-357@5.[189,1436,889,1911]" captionTargetPageId="5" captionText="Figures 810. Habitus of Scopaeus tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (8); S. posoanus, paratype, Sulawesi Tengah: Lake Poso (9); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (10)." figureDoi="http://doi.org/10.5281/zenodo.10724240" httpUri="https://zenodo.org/record/10724240/files/figure.png" pageId="12" pageNumber="35">Figs 10</figureCitation>
,
<figureCitation id="13256AE9FFCCFFF8BFCDFA960EA90184" box="[918,948,1390,1417]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="12" pageNumber="35">35</figureCitation>
,
<figureCitation id="13256AE9FFCCFFF8BFE4FA97090D0185" box="[959,1040,1390,1417]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="12" pageNumber="35">74 76</figureCitation>
,
<figureCitation id="13256AE9FFCCFFF8B847FA9709540184" box="[1052,1097,1391,1417]" captionStart="Figures 119133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119133. Abdominal sternite VII (126) and abdominal sternite VIII (119125, 127 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="12" pageNumber="35">128</figureCitation>
,
<figureCitation id="13256AE9FFCCFFF8B80EFA9609A80184" box="[1109,1205,1390,1417]" captionStart="Figures 155162" captionStartId="41.[189,262,1839,1861]" captionTargetBox="[260,1402,386,1672]" captionTargetId="figure-168@41.[189,1436,316,1820]" captionTargetPageId="41" captionText="Figures 155162. Sperm pump (155162) and bursa (155, 159, 161) of ♀ of Scopaeus posoanus, paratype, Sulawesi Tengah: Lake Poso (155, with lateral gonocoxal plates IX; 156); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (157, with bursal duct; 158); S. anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (159, with lateral gonocoxal plates folded laterally; 160); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (161, 162). Long scale bar: 156, 158, 160, 162; short scale bar: 155, 157, 159, 161. Abbreviations: ap apophysis of chamber segment, b (sclerotized end of) bursa, bd bursal duct, cs chamber segment, dp distal process of chamber segment, lt lateral gonocoxal plate IX, sd spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.10724311" httpUri="https://zenodo.org/record/10724311/files/figure.png" pageId="12" pageNumber="35">157, 158</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCCFFF9BF72FA4A0E090527" lastPageId="13" lastPageNumber="36" pageId="12" pageNumber="35" type="materials_examined">
<paragraph id="8BA1766CFFCCFFF9BF72FA4A0E090527" blockId="12.[786,1399,1458,1976]" lastBlockId="13.[189,800,241,1975]" lastPageId="13" lastPageNumber="36" pageId="12" pageNumber="35">
<emphasis id="B96AAA7EFFCCFFF8BF72FA4A0EF301C7" bold="true" box="[809,1006,1458,1482]" pageId="12" pageNumber="35">Type specimens:</emphasis>
<materialsCitation id="3B767C31FFCCFFF8BFA6FA4A086E0201" collectingDate="2017-05-09" collectionCode="MZB" collectorName="Frisch" country="Indonesia" county="Poso" elevation="550" latitude="-1.7997222" location="Lake Poso" longLatPrecision="21" longitude="120.63555" municipality="Tentena - Peura" pageId="12" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Tengah" typeStatus="holotype">
<collectingCountry id="F30936FCFFCCFFF8BFA6FA4A097701C7" box="[1021,1130,1458,1482]" name="Indonesia" pageId="12" pageNumber="35">Indonesia</collectingCountry>
:
<typeStatus id="54A5C8CEFFCCFFF8B820FA4A09C201C7" box="[1147,1247,1458,1482]" pageId="12" pageNumber="35" type="holotype">Holotype</typeStatus>
<specimenCount id="9D18BDE5FFCCFFF8B8B4FA4B081E01C7" box="[1263,1283,1459,1482]" count="1" pageId="12" pageNumber="35" type="male"></specimenCount>
,
<collectingRegion id="49DAB88EFFCCFFF8B94DFA4A0E7901E6" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="35">Sulawesi Tengah</collectingRegion>
,
<collectingCounty id="62C00EE0FFCCFFF8BF35FA2B0EB801E7" box="[878,933,1491,1514]" pageId="12" pageNumber="35">Poso</collectingCounty>
:
<collectingMunicipality id="6BC5EC16FFCCFFF8BFF5FA2B094701E7" box="[942,1114,1491,1514]" pageId="12" pageNumber="35">Tentena Peura</collectingMunicipality>
(
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), (
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,
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),
<quantity id="4CE6DB89FFCCFFF8BF9AFA0C090C0206" box="[961,1041,1524,1548]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="35" unit="m" value="550.0">
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</quantity>
,
<date id="FFA050ACFFCCFFF8B873FA0C099D0201" box="[1064,1152,1524,1548]" pageId="12" pageNumber="35" value="2017-05-09">
<collectingDate id="EFE4A944FFCCFFF8B873FA0C099D0201" box="[1064,1152,1524,1548]" pageId="12" pageNumber="35" value="2017-05-09">9.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8B894FA0C080F0201" box="[1231,1298,1524,1548]" pageId="12" pageNumber="35">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8B977FA0C08760201" box="[1324,1387,1524,1548]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="12" pageNumber="35" type="Museum">MZB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFCCFFF8BF49F9ED0E690243" collectingDate="2017-05-09" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Poso" elevation="550" latitude="-1.7997222" location="Lake Poso" longLatPrecision="21" longitude="120.63555" municipality="Tentena - Peura" pageId="12" pageNumber="35" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<typeStatus id="54A5C8CEFFCCFFF8BF49F9ED0E600220" box="[786,893,1557,1581]" pageId="12" pageNumber="35" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D18BDE5FFCCFFF8BFD4F9ED09340220" box="[911,1065,1557,1581]" count="16" pageId="12" pageNumber="35" type="generic">16 specimens</specimenCount>
):
<specimenCount id="9D18BDE5FFCCFFF8B865F9ED09740221" box="[1086,1129,1557,1581]" count="1" pageId="12" pageNumber="35" type="female">1 ♀</specimenCount>
, same data as holotype (
<collectionCode id="ED0FEEA9FFCCFFF8BF41F9CE0E710243" box="[794,876,1590,1614]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="35">MFNB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFCCFFF8BFD8F9CE0EDE0282" collectingDate="2017-05-09" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Poso" elevation="520" latitude="-1.8633333" location="Lake Poso" longLatPrecision="21" longitude="120.64972" municipality="Peura" pageId="12" pageNumber="35" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<collectingRegion id="49DAB88EFFCCFFF8BFD8F9CE095F0243" box="[899,1090,1590,1614]" country="Indonesia" name="Sulawesi Tengah" pageId="12" pageNumber="35">Sulawesi Tengah</collectingRegion>
:
<specimenCount id="9D18BDE5FFCCFFF8B80BF9CE09610240" box="[1104,1148,1590,1614]" count="1" pageId="12" pageNumber="35" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62C00EE0FFCCFFF8B8D5F9CE09D80240" box="[1166,1221,1590,1613]" pageId="12" pageNumber="35">Poso</collectingCounty>
: S
<collectingMunicipality id="6BC5EC16FFCCFFF8B8B5F9CE08360240" box="[1262,1323,1590,1613]" pageId="12" pageNumber="35">Peura</collectingMunicipality>
(
<location id="8EC120B7FFCCFFF8B964F9CE0E550262" LSID="urn:lsid:plazi:treatment:03B7C77AFFCCFFF9BF72FAB5082D0546:8EC120B7FFCCFFF8B964F9CE0E550262" country="Indonesia" county="Poso" latitude="-1.8633333" longLatPrecision="21" longitude="120.64972" municipality="Peura" name="Lake Poso" pageId="12" pageNumber="35" stateProvince="Sulawesi Tengah">Lake Poso</location>
), (
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,
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),
<quantity id="4CE6DB89FFCCFFF8B8C8F9AE09C60263" box="[1171,1243,1622,1646]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="35" unit="m" value="520.0">
<elevation id="0033915FFFCCFFF8B8C8F9AE09C60263" box="[1171,1243,1622,1646]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="35" unit="m" value="520.0">520 m</elevation>
</quantity>
,
<date id="FFA050ACFFCCFFF8B8B2F9AE085F0263" box="[1257,1346,1622,1646]" pageId="12" pageNumber="35" value="2017-05-09">
<collectingDate id="EFE4A944FFCCFFF8B8B2F9AE085F0263" box="[1257,1346,1622,1646]" pageId="12" pageNumber="35" value="2017-05-09">9.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8BF49F98F0E480282" box="[786,853,1655,1679]" pageId="12" pageNumber="35">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8BF32F98F0EA60282" box="[873,955,1655,1679]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="35">MFNB</collectionCode>
)
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,
<collectingMunicipality id="6BC5EC16FFCCFFF8B84FF98F099C0282" box="[1044,1153,1655,1679]" pageId="12" pageNumber="35">Morowali</collectingMunicipality>
:
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(
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),
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,
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<collectingDate id="EFE4A944FFCCFFF8B88AF960082502BD" box="[1233,1336,1688,1712]" pageId="12" pageNumber="35" value="2017-12-05">12.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8BF49F9410E4802DC" box="[786,853,1721,1745]" pageId="12" pageNumber="35">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8BF36F9410EA202DC" box="[877,959,1721,1745]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="35">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFCCFFF8BF81F9410EA1031E" collectingDate="2017-10-05" collectionCode="MFNB, MZB" collectorName="Frisch" country="Indonesia" elevation="880" latitude="-1.8111111" location="Tentena - Taripa" longLatPrecision="21" longitude="120.775" municipality="Poso" pageId="12" pageNumber="35" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCCFFF8BF81F941091602DC" box="[986,1035,1721,1745]" count="1" pageId="12" pageNumber="35" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFCCFFF8B87BF941094D02DC" box="[1056,1104,1721,1745]" count="5" pageId="12" pageNumber="35" type="female">5 ♀</specimenCount>
,
<collectingMunicipality id="6BC5EC16FFCCFFF8B83DF942098002DC" box="[1126,1181,1722,1745]" pageId="12" pageNumber="35">Poso</collectingMunicipality>
:
<location id="8EC120B7FFCCFFF8B8EBF942086B02DC" LSID="urn:lsid:plazi:treatment:03B7C77AFFCCFFF9BF72FAB5082D0546:8EC120B7FFCCFFF8B8EBF942086B02DC" box="[1200,1398,1721,1745]" country="Indonesia" latitude="-1.8111111" longLatPrecision="21" longitude="120.775" municipality="Poso" name="Tentena - Taripa" pageId="12" pageNumber="35" stateProvince="Sulawesi Tengah">Tentena Taripa</location>
(
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,
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),
<quantity id="4CE6DB89FFCCFFF8B81EF922099702FC" box="[1093,1162,1754,1778]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="12" pageNumber="35" unit="m" value="880.0">
<elevation id="0033915FFFCCFFF8B81EF922099702FC" box="[1093,1162,1754,1778]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="12" pageNumber="35" unit="m" value="880.0">880 m</elevation>
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,
<date id="FFA050ACFFCCFFF8B8CDF92209E602FF" box="[1174,1275,1754,1778]" pageId="12" pageNumber="35" value="2017-10-05">
<collectingDate id="EFE4A944FFCCFFF8B8CDF92209E602FF" box="[1174,1275,1754,1778]" pageId="12" pageNumber="35" value="2017-10-05">10.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8B968F922086B02FF" box="[1331,1398,1754,1778]" pageId="12" pageNumber="35">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8BF41F9030E71031E" box="[794,876,1787,1811]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="35">MFNB</collectionCode>
,
<collectionCode id="ED0FEEA9FFCCFFF8BF2CF9030EA8031E" box="[887,949,1787,1811]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="12" pageNumber="35" type="Museum">MZB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFCCFFF8BF9CF9030E690358" collectingDate="2017-05-09" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Poso" elevation="550" latitude="-1.7997222" location="Lake Poso" longLatPrecision="21" longitude="120.63555" municipality="Tentena - Peura" pageId="12" pageNumber="35" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCCFFF8BF9CF9030EF3031F" box="[967,1006,1787,1811]" count="1" pageId="12" pageNumber="35" type="female">1 ♀</specimenCount>
,
<collectingCounty id="62C00EE0FFCCFFF8BFA0F903092F031F" box="[1019,1074,1787,1810]" pageId="12" pageNumber="35">Poso</collectingCounty>
:
<collectingMunicipality id="6BC5EC16FFCCFFF8B867F90309F5031F" box="[1084,1256,1787,1810]" pageId="12" pageNumber="35">Tentena Peura</collectingMunicipality>
(
<location id="8EC120B7FFCCFFF8B8ACF9030876031E" LSID="urn:lsid:plazi:treatment:03B7C77AFFCCFFF9BF72FAB5082D0546:8EC120B7FFCCFFF8B8ACF9030876031E" box="[1271,1387,1787,1811]" country="Indonesia" county="Poso" latitude="-1.7997222" longLatPrecision="21" longitude="120.63555" municipality="Tentena - Peura" name="Lake Poso" pageId="12" pageNumber="35" stateProvince="Sulawesi Tengah">Lake Poso</location>
), (
<geoCoordinate id="EE2A10ABFFCCFFF8BF41F8E40E8A0339" box="[794,919,1820,1844]" degrees="01" direction="south" minutes="47" orientation="latitude" pageId="12" pageNumber="35" precision="15" seconds="59" value="-1.7997222">01°4759S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFCCFFF8BFFFF8E409280339" box="[932,1077,1820,1844]" degrees="120" direction="east" minutes="38" orientation="longitude" pageId="12" pageNumber="35" precision="15" seconds="08" value="120.63555">120°3808E</geoCoordinate>
),
<quantity id="4CE6DB89FFCCFFF8B811F8E4098C033E" box="[1098,1169,1820,1844]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="35" unit="m" value="550.0">
<elevation id="0033915FFFCCFFF8B811F8E4098C033E" box="[1098,1169,1820,1844]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="12" pageNumber="35" unit="m" value="550.0">550 m</elevation>
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,
<date id="FFA050ACFFCCFFF8B8C5F8E409EA0339" box="[1182,1271,1820,1844]" pageId="12" pageNumber="35" value="2017-05-09">
<collectingDate id="EFE4A944FFCCFFF8B8C5F8E409EA0339" box="[1182,1271,1820,1844]" pageId="12" pageNumber="35" value="2017-05-09">9.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8B968F8E4086B0339" box="[1331,1398,1820,1844]" pageId="12" pageNumber="35">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8BF41F8C50E710358" box="[794,876,1853,1877]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="35">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B767C31FFCCFFF8BFD8F8C50E69039B" collectingDate="2017-05-06" collectionCode="MFNB" collectorName="Frisch" country="Indonesia" county="Poso" elevation="1250" latitude="-1.3005555" location="Sedoa" longLatPrecision="21" longitude="120.27278" municipality="Palolo - Sedoa" pageId="12" pageNumber="35" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sulawesi Tengah" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCCFFF8BFD8F8C50EB20359" box="[899,943,1853,1877]" count="1" pageId="12" pageNumber="35" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D18BDE5FFCCFFF8BFE4F8C50EF60359" box="[959,1003,1853,1877]" count="3" pageId="12" pageNumber="35" type="female">3 ♀</specimenCount>
,
<collectingCounty id="62C00EE0FFCCFFF8BFA7F8C5092E0359" box="[1020,1075,1853,1876]" pageId="12" pageNumber="35">Poso</collectingCounty>
:
<collectingMunicipality id="6BC5EC16FFCCFFF8B81AF8C509F30358" box="[1089,1262,1853,1877]" pageId="12" pageNumber="35">Palolo Sedoa</collectingMunicipality>
: NW
<location id="8EC120B7FFCCFFF8B96EF8C5086A0358" LSID="urn:lsid:plazi:treatment:03B7C77AFFCCFFF9BF72FAB5082D0546:8EC120B7FFCCFFF8B96EF8C5086A0358" box="[1333,1399,1853,1877]" country="Indonesia" county="Poso" latitude="-1.3005555" longLatPrecision="21" longitude="120.27278" municipality="Palolo - Sedoa" name="Sedoa" pageId="12" pageNumber="35" stateProvince="Sulawesi Tengah">Sedoa</location>
(
<geoCoordinate id="EE2A10ABFFCCFFF8BF41F8A60E8A037B" box="[794,919,1886,1910]" degrees="01" direction="south" minutes="18" orientation="latitude" pageId="12" pageNumber="35" precision="15" seconds="02" value="-1.3005555">01°1802S</geoCoordinate>
,
<geoCoordinate id="EE2A10ABFFCCFFF8BFF9F8A5092D0378" box="[930,1072,1885,1909]" degrees="120" direction="east" minutes="16" orientation="longitude" pageId="12" pageNumber="35" precision="15" seconds="22" value="120.27278">120°1622E</geoCoordinate>
),
<quantity id="4CE6DB89FFCCFFF8B81FF8A5098B0378" box="[1092,1174,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="12" pageNumber="35" unit="m" value="1250.0">
<elevation id="0033915FFFCCFFF8B81FF8A5098B0378" box="[1092,1174,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="12" pageNumber="35" unit="m" value="1250.0">1250 m</elevation>
</quantity>
,
<date id="FFA050ACFFCCFFF8B8FAF8A509E60378" box="[1185,1275,1885,1909]" pageId="12" pageNumber="35" value="2017-05-06">
<collectingDate id="EFE4A944FFCCFFF8B8FAF8A509E60378" box="[1185,1275,1885,1909]" pageId="12" pageNumber="35" value="2017-05-06">6.5.2017</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8B968F8A5086B0378" box="[1331,1398,1885,1909]" pageId="12" pageNumber="35">Frisch</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8BF41F8860E71039B" box="[794,876,1918,1942]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="35">MFNB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFCCFFF8BFDEF886093C03BA" collectingDate="1982-07-17" collectionCode="HECO" collectorName="Rougemont" country="Indonesia" location="Makale" pageId="12" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Selatan" typeStatus="paratype">
<collectingRegion id="49DAB88EFFCCFFF8BFDEF8860955039B" box="[901,1096,1918,1942]" country="Indonesia" name="Sulawesi Selatan" pageId="12" pageNumber="35">Sulawesi Selatan</collectingRegion>
:
<specimenCount id="9D18BDE5FFCCFFF8B802F8860995039B" box="[1113,1160,1918,1942]" count="1" pageId="12" pageNumber="35" type="male">1 ♂</specimenCount>
,
<location id="8EC120B7FFCCFFF8B8C0F88609F2039B" LSID="urn:lsid:plazi:treatment:03B7C77AFFCCFFF9BF72FAB5082D0546:8EC120B7FFCCFFF8B8C0F88609F2039B" box="[1179,1263,1918,1942]" country="Indonesia" name="Makale" pageId="12" pageNumber="35" stateProvince="Sulawesi Selatan">Makale</location>
,
<date id="FFA050ACFFCCFFF8B95AF886086F039B" box="[1281,1394,1918,1942]" pageId="12" pageNumber="35" value="1982-07-17">
<collectingDate id="EFE4A944FFCCFFF8B95AF886086F039B" box="[1281,1394,1918,1942]" pageId="12" pageNumber="35" value="1982-07-17">17.07.1982</collectingDate>
</date>
, leg.
<collectorName id="26EB13BAFFCCFFF8BF1BF8580EA103B5" box="[832,956,1952,1976]" pageId="12" pageNumber="35">Rougemont</collectorName>
(
<collectionCode id="ED0FEEA9FFCCFFF8BF90F867090403BA" box="[971,1049,1951,1975]" pageId="12" pageNumber="35">HECO</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B767C31FFCCFFF9B870F8670E0D0527" collectionCode="ISNB" country="Indonesia" lastPageId="13" lastPageNumber="36" location="Celebes" pageId="12" pageNumber="35" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D18BDE5FFCCFFF8B870F867094E03BA" box="[1067,1107,1951,1975]" count="1" pageId="12" pageNumber="35" type="male">1 ♂</specimenCount>
, labeled
<collectingRegion id="49DAB88EFFCCFFF8B8ECF867080F03BA" box="[1207,1298,1951,1975]" country="Indonesia" name="Sulawesi Utara" pageId="12" pageNumber="35">Célebès</collectingRegion>
,
<collectorName id="26EB13BAFFCCFFF8B945F867086A03BA" box="[1310,1399,1951,1975]" pageId="12" pageNumber="35">Kandari</collectorName>
7 (handwritten;
<collectorName id="26EB13BAFFCDFFF9BDCBFF090CFB0504" box="[400,486,241,265]" pageId="13" pageNumber="36">Kendari</collectorName>
in
<collectingRegion id="49DAB88EFFCDFFF9BE71FF090E170507" box="[554,778,241,266]" country="Indonesia" name="Sulawesi Tenggara" pageId="13" pageNumber="36">Sulawesi Tenggara</collectingRegion>
?),
<emphasis id="B96AAA7EFFCDFFF9BC9DFEEB0C5C0527" box="[198,321,275,298]" italics="true" pageId="13" pageNumber="36">apicipennis</emphasis>
<collectorName id="26EB13BAFFCDFFF9BD1CFEEA0C6C0527" box="[327,369,274,298]" pageId="13" pageNumber="36">Fvl</collectorName>
(handwritten; in litteris name) (
<collectionCode id="ED0FEEA9FFCDFFF9BE9DFEEA0E150527" box="[710,776,274,298]" pageId="13" pageNumber="36">ISNB</collectionCode>
)
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.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCDFFF9BC8FFECB0FF702FF" pageId="13" pageNumber="36" type="description">
<paragraph id="8BA1766CFFCDFFF9BC8FFECB0FE80734" blockId="13.[189,800,241,1975]" pageId="13" pageNumber="36">
<emphasis id="B96AAA7EFFCDFFF9BC8FFECB0C430546" bold="true" box="[212,350,307,331]" pageId="13" pageNumber="36">Description:</emphasis>
Habitus and coloring as in
<figureCitation id="13256AE9FFCDFFF9BED5FECB0FC00546" box="[654,733,307,331]" captionStart="Figures 810" captionStartId="5.[189,262,1930,1952]" captionTargetBox="[237,1406,902,1874]" captionTargetId="figure-357@5.[189,1436,889,1911]" captionTargetPageId="5" captionText="Figures 810. Habitus of Scopaeus tortuosiflagellatus, paratype, Jawa Barat, Sukabumi: Cisalimas (8); S. posoanus, paratype, Sulawesi Tengah: Lake Poso (9); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (10)." figureDoi="http://doi.org/10.5281/zenodo.10724240" httpUri="https://zenodo.org/record/10724240/files/figure.png" pageId="13" pageNumber="36">Fig. 10</figureCitation>
. Total body length 3.1 4.0 mm; forebody length
<quantity id="4CE6DB89FFCDFFF9BECEFEAC0E000561" box="[661,797,340,364]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="13" pageNumber="36" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.9 2.2 mm</quantity>
. Head subcircular. Antennae with penultimate segment elongate. Mesotibia strongly thickened. Forebody surface with extremely spacious, fine, shallow, setose punctation, strongly shiny; abdomen also very finely, but more densely punctate than forebody, thus somewhat matter; body surface without microreticulation. Forebody setation comparatively long and strongly raised, including numerous, long macrosetae. Body black-brown except for yellow-brown posterior fifth of elytra, tip of abdomen, and legs with dark brown basal half of profemora becoming evenly lighter towards yellow-brown distal half; maxillary palpi darkened; antennae with scapus medium brown, pedicellus slightly darkened, segment 3 darkest, then evenly lighter towards light yellow-brown terminal segments.
</paragraph>
<paragraph id="8BA1766CFFCDFFF9BC8FFCB90F480220" blockId="13.[189,800,241,1975]" pageId="13" pageNumber="36">
Male:Abdominal sternite VII with unmodified posterior margin. Abdominal sternite VIII widely emarginate in posterior fifth with median third of emargination almost semicircularly projecting posteriad (
<figureCitation id="13256AE9FFCDFFF9BEF6FC5C0E0F07B1" box="[685,786,932,956]" captionStart="Figures 119133" captionStartId="38.[151,224,1796,1818]" captionTargetBox="[224,1369,297,1686]" captionTargetId="figure-201@38.[151,1398,236,1777]" captionTargetPageId="38" captionText="Figures 119133. Abdominal sternite VII (126) and abdominal sternite VIII (119125, 127 133) of ♂ of Scopaeus javanus, lectotype, Jawa Barat, N Bandung: Mt Tangkuban Perahu (119); S. batukaruensis, holotype, Bali, Tabanan: Bedugul (120); S. riedeli, holotype, Sumatra, South Lampung: Rajabasa Mts (121); S. jacobsoni, Jawa Barat: Mt Salak (122); S. mixtus, Malaysia, Sabah: Lokan (123); S. halimunsalakensis, holotype, Jawa Barat, Sukabumi: Sirnarasa (124); S. tortuosiflagellatus, holotype, Jawa Barat, Sukabumi: Cisalimas (125); S. posoanus, holotype, Sulawesi Tengah: Lake Poso (126, 127); S. spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (128); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi:Cidahu (129); S. pulcher, holotype, Jawa Barat, Sukabumi:Cikaniki (130); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (131); S. heronifer, holotype, Banten, Lebak: Majasari (132); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (133)." figureDoi="http://doi.org/10.5281/zenodo.10724304" httpUri="https://zenodo.org/record/10724304/files/figure.png" pageId="13" pageNumber="36">Fig. 128</figureCitation>
); macrosetae of sternite VIII long, conspicuous. Aedeagus (
<figureCitation id="13256AE9FFCDFFF9BC9FFC1E0C0107F3" box="[196,284,998,1022]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 35</figureCitation>
,
<figureCitation id="13256AE9FFCDFFF9BD77FC1E0C6B07F3" box="[300,374,998,1022]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">74 76</figureCitation>
) about
<quantity id="4CE6DB89FFCDFFF9BD8BFC1E0F3C07F0" box="[464,545,998,1022]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="36" unit="mm" value="1.2">1.2 mm</quantity>
long with distal lobes strongly asymmetrical; apical lobes remarkably enlarged apically with short, right-angled dorsad curved apices; dextral apical lobe ventroproximad extended, ending in dorsad pointing hook (
<figureCitation id="13256AE9FFCDFFF9BDBBFB910F25008C" box="[480,568,1129,1153]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 35</figureCitation>
,
<figureCitation id="13256AE9FFCDFFF9BE12FB910F7A008C" box="[585,615,1129,1153]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">74</figureCitation>
); ends of apical lobes, in ventral and dorsal view, curved towards each other, together forming triangular end (
<figureCitation id="13256AE9FFCDFFF9BEDDFB530E0F00CE" box="[646,786,1195,1219]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Figs 75, 76</figureCitation>
); dorsal lobe subrectangular in lateral view (
<figureCitation id="13256AE9FFCDFFF9BED0FB340FC100E9" box="[651,732,1228,1252]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Fig. 74</figureCitation>
), with transverse, slightly oblique, truncate end in dorsal view (
<figureCitation id="13256AE9FFCDFFF9BC9EFAF60C09012B" box="[197,276,1294,1318]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Fig. 76</figureCitation>
); flagellum extremely lengthened, whip-like, thin proximally, spiral with two windings at dextral side of aedeagus (
<figureCitation id="13256AE9FFCDFFF9BD74FAB70C9E016A" box="[303,387,1359,1383]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 35</figureCitation>
,
<figureCitation id="13256AE9FFCDFFF9BDCBFAA80CC0016A" box="[400,477,1359,1384]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">74 76</figureCitation>
); ventral lobe elongate, about four times as long as wide, pointing ventrodistad (
<figureCitation id="13256AE9FFCDFFF9BE92FA880E060185" box="[713,795,1392,1416]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 35</figureCitation>
,
<figureCitation id="13256AE9FFCDFFF9BCE6FA690DC601A4" box="[189,219,1425,1449]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">74</figureCitation>
); setose lateral lobes short, little projecting, strongly shifted distad to different extents (
<figureCitation id="13256AE9FFCDFFF9BE0DFA4A0FAF01C7" box="[598,690,1458,1482]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 35</figureCitation>
,
<figureCitation id="13256AE9FFCDFFF9BE9DFA4A0E0F01C7" box="[710,786,1458,1482]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">74 76</figureCitation>
); median foramen large, asymmetrically extended distad with concave, distolateral enlargement at either side, without transverse ridge (
<figureCitation id="13256AE9FFCDFFF9BD96F9ED0F540220" box="[461,585,1557,1581]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">Figs 74, 75</figureCitation>
).
</paragraph>
<paragraph id="8BA1766CFFCDFFF9BC8FF9CE0C5302BD" blockId="13.[189,800,241,1975]" pageId="13" pageNumber="36">
Female: Sperm pump with apophysis adjacent to bursal duct; bursal duct strongly sclerotized in distal half, proximal half membranous (
<figureCitation id="13256AE9FFCDFFF9BE6AF98F0FD30282" box="[561,718,1655,1679]" captionStart="Figures 155162" captionStartId="41.[189,262,1839,1861]" captionTargetBox="[260,1402,386,1672]" captionTargetId="figure-168@41.[189,1436,316,1820]" captionTargetPageId="41" captionText="Figures 155162. Sperm pump (155162) and bursa (155, 159, 161) of ♀ of Scopaeus posoanus, paratype, Sulawesi Tengah: Lake Poso (155, with lateral gonocoxal plates IX; 156); S. spiraliflagellatus, paratype, Sulawesi Tengah: Lake Poso (157, with bursal duct; 158); S. anuliflagellatus, paratype, Jawa Barat, Sukabumi: Cikaniki (159, with lateral gonocoxal plates folded laterally; 160); S. pulcher, paratype, Jawa Barat, Sukabumi: Cikaniki (161, 162). Long scale bar: 156, 158, 160, 162; short scale bar: 155, 157, 159, 161. Abbreviations: ap apophysis of chamber segment, b (sclerotized end of) bursa, bd bursal duct, cs chamber segment, dp distal process of chamber segment, lt lateral gonocoxal plate IX, sd spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.10724311" httpUri="https://zenodo.org/record/10724311/files/figure.png" pageId="13" pageNumber="36">Figs 157, 158</figureCitation>
); bursa membranous.
</paragraph>
<paragraph id="8BA1766CFFCDFFF9BC8FF9410FF702FF" blockId="13.[189,800,241,1975]" pageId="13" pageNumber="36">
<emphasis id="B96AAA7EFFCDFFF9BC8FF9410C4E02DC" bold="true" box="[212,339,1721,1745]" pageId="13" pageNumber="36">Phylogeny:</emphasis>
<taxonomicName id="4C1E0DEFFFCDFFF9BD01F9410F6702DC" authorityName="Frisch" authorityYear="2022" box="[346,634,1721,1745]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="spiraliflagellatus">
<emphasis id="B96AAA7EFFCDFFF9BD01F9410F6702DC" box="[346,634,1721,1745]" italics="true" pageId="13" pageNumber="36">Scopaeus spiraliflagellatus</emphasis>
</taxonomicName>
is a member of the
<taxonomicName id="4C1E0DEFFFCDFFF9BCBFF9220C4D02FF" authorityName="Oke" authorityYear="1933" box="[228,336,1754,1778]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="B96AAA7EFFCDFFF9BCBFF9220C4D02FF" box="[228,336,1754,1778]" italics="true" pageId="13" pageNumber="36">S. gracilis</emphasis>
</taxonomicName>
species group (
<bibRefCitation id="EF8F0B9DFFCDFFF9BDACF9220FC302FF" author="Frisch, J. &amp; D. Burckhardt &amp; V. Wolters" box="[503,734,1754,1778]" pageId="13" pageNumber="36" pagination="27 - 53" refId="ref31920" refString="Frisch, J., D. Burckhardt &amp; V. Wolters (2002): Rove beetles of the subtribe Scopaeina Mulsant &amp; Rey (Coleoptera: Staphylinidae) in the West Palaearctic: Phylogeny, biogeography and species catalogue. - Organisms, Diversity &amp; Evolution 2: 27 - 53." type="journal article" year="2002">Frisch et al. 2002: 39</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCDFFF9BC8FF9030C3A037B" pageId="13" pageNumber="36" type="distribution">
<paragraph id="8BA1766CFFCDFFF9BC8FF9030C3A037B" blockId="13.[189,800,241,1975]" pageId="13" pageNumber="36">
<emphasis id="B96AAA7EFFCDFFF9BC8FF9030C75031E" bold="true" box="[212,360,1787,1811]" pageId="13" pageNumber="36">Distribution:</emphasis>
<taxonomicName id="4C1E0DEFFFCDFFF9BD20F9030FBA031E" authorityName="Frisch" authorityYear="2022" box="[379,679,1787,1811]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="spiraliflagellatus">
<emphasis id="B96AAA7EFFCDFFF9BD20F9030FBA031E" box="[379,679,1787,1811]" italics="true" pageId="13" pageNumber="36">Scopaeus spiraliflagellatus</emphasis>
</taxonomicName>
, probably widespread in
<collectingRegion id="49DAB88EFFCDFFF9BD06F8E40CDD0339" box="[349,448,1820,1844]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="36">Sulawesi</collectingRegion>
, is recorded in the surroundings of Lake Poso,
<collectingRegion id="49DAB88EFFCDFFF9BD0EF8C50F110358" box="[341,524,1853,1877]" country="Indonesia" name="Sulawesi Tengah" pageId="13" pageNumber="36">Sulawesi Tengah</collectingRegion>
, and in Kendari,
<collectingRegion id="49DAB88EFFCDFFF9BEE4F8C50C3E037B" country="Indonesia" name="Sulawesi Tenggara" pageId="13" pageNumber="36">Sulawesi Tenggara</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30425E7FFCDFFF9BC8FF886082D0546" pageId="13" pageNumber="36" type="etymology">
<paragraph id="8BA1766CFFCDFFF9BC8FF886082D0546" blockId="13.[189,800,241,1975]" lastBlockId="13.[824,1437,241,331]" pageId="13" pageNumber="36">
<emphasis id="B96AAA7EFFCDFFF9BC8FF8860C45039B" bold="true" box="[212,344,1918,1942]" pageId="13" pageNumber="36">Etymology:</emphasis>
The epithet
<taxonomicName id="4C1E0DEFFFCDFFF9BDB5F8860FBE039B" authorityName="Frisch" authorityYear="2022" box="[494,675,1918,1942]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="spiraliflagellatus">
<emphasis id="B96AAA7EFFCDFFF9BDB5F8860FBE039B" box="[494,675,1918,1942]" italics="true" pageId="13" pageNumber="36">spiraliflagellatus</emphasis>
</taxonomicName>
[adjective, Latin, composed of the noun
<emphasis id="B96AAA7EFFCDFFF9BE4EF8670F7803BA" box="[533,613,1951,1975]" italics="true" pageId="13" pageNumber="36">spiralis</emphasis>
(spiral) and the adjective
<emphasis id="B96AAA7EFFCDFFF9BFFDFF0909080504" box="[934,1045,241,265]" italics="true" pageId="13" pageNumber="36">flagellatus</emphasis>
(indicating a characteristic/feature of the flagellum)] refers to the spiral flagellum of the aedeagus of this new species (
<figureCitation id="13256AE9FFCDFFF9B822FECB09D10546" box="[1145,1228,307,331]" captionStart="Figures 3541" captionStartId="16.[151,224,1890,1912]" captionTargetBox="[184,1354,294,1780]" captionTargetId="figure-104@16.[151,1398,258,1871]" captionTargetPageId="16" captionText="Figures 3541. Aedeagus in lateral view of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (35); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (36); S. grandis, holotype, Jawa Barat, Sukabumi: Cidahu (37); S. cuspilobatus, holotype, Jawa Barat, Sukabumi: Sirnarasa (38); S. diversilobatus, holotype, Jawa Barat, Sukabumi: Djampang Kulon (39); S. pulcher, holotype, Jawa Barat, Sukabumi: Cikaniki (40); S. heronifer, holotype, Banten, Lebak: Majasari (41)." figureDoi="http://doi.org/10.5281/zenodo.10724258" httpUri="https://zenodo.org/record/10724258/files/figure.png" pageId="13" pageNumber="36">Figs 35</figureCitation>
,
<figureCitation id="13256AE9FFCDFFF9B88CFECB083E0546" box="[1239,1315,307,331]" captionStart="Figures 7479" captionStartId="24.[151,224,1866,1888]" captionTargetId="figure-134@24.[151,1398,277,1847]" captionTargetPageId="24" captionText="Figures 7479. Apex of aedeagus in lateral (74, 77), ventral (75, 78), dorsal view (76, 79) of Scopaeus spiraliflagellatus, holotype, Sulawesi Tengah: Lake Poso (7476); S. anuliflagellatus, holotype, Jawa Barat, Sukabumi: Cidahu (7779). Abbreviations: al apical lobes, al(d) dextral apical lobe, al(s) sinistral apical lobe, dl dorsal lobe, dl(d) dextral end of dorsal lobe, dl(s) sinistral end of dorsal lobe, f flagellum, ll lateral lobes, mf median foramen, tr transverse ridge of median foramen, vld dextral portion of ventral lobe, vls sinistral portion of ventral lobe." figureDoi="http://doi.org/10.5281/zenodo.10880227" httpUri="https://zenodo.org/record/10880227/files/figure.png" pageId="13" pageNumber="36">74 76</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>