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<document id="36DB744CB3916695FC5C8DC57869ED2B" ID-DOI="10.5281/zenodo.214740" ID-GBIF-Dataset="4ef4df5b-682e-45e2-a0c1-c3c51ea1047d" ID-ISSN="1175-5326" ID-Zenodo-Dep="214740" ID-ZooBank="ABBE345C-EE4B-46A2-8F4F-EE0719C5D7ED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460528846328" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2012" docId="03AE3D4F9120AF5E8EDF5B38DDEAFD1F" docLanguage="en" docName="zt03482p076.pdf" docOrigin="Zootaxa 3482" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Mastigus deustus Thunberg 1789" docType="treatment" docVersion="7" lastPageNumber="76" masterDocId="FF9745379124AF568E485827D865FF9B" masterDocTitle="New synonymy and redescription of Mastigus deustus (Thunberg) (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="76" masterPageNumber="68" pageNumber="72" updateTime="1698315462857" updateUser="plazi">
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<mods:title id="60B268E065D2335B14A89FA57397E62E">New synonymy and redescription of Mastigus deustus (Thunberg) (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="03AE3D4F9120AF5E8EDF5B38DDEAFD1F" ID-DOI="http://doi.org/10.5281/zenodo.6181691" ID-GBIF-Taxon="119526616" ID-Zenodo-Dep="6181691" LSID="urn:lsid:plazi:treatment:03AE3D4F9120AF5E8EDF5B38DDEAFD1F" httpUri="http://treatment.plazi.org/id/03AE3D4F9120AF5E8EDF5B38DDEAFD1F" lastPageId="8" lastPageNumber="76" pageId="4" pageNumber="72">
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<paragraph id="8BB88C599120AF528EDF5B38DA2CFCA2" blockId="4.[151,585,799,858]" box="[151,585,799,825]" pageId="4" pageNumber="72">
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<taxonomicName id="4C07F7DA9120AF528EDF5B38DA2CFCA2" ID-CoL="3YD7G" authority="Thunberg, 1789" authorityName="Thunberg" authorityYear="1789" box="[151,585,799,825]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9120AF528EDF5B38DA2CFCA2" bold="true" box="[151,585,799,825]" pageId="4" pageNumber="72">
<emphasis id="B973504B9120AF528EDF5B38D90DFCA2" bold="true" box="[151,360,799,825]" italics="true" pageId="4" pageNumber="72">Mastigus deustus</emphasis>
(
<bibRefCitation id="EF96F1A89120AF528F3F5B38DA24FCA2" author="Thunberg" box="[375,577,799,825]" pageId="4" pageNumber="72" refString="Thunberg, C. P. (1789) Dissertatio entomologica novas Insectorum species sistens. Pars 5. Upsaliae, Edmann. (1787), 85 - 106 pp." type="journal article" year="1789">Thunberg, 1789</bibRefCitation>
)
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<paragraph id="8BB88C599120AF528EDF5B65D945FCC1" blockId="4.[151,585,799,858]" box="[151,288,834,858]" pageId="4" pageNumber="72">
(
<figureCitation id="133C90DC9120AF528ED75B65D972FCC1" box="[159,279,834,858]" captionStart-0="FIGURES 1 7" captionStart-1="FIGURES 8 9" captionStart-2="FIGURES 10 13" captionStart-3="FIGURES 14 18" captionStartId-0="2.[151,264,1911,1934]" captionStartId-1="5.[151,264,1805,1828]" captionStartId-2="6.[151,264,1632,1655]" captionStartId-3="7.[151,264,1912,1935]" captionTargetBox-0="[188,1384,190,1895]" captionTargetBox-1="[174,1413,232,1762]" captionTargetBox-2="[157,1421,214,1601]" captionTargetBox-3="[158,1428,1044,1883]" captionTargetId-0="figure@2.[172,1416,185,1903]" captionTargetId-1="figure@5.[151,1436,193,1782]" captionTargetId-2="figure@6.[151,1436,193,1611]" captionTargetId-3="figure@7.[151,1436,1039,1896]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 1 7. Mastigus deustus (Thunberg). Lectotype female of M. deustus in dorsal (1) and lateral (2) views; microscope slide of antenna and maxillary palp of the lectotype female of M. deustus (3); labels of the lectotype female of M. deustus (4). Lectotype male of M. glabratus Klug in dorsal (5) and lateral (6) views; labels of the lectotype male of M. glabratus (7). Scale bars: 2 mm." captionText-1="FIGURES 8 9. Dorsal habitus of male (8) and female (9) of Mastigus deustus (Thunberg). Scale bar: 2 mm." captionText-2="FIGURES 10 13. Female of Mastigus deustus (Thunberg). Head in dorsal (10), lateral (11) and ventral (12) views; pterothorax and abdomen in ventral view (13). Scale bars: 10 12 — 0.5 mm, 13 — 1 mm." captionText-3="FIGURES 14 18. Mastigus deustus (Thunberg). Aedeagus in abparameral (14), right lateral (15), parameral (16) and left lateral (17) views, and spermatheca (18). Abbreviations: ag, accessory gland; cp, copulatory piece; ds, ductus spermathecae; f, flagellum; lp, long paramere; sp, short paramere; st, subapical tooth. Scale bars: 14 17 — 0.5 mm, 18 — 0.1 mm." httpUri-0="https://zenodo.org/record/214741/files/figure.png" httpUri-1="https://zenodo.org/record/214742/files/figure.png" httpUri-2="https://zenodo.org/record/214743/files/figure.png" httpUri-3="https://zenodo.org/record/214744/files/figure.png" pageId="4" pageNumber="72">Figs. 118</figureCitation>
)
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<paragraph id="8BB88C599120AF528EDF5BACDA7BFC3B" blockId="4.[151,1436,906,1051]" box="[151,542,906,928]" pageId="4" pageNumber="72">
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<taxonomicName id="4C07F7DA9120AF528EDF5BACDA7FFC3B" authority="Thunberg, 1789: 101" authorityName="Thunberg" authorityPageNumber="101" authorityYear="1789" box="[151,538,906,928]" class="Insecta" family="Anthicidae" genus="Notoxus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9120AF528EDF5BACD959FC3B" box="[151,316,906,928]" italics="true" pageId="4" pageNumber="72">Notoxus deustus</emphasis>
<treatmentCitation id="0AA6AA489120AF528F0A5BADDA7FFC3B" author="Thunberg" box="[322,538,906,928]" page="101" pageId="4" pageNumber="72" year="1789">
<bibRefCitation id="EF96F1A89120AF528F0A5BADD982FC3B" author="Thunberg" box="[322,487,906,928]" pageId="4" pageNumber="72" refString="Thunberg, C. P. (1789) Dissertatio entomologica novas Insectorum species sistens. Pars 5. Upsaliae, Edmann. (1787), 85 - 106 pp." type="journal article" year="1789">Thunberg, 1789</bibRefCitation>
: 101
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.
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</paragraph>
<paragraph id="8BB88C599120AF528EDF5B8DDA3DFC24" blockId="4.[151,1436,906,1051]" box="[151,600,937,959]" pageId="4" pageNumber="72">
<treatmentCitationGroup id="AB17AB779120AF528EDF5B8DDA3DFC24" box="[151,600,937,959]" pageId="4" pageNumber="72">
<taxonomicName id="4C07F7DA9120AF528EDF5B8DDA31FC24" ID-CoL="3YD87" authority="Peringuey, 1908: 298" authorityName="Peringuey" authorityPageNumber="298" authorityYear="1908" box="[151,596,937,959]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="promontorii">
<emphasis id="B973504B9120AF528EDF5B8DD916FC24" box="[151,371,938,959]" italics="true" pageId="4" pageNumber="72">Mastigus promontorii</emphasis>
<treatmentCitation id="0AA6AA489120AF528F325B8EDA31FC24" author="Peringuey" box="[378,596,937,959]" page="298" pageId="4" pageNumber="72" year="1908">
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: 298
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.
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</paragraph>
<paragraph id="8BB88C599120AF528EDF5BEFDB1CFC46" blockId="4.[151,1436,906,1051]" box="[151,889,968,990]" pageId="4" pageNumber="72">
<treatmentCitationGroup id="AB17AB779120AF528EDF5BEFDB1CFC46" box="[151,889,968,990]" pageId="4" pageNumber="72">
<taxonomicName id="4C07F7DA9120AF528EDF5BEFDA52FC45" authority="Thunberg, 1789" authorityName="Thunberg" authorityYear="1789" box="[151,567,968,990]" class="Insecta" family="Staphylinidae" genus="Australostigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9120AF528EDF5BEFD91EFC45" box="[151,379,968,990]" italics="true" pageId="4" pageNumber="72">Australostigus deustus</emphasis>
(
<bibRefCitation id="EF96F1A89120AF528FC05BEFDA55FC45" author="Thunberg" box="[392,560,968,990]" pageId="4" pageNumber="72" refString="Thunberg, C. P. (1789) Dissertatio entomologica novas Insectorum species sistens. Pars 5. Upsaliae, Edmann. (1787), 85 - 106 pp." type="journal article" year="1789">Thunberg, 1789</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="0AA6AA489120AF528C095BEFDA8AFC46" author="Leleup" box="[577,751,968,990]" page="47" pageId="4" pageNumber="72" year="1968">
<bibRefCitation id="EF96F1A89120AF528C095BEFDAAFFC45" author="Leleup" box="[577,714,968,990]" pageId="4" pageNumber="72" refString="Leleup, N. (1968) Revision des Mastigini de l'Afrique du Sud. Annales du Musee Royal de l'Afrique Centrale, Tervuren (series 8: Sciences Zoologiques) 166, 1 - 107." type="journal article" year="1968">Leleup, 1968</bibRefCitation>
: 47
</treatmentCitation>
, Figs. 3738.
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</paragraph>
<paragraph id="8BB88C599120AF528EDF5BCFD90CFB80" blockId="4.[151,1436,906,1051]" pageId="4" pageNumber="72">
<treatmentCitationGroup id="AB17AB779120AF528EDF5BCFD90CFB80" pageId="4" pageNumber="72">
<taxonomicName id="4C07F7DA9120AF528EDF5BCFDA4DFC67" ID-CoL="3YD7K" authority="Klug, 1824: 166" authorityName="Klug" authorityPageNumber="166" authorityYear="1824" box="[151,552,998,1021]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis id="B973504B9120AF528EDF5BCFD906FC66" box="[151,355,999,1021]" italics="true" pageId="4" pageNumber="72">Mastigus glabratus</emphasis>
<treatmentCitation id="0AA6AA489120AF528F395BC1DA4DFC67" author="Klug" box="[369,552,998,1020]" page="166" pageId="4" pageNumber="72" year="1824">
<bibRefCitation id="EF96F1A89120AF528F395BC1D98AFC67" author="Klug" box="[369,495,998,1020]" pageId="4" pageNumber="72" refString="Klug, J. C. F. (1824) Entomologische Monographieen. G. Reimer, Berlin. 242 pp." type="book" year="1824">Klug, 1824</bibRefCitation>
: 166
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</taxonomicName>
,
<emphasis id="B973504B9120AF528C735BC0DAC5FC67" bold="true" box="[571,672,999,1020]" pageId="4" pageNumber="72">
<taxonomicNameLabel id="A240ED309120AF528C735BC0DAC5FC67" box="[571,672,999,1020]" pageId="4" pageNumber="72">syn. nov.</taxonomicNameLabel>
</emphasis>
;
<treatmentCitation id="0AA6AA489120AF528CFE5BC1DBE7FC67" author="Leleup" box="[694,898,998,1020]" page="102" pageId="4" pageNumber="72" year="1968">
<bibRefCitation id="EF96F1A89120AF528CFE5BC1DB2DFC67" author="Leleup" box="[694,840,998,1020]" pageId="4" pageNumber="72" refString="Leleup, N. (1968) Revision des Mastigini de l'Afrique du Sud. Annales du Musee Royal de l'Afrique Centrale, Tervuren (series 8: Sciences Zoologiques) 166, 1 - 107." type="journal article" year="1968">Leleup, 1968</bibRefCitation>
: 102
</treatmentCitation>
;
<treatmentCitation id="0AA6AA489120AF528DDC5BC1DC98FC67" author="Bordoni" box="[916,1277,998,1020]" page="321" pageId="4" pageNumber="72" year="1973">
<bibRefCitation id="EF96F1A89120AF528DDC5BC1DCA1FC67" author="Bordoni" box="[916,1220,998,1020]" pageId="4" pageNumber="72" refString="Bordoni, A., Castellini, G. (1973) Sulle specie Paleartiche del genere Mastigus Latreille con osservazioni su due specie dell'Africa Australe (Coleoptera Scydmaenidae). Redia, 54, 295 - 323." type="journal article" year="1973">Bordoni &amp; Castellini, 1973</bibRefCitation>
: 321
</treatmentCitation>
,
<figureCitation id="133C90DC9120AF528B475BC1DD3AFC67" box="[1295,1375,998,1020]" captionStart="FIGURES 14 18" captionStartId="7.[151,264,1912,1935]" captionTargetBox="[158,1428,1044,1883]" captionTargetId="figure@7.[151,1436,1039,1896]" captionTargetPageId="7" captionText="FIGURES 14 18. Mastigus deustus (Thunberg). Aedeagus in abparameral (14), right lateral (15), parameral (16) and left lateral (17) views, and spermatheca (18). Abbreviations: ag, accessory gland; cp, copulatory piece; ds, ductus spermathecae; f, flagellum; lp, long paramere; sp, short paramere; st, subapical tooth. Scale bars: 14 17 — 0.5 mm, 18 — 0.1 mm." httpUri="https://zenodo.org/record/214744/files/figure.png" pageId="4" pageNumber="72">Fig. 15</figureCitation>
B (as
<taxonomicName id="4C07F7DA9120AF528E8F5C22D93CFB80" ID-CoL="8XJTK" box="[199,345,1029,1051]" class="Insecta" family="Staphylinidae" genus="Australostigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B973504B9120AF528E8F5C22D93CFB80" box="[199,345,1029,1051]" italics="true" pageId="4" pageNumber="72">Australostigus</emphasis>
</taxonomicName>
).
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</paragraph>
</subSubSection>
<subSubSection id="C31DDFD29120AF528EDF5C6FDA23F897" pageId="4" pageNumber="72" type="materials_examined">
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<typeStatus id="54BC32FB9120AF528EDF5C6FD8B7FBFB" box="[151,210,1096,1120]" pageId="4" pageNumber="72">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54BC32FB9120AF528F1E5C6FD9ACFBFB" box="[342,457,1096,1120]" pageId="4" pageNumber="72" type="lectotype">Lectotype</typeStatus>
, designated here: Ƥ: dry-mounted specimen (
<figureCitation id="133C90DC9120AF528DAF5C60DC33FBC4" box="[999,1110,1095,1119]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[188,1384,190,1895]" captionTargetId="figure@2.[172,1416,185,1903]" captionTargetPageId="2" captionText="FIGURES 1 7. Mastigus deustus (Thunberg). Lectotype female of M. deustus in dorsal (1) and lateral (2) views; microscope slide of antenna and maxillary palp of the lectotype female of M. deustus (3); labels of the lectotype female of M. deustus (4). Lectotype male of M. glabratus Klug in dorsal (5) and lateral (6) views; labels of the lectotype male of M. glabratus (7). Scale bars: 2 mm." httpUri="https://zenodo.org/record/214741/files/figure.png" pageId="4" pageNumber="72">Figs. 12</figureCitation>
) with parts of antenna and maxillary palp mounted in euparal on microscopic slide (
<figureCitation id="133C90DC9120AF528D085C4BDBECFB1F" box="[832,905,1132,1156]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[188,1384,190,1895]" captionTargetId="figure@2.[172,1416,185,1903]" captionTargetPageId="2" captionText="FIGURES 1 7. Mastigus deustus (Thunberg). Lectotype female of M. deustus in dorsal (1) and lateral (2) views; microscope slide of antenna and maxillary palp of the lectotype female of M. deustus (3); labels of the lectotype female of M. deustus (4). Lectotype male of M. glabratus Klug in dorsal (5) and lateral (6) views; labels of the lectotype male of M. glabratus (7). Scale bars: 2 mm." httpUri="https://zenodo.org/record/214741/files/figure.png" pageId="4" pageNumber="72">Fig. 3</figureCitation>
), labels under the pinned specimen (
<figureCitation id="133C90DC9120AF528B0B5C4BDDEBFB1F" box="[1347,1422,1132,1156]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[188,1384,190,1895]" captionTargetId="figure@2.[172,1416,185,1903]" captionTargetPageId="2" captionText="FIGURES 1 7. Mastigus deustus (Thunberg). Lectotype female of M. deustus in dorsal (1) and lateral (2) views; microscope slide of antenna and maxillary palp of the lectotype female of M. deustus (3); labels of the lectotype female of M. deustus (4). Lectotype male of M. glabratus Klug in dorsal (5) and lateral (6) views; labels of the lectotype male of M. glabratus (7). Scale bars: 2 mm." httpUri="https://zenodo.org/record/214741/files/figure.png" pageId="4" pageNumber="72">Fig. 4</figureCitation>
): &quot;Uppsala Univ. Zool.
<taxonomicName id="4C07F7DA9120AF528FDA5CB7D9A2FB3C" box="[402,455,1168,1191]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="72" phylum="Chordata" rank="genus">Mus</taxonomicName>
. / Thunbergsaml. nr. 6597 /
<taxonomicName id="4C07F7DA9120AF528D475CB6DBB6FB33" box="[783,979,1168,1192]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9120AF528D475CB6DBB6FB33" box="[783,979,1168,1192]" italics="true" pageId="4" pageNumber="72">Mastigus deustus</emphasis>
</taxonomicName>
/
<emphasis id="B973504B9120AF528DA55CB6DC18FB33" box="[1005,1149,1168,1192]" italics="true" pageId="4" pageNumber="72">
<taxonomicName id="4C07F7DA9120AF528DA55CB6DC44FB33" box="[1005,1057,1169,1192]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="72" phylum="Chordata" rank="genus">Mus</taxonomicName>
. Thunb.
</emphasis>
TYP&quot; [red, printed], &quot;N. Leleup / vid. 1964&quot; [white, handwritten], &quot;
<typeStatus id="54BC32FB9120AF528C355C93DB4CFB57" box="[637,809,1204,1228]" pageId="4" pageNumber="72" type="holotype">HOLOTYPUS</typeStatus>
&quot; [pink, printed], additional label was added during the present study: &quot;
<taxonomicName id="4C07F7DA9120AF528F185CFFD9B3FB6B" box="[336,470,1240,1264]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B973504B9120AF528F185CFFD9B3FB6B" box="[336,470,1240,1264]" italics="true" pageId="4" pageNumber="72">MASTIGUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528FBE5CFFDA2FFB6B" box="[502,586,1240,1264]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9120AF528FBE5CFFDA2FFB6B" box="[502,586,1240,1264]" italics="true" pageId="4" pageNumber="72">deustus</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EF96F1A89120AF528C3A5CF0DB49FB74" author="Thunberg" box="[626,812,1239,1263]" pageId="4" pageNumber="72" refString="Thunberg, C. P. (1789) Dissertatio entomologica novas Insectorum species sistens. Pars 5. Upsaliae, Edmann. (1787), 85 - 106 pp." type="journal article" year="1789">Thunberg, 1789</bibRefCitation>
) /
<typeStatus id="54BC32FB9120AF528D1D5CF0DC62FB74" box="[853,1031,1239,1263]" pageId="4" pageNumber="72" type="lectotype">LECTOTYPUS</typeStatus>
/ des. P. Jałoszyński, '12&quot; [white, printed]; labels on the microscope slide (
<figureCitation id="133C90DC9120AF528C215CDBDAD5FA8F" box="[617,688,1276,1300]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[188,1384,190,1895]" captionTargetId="figure@2.[172,1416,185,1903]" captionTargetPageId="2" captionText="FIGURES 1 7. Mastigus deustus (Thunberg). Lectotype female of M. deustus in dorsal (1) and lateral (2) views; microscope slide of antenna and maxillary palp of the lectotype female of M. deustus (3); labels of the lectotype female of M. deustus (4). Lectotype male of M. glabratus Klug in dorsal (5) and lateral (6) views; labels of the lectotype male of M. glabratus (7). Scale bars: 2 mm." httpUri="https://zenodo.org/record/214741/files/figure.png" pageId="4" pageNumber="72">Fig. 3</figureCitation>
): &quot;Zool.
<taxonomicName id="4C07F7DA9120AF528D505CDADB28FA8F" box="[792,845,1277,1300]" class="Mammalia" family="Muridae" genus="Mus" kingdom="Animalia" order="Rodentia" pageId="4" pageNumber="72" phylum="Chordata" rank="genus">Mus</taxonomicName>
. / Uppsala /
<collectionCode id="ED16149C9120AF528D975CDBDC77FA8F" LSID="urn:lsid:biocol.org:col:15542" box="[991,1042,1276,1300]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15542" name="Uppsala University, Museum of Evolution, Botany Section (Fytoteket)" pageId="4" pageNumber="72">UPS</collectionCode>
<collectionCode id="ED16149C9120AF528A535CDBDC31FA8F" box="[1051,1108,1276,1300]" pageId="4" pageNumber="72">ZTY</collectionCode>
99Z78&quot; [white, handwritten], &quot;antennae + / premiers arti- / cles palpes max&quot; [yellowish, handwritten], &quot;
<taxonomicName id="4C07F7DA9120AF528DAF5D38DCE9FAAC" box="[999,1164,1311,1335]" class="Insecta" family="Staphylinidae" genus="Australostigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">Australostigus</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528AEE5D38DD03FAAC" authority="Thunb." authorityName="Thunb." box="[1190,1382,1311,1335]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">deustus (Thunb.)</taxonomicName>
/ N. Leleup det., 1966&quot; [white, handwritten and printed], &quot;
<typeStatus id="54BC32FB9120AF528CA65D63DBFEFAC7" box="[750,923,1348,1372]" pageId="4" pageNumber="72" type="holotype">HOLOTYPUS</typeStatus>
&quot; [pink, printed] (
<collectionCode id="ED16149C9120AF528A125D63DCD6FAC7" box="[1114,1203,1348,1372]" httpUri="http://grbio.org/cool/80z8-1h9s" name="Uppsala University, Zoological Museum" pageId="4" pageNumber="72">UUZM</collectionCode>
).
</paragraph>
<paragraph id="8BB88C599120AF528E8F5D40DA23F897" blockId="4.[151,1436,1095,2020]" pageId="4" pageNumber="72">
<emphasis id="B973504B9120AF528E8F5D40D9DFFA1B" bold="true" box="[199,442,1383,1408]" pageId="4" pageNumber="72">Additional material.</emphasis>
<typeStatus id="54BC32FB9120AF528F8A5D4FDA57FA1B" box="[450,562,1384,1408]" pageId="4" pageNumber="72" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C07F7DA9120AF528C105D4EDB11FAE4" authority="Klug" authorityName="Klug" box="[600,884,1383,1408]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis id="B973504B9120AF528C105D4EDB56FA1B" box="[600,819,1384,1408]" italics="true" pageId="4" pageNumber="72">Mastigus glabratus</emphasis>
Klug
</taxonomicName>
, designated here (
<figureCitation id="133C90DC9120AF528A085D40DCCBFAE4" box="[1088,1198,1383,1407]" captionStart="FIGURES 1 7" captionStartId="2.[151,264,1911,1934]" captionTargetBox="[188,1384,190,1895]" captionTargetId="figure@2.[172,1416,185,1903]" captionTargetPageId="2" captionText="FIGURES 1 7. Mastigus deustus (Thunberg). Lectotype female of M. deustus in dorsal (1) and lateral (2) views; microscope slide of antenna and maxillary palp of the lectotype female of M. deustus (3); labels of the lectotype female of M. deustus (4). Lectotype male of M. glabratus Klug in dorsal (5) and lateral (6) views; labels of the lectotype male of M. glabratus (7). Scale bars: 2 mm." httpUri="https://zenodo.org/record/214741/files/figure.png" pageId="4" pageNumber="72">Figs. 57</figureCitation>
): 3: &quot;Pr.b.sp. Krebs&quot; [i.e., Promontorium bonae spei, the Cape of Good Hope; blue-greenish, handwritten], &quot;N. Leleup det., 1974 /
<taxonomicName id="4C07F7DA9120AF528EDF5D88D95CFA5C" box="[151,313,1455,1479]" class="Insecta" family="Staphylinidae" genus="Australostigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">Australostigus</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528F185D88DA74FA5C" authority="Thunb." authorityName="Thunb." box="[336,529,1455,1479]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">deustus (Thunb.)</taxonomicName>
&quot; [white, printed and handrwitten], &quot;5111&quot; [yellowish, printed], &quot;
<taxonomicName id="4C07F7DA9120AF528AAF5D88DDE9FA5C" box="[1255,1420,1455,1479]" class="Insecta" family="Staphylinidae" genus="Australostigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">Australostigus</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528EDF5DF3D930FA77" authority="Klug" authorityName="Klug" box="[151,341,1492,1516]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="glabratus">
glabratus
<collectionCode id="ED16149C9120AF528F4F5DF3D930FA77" box="[263,341,1492,1516]" pageId="4" pageNumber="72">KLUG</collectionCode>
</taxonomicName>
/
<typeStatus id="54BC32FB9120AF528F385DF3D980FA77" box="[368,485,1492,1516]" pageId="4" pageNumber="72" type="holotype">Holotypus</typeStatus>
3 / Bordoni &amp; Castellini / det. 1973&quot; [white, handwritten and printed], additional label was added during the present study: &quot;
<taxonomicName id="4C07F7DA9120AF528CEA5DDFDB42F98B" box="[674,807,1528,1552]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis id="B973504B9120AF528CEA5DDFDB42F98B" box="[674,807,1528,1552]" italics="true" pageId="4" pageNumber="72">MASTIGUS</emphasis>
</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528D015DDFDBD0F98B" box="[841,949,1528,1552]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis id="B973504B9120AF528D015DDFDBD0F98B" box="[841,949,1528,1552]" italics="true" pageId="4" pageNumber="72">glabratus</emphasis>
</taxonomicName>
/
<bibRefCitation id="EF96F1A89120AF528D9E5DD0DC3FF994" author="Klug" box="[982,1114,1527,1551]" pageId="4" pageNumber="72" refString="Klug, J. C. F. (1824) Entomologische Monographieen. G. Reimer, Berlin. 242 pp." type="book" year="1824">Klug, 1824</bibRefCitation>
/
<typeStatus id="54BC32FB9120AF528A335DD0DD48F994" box="[1147,1325,1527,1551]" pageId="4" pageNumber="72" type="lectotype">LECTOTYPUS</typeStatus>
/ des. P. Jałoszyński, '12&quot; [white, printed] (
<collectionCode id="ED16149C9120AF528C635E3BDAFFF9AF" box="[555,666,1564,1588]" pageId="4" pageNumber="72">MNHUB</collectionCode>
); 3, &quot;Swedish
<collectingCountry id="F310CCC99120AF528D195E3BDB83F9AF" box="[849,998,1564,1588]" name="South Africa" pageId="4" pageNumber="72">South Africa</collectingCountry>
/ Expedition /
<date id="FFB9AA999120AF528AE95E3BDD7EF9AF" box="[1185,1307,1564,1588]" pageId="4" pageNumber="72" value="1950" valueMax="1951">1950-1951</date>
/ Brinck - Rudebeck&quot; [white, printed], &quot;S. Afr. Cape Prov. / Langebaan /
<date id="FFB9AA999120AF528D365E18DBB9F9CC" box="[894,988,1599,1623]" pageId="4" pageNumber="72" value="1950-10-31">31.X.50</date>
. No. 20&quot; [white, printed], &quot;
<taxonomicName id="4C07F7DA9120AF528B575E18DDEDF9CC" box="[1311,1416,1599,1623]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="76" phylum="Arthropoda" rank="genus">Mastigus</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528EDF5E43D978F9E7" box="[151,285,1636,1660]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="promontorii">promontorii</taxonomicName>
/ Perring. / det. H. Franz&quot; [white, printed and handwritten], &quot;
<taxonomicName id="4C07F7DA9120AF528DAD5E43DCEFF9E7" box="[997,1162,1636,1660]" class="Insecta" family="Staphylinidae" genus="Australostigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="genus">Australostigus</taxonomicName>
/
<taxonomicName id="4C07F7DA9120AF528AED5E43DD03F9E7" authority="Thunb." authorityName="Thunb." box="[1189,1382,1636,1660]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="72" phylum="Arthropoda" rank="species" species="deustus">deustus (Thunb.)</taxonomicName>
/ N. Leleup det., 1966&quot; [white, printed and handwritten], &quot;ALLOTYPUS&quot; [pink, printed] (
<collectionCode id="ED16149C9120AF528AD15EA0DC8AF904" LSID="urn:lsid:biocol.org:col:34994" box="[1177,1263,1671,1695]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" name="Lund University" pageId="4" pageNumber="72">MZLU</collectionCode>
); 2 ƤƤ, same collecting data (
<collectionCode id="ED16149C9120AF528F075E8BD9C1F95F" LSID="urn:lsid:biocol.org:col:34994" box="[335,420,1708,1732]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34994" name="Lund University" pageId="4" pageNumber="72">MZLU</collectionCode>
); 1 3, 4 ƤƤ, &quot;S.Afr., S.W. Cape / Langebaanweg /
<geoCoordinate id="EE33EA9E9120AF528A495E8BDC3FF95F" box="[1025,1114,1708,1732]" direction="south" orientation="latitude" pageId="4" pageNumber="72" precision="555" value="-29.57">29.57 S</geoCoordinate>
-
<geoCoordinate id="EE33EA9E9120AF528A3E5E8BDCB4F95F" box="[1142,1233,1708,1732]" direction="east" orientation="longitude" pageId="4" pageNumber="72" precision="555" value="18.08">18.08 E</geoCoordinate>
&quot; [white, printed], &quot;
<date id="FFB9AA999120AF528ED75EE8D970F97C" box="[159,277,1743,1767]" pageId="4" pageNumber="72" value="1983-08-24">24.8.1983</date>
; E-Y: 1970 / groundtraps, 71 days / leg. Endrödy, Penrith&quot; [white, printed], &quot;singled around / traps at setting&quot; [white, printed] (
<collectionCode id="ED16149C9120AF528FF95ED3DA66F897" box="[433,515,1780,1804]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="4" pageNumber="72">TMSA</collectionCode>
, cPJ).
</paragraph>
</subSubSection>
<subSubSection id="C31DDFD29120AF528E8F5F30DBF3F8EC" pageId="4" pageNumber="72" type="diagnosis">
<paragraph id="8BB88C599120AF528E8F5F30DBF3F8EC" blockId="4.[151,1436,1095,2020]" pageId="4" pageNumber="72">
<emphasis id="B973504B9120AF528E8F5F30D924F8AB" bold="true" box="[199,321,1815,1840]" pageId="4" pageNumber="72">Diagnosis.</emphasis>
This species can be distinguished from all other congeners on the basis of brown elytra with darkened apices, nearly black head and pronotum, often with reddish hue; fine transverse microsculpture of elytra; and moderately elongate aedeagus with rudimentary short paramere.
</paragraph>
</subSubSection>
<subSubSection id="C31DDFD29120AF5E8E8F5FA4D9E2FE37" lastPageId="8" lastPageNumber="76" pageId="4" pageNumber="72" type="description">
<paragraph id="8BB88C599120AF528E8F5FA4DCC2F87F" blockId="4.[151,1436,1095,2020]" pageId="4" pageNumber="72">
<emphasis id="B973504B9120AF528E8F5FA4D911F807" bold="true" box="[199,372,1923,1948]" pageId="4" pageNumber="72">Redescription.</emphasis>
Body of male (
<figureCitation id="133C90DC9120AF528C625FA3DA0BF807" box="[554,622,1924,1948]" captionStart="FIGURES 8 9" captionStartId="5.[151,264,1805,1828]" captionTargetBox="[174,1413,232,1762]" captionTargetId="figure@5.[151,1436,193,1782]" captionTargetPageId="5" captionText="FIGURES 8 9. Dorsal habitus of male (8) and female (9) of Mastigus deustus (Thunberg). Scale bar: 2 mm." httpUri="https://zenodo.org/record/214742/files/figure.png" pageId="4" pageNumber="72">Fig. 8</figureCitation>
) strongly elongate and strongly convex, BL
<quantity id="4CFF21BC9120AF528A275FA3DD71F807" box="[1135,1300,1924,1948]" metricMagnitude="-3" metricUnit="m" metricValue="5.665" metricValueMax="5.75" metricValueMin="5.58" pageId="4" pageNumber="72" unit="mm" value="5.665" valueMax="5.75" valueMin="5.58">5.585.75 mm</quantity>
(mean
<quantity id="4CFF21BC9120AF528B235FA3D8A2F85B" metricMagnitude="-3" metricUnit="m" metricValue="5.64" pageId="4" pageNumber="72" unit="mm" value="5.64">5.64 mm</quantity>
); dorsum distinctly bicolorous, head and appendages nearly black, pronotum dark brown, nearly black but with indistinct reddish hue, elytra in color of raw umber, with darkened apices; vestiture silverish.
</paragraph>
<caption id="DF78DCD19121AF538EDF5F2ADC82F8B8" box="[151,1255,1805,1828]" httpUri="https://zenodo.org/record/214742/files/figure.png" pageId="5" pageNumber="73" targetBox="[174,1413,232,1762]" targetPageId="5">
<paragraph id="8BB88C599121AF538EDF5F2ADC82F8B8" blockId="5.[151,1255,1805,1828]" box="[151,1255,1805,1828]" pageId="5" pageNumber="73">
<emphasis id="B973504B9121AF538EDF5F2AD95FF8B9" bold="true" box="[151,314,1805,1828]" pageId="5" pageNumber="73">FIGURES 89.</emphasis>
Dorsal habitus of male (8) and female (9) of
<taxonomicName id="4C07F7DA9121AF538D4D5F29DC54F8B8" authority="Thunberg" authorityName="Thunberg" box="[773,1073,1805,1827]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="73" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9121AF538D4D5F29DBD0F8B8" box="[773,949,1805,1827]" italics="true" pageId="5" pageNumber="73">Mastigus deustus</emphasis>
(Thunberg)
</taxonomicName>
. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph id="8BB88C599121AF518E8F5F74DB66FE37" blockId="5.[151,1436,1875,2009]" lastBlockId="7.[151,1436,151,1004]" lastPageId="7" lastPageNumber="75" pageId="5" pageNumber="73">
Head (
<figureCitation id="133C90DC9121AF538F5C5F74D9F8F8F0" box="[276,413,1875,1899]" captionStart="FIGURES 10 13" captionStartId="6.[151,264,1632,1655]" captionTargetBox="[157,1421,214,1601]" captionTargetId="figure@6.[151,1436,193,1611]" captionTargetPageId="6" captionText="FIGURES 10 13. Female of Mastigus deustus (Thunberg). Head in dorsal (10), lateral (11) and ventral (12) views; pterothorax and abdomen in ventral view (13). Scale bars: 10 12 — 0.5 mm, 13 — 1 mm." httpUri="https://zenodo.org/record/214743/files/figure.png" pageId="5" pageNumber="73">Figs. 1012</figureCitation>
) broadest at eyes, HL
<quantity id="4CFF21BC9121AF538CEE5F74DB2EF8F0" box="[678,843,1875,1899]" metricMagnitude="-4" metricUnit="m" metricValue="8.25" metricValueMax="8.5" metricValueMin="8.0" pageId="5" pageNumber="73" unit="mm" value="0.825" valueMax="0.85" valueMin="0.8">0.800.85 mm</quantity>
(mean
<quantity id="4CFF21BC9121AF538DEC5F74DC6BF8F0" box="[932,1038,1875,1899]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="5" pageNumber="73" unit="mm" value="0.83">0.83 mm</quantity>
), HW
<quantity id="4CFF21BC9121AF538A145F74DD64F8F0" box="[1116,1281,1875,1899]" metricMagnitude="-4" metricUnit="m" metricValue="9.55" metricValueMax="9.8" metricValueMin="9.3" pageId="5" pageNumber="73" unit="mm" value="0.955" valueMax="0.98" valueMin="0.93">0.930.98 mm</quantity>
(mean 0.94); occipital constriction distinctly broader than 0.5
<emphasis id="B973504B9121AF538CF85F5EDADAF80B" box="[688,703,1913,1936]" italics="true" pageId="5" pageNumber="73">×</emphasis>
HW; vertex more than twice as broad as long, distinctly concave in middle, with posterior margin bearing narrow median notch extended anterad by indistinct and shallow median longitudinal groove not reaching frons, lateral parts of vertex convex; frons transverse, impressed between weakly convex supraantennal tubercles, anterior part located between antennal insertions convex and bulging anteriorly to form rounded median projection; vertex and frons uniformly covered with fine and moderately shallow punctures separated by spaces subequal to puncture diameters, without microsculpture; vestiture extremely short but dense, composed of recumbent setae; clypeus nearly four times as broad as long, with rounded anterior margin and glossy surface and transverse row of several very long, thick and erect macrosetae directed anterad; labrum 2.5
<emphasis id="B973504B9122AF508B6F5F74DD53F8F1" box="[1319,1334,1875,1898]" italics="true" pageId="6" pageNumber="74">×</emphasis>
as broad as long, with distinct median emargination and glossy surface bearing row of several very long and thick macrosetae directed anteriorly and curved ventrally. Eyes large and moderately coarsely faceted, each in lateral view oval and elongate, distinctly shorter than temple. Antennae slender, shorter than body, AeL
<quantity id="4CFF21BC9122AF508A915F99DD5AF84D" box="[1241,1343,1982,2006]" metricMagnitude="-3" metricUnit="m" metricValue="3.34" pageId="6" pageNumber="74" unit="mm" value="3.34">3.34 mm</quantity>
, lengths of antennomeres (from I to XI):
<quantity id="4CFF21BC9123AF518C4058B0DA0AFF34" box="[520,623,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="7" pageNumber="75" unit="mm" value="0.95">0.95 mm</quantity>
,
<quantity id="4CFF21BC9123AF518C3358B0DA87FF34" box="[635,738,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="7" pageNumber="75" unit="mm" value="0.43">0.43 mm</quantity>
,
<quantity id="4CFF21BC9123AF518CA758B0DB33FF34" box="[751,854,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="75" unit="mm" value="0.23">0.23 mm</quantity>
,
<quantity id="4CFF21BC9123AF518D2A58B0DBACFF34" box="[866,969,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="75" unit="mm" value="0.22">0.22 mm</quantity>
,
<quantity id="4CFF21BC9123AF518D9E58B0DC58FF34" box="[982,1085,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="75" unit="mm" value="0.23">0.23 mm</quantity>
,
<quantity id="4CFF21BC9123AF518A0158B0DCD5FF34" box="[1097,1200,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="7" pageNumber="75" unit="mm" value="0.21">0.21 mm</quantity>
,
<quantity id="4CFF21BC9123AF518AF558B0DD41FF34" box="[1213,1316,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="75" unit="mm" value="0.2">0.20 mm</quantity>
,
<quantity id="4CFF21BC9123AF518B7858B0DDF2FF34" box="[1328,1431,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="75" unit="mm" value="0.2">0.20 mm</quantity>
,
<quantity id="4CFF21BC9123AF518EDF589BD964FF4F" box="[151,257,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="75" unit="mm" value="0.23">0.23 mm</quantity>
,
<quantity id="4CFF21BC9123AF518F59589BD919FF4F" box="[273,380,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="75" unit="mm" value="0.2">0.20 mm</quantity>
,
<quantity id="4CFF21BC9123AF518FC4589BD99DFF4F" box="[396,504,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="75" unit="mm" value="0.25">0.25 mm</quantity>
; scape strongly, pedicel slightly thickened, all flagellomeres slender and hardly noticeably thickening towards antennal apex; scape about 6
<emphasis id="B973504B9123AF518D1C58C6DB06FF63" box="[852,867,225,248]" italics="true" pageId="7" pageNumber="75">×</emphasis>
as long as broad, distinctly broadening distally, broadest near apex, with two longitudinal ventral setal rows each composed of 45 moderately long and thick, strongly erect bristles and surface covered uniformly with moderately sparse, short suberect setae; pedicel 5
<emphasis id="B973504B9123AF518B24590EDD1EFEDB" box="[1388,1403,297,320]" italics="true" pageId="7" pageNumber="75">×</emphasis>
as long as broad, covered with moderately sparse, short suberect setae and with 12 strongly erect but very thin and long ventral setae; all flagellomeres much narrower than scape but comparable in width to pedicel, each covered with short and nearly recumbent setae, without bristles.
</paragraph>
<caption id="DF78DCD19122AF508EDF5E47DBCBF90E" httpUri="https://zenodo.org/record/214743/files/figure.png" pageId="6" pageNumber="74" targetBox="[157,1421,214,1601]" targetPageId="6">
<paragraph id="8BB88C599122AF508EDF5E47DBCBF90E" blockId="6.[151,1436,1632,1686]" pageId="6" pageNumber="74">
<emphasis id="B973504B9122AF508EDF5E47D936F9ED" bold="true" box="[151,339,1632,1655]" pageId="6" pageNumber="74">FIGURES 1013.</emphasis>
Female of
<taxonomicName id="4C07F7DA9122AF508F8C5E45DA8BF9EC" authority="Thunberg" authorityName="Thunberg" box="[452,750,1633,1655]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="74" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9122AF508F8C5E45DA17F9EC" box="[452,626,1633,1655]" italics="true" pageId="6" pageNumber="74">Mastigus deustus</emphasis>
(Thunberg)
</taxonomicName>
. Head in dorsal (10), lateral (11) and ventral (12) views; pterothorax and abdomen in ventral view (13). Scale bars: 1012 — 0.5 mm, 13 — 1 mm.
</paragraph>
</caption>
<paragraph id="8BB88C599123AF518E8F599FDBC3FDC4" blockId="7.[151,1436,151,1004]" pageId="7" pageNumber="75">
Pronotum inversely pear-shaped in dorsal view, strongly elongate and broadest near anterior third, PL
<quantity id="4CFF21BC9123AF518B155990D898FE6F" metricMagnitude="-3" metricUnit="m" metricValue="1.455" metricValueMax="1.48" metricValueMin="1.43" pageId="7" pageNumber="75" unit="mm" value="1.455" valueMax="1.48" valueMin="1.43">1.43 1.48 mm</quantity>
(mean
<quantity id="4CFF21BC9123AF518F1C59FBD9DBFE6F" box="[340,446,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="7" pageNumber="75" unit="mm" value="1.45">1.45 mm</quantity>
), PW
<quantity id="4CFF21BC9123AF518C4D59FBDACFFE6F" box="[517,682,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" metricValueMax="1.08" metricValueMin="0.98" pageId="7" pageNumber="75" unit="mm" value="1.03" valueMax="1.08" valueMin="0.98">0.981.08 mm</quantity>
(mean
<quantity id="4CFF21BC9123AF518D4959FBDB0EFE6F" box="[769,875,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="7" pageNumber="75" unit="mm" value="1.03">1.03 mm</quantity>
); anterior margin weakly arcuate; lateral margins strongly rounded in anterior half, broadly constricted posterior to middle and weakly rounded in posterior fourth; posterior margin strongly arcuate; pronotal disc without median longitudinal carina or impunctate line, uniformly covered with punctures and setae similar to those on vertex and frons.
</paragraph>
<paragraph id="8BB88C599123AF518E8F5A4BDA14FC8F" blockId="7.[151,1436,151,1004]" pageId="7" pageNumber="75">
Elytra oval and more convex than pronotum, broadest near middle, EL
<quantity id="4CFF21BC9123AF518DBF5A4BDCFEFD1F" box="[1015,1179,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="7" pageNumber="75" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">3.303.50 mm</quantity>
(mean
<quantity id="4CFF21BC9123AF518ABA5A4BDD39FD1F" box="[1266,1372,620,644]" metricMagnitude="-3" metricUnit="m" metricValue="3.37" pageId="7" pageNumber="75" unit="mm" value="3.37">3.37 mm</quantity>
), EW 1.832.00 mm (mean
<quantity id="4CFF21BC9123AF518FC65AA8D993FD3C" box="[398,502,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="7" pageNumber="75" unit="mm" value="1.9">1.90 mm</quantity>
), EI 1.751.81; basal part of elytra without impressions, apices separately rounded; each elytron with several indistinct and irregular longitudinal rows of small and shallow punctures and slightly raised suture; surface of elytra covered with fine, indistinct microsculpture composed of transverse rows of flat granules; setae fine, dense and recumbent.
</paragraph>
<paragraph id="8BB88C599123AF518E8F5B07DAF4FCC7" blockId="7.[151,1436,151,1004]" pageId="7" pageNumber="75">Legs long and slender; protrochanters unmodified; protibiae distinctly curved, meso- and metatibiae nearly straight; all tibiae with two long apical spurs.</paragraph>
<paragraph id="8BB88C599123AF518E8F5B4FDC38FCE4" blockId="7.[151,1436,151,1004]" box="[199,1117,871,895]" pageId="7" pageNumber="75">
Venter (
<figureCitation id="133C90DC9123AF518F575B40D90AFCE4" box="[287,367,871,895]" captionStart="FIGURES 10 13" captionStartId="6.[151,264,1632,1655]" captionTargetBox="[157,1421,214,1601]" captionTargetId="figure@6.[151,1436,193,1611]" captionTargetPageId="6" captionText="FIGURES 10 13. Female of Mastigus deustus (Thunberg). Head in dorsal (10), lateral (11) and ventral (12) views; pterothorax and abdomen in ventral view (13). Scale bars: 10 12 — 0.5 mm, 13 — 1 mm." httpUri="https://zenodo.org/record/214743/files/figure.png" pageId="7" pageNumber="75">Fig. 13</figureCitation>
) largely brown, with median part of abdominal sternites darkened.
</paragraph>
<paragraph id="8BB88C599123AF518E8F5BABD92DFC77" blockId="7.[151,1436,151,1004]" pageId="7" pageNumber="75">
Aedeagus (
<figureCitation id="133C90DC9123AF518F0D5BABD9A9FC3F" box="[325,460,908,932]" captionStart="FIGURES 14 18" captionStartId="7.[151,264,1912,1935]" captionTargetBox="[158,1428,1044,1883]" captionTargetId="figure@7.[151,1436,1039,1896]" captionTargetPageId="7" captionText="FIGURES 14 18. Mastigus deustus (Thunberg). Aedeagus in abparameral (14), right lateral (15), parameral (16) and left lateral (17) views, and spermatheca (18). Abbreviations: ag, accessory gland; cp, copulatory piece; ds, ductus spermathecae; f, flagellum; lp, long paramere; sp, short paramere; st, subapical tooth. Scale bars: 14 17 — 0.5 mm, 18 — 0.1 mm." httpUri="https://zenodo.org/record/214744/files/figure.png" pageId="7" pageNumber="75">Figs. 1417</figureCitation>
) moderately slender, AeL
<quantity id="4CFF21BC9123AF518CB15BABDB05FC3F" box="[761,864,908,932]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="7" pageNumber="75" unit="mm" value="1.13">1.13 mm</quantity>
, in abparameral view with oval median lobe, short paramere rudimentary, subtriangular, long paramere stout, subtriangular; copulatory piece recurved, with single subapical tooth.
</paragraph>
<caption id="DF78DCD19123AF518EDF5F5FDD7DF850" httpUri="https://zenodo.org/record/214744/files/figure.png" pageId="7" pageNumber="75" targetBox="[158,1428,1044,1883]" targetPageId="7">
<paragraph id="8BB88C599123AF518EDF5F5FDD7DF850" blockId="7.[151,1436,1912,1995]" pageId="7" pageNumber="75">
<emphasis id="B973504B9123AF518EDF5F5FD932F816" bold="true" box="[151,343,1912,1935]" pageId="7" pageNumber="75">FIGURES 1418.</emphasis>
<taxonomicName id="4C07F7DA9123AF518F285F5EDAF6F815" authority="Thunberg" authorityName="Thunberg" box="[352,659,1912,1934]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="75" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B9123AF518F285F5EDA76F815" box="[352,531,1912,1934]" italics="true" pageId="7" pageNumber="75">Mastigus deustus</emphasis>
(Thunberg)
</taxonomicName>
. Aedeagus in abparameral (14), right lateral (15), parameral (16) and left lateral (17) views, and spermatheca (18). Abbreviations: ag, accessory gland; cp, copulatory piece; ds, ductus spermathecae; f, flagellum; lp, long paramere; sp, short paramere; st, subapical tooth. Scale bars: 1417 — 0.5 mm, 18 — 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB88C59912CAF5E8E8F58BFD992FEFF" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="76">
<emphasis id="B973504B912CAF5E8E8F58BFD940FF2B" bold="true" box="[199,293,152,176]" pageId="8" pageNumber="76">Female.</emphasis>
Similar to males but distinctly larger and with indistinct pair of circumsutural impressions in anterior third of elytra; protibiae nearly straight; BL
<quantity id="4CFF21BC912CAF5E8CC2589BDB4BFF4F" box="[650,814,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="6.1899999999999995" metricValueMax="6.8" metricValueMin="5.58" pageId="8" pageNumber="76" unit="mm" value="6.1899999999999995" valueMax="6.8" valueMin="5.58">5.586.80 mm</quantity>
(mean
<quantity id="4CFF21BC912CAF5E8DCB589BDB89FF4F" box="[899,1004,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="8" pageNumber="76" unit="mm" value="6.4">6.40 mm</quantity>
); HL
<quantity id="4CFF21BC912CAF5E8A66589BDCB7FF4F" box="[1070,1234,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="8" pageNumber="76" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.800.90 mm</quantity>
(mean
<quantity id="4CFF21BC912CAF5E8B6F589BDDF5FF4F" box="[1319,1424,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="8" pageNumber="76" unit="mm" value="0.87">0.87 mm</quantity>
), HW
<quantity id="4CFF21BC912CAF5E8E8558F8D914FF6C" box="[205,369,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.015" metricValueMax="1.05" metricValueMin="0.98" pageId="8" pageNumber="76" unit="mm" value="1.015" valueMax="1.05" valueMin="0.98">0.981.05 mm</quantity>
(mean
<quantity id="4CFF21BC912CAF5E8F8F58F8DA55FF6C" box="[455,560,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="8" pageNumber="76" unit="mm" value="1.03">1.03 mm</quantity>
), AnL
<quantity id="4CFF21BC912CAF5E8C3758F8DA83FF6C" box="[639,742,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="8" pageNumber="76" unit="mm" value="3.8">3.80 mm</quantity>
, lengths of antennomeres (from I to XI):
<quantity id="4CFF21BC912CAF5E8AF558F8DD41FF6C" box="[1213,1316,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="76" unit="mm" value="1.1">1.10 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8B7858F8DDF2FF6C" box="[1328,1431,223,247]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="76" unit="mm" value="0.5">0.50 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8EDF5923D965FE87" box="[151,256,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="76" unit="mm" value="0.28">0.28 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8F475923D912FE87" box="[271,375,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="76" unit="mm" value="0.25">0.25 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8FCE5923D98AFE87" box="[390,495,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="76" unit="mm" value="0.23">0.23 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8FB65923DA02FE87" box="[510,615,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="76" unit="mm" value="0.21">0.21 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8C3D5923DABBFE87" box="[629,734,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="76" unit="mm" value="0.22">0.22 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8CA55923DB33FE87" box="[749,854,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="76" unit="mm" value="0.21">0.21 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8D2C5923DBA8FE87" box="[868,973,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="76" unit="mm" value="0.25">0.25 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8D945923DC20FE87" box="[988,1093,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="76" unit="mm" value="0.25">0.25 mm</quantity>
,
<quantity id="4CFF21BC912CAF5E8A1B5923DCDBFE87" box="[1107,1214,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="76" unit="mm" value="0.3">0.30 mm</quantity>
; PL
<quantity id="4CFF21BC912CAF5E8ABE5923DDF9FE87" box="[1270,1436,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="8" pageNumber="76" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.501.70 mm</quantity>
(mean
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), PW
<quantity id="4CFF21BC912CAF5E8FD25900DA5AFEA4" box="[410,575,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="1.165" metricValueMax="1.2" metricValueMin="1.13" pageId="8" pageNumber="76" unit="mm" value="1.165" valueMax="1.2" valueMin="1.13">1.131.20 mm</quantity>
(mean
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); EL
<quantity id="4CFF21BC912CAF5E8D0C5900DB8CFEA4" box="[836,1001,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="3.89" metricValueMax="4.2" metricValueMin="3.58" pageId="8" pageNumber="76" unit="mm" value="3.89" valueMax="4.2" valueMin="3.58">3.584.20 mm</quantity>
(mean
<quantity id="4CFF21BC912CAF5E8A0A5900DCC9FEA4" box="[1090,1196,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="3.93" pageId="8" pageNumber="76" unit="mm" value="3.93">3.93 mm</quantity>
), EW
<quantity id="4CFF21BC912CAF5E8ABF5900DDF9FEA4" box="[1271,1436,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="2.2649999999999997" metricValueMax="2.38" metricValueMin="2.15" pageId="8" pageNumber="76" unit="mm" value="2.2649999999999997" valueMax="2.38" valueMin="2.15">2.152.38 mm</quantity>
(mean
<quantity id="4CFF21BC912CAF5E8EAC596BD929FEFF" box="[228,332,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="8" pageNumber="76" unit="mm" value="2.33">2.33 mm</quantity>
), EI 1.531.87.
</paragraph>
<paragraph id="8BB88C59912CAF5E8E8F5948D9E2FE37" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="76">
Spermatheca (
<figureCitation id="133C90DC912CAF5E8F215948D9D8FE1C" box="[361,445,367,391]" captionStart="FIGURES 14 18" captionStartId="7.[151,264,1912,1935]" captionTargetBox="[158,1428,1044,1883]" captionTargetId="figure@7.[151,1436,1039,1896]" captionTargetPageId="7" captionText="FIGURES 14 18. Mastigus deustus (Thunberg). Aedeagus in abparameral (14), right lateral (15), parameral (16) and left lateral (17) views, and spermatheca (18). Abbreviations: ag, accessory gland; cp, copulatory piece; ds, ductus spermathecae; f, flagellum; lp, long paramere; sp, short paramere; st, subapical tooth. Scale bars: 14 17 — 0.5 mm, 18 — 0.1 mm." httpUri="https://zenodo.org/record/214744/files/figure.png" pageId="8" pageNumber="76">Fig. 18</figureCitation>
) subglobose, diameter
<quantity id="4CFF21BC912CAF5E8C825948DB56FE1C" box="[714,819,367,391]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="76" unit="mm" value="0.14">0.14 mm</quantity>
, thick-walled, with small accessory gland and simple ductus spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="C31DDFD2912CAF5E8E8F5990DAD4FE6F" pageId="8" pageNumber="76" type="distribution">
<paragraph id="8BB88C59912CAF5E8E8F5990DAD4FE6F" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="76">
<emphasis id="B973504B912CAF5E8E8F5990D905FE4B" bold="true" box="[199,352,439,464]" pageId="8" pageNumber="76">Distribution.</emphasis>
<collectingCountry id="F310CCC9912CAF5E8F2F5990DAE0FE54" box="[359,645,439,463]" name="South Africa" pageId="8" pageNumber="76">Republic of South Africa</collectingCountry>
, Western Cape Province, along Atlantic coast from near Cape Town in the south to the Langebaan Lagoon to the north.
</paragraph>
</subSubSection>
<subSubSection id="C31DDFD2912CAF5E8E8F5A27DDEAFD1F" pageId="8" pageNumber="76" type="discussion">
<paragraph id="8BB88C59912CAF5E8E8F5A27DDEAFD1F" blockId="8.[151,1436,151,644]" pageId="8" pageNumber="76">
<emphasis id="B973504B912CAF5E8E8F5A27D95EFD83" bold="true" box="[199,315,512,536]" pageId="8" pageNumber="76">Remarks.</emphasis>
The aedeagus of
<taxonomicName id="4C07F7DA912CAF5E8C495A26DA1FFD83" box="[513,634,512,536]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B912CAF5E8C495A26DA1FFD83" box="[513,634,512,536]" italics="true" pageId="8" pageNumber="76">M. deustus</emphasis>
</taxonomicName>
is most similar to those of
<taxonomicName id="4C07F7DA912CAF5E8DE45A26DCE2FD8C" authority="Lhoste" authorityName="Lhoste" box="[940,1159,511,536]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="andrae">
<emphasis id="B973504B912CAF5E8DE45A26DC45FD83" box="[940,1056,512,536]" italics="true" pageId="8" pageNumber="76">M. andrae</emphasis>
(Lhoste)
</taxonomicName>
(
<bibRefCitation id="EF96F1A8912CAF5E8ADE59D8D8D1FDA7" author="Leleup" pageId="8" pageNumber="76" refString="Leleup, N. (1968) Revision des Mastigini de l'Afrique du Sud. Annales du Musee Royal de l'Afrique Centrale, Tervuren (series 8: Sciences Zoologiques) 166, 1 - 107." type="journal article" year="1968">Leleup 1968; Figs. 42 43</bibRefCitation>
) and
<taxonomicName id="4C07F7DA912CAF5E8EBD5A02DA79FDA7" authority="Fabricius" authorityName="Fabricius" box="[245,540,548,572]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="B973504B912CAF5E8EBD5A02D9FCFDA7" box="[245,409,549,572]" italics="true" pageId="8" pageNumber="76">M. spinicornis</emphasis>
(Fabricius)
</taxonomicName>
(
<bibRefCitation id="EF96F1A8912CAF5E8C645A03DB33FDA7" author="Leleup" box="[556,854,548,572]" pageId="8" pageNumber="76" refString="Leleup, N. (1968) Revision des Mastigini de l'Afrique du Sud. Annales du Musee Royal de l'Afrique Centrale, Tervuren (series 8: Sciences Zoologiques) 166, 1 - 107." type="journal article" year="1968">Leleup 1968; Figs. 4647</bibRefCitation>
). However, the aedeagus of
<taxonomicName id="4C07F7DA912CAF5E8AD25A02DD6AFDA7" box="[1178,1295,548,572]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="andrae">
<emphasis id="B973504B912CAF5E8AD25A02DD6AFDA7" box="[1178,1295,548,572]" italics="true" pageId="8" pageNumber="76">M. andrae</emphasis>
</taxonomicName>
is distinctly stouter and its parameres have clearly different proportions than in
<taxonomicName id="4C07F7DA912CAF5E8D855A6EDC2BFDFB" box="[973,1102,584,608]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="deustus">
<emphasis id="B973504B912CAF5E8D855A6EDC2BFDFB" box="[973,1102,584,608]" italics="true" pageId="8" pageNumber="76">M. deustus</emphasis>
</taxonomicName>
. The short paramere of
<taxonomicName id="4C07F7DA912CAF5E8B355A6ED976FD1F" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="spinicornis">
<emphasis id="B973504B912CAF5E8B355A6ED976FD1F" italics="true" pageId="8" pageNumber="76">M. spinicornis</emphasis>
</taxonomicName>
is well-developed and cannot be mistaken compared to the rudimentary short paramere of
<emphasis id="B973504B912CAF5E8B465A4ADDEAFD1F" box="[1294,1423,620,644]" italics="true" pageId="8" pageNumber="76">
<taxonomicName id="4C07F7DA912CAF5E8B465A4ADDEFFD1F" box="[1294,1418,620,644]" class="Insecta" family="Staphylinidae" genus="Mastigus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="76" phylum="Arthropoda" rank="species" species="deustus">M. deustus</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>