treatments-xml/data/03/A4/82/03A4827A9D13FFBE3E975C36FAFCFD5B.xml
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<document id="10E2BFFC69F4FF2ACC66D8472A3B9393" ID-DOI="10.11646/zootaxa.5016.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222372" ID-ZooBank="C129938C-C9AA-4874-8DEF-CA8191D02A5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629381479438" checkinUser="plazi" docAuthor="Huerta, Heron &amp; Ibáñez-Bernal, Sergio" docDate="2021" docId="03A4827A9D13FFBE3E975C36FAFCFD5B" docLanguage="en" docName="zootaxa.5016.3.8.pdf" docOrigin="Zootaxa 5016 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dixella atitla Huerta &amp; Ibanez-Bernal 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="436" masterDocId="FF9DFA029D11FFB83E005846FFEDFFF3" masterDocTitle="A new species of Dixella, the first Mexican record of Dixella shannoni, and a summary of dixid records from Mexico (Diptera: Dixidae)" masterLastPageNumber="440" masterPageNumber="430" pageNumber="432" updateTime="1699070574525" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FF051AF11FF3F8DEA363FE39247F8AED">A new species of Dixella, the first Mexican record of Dixella shannoni, and a summary of dixid records from Mexico (Diptera: Dixidae)</mods:title>
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<mods:name id="EED50FD0E40737B40C740040991CA217" type="personal">
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<mods:namePart id="2FB24DB3F1F9522CDD94FA307AD21867">Huerta, Heron</mods:namePart>
<mods:affiliation id="6E3997D98619808963246C0BF98AE66A">Laboratorio de Entomología, Instituto de Diagnóstico y Referencia Epidemiológicos (InDRE), Francisco P. Miranda # 177, Unidad Lomas de Plateros, DT. Álvaro Obregón, 01480, Ciudad de México, México.</mods:affiliation>
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</mods:role>
<mods:namePart id="15BC6DD5F538E6C052B0B7727D4DFEFF">Ibáñez-Bernal, Sergio</mods:namePart>
<mods:affiliation id="901F3DD23EB5A08868F91C528D8F9E4B">Instituto de Ecología, A. C., Red Ambiente y Sustentabilidad, Antigua Carretera a Coatepec No. 351, El Haya, Xalapa, 91073, Veracruz, México.</mods:affiliation>
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<mods:title id="43712B75D9F8CEAFE73538EAFAFC16D3">Zootaxa</mods:title>
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<mods:date id="BEEB49B6C1391BCE1D23DE0D4DE1B46D">2021</mods:date>
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<mods:number id="F1D0C6294AB0EE8368DA701A5B394F82">2021-08-06</mods:number>
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<mods:number id="91D363419F57AC9427E9BB0CAA5B2599">5016</mods:number>
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<mods:identifier id="2E1E6A23F3F7756B3D430862A68086B5" type="DOI">10.11646/zootaxa.5016.3.8</mods:identifier>
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<treatment id="03A4827A9D13FFBE3E975C36FAFCFD5B" LSID="urn:lsid:plazi:treatment:03A4827A9D13FFBE3E975C36FAFCFD5B" httpUri="http://treatment.plazi.org/id/03A4827A9D13FFBE3E975C36FAFCFD5B" lastPageId="6" lastPageNumber="436" pageId="2" pageNumber="432">
<subSubSection id="C31760E79D13FFBA3E975C36FD31FB78" box="[151,732,1136,1163]" pageId="2" pageNumber="432" type="nomenclature">
<paragraph id="8BB2336C9D13FFBA3E975C36FD31FB78" blockId="2.[151,732,1136,1199]" box="[151,732,1136,1163]" pageId="2" pageNumber="432">
<heading id="D0FA84009D13FFBA3E975C36FD31FB78" bold="true" box="[151,732,1136,1163]" fontSize="11" level="1" pageId="2" pageNumber="432" reason="1">
<emphasis id="B979EF7E9D13FFBA3E975C36FD31FB78" bold="true" box="[151,732,1136,1163]" pageId="2" pageNumber="432">
<taxonomicName id="4C0D48EF9D13FFBA3E975C36FD9CFB78" ID-CoL="9HWQC" authority="Huerta &amp; Ibanez-Bernal" authorityName="Huerta &amp; Ibanez-Bernal" authorityYear="2021" box="[151,625,1136,1163]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D13FFBA3E975C36FED8FB78" bold="true" box="[151,309,1136,1163]" italics="true" pageId="2" pageNumber="432">Dixella atitla</emphasis>
Huerta &amp; Ibáñez-Bernal
</taxonomicName>
,
<taxonomicNameLabel id="A24A52059D13FFBA3C7E5C37FD31FB78" box="[638,732,1137,1163]" pageId="2" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31760E79D13FFBA3E975CD3FEFFFB5C" box="[151,274,1173,1199]" pageId="2" pageNumber="432" type="description">
<paragraph id="8BB2336C9D13FFBA3E975CD3FEFFFB5C" blockId="2.[151,732,1136,1199]" box="[151,274,1173,1199]" pageId="2" pageNumber="432">
(
<figureCitation id="13362FE99D13FFBA3E9F5CD3FEE4FB5C" box="[159,265,1173,1199]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="3.[151,250,926,950]" captionStartId-1="3.[151,250,1608,1632]" captionStartId-2="4.[151,250,1803,1827]" captionStartId-3="5.[151,250,1943,1967]" captionStartId-4="6.[151,250,1969,1993]" captionStartId-5="7.[151,250,1151,1175]" captionStartId-6="7.[151,250,2015,2039]" captionTargetBox-0="[162,1430,191,899]" captionTargetBox-1="[157,1427,1013,1581]" captionTargetBox-2="[151,1421,466,1764]" captionTargetBox-3="[297,1282,185,1886]" captionTargetBox-4="[171,1412,726,1927]" captionTargetBox-5="[228,1357,190,1119]" captionTargetBox-6="[228,1358,1295,1987]" captionTargetId-0="figure-25@3.[151,1436,181,903]" captionTargetId-1="figure-69@3.[151,1436,1011,1584]" captionTargetId-2="figure-139@4.[151,1436,453,1779]" captionTargetId-3="figure-25@5.[297,1290,181,1919]" captionTargetId-4="figure-258@6.[159,1429,704,1945]" captionTargetId-5="figure-25@7.[225,1363,181,1127]" captionTargetId-6="figure-83@7.[223,1363,1271,1991]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="7" captionText-0="FIGURE 1. Dixella atitla sp. nov. AB. Head, dorsal view. C. Antenna, lateral view. A. Male; BC. Female. Abbreviations: flgm I to V= first to fifth flagellomere; pe= pedicel; scp= scape. Scale = 0.1 mm." captionText-1="FIGURE 2. Dixella atitla sp. nov. AB. Thorax, male. A. Dorsal view (pinned specimen); B. Lateral view (mounted in slide). C. Hind tibia, distal portion. D. Fifth tarsomere and claw of foreleg (dark arrow points to the seta). Scale: AB= 0.5 mm; CD= 0.1 mm." captionText-2="FIGURE 3. Wing, dorsal view. A. Male, Dixella atitla sp. nov. B. Female, Dixella atitla sp. nov. C. Male, Dixella shannoni. Abbreviations: C= costal vein; CuA= anterior branch of cubital vein; CuP= posterior branch of cubital vein; M 1 = first branch of media; M 2 = second branch of media; M 3 + 4 = third and fourth branch of media;; m-cu= medial-cubital crossvein; m-m= medial crossvein; Rs= radial sector; R 1 = anterior branch of radius; R 2 = upper branch of second branch of radius; R 3 = lower branch of second branch of radius; R 2+3 = second branch of radius; R = third branch of radius; r-m= radial-medial crossvein; Sc = sub4+5 costal vein. Scale = 0.5 mm." captionText-3="FIGURE 4. Dixella atitla sp. nov. AB. Male genitalia. C. Female genitalia. A. Tergite 9, dorsal view; B. Sternite 9, ventral view. C. Ventral view. Scale = 0.1 mm." captionText-4="FIGURE 5. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Tergum 10, ventral view. C. Gonostyle, top view. D. Dorsal view. Scale: AD= 0.1 mm" captionText-5="FIGURE 6. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Gonostyle, dorsal view, C, E. Lateral view. D. Dorsal view. Abbreviations: ae ap= aedeagal apodeme; ae com= aedeagal complex; a lob= anterior lobe of gonocoxite; goncx= gonocoxite; gonst= gonostyle. Scale: AD= 0.1 mm; E= 0.064 mm." captionText-6="FIGURE 7. Distribution of Dixella atitla sp. nov. in state of Veracruz and type locality." figureDoi-0="http://doi.org/10.5281/zenodo.5222374" figureDoi-1="http://doi.org/10.5281/zenodo.5222376" figureDoi-2="http://doi.org/10.5281/zenodo.5222378" figureDoi-3="http://doi.org/10.5281/zenodo.5222380" figureDoi-4="http://doi.org/10.5281/zenodo.5222382" figureDoi-5="http://doi.org/10.5281/zenodo.5222384" figureDoi-6="http://doi.org/10.5281/zenodo.5222386" httpUri-0="https://zenodo.org/record/5222374/files/figure.png" httpUri-1="https://zenodo.org/record/5222376/files/figure.png" httpUri-2="https://zenodo.org/record/5222378/files/figure.png" httpUri-3="https://zenodo.org/record/5222380/files/figure.png" httpUri-4="https://zenodo.org/record/5222382/files/figure.png" httpUri-5="https://zenodo.org/record/5222384/files/figure.png" httpUri-6="https://zenodo.org/record/5222386/files/figure.png" pageId="2" pageNumber="432">Figs. 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31760E79D13FFBA3E975C9BFDCEFACC" pageId="2" pageNumber="432" type="materials_examined">
<paragraph id="8BB2336C9D13FFBA3E975C9BFDCEFACC" blockId="2.[151,1437,1245,2028]" pageId="2" pageNumber="432">
<emphasis id="B979EF7E9D13FFBA3E975C9BFEB8FB04" bold="true" box="[151,341,1245,1271]" pageId="2" pageNumber="432">Type specimens</emphasis>
.
<materialsCitation id="3B6539319D13FFBA3F665C9BFC26FAE8" collectionCode="CAIM" collectorName="Ibanez-Bernal, S." country="Mexico" county="Huatusco" location="February" municipality="Laguna de Santo Domingo" pageId="2" pageNumber="432" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54B68DCE9D13FFBA3F665C9BFE17FB04" box="[358,506,1245,1271]" pageId="2" pageNumber="432" type="holotype">HOLOTYPE</typeStatus>
Male.
<collectingCountry id="F31A73FC9D13FFBA3C4F5C9BFD2DFB04" box="[591,704,1245,1271]" name="Mexico" pageId="2" pageNumber="432">MEXICO</collectingCountry>
,
<collectingRegion id="49C9FD8E9D13FFBA3CCE5C98FCD8FB0B" box="[718,821,1246,1272]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="2" pageNumber="432">Veracruz</collectingRegion>
,
<collectingCounty id="62D34BE09D13FFBA3D435C98FC5DFB0B" box="[835,944,1246,1272]" pageId="2" pageNumber="432">Huatusco</collectingCounty>
,
<collectingMunicipality id="6BD6A9169D13FFBA3DBD5C98FB1AFB04" box="[957,1271,1245,1272]" pageId="2" pageNumber="432">Laguna de Santo Domingo</collectingMunicipality>
, 22,
<location id="8ED265B79D13FFBA3B325C9BFA75FB04" LSID="urn:lsid:plazi:treatment:03A4827A9D13FFBE3E975C36FAFCFD5B:8ED265B79D13FFBA3B325C9BFA75FB04" box="[1330,1432,1245,1271]" country="Mexico" county="Huatusco" municipality="Laguna de Santo Domingo" name="February" pageId="2" pageNumber="432" stateProvince="Veracruz de Ignacio de la Llave">February</location>
, 1992; leg.
<collectorName id="26F856BA9D13FFBA3F155D47FE36FAE8" box="[277,475,1281,1307]" pageId="2" pageNumber="432">Ibáñez-Bernal, S.</collectorName>
(mounted on microscope slide, in
<collectionCode id="ED1CABA99D13FFBA3D755D47FC2EFAE8" box="[885,963,1281,1307]" country="Mexico" httpUri="http://grbio.org/cool/z08b-033r" name="Collection of Aquatic Important Microorganisms" pageId="2" pageNumber="432">CAIM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B6539319D13FFBA3DD95D44FA8DFAE8" box="[985,1376,1281,1308]" collectionCode="CAIM" collectorName="Ibanez-Bernal, S." country="Mexico" county="Huatusco" location="February" municipality="Laguna de Santo Domingo" pageId="2" pageNumber="432" specimenCount="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<typeStatus id="54B68DCE9D13FFBA3DD95D44FBA7FAEF" box="[985,1098,1282,1308]" pageId="2" pageNumber="432" type="paratype">Paratypes</typeStatus>
: Same data as holotype
</materialsCitation>
,
<materialsCitation id="3B6539319D13FFBA3B6F5D47FF02FACC" collectionCode="IEXA" country="Mexico" pageId="2" pageNumber="432" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D0BF8E59D13FFBA3B6F5D47FA71FAEF" box="[1391,1436,1281,1308]" pageId="2" pageNumber="432" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED1CABA99D13FFBA3E9F5D63FF0AFACC" box="[159,231,1317,1343]" pageId="2" pageNumber="432">IEXA</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B6539319D13FFBA3EFB5D63FE67FACC" box="[251,394,1317,1344]" collectionCode="CAIM" country="Mexico" pageId="2" pageNumber="432" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D0BF8E59D13FFBA3EFB5D63FEC8FAB3" box="[251,293,1317,1344]" pageId="2" pageNumber="432" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1CABA99D13FFBA3F345D63FE6EFACC" box="[308,387,1317,1343]" country="Mexico" httpUri="http://grbio.org/cool/z08b-033r" name="Collection of Aquatic Important Microorganisms" pageId="2" pageNumber="432">CAIM</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B6539319D13FFBA3F955D63FDCDFACC" box="[405,544,1317,1344]" collectionCode="IEXA" country="Mexico" pageId="2" pageNumber="432" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="paratype">
<specimenCount id="9D0BF8E59D13FFBA3F955D63FE52FAB3" box="[405,447,1317,1344]" pageId="2" pageNumber="432" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED1CABA99D13FFBA3FCE5D63FDFAFACC" box="[462,535,1317,1343]" pageId="2" pageNumber="432">IEXA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31760E79D13FFBA3EC75D0FFB55FA5F" pageId="2" pageNumber="432" type="diagnosis">
<paragraph id="8BB2336C9D13FFBA3EC75D0FFB55FA5F" blockId="2.[151,1437,1245,2028]" pageId="2" pageNumber="432">
<emphasis id="B979EF7E9D13FFBA3EC75D0FFEAFFA90" bold="true" box="[199,322,1353,1379]" pageId="2" pageNumber="432">Diagnosis.</emphasis>
<taxonomicName id="4C0D48EF9D13FFBA3F4A5D0FFE0CFA90" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[330,481,1353,1379]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="432" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D13FFBA3F4A5D0FFE0CFA90" box="[330,481,1353,1379]" italics="true" pageId="2" pageNumber="432">Dixella atitla</emphasis>
</taxonomicName>
<emphasis id="B979EF7E9D13FFBA3FEA5D0FFDABFA90" bold="true" box="[490,582,1353,1379]" pageId="2" pageNumber="432">
<taxonomicNameLabel id="A24A52059D13FFBA3FEA5D0FFDABFA90" box="[490,582,1353,1379]" pageId="2" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is unique among species of the Neotropical region with apical lobe of the gonocoxite hammer-like (
<figureCitation id="13362FE99D13FFBA3FBA5D2BFD8CFA74" box="[442,609,1389,1416]" captionStart="FIGURE 5" captionStartId="6.[151,250,1969,1993]" captionTargetBox="[171,1412,726,1927]" captionTargetId="figure-258@6.[159,1429,704,1945]" captionTargetPageId="6" captionText="FIGURE 5. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Tergum 10, ventral view. C. Gonostyle, top view. D. Dorsal view. Scale: AD= 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.5222382" httpUri="https://zenodo.org/record/5222382/files/figure.png" pageId="2" pageNumber="432">Figs. 5A, CD</figureCitation>
) and gonostylus wider at base and dorsoventrally flattened, ending hookshaped, and at middle with a pair of short internal processes posteriorly directed (
<figureCitation id="13362FE99D13FFBA3A275DD7FB40FA5F" box="[1063,1197,1425,1452]" captionStart="FIGURE 6" captionStartId="7.[151,250,1151,1175]" captionTargetBox="[228,1357,190,1119]" captionTargetId="figure-25@7.[225,1363,181,1127]" captionTargetPageId="7" captionText="FIGURE 6. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Gonostyle, dorsal view, C, E. Lateral view. D. Dorsal view. Abbreviations: ae ap= aedeagal apodeme; ae com= aedeagal complex; a lob= anterior lobe of gonocoxite; goncx= gonocoxite; gonst= gonostyle. Scale: AD= 0.1 mm; E= 0.064 mm." figureDoi="http://doi.org/10.5281/zenodo.5222384" httpUri="https://zenodo.org/record/5222384/files/figure.png" pageId="2" pageNumber="432">Figs. 6AE</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31760E79D13FFBC3EC75DF3FB15FE5E" lastPageId="4" lastPageNumber="434" pageId="2" pageNumber="432" type="description">
<paragraph id="8BB2336C9D13FFBA3EC75DF3FC0EF9CF" blockId="2.[151,1437,1245,2028]" pageId="2" pageNumber="432">
<emphasis id="B979EF7E9D13FFBA3EC75DF3FEB4FA3C" bold="true" box="[199,345,1461,1487]" pageId="2" pageNumber="432">Description.</emphasis>
<emphasis id="B979EF7E9D13FFBA3F615DF3FE76FA3C" box="[353,411,1461,1487]" italics="true" pageId="2" pageNumber="432">Male</emphasis>
.
<emphasis id="B979EF7E9D13FFBA3FA95DF3FE04FA3C" bold="true" box="[425,489,1461,1487]" pageId="2" pageNumber="432">Head</emphasis>
(
<figureCitation id="13362FE99D13FFBA3FFA5DF3FDBCFA3C" box="[506,593,1461,1487]" captionStart="FIGURE 1" captionStartId="3.[151,250,926,950]" captionTargetBox="[162,1430,191,899]" captionTargetId="figure-25@3.[151,1436,181,903]" captionTargetPageId="3" captionText="FIGURE 1. Dixella atitla sp. nov. AB. Head, dorsal view. C. Antenna, lateral view. A. Male; BC. Female. Abbreviations: flgm I to V= first to fifth flagellomere; pe= pedicel; scp= scape. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222374" httpUri="https://zenodo.org/record/5222374/files/figure.png" pageId="2" pageNumber="432">Fig. 1A</figureCitation>
) slightly wider than high in frontal view, dark brown; clypeus longer than width, pale yellow, with about 9 setae at disk. Antenna with scape, pedicel and flagellomeres dark brown, length
<quantity id="4CF59E899D13FFBA3B795D9FFF2BF9EB" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="432" unit="mm" value="2.1">2.1 mm</quantity>
; all flagellomeres cylindrical, the first flagellomere length/ wide= 12.1, two times longer than second flagellomere; palpus brown, length
<quantity id="4CF59E899D13FFBA3FE65E67FDA0F9CF" box="[486,589,1569,1596]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="432" unit="mm" value="0.4">0.40 mm</quantity>
; palpomere ratios: 1.0: 3.0: 3.0: 3.5.
</paragraph>
<paragraph id="8BB2336C9D13FFBA3EC75E03FA94F938" blockId="2.[151,1437,1245,2028]" pageId="2" pageNumber="432">
<emphasis id="B979EF7E9D13FFBA3EC75E03FECDF9AC" bold="true" box="[199,288,1605,1631]" pageId="2" pageNumber="432">Thorax</emphasis>
(
<figureCitation id="13362FE99D13FFBA3F2F5E03FE5EF993" box="[303,435,1605,1632]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[157,1427,1013,1581]" captionTargetId="figure-69@3.[151,1436,1011,1584]" captionTargetPageId="3" captionText="FIGURE 2. Dixella atitla sp. nov. AB. Thorax, male. A. Dorsal view (pinned specimen); B. Lateral view (mounted in slide). C. Hind tibia, distal portion. D. Fifth tarsomere and claw of foreleg (dark arrow points to the seta). Scale: AB= 0.5 mm; CD= 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222376" httpUri="https://zenodo.org/record/5222376/files/figure.png" pageId="2" pageNumber="432">Figs. 2A, B</figureCitation>
). Scutum with wide dark brown medial vitta extending from anterior margin to middle of scutum, anteriorly narrowly continuing laterally to supra-alar area and lateral dark brown vitta. Scutellum (with six setae) and mediotergite dark brown. Pleura with pronotum, propleuron, anepisternum, katepisternum, anepimeron, and metanepisternum dark brown, except the distal part of anepimeron which is yellowish. Katepisternum bare.
</paragraph>
<paragraph id="8BB2336C9D13FFBA3EC75E93FE0CF8A8" blockId="2.[151,1437,1245,2028]" pageId="2" pageNumber="432">
<emphasis id="B979EF7E9D13FFBA3EC75E93FEE5F91C" bold="true" box="[199,264,1749,1775]" pageId="2" pageNumber="432">Wing</emphasis>
(
<figureCitation id="13362FE99D13FFBA3F175E93FE9CF91C" box="[279,369,1749,1775]" captionStart="FIGURE 3" captionStartId="4.[151,250,1803,1827]" captionTargetBox="[151,1421,466,1764]" captionTargetId="figure-139@4.[151,1436,453,1779]" captionTargetPageId="4" captionText="FIGURE 3. Wing, dorsal view. A. Male, Dixella atitla sp. nov. B. Female, Dixella atitla sp. nov. C. Male, Dixella shannoni. Abbreviations: C= costal vein; CuA= anterior branch of cubital vein; CuP= posterior branch of cubital vein; M 1 = first branch of media; M 2 = second branch of media; M 3 + 4 = third and fourth branch of media;; m-cu= medial-cubital crossvein; m-m= medial crossvein; Rs= radial sector; R 1 = anterior branch of radius; R 2 = upper branch of second branch of radius; R 3 = lower branch of second branch of radius; R 2+3 = second branch of radius; R = third branch of radius; r-m= radial-medial crossvein; Sc = sub4+5 costal vein. Scale = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222378" httpUri="https://zenodo.org/record/5222378/files/figure.png" pageId="2" pageNumber="432">Fig. 3A</figureCitation>
). Length,
<quantity id="4CF59E899D13FFBA3FE25E93FDD7F903" box="[482,570,1749,1776]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="432" unit="mm" value="2.8">2.8 mm</quantity>
, width, 1.0 mm; membrane pale without dark spot over r-m; veins brown, setose except r-m and m-m; Sc elongate, extending more than 0.33 the length of the wing, ending in C, slightly before at level of the origin of the Rs; R
<subScript id="178931299D13FFBA3C0B5F69FDC9F8CE" attach="left" box="[523,548,1839,1853]" fontSize="6" pageId="2" pageNumber="432">2+3</subScript>
, arched, originating distal to r-m; stem of R
<subScript id="178931299D13FFBA3A155F69FBC3F8CE" attach="left" box="[1045,1070,1839,1853]" fontSize="6" pageId="2" pageNumber="432">2+3</subScript>
0.25 length of R
<subScript id="178931299D13FFBA3AEA5F69FB1FF8CE" attach="left" box="[1258,1266,1839,1853]" fontSize="6" pageId="2" pageNumber="432">3</subScript>
. Halter brown, 0.34 the length of fore femur.
</paragraph>
<paragraph id="8BB2336C9D13FFBA3EC75F23FD86F818" blockId="2.[151,1437,1245,2028]" pageId="2" pageNumber="432">
<emphasis id="B979EF7E9D13FFBA3EC75F23FF12F88C" bold="true" box="[199,255,1893,1919]" pageId="2" pageNumber="432">Legs</emphasis>
. Coxae and trochanters pale yellow, except the base of fore coxa which is dark brown; femora yellowish with brown apices; tibiae pale brown with brown apices, more distinctive in hind tibia; hind tibia with a narrow longitudinal ridge proximad of comb near dorsal margin, apparently consisting of fused spicules (
<figureCitation id="13362FE99D13FFBA3AD35FEBFAC6F834" box="[1235,1323,1965,1991]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[157,1427,1013,1581]" captionTargetId="figure-69@3.[151,1436,1011,1584]" captionTargetPageId="3" captionText="FIGURE 2. Dixella atitla sp. nov. AB. Thorax, male. A. Dorsal view (pinned specimen); B. Lateral view (mounted in slide). C. Hind tibia, distal portion. D. Fifth tarsomere and claw of foreleg (dark arrow points to the seta). Scale: AB= 0.5 mm; CD= 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222376" httpUri="https://zenodo.org/record/5222376/files/figure.png" pageId="2" pageNumber="432">Fig. 2C</figureCitation>
); claws of all legs with external basal seta (
<figureCitation id="13362FE99D13FFBA3C045F97FDB3F818" box="[516,606,2001,2027]" captionStart="FIGURE 2" captionStartId="3.[151,250,1608,1632]" captionTargetBox="[157,1427,1013,1581]" captionTargetId="figure-69@3.[151,1436,1011,1584]" captionTargetPageId="3" captionText="FIGURE 2. Dixella atitla sp. nov. AB. Thorax, male. A. Dorsal view (pinned specimen); B. Lateral view (mounted in slide). C. Hind tibia, distal portion. D. Fifth tarsomere and claw of foreleg (dark arrow points to the seta). Scale: AB= 0.5 mm; CD= 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222376" httpUri="https://zenodo.org/record/5222376/files/figure.png" pageId="2" pageNumber="432">Fig. 2D</figureCitation>
).
</paragraph>
<caption id="DF7263E49D12FFBB3E975BD8FC27FC29" ID-DOI="http://doi.org/10.5281/zenodo.5222374" ID-Zenodo-Dep="5222374" httpUri="https://zenodo.org/record/5222374/files/figure.png" pageId="3" pageNumber="433" startId="3.[151,250,926,950]" targetBox="[162,1430,191,899]" targetPageId="3">
<paragraph id="8BB2336C9D12FFBB3E975BD8FC27FC29" blockId="3.[151,1437,926,987]" pageId="3" pageNumber="433">
<emphasis id="B979EF7E9D12FFBB3E975BD8FEF8FC45" bold="true" box="[151,277,926,950]" pageId="3" pageNumber="433">FIGURE 1.</emphasis>
<taxonomicName id="4C0D48EF9D12FFBB3F1C5BD9FE48FC44" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[284,421,927,951]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D12FFBB3F1C5BD9FE48FC44" box="[284,421,927,951]" italics="true" pageId="3" pageNumber="433">Dixella atitla</emphasis>
</taxonomicName>
<emphasis id="B979EF7E9D12FFBB3FAC5BD9FE13FC44" bold="true" box="[428,510,927,951]" pageId="3" pageNumber="433">
<taxonomicNameLabel id="A24A52059D12FFBB3FAC5BD9FE13FC44" box="[428,510,927,951]" pageId="3" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB. Head, dorsal view. C. Antenna, lateral view. A. Male; BC. Female. Abbreviations: flgm I to V= first to fifth flagellomere; pe= pedicel; scp= scape. Scale = 0.1 mm.
</paragraph>
</caption>
<caption id="DF7263E49D12FFBB3E975E0EFF01F95B" ID-DOI="http://doi.org/10.5281/zenodo.5222376" ID-Zenodo-Dep="5222376" httpUri="https://zenodo.org/record/5222376/files/figure.png" pageId="3" pageNumber="433" startId="3.[151,250,1608,1632]" targetBox="[157,1427,1013,1581]" targetPageId="3">
<paragraph id="8BB2336C9D12FFBB3E975E0EFF01F95B" blockId="3.[151,1437,1608,1704]" pageId="3" pageNumber="433">
<emphasis id="B979EF7E9D12FFBB3E975E0EFEFEF993" bold="true" box="[151,275,1608,1632]" pageId="3" pageNumber="433">FIGURE 2.</emphasis>
<taxonomicName id="4C0D48EF9D12FFBB3F1A5E0EFE4CF993" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[282,417,1608,1632]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="433" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D12FFBB3F1A5E0EFE4CF993" box="[282,417,1608,1632]" italics="true" pageId="3" pageNumber="433">Dixella atitla</emphasis>
</taxonomicName>
<emphasis id="B979EF7E9D12FFBB3FA75E0EFE15F993" bold="true" box="[423,504,1608,1632]" pageId="3" pageNumber="433">
<taxonomicNameLabel id="A24A52059D12FFBB3FA75E0EFE15F993" box="[423,504,1608,1632]" pageId="3" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB. Thorax, male. A. Dorsal view (pinned specimen); B. Lateral view (mounted in slide). C. Hind tibia, distal portion. D. Fifth tarsomere and claw of foreleg (dark arrow points to the seta). Scale: AB= 0.5 mm; CD= 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BB2336C9D12FFBC3EC75E90FE38FF0A" blockId="3.[151,1437,1750,2029]" lastBlockId="4.[151,1436,150,429]" lastPageId="4" lastPageNumber="434" pageId="3" pageNumber="433">
<emphasis id="B979EF7E9D12FFBB3EC75E90FEA7F903" bold="true" box="[199,330,1750,1776]" pageId="3" pageNumber="433">Terminalia</emphasis>
(
<figureCitation id="13362FE99D12FFBB3F5B5E90FE24F902" box="[347,457,1750,1777]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="5.[151,250,1943,1967]" captionStartId-1="6.[151,250,1969,1993]" captionStartId-2="7.[151,250,1151,1175]" captionTargetBox-0="[297,1282,185,1886]" captionTargetBox-1="[171,1412,726,1927]" captionTargetBox-2="[228,1357,190,1119]" captionTargetId-0="figure-25@5.[297,1290,181,1919]" captionTargetId-1="figure-258@6.[159,1429,704,1945]" captionTargetId-2="figure-25@7.[225,1363,181,1127]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURE 4. Dixella atitla sp. nov. AB. Male genitalia. C. Female genitalia. A. Tergite 9, dorsal view; B. Sternite 9, ventral view. C. Ventral view. Scale = 0.1 mm." captionText-1="FIGURE 5. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Tergum 10, ventral view. C. Gonostyle, top view. D. Dorsal view. Scale: AD= 0.1 mm" captionText-2="FIGURE 6. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Gonostyle, dorsal view, C, E. Lateral view. D. Dorsal view. Abbreviations: ae ap= aedeagal apodeme; ae com= aedeagal complex; a lob= anterior lobe of gonocoxite; goncx= gonocoxite; gonst= gonostyle. Scale: AD= 0.1 mm; E= 0.064 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5222380" figureDoi-1="http://doi.org/10.5281/zenodo.5222382" figureDoi-2="http://doi.org/10.5281/zenodo.5222384" httpUri-0="https://zenodo.org/record/5222380/files/figure.png" httpUri-1="https://zenodo.org/record/5222382/files/figure.png" httpUri-2="https://zenodo.org/record/5222384/files/figure.png" pageId="3" pageNumber="433">Figs. 46</figureCitation>
). Tergum 9 (
<figureCitation id="13362FE99D12FFBB3C5B5E90FD59F903" box="[603,692,1750,1776]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[297,1282,185,1886]" captionTargetId="figure-25@5.[297,1290,181,1919]" captionTargetPageId="5" captionText="FIGURE 4. Dixella atitla sp. nov. AB. Male genitalia. C. Female genitalia. A. Tergite 9, dorsal view; B. Sternite 9, ventral view. C. Ventral view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222380" httpUri="https://zenodo.org/record/5222380/files/figure.png" pageId="3" pageNumber="433">Fig. 4A</figureCitation>
) with anterior margin slightly emarginated, posterior margin pro- duced medially, with lateral setae. Sternum 9 (
<figureCitation id="13362FE99D12FFBB3CA65EBCFD10F8E7" box="[678,765,1786,1812]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[297,1282,185,1886]" captionTargetId="figure-25@5.[297,1290,181,1919]" captionTargetPageId="5" captionText="FIGURE 4. Dixella atitla sp. nov. AB. Male genitalia. C. Female genitalia. A. Tergite 9, dorsal view; B. Sternite 9, ventral view. C. Ventral view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222380" httpUri="https://zenodo.org/record/5222380/files/figure.png" pageId="3" pageNumber="433">Fig. 4B</figureCitation>
) with posterior margin slightly emarginated, with scattered setae. Gonocoxite slightly longer than wide; gonocoxal apodeme small and capitate; basal lobe not visible; apical lobe similar to the length of gonostylus, hammer-like, stem curved subapically with a long seta, apically with two opposite processes. Gonostylus dorsoventrally compressed, wider basally with scattered setae on ventral surface, with a pair of processes, one long than the other, the medial process with a setose rounded tubercle. Apodemes dark brown, slim and pointed (not completely visible in figures as in the slides they are ventrally oriented) (
<figureCitation id="13362FE99D12FFBB3B335FE8FA63F83A" box="[1331,1422,1966,1993]" captionStart="FIGURE 6" captionStartId="7.[151,250,1151,1175]" captionTargetBox="[228,1357,190,1119]" captionTargetId="figure-25@7.[225,1363,181,1127]" captionTargetPageId="7" captionText="FIGURE 6. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Gonostyle, dorsal view, C, E. Lateral view. D. Dorsal view. Abbreviations: ae ap= aedeagal apodeme; ae com= aedeagal complex; a lob= anterior lobe of gonocoxite; goncx= gonocoxite; gonst= gonostyle. Scale: AD= 0.1 mm; E= 0.064 mm." figureDoi="http://doi.org/10.5281/zenodo.5222384" httpUri="https://zenodo.org/record/5222384/files/figure.png" pageId="3" pageNumber="433">Fig. 6A</figureCitation>
); parameres triangular at base, hook-shaped at apex (
<figureCitation id="13362FE99D12FFBB3CDA5F94FCD9F81E" box="[730,820,2002,2029]" captionStart="FIGURE 6" captionStartId="7.[151,250,1151,1175]" captionTargetBox="[228,1357,190,1119]" captionTargetId="figure-25@7.[225,1363,181,1127]" captionTargetPageId="7" captionText="FIGURE 6. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Gonostyle, dorsal view, C, E. Lateral view. D. Dorsal view. Abbreviations: ae ap= aedeagal apodeme; ae com= aedeagal complex; a lob= anterior lobe of gonocoxite; goncx= gonocoxite; gonst= gonostyle. Scale: AD= 0.1 mm; E= 0.064 mm." figureDoi="http://doi.org/10.5281/zenodo.5222384" httpUri="https://zenodo.org/record/5222384/files/figure.png" pageId="3" pageNumber="433">Fig. 6A</figureCitation>
). Aedeagus basally a blunt triangle anteriorly directed, filiform posteriorly with nearly imperceptible spicules (
<figureCitation id="13362FE99D15FFBC3CFE58D1FCBAFF41" box="[766,855,151,178]" captionStart="FIGURE 6" captionStartId="7.[151,250,1151,1175]" captionTargetBox="[228,1357,190,1119]" captionTargetId="figure-25@7.[225,1363,181,1127]" captionTargetPageId="7" captionText="FIGURE 6. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Gonostyle, dorsal view, C, E. Lateral view. D. Dorsal view. Abbreviations: ae ap= aedeagal apodeme; ae com= aedeagal complex; a lob= anterior lobe of gonocoxite; goncx= gonocoxite; gonst= gonostyle. Scale: AD= 0.1 mm; E= 0.064 mm." figureDoi="http://doi.org/10.5281/zenodo.5222384" httpUri="https://zenodo.org/record/5222384/files/figure.png" pageId="4" pageNumber="434">Fig. 6D</figureCitation>
). Tergum 10 (
<figureCitation id="13362FE99D15FFBC3DEF58D1FBAEFF42" box="[1007,1091,150,177]" captionStart="FIGURE 5" captionStartId="6.[151,250,1969,1993]" captionTargetBox="[171,1412,726,1927]" captionTargetId="figure-258@6.[159,1429,704,1945]" captionTargetPageId="6" captionText="FIGURE 5. Dixella atitla sp. nov. Male genitalia. A. Ventral view. B. Tergum 10, ventral view. C. Gonostyle, top view. D. Dorsal view. Scale: AD= 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.5222382" httpUri="https://zenodo.org/record/5222382/files/figure.png" pageId="4" pageNumber="434">Fig. 5B</figureCitation>
) wider than long, anterior margin with distinctive basolateral process, narrowly posteriorly, the posterior margin membranous, with two patches of submedial elongate setae.
</paragraph>
<paragraph id="8BB2336C9D15FFBC3EC75945FB15FE5E" blockId="4.[151,1436,150,429]" pageId="4" pageNumber="434">
<emphasis id="B979EF7E9D15FFBC3EC75945FECEFEEE" box="[199,291,259,285]" italics="true" pageId="4" pageNumber="434">Female.</emphasis>
As in male but with the following differences: Head (
<figureCitation id="13362FE99D15FFBC3D8B5945FC0EFEEE" box="[907,995,259,285]" captionStart="FIGURE 1" captionStartId="3.[151,250,926,950]" captionTargetBox="[162,1430,191,899]" captionTargetId="figure-25@3.[151,1436,181,903]" captionTargetPageId="3" captionText="FIGURE 1. Dixella atitla sp. nov. AB. Head, dorsal view. C. Antenna, lateral view. A. Male; BC. Female. Abbreviations: flgm I to V= first to fifth flagellomere; pe= pedicel; scp= scape. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222374" httpUri="https://zenodo.org/record/5222374/files/figure.png" pageId="4" pageNumber="434">Fig. 1B</figureCitation>
). Clypeus pale yellow with 10 setae at disk. Antenna (
<figureCitation id="13362FE99D15FFBC3F425961FE75FEB2" box="[322,408,295,321]" captionStart="FIGURE 1" captionStartId="3.[151,250,926,950]" captionTargetBox="[162,1430,191,899]" captionTargetId="figure-25@3.[151,1436,181,903]" captionTargetPageId="3" captionText="FIGURE 1. Dixella atitla sp. nov. AB. Head, dorsal view. C. Antenna, lateral view. A. Male; BC. Female. Abbreviations: flgm I to V= first to fifth flagellomere; pe= pedicel; scp= scape. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222374" httpUri="https://zenodo.org/record/5222374/files/figure.png" pageId="4" pageNumber="434">Fig. 1C</figureCitation>
) dark brown. Wing (3B), length 3.0 mm. Terminalia (
<figureCitation id="13362FE99D15FFBC3DFC5961FBBBFEB2" box="[1020,1110,295,321]" captionStart="FIGURE 4" captionStartId="5.[151,250,1943,1967]" captionTargetBox="[297,1282,185,1886]" captionTargetId="figure-25@5.[297,1290,181,1919]" captionTargetPageId="5" captionText="FIGURE 4. Dixella atitla sp. nov. AB. Male genitalia. C. Female genitalia. A. Tergite 9, dorsal view; B. Sternite 9, ventral view. C. Ventral view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5222380" httpUri="https://zenodo.org/record/5222380/files/figure.png" pageId="4" pageNumber="434">Fig. 4C</figureCitation>
): sternum 8 wider than long; tergum 9 dark brown, with anterior margin slightly emarginated, posterior margin rounded; sternum 9 forming a band with anterolateral projections; apically rounded, with dense minute spicules and moderately sized setae present. Segment 10 shorter than cerci, long than wider, V-shaped; cercus dark brown, longer than wide.
</paragraph>
</subSubSection>
<caption id="DF7263E49D15FFBC3E975F4DFE5AF825" ID-DOI="http://doi.org/10.5281/zenodo.5222378" ID-Zenodo-Dep="5222378" httpUri="https://zenodo.org/record/5222378/files/figure.png" pageId="4" pageNumber="434" startId="4.[151,250,1803,1827]" targetBox="[151,1421,466,1764]" targetPageId="4">
<paragraph id="8BB2336C9D15FFBC3E975F4DFE5AF825" blockId="4.[151,1437,1803,2007]" pageId="4" pageNumber="434">
<emphasis id="B979EF7E9D15FFBC3E975F4DFEF9F8D0" bold="true" box="[151,276,1803,1827]" pageId="4" pageNumber="434">FIGURE 3.</emphasis>
Wing, dorsal view. A. Male,
<taxonomicName id="4C0D48EF9D15FFBC3C415F4DFD24F8D0" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[577,713,1803,1827]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D15FFBC3C415F4DFD24F8D0" box="[577,713,1803,1827]" italics="true" pageId="4" pageNumber="434">Dixella atitla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A52059D15FFBC3CCF5F4DFCCFF8D1" box="[719,802,1803,1827]" pageId="4" pageNumber="434" rank="species">
<emphasis id="B979EF7E9D15FFBC3CCF5F4DFCF6F8D0" bold="true" box="[719,795,1803,1827]" pageId="4" pageNumber="434">sp. nov</emphasis>
.
</taxonomicNameLabel>
B. Female,
<taxonomicName id="4C0D48EF9D15FFBC3D9F5F4DFBCAF8D0" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[927,1063,1803,1827]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D15FFBC3D9F5F4DFBCAF8D0" box="[927,1063,1803,1827]" italics="true" pageId="4" pageNumber="434">Dixella atitla</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24A52059D15FFBC3A2E5F4DFB6DF8D1" box="[1070,1152,1803,1827]" pageId="4" pageNumber="434" rank="species">
<emphasis id="B979EF7E9D15FFBC3A2E5F4DFB97F8D0" bold="true" box="[1070,1146,1803,1827]" pageId="4" pageNumber="434">sp. nov</emphasis>
.
</taxonomicNameLabel>
C. Male,
<taxonomicName id="4C0D48EF9D15FFBC3AE75F4DFA7BF8D0" baseAuthorityName="Lane" baseAuthorityYear="1942" box="[1255,1430,1803,1827]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="434" phylum="Arthropoda" rank="species" species="shannoni">
<emphasis id="B979EF7E9D15FFBC3AE75F4DFA7BF8D0" box="[1255,1430,1803,1827]" italics="true" pageId="4" pageNumber="434">Dixella shannoni</emphasis>
</taxonomicName>
. Abbreviations: C= costal vein; CuA= anterior branch of cubital vein; CuP= posterior branch of cubital vein; M
<subScript id="178931299D15FFBC3AF25F79FB14F8BE" attach="left" box="[1266,1273,1855,1869]" fontSize="6" pageId="4" pageNumber="434">1</subScript>
= first branch of media; M
<subScript id="178931299D15FFBC3EFA5F25FEECF882" attach="left" box="[250,257,1891,1905]" fontSize="6" pageId="4" pageNumber="434">2</subScript>
= second branch of media; M
<subScript id="178931299D15FFBC3C2E5F25FDD8F882" attach="left" box="[558,565,1891,1905]" fontSize="6" pageId="4" pageNumber="434">3</subScript>
+
<subScript id="178931299D15FFBC3C435F25FDA7F882" attach="none" box="[579,586,1891,1905]" fontSize="6" pageId="4" pageNumber="434">4</subScript>
= third and fourth branch of media;; m-cu= medial-cubital crossvein; m-m= medial crossvein; Rs= radial sector; R
<subScript id="178931299D15FFBC3FD15FC1FE35F866" attach="left" box="[465,472,1927,1941]" fontSize="6" pageId="4" pageNumber="434">1</subScript>
= anterior branch of radius; R
<subScript id="178931299D15FFBC3D075FC1FCE3F866" attach="left" box="[775,782,1927,1941]" fontSize="6" pageId="4" pageNumber="434">2</subScript>
= upper branch of second branch of radius; R
<subScript id="178931299D15FFBC3ADE5FC1FB08F866" attach="left" box="[1246,1253,1927,1941]" fontSize="6" pageId="4" pageNumber="434">3</subScript>
= lower branch of second branch of radius; R
<subScript id="178931299D15FFBC3FAA5FEDFE2CF84A" attach="left" box="[426,449,1963,1977]" fontSize="6" pageId="4" pageNumber="434">2+3</subScript>
= second branch of radius; R = third branch of radius; r-m= radial-medial crossvein; Sc =
<subScript id="178931299D15FFBC3B715FDDFCEDF849" attach="left" fontSize="6" pageId="4" pageNumber="434">sub-4+5</subScript>
costal vein. Scale = 0.5 mm.
</paragraph>
</caption>
<caption id="DF7263E49D14FFBD3E975FD1FDCEF820" ID-DOI="http://doi.org/10.5281/zenodo.5222380" ID-Zenodo-Dep="5222380" httpUri="https://zenodo.org/record/5222380/files/figure.png" pageId="5" pageNumber="435" startId="5.[151,250,1943,1967]" targetBox="[297,1282,185,1886]" targetPageId="5">
<paragraph id="8BB2336C9D14FFBD3E975FD1FDCEF820" blockId="5.[151,1436,1943,2004]" pageId="5" pageNumber="435">
<emphasis id="B979EF7E9D14FFBD3E975FD1FEF9F85C" bold="true" box="[151,276,1943,1967]" pageId="5" pageNumber="435">FIGURE 4.</emphasis>
<taxonomicName id="4C0D48EF9D14FFBD3F1C5FDEFE49F843" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[284,420,1944,1968]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="435" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D14FFBD3F1C5FDEFE49F843" box="[284,420,1944,1968]" italics="true" pageId="5" pageNumber="435">Dixella atitla</emphasis>
</taxonomicName>
<emphasis id="B979EF7E9D14FFBD3FAB5FDEFE10F843" bold="true" box="[427,509,1944,1968]" pageId="5" pageNumber="435">
<taxonomicNameLabel id="A24A52059D14FFBD3FAB5FDEFE10F843" box="[427,509,1944,1968]" pageId="5" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
AB. Male genitalia. C. Female genitalia. A. Tergite 9, dorsal view; B. Sternite 9, ventral view. C. Ventral view. Scale = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C31760E79D17FFBE3EC758D0FE60FF26" pageId="6" pageNumber="436" type="distribution">
<paragraph id="8BB2336C9D17FFBE3EC758D0FE60FF26" blockId="6.[151,1437,150,681]" pageId="6" pageNumber="436">
<emphasis id="B979EF7E9D17FFBE3EC758D0FE8CFF43" bold="true" box="[199,353,150,176]" pageId="6" pageNumber="436">Distribution.</emphasis>
This species is known only from the
<typeStatus id="54B68DCE9D17FFBE3D1158D1FCACFF42" box="[785,833,151,177]" pageId="6" pageNumber="436">type</typeStatus>
locality Laguna de Santo Domingo, Huatusco,
<collectingRegion id="49C9FD8E9D17FFBE3B6058D1FF27FF26" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="6" pageNumber="436">Veracruz</collectingRegion>
,
<collectingCountry id="F31A73FC9D17FFBE3ED558FCFEC6FF27" box="[213,299,186,212]" name="Mexico" pageId="6" pageNumber="436">Mexico</collectingCountry>
(
<figureCitation id="13362FE99D17FFBE3F3A58FCFE92FF26" box="[314,383,186,213]" captionStart="FIGURE 7" captionStartId="7.[151,250,2015,2039]" captionTargetBox="[228,1358,1295,1987]" captionTargetId="figure-83@7.[223,1363,1271,1991]" captionTargetPageId="7" captionText="FIGURE 7. Distribution of Dixella atitla sp. nov. in state of Veracruz and type locality." figureDoi="http://doi.org/10.5281/zenodo.5222386" httpUri="https://zenodo.org/record/5222386/files/figure.png" pageId="6" pageNumber="436">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31760E79D17FFBE3EC75898FD72FEEF" pageId="6" pageNumber="436" type="etymology">
<paragraph id="8BB2336C9D17FFBE3EC75898FD72FEEF" blockId="6.[151,1437,150,681]" pageId="6" pageNumber="436">
<emphasis id="B979EF7E9D17FFBE3EC75898FEA3FF0B" bold="true" box="[199,334,222,248]" pageId="6" pageNumber="436">Etymology.</emphasis>
The specific epithet is derived from Nahualt language, abbreviation of the word “
<emphasis id="B979EF7E9D17FFBE3ADF5898FACFFF0B" box="[1247,1314,222,248]" italics="true" pageId="6" pageNumber="436">atitlaj</emphasis>
”, meaning “in water”, referring to the collection location.
</paragraph>
</subSubSection>
<subSubSection id="C31760E79D17FFBE3EC75960FAFCFD5B" pageId="6" pageNumber="436" type="discussion">
<paragraph id="8BB2336C9D17FFBE3EC75960FC19FD92" blockId="6.[151,1437,150,681]" pageId="6" pageNumber="436">
<emphasis id="B979EF7E9D17FFBE3EC75960FED6FEB3" bold="true" box="[199,315,294,320]" pageId="6" pageNumber="436">Remarks.</emphasis>
This new species is distinguished from all other known species of
<taxonomicName id="4C0D48EF9D17FFBE3A215960FB9EFEB3" authorityName="Dyar &amp; Shannon" authorityYear="1924" box="[1057,1139,294,320]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="B979EF7E9D17FFBE3A215960FB9EFEB3" box="[1057,1139,294,320]" italics="true" pageId="6" pageNumber="436">Dixella</emphasis>
</taxonomicName>
by the hammer-like apical lobe of the gonocoxite and flattened basally expanded gonostylus bearing a pair of apical processes. This species is most closely related to
<taxonomicName id="4C0D48EF9D17FFBE3F9D5928FCE3FE7B" authority="Peters &amp; Cook" authorityName="Peters &amp; Cook" authorityYear="1966" box="[413,782,366,393]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="cumbrica">
<emphasis id="B979EF7E9D17FFBE3F9D5928FD8CFE7B" box="[413,609,366,392]" italics="true" pageId="6" pageNumber="436">Dixella cumbrica</emphasis>
Peters &amp; Cook
</taxonomicName>
from
<collectingRegion id="49C9FD8E9D17FFBE3D535928FBC3FE7B" box="[851,1070,366,392]" country="Mexico" name="Baja California Sur" pageId="6" pageNumber="436">Baja California Sur</collectingRegion>
(Sierra Laguna), sharing similar characteristics of body color, the shape of the apical lobe of the gonocoxite and flattened gonostylus.
<taxonomicName id="4C0D48EF9D17FFBE3B0859D4FA71FE5F" authority="Huerta &amp; Ibáñez-Bernal, 2021" authorityName="Huerta &amp; Ibáñez-Bernal" authorityYear="2021" box="[1288,1436,402,428]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="atitla" status="sp. nov.">
<emphasis id="B979EF7E9D17FFBE3B0859D4FA71FE5F" box="[1288,1436,402,428]" italics="true" pageId="6" pageNumber="436">Dixella atitla</emphasis>
</taxonomicName>
<emphasis id="B979EF7E9D17FFBE3E9759F0FF1CFE23" bold="true" box="[151,241,438,464]" pageId="6" pageNumber="436">
<taxonomicNameLabel id="A24A52059D17FFBE3E9759F0FF1CFE23" box="[151,241,438,464]" pageId="6" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C0D48EF9D17FFBE3F8459F1FDFDFE23" authorityName="Peters &amp; Cook" authorityYear="1966" box="[388,528,438,464]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="cumbrica">
<emphasis id="B979EF7E9D17FFBE3F8459F1FDFDFE23" box="[388,528,438,464]" italics="true" pageId="6" pageNumber="436">D. cumbrica</emphasis>
</taxonomicName>
(characters in parenthesis) in having: (1) fore coxa dark brown basally (hyaline); (2) femora yellowish with brown apices and tibiae pale brown, with brown apices (femora and tibiae light brown); (3) halter brown (hyaline); (4) wing not spotted (wing slightly spotted on r-m and base R
<subScript id="178931299D17FFBE3A765A57FB62FDEC" attach="left" box="[1142,1167,529,543]" fontSize="6" pageId="6" pageNumber="436">2+3</subScript>
); (5) scutellum with six setae (two weak setae); (6) apical lobe of the gonocoxite hammer-like (moderately long, thick, with stellate tip), and (7) gonostylus flattened, wider basally with a pair of processes (flat, simple).
</paragraph>
<paragraph id="8BB2336C9D17FFBE3EC75A2CFAFCFD5B" blockId="6.[151,1437,150,681]" pageId="6" pageNumber="436">
The female is similar to male, with segment 10 longer than wide, V-shaped and shorter than cercus, which distinguishes it from other species for which females are known. The female of
<taxonomicName id="4C0D48EF9D17FFBE3DF45AC9FB6DFD5B" authorityName="Peters &amp; Cook" authorityYear="1966" box="[1012,1152,654,680]" class="Insecta" family="Dixidae" genus="Dixella" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="436" phylum="Arthropoda" rank="species" species="cumbrica">
<emphasis id="B979EF7E9D17FFBE3DF45AC9FB6DFD5B" box="[1012,1152,654,680]" italics="true" pageId="6" pageNumber="436">D. cumbrica</emphasis>
</taxonomicName>
is unknown.
</paragraph>
</subSubSection>
</treatment>
</document>