571 lines
76 KiB
XML
571 lines
76 KiB
XML
<document id="634FEECA126C4EA21878A11724D53DC4" ID-CLB-Dataset="7166" ID-DOI="10.1093/jmammal/gyab023" ID-GBIF-Dataset="d1c8718a-3b16-45c7-867d-c8ba4e9ff482" ID-ISSN="1545-1542" ID-Zenodo-Dep="5613298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619530923730" checkinUser="felipe" docAuthor="Ochoa, Janine, Mijares, Armand S. B., Piper, Philip J., Reyes, Marian C. & Heaney, Lawrence R." docDate="2021" docId="03A41512FFFB482EFC6F2EDD44DE01A3" docLanguage="en" docName="jMammalogy.102.2.1-22.pdf.imf" docOrigin="Journal of Mammalogy 102 (2)" docStyle="DocumentStyle:8488A0FD6434D222CC79ECAD85E66EFE.2:JourMammal.2015-.journal_article" docStyleId="8488A0FD6434D222CC79ECAD85E66EFE" docStyleName="JourMammal.2015-.journal_article" docStyleVersion="2" docTitle="Carpomys dakal Ochoa & Mijares & Piper & Reyes & Heaney 2021, new species" docType="treatment" docUuid="6A52EA3E-46A4-4563-8DB6-8E842352C9A0" docUuidSource="ZooBank" docVersion="9" lastPageNumber="12" masterDocId="FF9D6D6AFFF24825FF8C2A7447020664" masterDocTitle="Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini)" masterLastPageNumber="22" masterPageNumber="1" pageNumber="10" updateTime="1698937702425" updateUser="ExternalLinkService">
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<mods:title id="3E2B2BACDF9F6C2792E68BE96B0F284D">Three new extinct species from the endemic Philippine cloud rat radiation (Rodentia, Muridae, Phloeomyini)</mods:title>
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<mods:name id="CE02AA006E42D756C02CADF9A18C4E94" type="personal">
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<mods:namePart id="B4ED71AD9229C09006E544F107C859DC">Ochoa, Janine</mods:namePart>
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<mods:affiliation id="F52DE51E9FC90DE69A28785FBB3F6026">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:nameIdentifier id="9D98576E7A5D19E980A3F284006CCD02" type="email">jochoa@up.edu.ph</mods:nameIdentifier>
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<mods:namePart id="91D3265DD3F11F3DBEBB00A5121C4238">Mijares, Armand S. B.</mods:namePart>
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<mods:affiliation id="E6DFF3A67E99EACA5AF738E8C8492876">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="78254682C6FE581554B3FE5111A3FDE3">Piper, Philip J.</mods:namePart>
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<mods:affiliation id="8E32E9F1938C9E7C3B8DF5BD486333C0">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="4A9B8CBBC21C9E24FC170D987910082E">Reyes, Marian C.</mods:namePart>
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<mods:affiliation id="CD2D979B1CEB1179A6359B78A844E999">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:namePart id="B8459D43AC274C5A626A61A4CC1C7127">Heaney, Lawrence R.</mods:namePart>
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<mods:affiliation id="4E149EDCB81188473C1BB3BB04A7D905">Department of Anthropology, University of the Philippines, Diliman, Quezon City 1101, Philippines (JO) Archaeological Studies Program, University of the Philippines, Diliman, Quezon City 1101, Philippines (ASBM, PJP) National Museum of the Philippines, Padre Burgos Street, Manila 1000, Philippines (ASBM, PJP, MCR, LRH) Field Museum of Natural History, 1400 S Lake Shore Drive, Chicago, IL 60605, USA (LRH) Present address of PJP: School of Archaeology and Anthropology, Australian National University, Canberra, ACT 2601, Australia</mods:affiliation>
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<mods:title id="60EEFFA2D192B3DF752623B15CA40AC5">Journal of Mammalogy</mods:title>
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<mods:part id="BC522D1558B1EE83A962ACA0650E3CBB">
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<mods:date id="98B37B2EE06EBC121FFF139110A05759">2021</mods:date>
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<mods:number id="91CF6C1CFB226F6780EB5731B5B33C0F">2021-04-23</mods:number>
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<mods:number id="5064AA8838C4E99462B7D7B92E4492D2">102</mods:number>
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<mods:identifier id="46293FB0D69886FC042E1850ADFA9E5F" type="DOI">10.1093/jmammal/gyab023</mods:identifier>
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<treatment id="03A41512FFFB482EFC6F2EDD44DE01A3" ID-DOI="http://doi.org/10.5281/zenodo.4723434" ID-GBIF-Taxon="181653062" ID-Zenodo-Dep="4723434" LSID="urn:lsid:zoobank.org:act:6A52EA3E-46A4-4563-8DB6-8E842352C9A0" httpUri="http://treatment.plazi.org/id/03A41512FFFB482EFC6F2EDD44DE01A3" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection id="C317F78FFFFB482CFC6F2EDD422202A4" box="[995,1312,1193,1216]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BB2A404FFFB482CFC6F2EDD422202A4" blockId="9.[819,1488,1193,1888]" box="[995,1312,1193,1216]" pageId="9" pageNumber="10">
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<heading id="D0FA1368FFFB482CFC6F2EDD422202A4" box="[995,1312,1193,1216]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="10" reason="4">
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<taxonomicName id="4C0DDF87FFFB482CFC6F2EDD439102A4" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[995,1171,1193,1216]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="dakal" status="sp. nov.">
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<emphasis id="B9797816FFFB482CFC6F2EDD439102A4" box="[995,1171,1193,1216]" italics="true" pageId="9" pageNumber="10">Carpomys dakal</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24AC56DFFFB482CFB2C2EDD422202A4" box="[1184,1312,1193,1216]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C317F78FFFFB482CFC152EBD420C0364" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BB2A404FFFB482CFC152EBD426B0284" blockId="9.[819,1488,1193,1888]" box="[921,1385,1225,1248]" pageId="9" pageNumber="10">
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<figureCitation id="1336B881FFFB482CFC152EBD43240284" box="[921,1062,1225,1248]" captionStart="Fig" captionStartId="8.[144,179,1893,1914]" captionTargetBox="[271,1405,888,1880]" captionTargetId="figure-359@8.[271,1405,888,1880]" captionTargetPageId="8" captionText="Fig. 3.—Holotype of Carpomys dakal n. sp. (PNM II-1977-J11A-16301), left dentary: (A) labial view of mandible; (B) lingual view of mandible (mirrored)." figureDoi="http://doi.org/10.5281/zenodo.5613308" httpUri="https://zenodo.org/record/5613308/files/figure.png" pageId="9" pageNumber="10">Figs. 3A, 3B</figureCitation>
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,
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<figureCitation id="1336B881FFFB482CFBB82EBD43540284" box="[1076,1110,1225,1248]" captionStart="Fig" captionStartId="9.[101,136,1865,1886]" captionTargetBox="[106,767,201,1850]" captionTargetId="figure-521@9.[105,767,200,1851]" captionTargetPageId="9" captionText="Fig. 4.—Occlusal view of maxillary molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-15997), left side; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus(FMNH193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-J3-10389), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), right side (mirrored); (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="9" pageNumber="10">4A</figureCitation>
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, and
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<figureCitation id="1336B881FFFB482CFB1F2EBD43B60284" box="[1171,1204,1225,1248]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="9" pageNumber="10">5A</figureCitation>
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;
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||
<tableCitation id="C68F91BFFFFB482CFB4E2EBD421C0284" box="[1218,1310,1225,1248]" captionStart="Table 1" captionStartId="3.[125,181,1354,1375]" captionTargetPageId="3" captionText="Table 1.—Reported radiometric ages for provenance and study sites of fossil cloud rat specimens. Holotypes are underlined. *Minimum uranium series ages for Callao Layer 14. **Depth of specimen is above the last recorded occurrence of pottery; specimen is not directly associated with dated sample. 14C calibration was done using OxCal 4.3 (Bronk Ramsey 2017) and ranges shown are at 95.4% probability. Numbers in the last three columns are specimen numbers; see lists of holotypes and paratypes in text." httpUri="http://table.plazi.org/id/DF72F48CFFF14826FFF12F3E449803D3" pageId="9" pageNumber="10" tableUuid="DF72F48CFFF14826FFF12F3E449803D3">Tables 1</tableCitation>
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and
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<tableCitation id="C68F91BFFFFB482CFAD82EBD42630284" box="[1364,1377,1225,1248]" captionStart="Table 2" captionStartId="6.[124,180,651,672]" captionTargetPageId="6" captionText="Table 2.—Measurements (in mm) of maxillary and mandibular teeth of Carpomys and Musseromys." httpUri="http://table.plazi.org/id/DF72F48CFFF44823FFF028FF433D04C4" pageId="9" pageNumber="10" tableUuid="DF72F48CFFF44823FFF028FF433D04C4">2</tableCitation>
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;
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</paragraph>
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<paragraph id="8BB2A404FFFB482CFC792E9D420C0364" blockId="9.[819,1488,1193,1888]" box="[1013,1294,1257,1280]" pageId="9" pageNumber="10">Supplementary Data SD1</paragraph>
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</subSubSection>
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<subSubSection id="C317F78FFFFB482FFCC12F7D42670444" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
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<paragraph id="8BB2A404FFFB482CFCC12F7D44C503C4" blockId="9.[819,1488,1193,1888]" pageId="9" pageNumber="10">
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<materialsCitation id="3B65AE59FFFB482CFCC12F7D44C503C4" ID-GBIF-Occurrence="3091612309" collectionCode="PNM" country="Philippines" county="Minori Cave" elevation="200" latitude="17.72139" location="Minori Cave" longLatPrecision="21" longitude="121.82833" municipality="Penablanca Municipality" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J11A-16301" specimenCount="1" stateProvince="Cagayan Province" typeStatus="holotype">
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<emphasis id="B9797816FFFB482CFCC12F7D44CF0344" box="[845,973,1289,1312]" italics="true" pageId="9" pageNumber="10">
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<typeStatus id="54B61AA6FFFB482CFCC12F7D44AC0344" box="[845,942,1289,1312]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
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.—
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</emphasis>
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<specimenCode id="DBAB0C7FFFFB482CFC412F7D43EC0344" box="[973,1262,1289,1312]" pageId="9" pageNumber="10">PNM II-1977-J11A-16301</specimenCode>
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, left dentary with incisor and m1–3; only a small portion of the dentary posterior to m3 is present (
|
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<figureCitation id="1336B881FFFB482CFC612F3D439E0304" box="[1005,1180,1353,1376]" captionStart="Fig" captionStartId="8.[144,179,1893,1914]" captionTargetBox="[271,1405,888,1880]" captionTargetId="figure-359@8.[271,1405,888,1880]" captionTargetPageId="8" captionText="Fig. 3.—Holotype of Carpomys dakal n. sp. (PNM II-1977-J11A-16301), left dentary: (A) labial view of mandible; (B) lingual view of mandible (mirrored)." figureDoi="http://doi.org/10.5281/zenodo.5613308" httpUri="https://zenodo.org/record/5613308/files/figure.png" pageId="9" pageNumber="10">Figs. 3A and 3B</figureCitation>
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). The m1–3 are lightly worn (
|
||
<figureCitation id="1336B881FFFB482CFCB02F1D448D03E4" box="[828,911,1385,1408]" captionStart="Fig" captionStartId="10.[144,178,1865,1886]" captionTargetBox="[148,809,201,1850]" captionTargetId="figure-534@10.[147,809,200,1851]" captionTargetPageId="10" captionText="Fig. 5.—Occlusal view of mandibular molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-16301, holotype), leftside; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus (FMNH 193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-I8-402, holotype), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), left side; (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys cagayanensis n. sp. (PNM II-1977-J3-7554, holotype), right side (mirrored); (H) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="9" pageNumber="10">Fig. 5A</figureCitation>
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), indicating that the animal was a young adult at the time of death.
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||
</materialsCitation>
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</paragraph>
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<paragraph id="8BB2A404FFFB482CFCC12FDD43AD0024" blockId="9.[819,1488,1193,1888]" pageId="9" pageNumber="10">
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<emphasis id="B9797816FFFB482CFCC12FDD44F703A4" box="[845,1013,1449,1472]" italics="true" pageId="9" pageNumber="10">Type locality.—</emphasis>
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<location id="8ED2F2DFFFFB482CFC792FDD438003A4" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFFB482CFC792FDD438003A4" box="[1013,1154,1449,1472]" country="Philippines" county="Minori Cave" latitude="17.72139" longLatPrecision="21" longitude="121.82833" municipality="Penablanca Municipality" name="Minori Cave" pageId="9" pageNumber="10" stateProvince="Cagayan Province">Minori Cave</location>
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,
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<location id="8ED2F2DFFFFB482CFB032FDD424B03A4" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFFB482CFB032FDD424B03A4" box="[1167,1353,1449,1472]" country="Philippines" county="Minori Cave" latitude="17.72139" longLatPrecision="21" longitude="121.82833" municipality="Penablanca Municipality" name="Barangay Quibal" pageId="9" pageNumber="10" stateProvince="Cagayan Province">Barangay Quibal</location>
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,
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||
<collectingMunicipality id="6BD63E7EFFFB482CFADB2FDD44BC0384" pageId="9" pageNumber="10">Peñablanca Municipality</collectingMunicipality>
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,
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<collectingRegion id="49C96AE6FFFB482CFC5C2FBD439F0384" box="[976,1181,1481,1504]" country="Philippines" name="Cagayan" pageId="9" pageNumber="10">Cagayan Province</collectingRegion>
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,
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<collectingRegion id="49C96AE6FFFB482CFB232FBD42440384" box="[1199,1350,1481,1504]" pageId="9" pageNumber="10">Luzon Island</collectingRegion>
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,
|
||
<collectingCountry id="F31AE494FFFB482CFAD42FBD42D20384" box="[1368,1488,1481,1504]" name="Philippines" pageId="9" pageNumber="10">Philippines</collectingCountry>
|
||
(
|
||
<figureCitation id="1336B881FFFB482CFCB02F9D44780064" box="[828,890,1513,1536]" captionStart="Fig" captionStartId="2.[144,179,2005,2026]" captionTargetBox="[228,1445,202,1991]" captionTargetId="figure-13@2.[226,1448,200,1992]" captionTargetPageId="2" captionText="Fig. 1.—Map of the Callao Cave Complex, showing location of cave sites: Callao (1), Eme (2), Dalan Serkot (3), Minori (4), and Musang (5). Inset shows the location within Luzon Island." figureDoi="http://doi.org/10.5281/zenodo.5613302" httpUri="https://zenodo.org/record/5613302/files/figure.png" pageId="9" pageNumber="10">Fig. 1</figureCitation>
|
||
),
|
||
<quantity id="4CF509E1FFFB482CFC1D2F9D44D70064" box="[913,981,1513,1536]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="m" value="200.0">
|
||
<elevation id="00204337FFFB482CFC1D2F9D44D70064" box="[913,981,1513,1536]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="10" unit="m" value="200.0">200 m</elevation>
|
||
</quantity>
|
||
elevation,
|
||
<geoCoordinate id="EE39C2C3FFFB482CFBC62F9D43CE0064" box="[1098,1228,1511,1536]" degrees="17" direction="north" minutes="43" orientation="latitude" pageId="9" pageNumber="10" precision="15" seconds="17" value="17.72139">
|
||
17°43
|
||
<emphasis id="B9797816FFFB482CFB062F9343930064" box="[1162,1169,1511,1536]" italics="true" pageId="9" pageNumber="10">′</emphasis>
|
||
17
|
||
<emphasis id="B9797816FFFB482CFB272F9343B40064" box="[1195,1206,1511,1536]" italics="true" pageId="9" pageNumber="10">″</emphasis>
|
||
N
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE39C2C3FFFB482CFB5B2F9D42610064" box="[1239,1379,1511,1536]" degrees="121" direction="east" minutes="49" orientation="longitude" pageId="9" pageNumber="10" precision="15" seconds="42" value="121.82833">
|
||
121°49
|
||
<emphasis id="B9797816FFFB482CFAA82F9342290064" box="[1316,1323,1511,1536]" italics="true" pageId="9" pageNumber="10">′</emphasis>
|
||
42
|
||
<emphasis id="B9797816FFFB482CFACA2F9342530064" box="[1350,1361,1511,1536]" italics="true" pageId="9" pageNumber="10">″</emphasis>
|
||
E
|
||
</geoCoordinate>
|
||
, Chamber Dof Minori Cave, Square 27; derives from Holocene aceramic level
|
||
<quantity id="4CF509E1FFFB482CFCE02C5D44D40024" box="[876,982,1577,1600]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="9" pageNumber="10" unit="cm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 cm</quantity>
|
||
below cave surface.
|
||
</paragraph>
|
||
<paragraph id="8BB2A404FFFB482FFCC12C3D422A0744" blockId="9.[819,1488,1193,1888]" lastBlockId="10.[861,1531,201,1856]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
<materialsCitation id="3B65AE59FFFB482CFCC12C3D42D200E4" ID-GBIF-Occurrence="3091612302" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J3-10196" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B9797816FFFB482CFCC12C3D44D40004" box="[845,982,1609,1632]" italics="true" pageId="9" pageNumber="10">
|
||
<typeStatus id="54B61AA6FFFB482CFCC12C3D44B40004" box="[845,950,1609,1632]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
|
||
.—
|
||
</emphasis>
|
||
(n = 10; Supplementary Data SD1).
|
||
<location id="8ED2F2DFFFFB482CFA062C3D446F00E4" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFFB482CFA062C3D446F00E4" county="Cave Chamber" municipality="Minori Cave" name="Callao Cave" pageId="9" pageNumber="10">Callao Cave</location>
|
||
: partial right dentary with m2 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFB4C2C1D42C600E4" box="[1216,1476,1641,1664]" pageId="9" pageNumber="10">PNM II-1977-J3-10196</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFCBF2CFD428000C4" ID-GBIF-Occurrence="3091612312" box="[819,1410,1673,1696]" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J3-10544" specimenCount="1" typeStatus="paratype">
|
||
partial left dentary with m1 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFBFD2CFD427400C4" box="[1137,1398,1673,1696]" pageId="9" pageNumber="10">PNM II-1977-J3-10544</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFA002CFD427900A4" ID-GBIF-Occurrence="3091612305" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J3-11075/3" specimenCount="1" typeStatus="paratype">
|
||
partial right maxillary with M2 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFBDF2CDD426D00A4" box="[1107,1391,1705,1728]" pageId="9" pageNumber="10">PNM II-1977-J3-11075/3</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFA0A2CDD44BA0164" ID-GBIF-Occurrence="3091612313" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Minori Cave Chamber A" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977- J11B-2986" specimenCount="1" typeStatus="paratype">
|
||
<location id="8ED2F2DFFFFB482CFA0A2CDD44E80084" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFFB482CFA0A2CDD44E80084" county="Cave Chamber" municipality="Minori Cave" name="Minori Cave Chamber A" pageId="9" pageNumber="10">Minori Cave Chamber A</location>
|
||
: partial leftdentary with m1–2 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFABB2CBD44AE0164" pageId="9" pageNumber="10">PNM II-1977- J11B-2986</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFC332C9D43B70144" ID-GBIF-Occurrence="3091612303" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Minori Cave Chamber D" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J11A-15997" specimenCount="1" typeStatus="paratype">
|
||
<location id="8ED2F2DFFFFB482CFC332C9D43C90164" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFFB482CFC332C9D43C90164" box="[959,1227,1769,1792]" county="Cave Chamber" municipality="Minori Cave" name="Minori Cave Chamber D" pageId="9" pageNumber="10">Minori Cave Chamber D</location>
|
||
: partial left maxilla with M1–3 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFC0F2D7D43AA0144" box="[899,1192,1801,1824]" pageId="9" pageNumber="10">PNM II-1977-J11A-15997</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFB322D7D43B80124" ID-GBIF-Occurrence="3091612301" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J11A-16447" specimenCount="1" typeStatus="paratype">
|
||
partial right maxilla with M1–2 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFC092D5D43AF0124" box="[901,1197,1833,1856]" pageId="9" pageNumber="10">PNM II-1977-J11A-16447</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFB4A2D5D43BB0104" ID-GBIF-Occurrence="3091612307" collectingDate="1977-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J11A-16481" specimenCount="1" typeStatus="paratype">
|
||
partial left maxilla with M1–2 (
|
||
<specimenCode id="DBAB0C7FFFFB482CFC092D3D43AC0104" box="[901,1198,1865,1888]" pageId="9" pageNumber="10">PNM II-1977-J11A-16481</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFFB482CFB4A2D3D42D20104" ID-GBIF-Occurrence="3091612308" box="[1222,1488,1865,1888]" collectingDate="1977-02" collectingDateMax="1977-02" collectingDateMin="1976-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="9" pageNumber="10" specimenCode="PNM II-1977-J11A-16825" specimenCount="1" typeStatus="paratype">partial left maxilla with</materialsCitation>
|
||
M1–2 (
|
||
<specimenCode id="DBAB0C7FFFF8482FFC212ABD43D10684" box="[941,1235,201,224]" pageId="10" pageNumber="11">PNM II-1977-J11A-16825</specimenCode>
|
||
);
|
||
<materialsCitation id="3B65AE59FFF8482FFB652ABD43DE0764" ID-GBIF-Occurrence="3091612317" collectingDate="1977-02" collectingDateMax="1977-02" collectingDateMin="1976-02" collectionCode="PNM" county="Cave Chamber" location="Callao Cave" municipality="Minori Cave" pageId="10" pageNumber="11" specimenCode="PNM II-1977-J11A-16951" specimenCount="1" typeStatus="paratype">
|
||
partial right dentary with m2–3 (
|
||
<specimenCode id="DBAB0C7FFFF8482FFC262A9D43D20764" box="[938,1232,233,256]" pageId="10" pageNumber="11">PNM II-1977-J11A-16951</specimenCode>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="3B65AE59FFF8482FFB692A9D42270744" ID-GBIF-Occurrence="3091612314" collectingDate="1977-02" collectingDateMax="1977-02" collectingDateMin="1976-02" collectionCode="PNM" county="Cave Chamber" location="Musang Cave" municipality="Minori Cave" pageId="10" pageNumber="11" specimenCode="PNM II-1976-M-101" specimenCount="1" typeStatus="paratype">
|
||
<location id="8ED2F2DFFFF8482FFB692A9D427C0764" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFF8482FFB692A9D427C0764" box="[1253,1406,233,256]" county="Cave Chamber" municipality="Minori Cave" name="Musang Cave" pageId="10" pageNumber="11">Musang Cave</location>
|
||
: partial left dentary with m1–3 (
|
||
<specimenCode id="DBAB0C7FFFF8482FFBB92B7D421F0744" box="[1077,1309,265,288]" pageId="10" pageNumber="11">PNM II-1976-M-101</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF72F48CFFFB482CFFE92D3D43CD01B7" ID-DOI="http://doi.org/10.5281/zenodo.5613310" ID-Zenodo-Dep="5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="9" pageNumber="10" startId="9.[101,136,1865,1886]" targetBox="[106,767,201,1850]" targetPageId="9">
|
||
<paragraph id="8BB2A404FFFB482CFFE92D3D43CD01B7" blockId="9.[101,771,1865,2003]" lastBlockId="9.[819,1488,1952,2003]" pageId="9" pageNumber="10">
|
||
<emphasis id="B9797816FFFB482CFFE92D3D47A5013A" bold="true" box="[101,167,1865,1886]" pageId="9" pageNumber="10">Fig. 4.</emphasis>
|
||
—Occlusal view of maxillary molars of (A)
|
||
<taxonomicName id="4C0DDF87FFFB482CFDEE2D3D4400013A" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[610,770,1865,1886]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="dakal" status="sp. nov.">
|
||
<emphasis id="B9797816FFFB482CFDEE2D3D4400013A" box="[610,770,1865,1886]" italics="true" pageId="9" pageNumber="10">Carpomys dakal</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFFB482CFFE92D124798011F" box="[101,154,1894,1915]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM II-1977-J11A-15997), left side; (B)
|
||
<taxonomicName id="4C0DDF87FFFB482CFDCC2D124400011F" authorityName="Thomas" authorityYear="1895" box="[576,770,1894,1915]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFFB482CFDCC2D124400011F" box="[576,770,1894,1915]" italics="true" pageId="9" pageNumber="10">Carpomys phaeurus</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 62291), leftside; (C)
|
||
<taxonomicName id="4C0DDF87FFFB482CFEF62DF7455B01FC" authorityName="Heaney, Balete, Rickart, Veluz & Jansa" authorityYear="2014" box="[378,601,1923,1944]" class="Mammalia" family="Muridae" genus="Musseromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="inopinatus">
|
||
<emphasis id="B9797816FFFB482CFEF62DF7455B01FC" box="[378,601,1923,1944]" italics="true" pageId="9" pageNumber="10">Musseromysinopinatus</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 193840), left side; (D)
|
||
<taxonomicName id="4C0DDF87FFFB482CFF6B2DD5469701D2" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[231,405,1953,1974]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="ballik" status="sp. nov.">
|
||
<emphasis id="B9797816FFFB482CFF6B2DD5469701D2" box="[231,405,1953,1974]" italics="true" pageId="9" pageNumber="10">Crateromys ballik</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFFB482CFE172DD546CD01D2" box="[411,463,1953,1974]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM II-1977-J3-10389), right side (mirrored); (E)
|
||
<taxonomicName id="4C0DDF87FFFB482CFEA22DCA452401B7" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[302,550,1982,2003]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFFB482CFEA22DCA452401B7" box="[302,550,1982,2003]" italics="true" pageId="9" pageNumber="10">Crateromys schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 62291), right side (mirrored); (F)
|
||
<taxonomicName id="4C0DDF87FFFB482CFC7B2DD443BA01D1" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1015,1208,1952,1973]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFFB482CFC7B2DD443BA01D1" box="[1015,1208,1952,1973]" italics="true" pageId="9" pageNumber="10">Crateromys heaneyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED1C3CC1FFFB482CFB4A2DD443FF01D1" box="[1222,1277,1952,1973]" country="New Zealand" httpUri="http://grbio.org/cool/x09h-x3qj" name="Canterbury Museum" pageId="9" pageNumber="10">CMC</collectionCode>
|
||
M626), left side; (G)
|
||
<taxonomicName id="4C0DDF87FFFB482CFCBF2DCA44C701B7" box="[819,965,1982,2003]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="granti">
|
||
<emphasis id="B9797816FFFB482CFCBF2DCA44C701B7" box="[819,965,1982,2003]" italics="true" pageId="9" pageNumber="10">Batomys granti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="ED1C3CC1FFFB482CFC5E2DCA431B01B7" box="[978,1049,1982,2003]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="9" pageNumber="10" type="Museum">FMNH</collectionCode>
|
||
214412), left side.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2A404FFF8482FFCF42B5D42670444" blockId="10.[861,1531,201,1856]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9797816FFF8482FFCF42B5D43490724" box="[888,1099,297,320]" italics="true" pageId="10" pageNumber="11">Age of specimens.—</emphasis>
|
||
<location id="8ED2F2DFFFF8482FFBC02B5D43CE0724" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFF8482FFBC02B5D43CE0724" box="[1100,1228,297,320]" county="Musang Cave" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Callao Cave" pageId="10" pageNumber="11">Callao Cave</location>
|
||
specimens 10544 and 10196 originated in a layer dated at least ca. 67,000 BP; specimen 11075 is from
|
||
<location id="8ED2F2DFFFF8482FFC752B1D433507E4" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFF8482FFC752B1D433507E4" box="[1017,1079,361,384]" county="Musang Cave" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Layer" pageId="10" pageNumber="11">Layer</location>
|
||
11 of
|
||
<location id="8ED2F2DFFFF8482FFBF02B1D420307E4" LSID="urn:lsid:plazi:treatment:03A41512FFFB482EFC6F2EDD44DE01A3:8ED2F2DFFFF8482FFBF02B1D420307E4" box="[1148,1281,361,384]" county="Musang Cave" latitude="4.5" longLatPrecision="78" longitude="-4.0" municipality="Minori Cave" name="Callao Cave" pageId="10" pageNumber="11">Callao Cave</location>
|
||
, which is older than ca. 30,000 cal BP but younger than 52,000 BP;
|
||
<specimenCount id="9D0B6F8DFFF8482FFACB2BFD42F907C4" box="[1351,1531,393,416]" pageId="10" pageNumber="11" type="generic">seven specimens</specimenCount>
|
||
from
|
||
<collectingMunicipality id="6BD63E7EFFF8482FFC162BDD432707A4" box="[922,1061,425,448]" pageId="10" pageNumber="11">Minori Cave</collectingMunicipality>
|
||
, including the
|
||
<typeStatus id="54B61AA6FFF8482FFB442BDD422A07A4" box="[1224,1320,425,448]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
, are from aceramic
|
||
<collectorName id="26F8C1D2FFF8482FFCD12BBD44C10784" box="[861,963,457,480]" pageId="10" pageNumber="11">Holocene</collectorName>
|
||
levels (prior to the introduction of pottery on
|
||
<collectorName id="26F8C1D2FFF8482FFA3B2BBD42F90784" box="[1463,1531,457,480]" pageId="10" pageNumber="11">Luzon</collectorName>
|
||
at
|
||
<geoCoordinate id="EE39C2C3FFF8482FFCF72B9D44B50464" box="[891,951,489,512]" degrees="4,500" direction="north" orientation="latitude" pageId="10" pageNumber="11" precision="55" value="4.5">4,500</geoCoordinate>
|
||
<geoCoordinate id="EE39C2C3FFF8482FFC3B2B9D43020464" box="[951,1024,489,512]" degrees="4,000" direction="west" orientation="longitude" pageId="10" pageNumber="11" precision="55" value="-4.0">–4,000</geoCoordinate>
|
||
cal BP);
|
||
<collectingCounty id="62D3DC88FFF8482FFBE22B9D42060464" box="[1134,1284,489,512]" pageId="10" pageNumber="11">Musang Cave</collectingCounty>
|
||
specimen 101 derives from aceramic-bearing level (
|
||
<collectorName id="26F8C1D2FFF8482FFB13287D43EC0444" box="[1183,1262,521,544]" pageId="10" pageNumber="11">Level I</collectorName>
|
||
3;
|
||
<tableCitation id="C68F91BFFFF8482FFA8B287D42570444" box="[1287,1365,521,544]" captionStart="Table 1" captionStartId="3.[125,181,1354,1375]" captionTargetPageId="3" captionText="Table 1.—Reported radiometric ages for provenance and study sites of fossil cloud rat specimens. Holotypes are underlined. *Minimum uranium series ages for Callao Layer 14. **Depth of specimen is above the last recorded occurrence of pottery; specimen is not directly associated with dated sample. 14C calibration was done using OxCal 4.3 (Bronk Ramsey 2017) and ranges shown are at 95.4% probability. Numbers in the last three columns are specimen numbers; see lists of holotypes and paratypes in text." httpUri="http://table.plazi.org/id/DF72F48CFFF14826FFF12F3E449803D3" pageId="10" pageNumber="11" tableUuid="DF72F48CFFF14826FFF12F3E449803D3">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFF8482FFCF4285D44A40484" pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph id="8BB2A404FFF8482FFCF4285D44A40484" blockId="10.[861,1531,201,1856]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9797816FFF8482FFCF4285D43050424" box="[888,1031,553,576]" italics="true" pageId="10" pageNumber="11">Etymology.—</emphasis>
|
||
The new species name comes from the languages of the Dupaningan Agta, Itawes, and Ibanag of NE Luzon, an adjective meaning “large” in all three languages (see Tharp and Natividad 1976;
|
||
<bibRefCitation id="EF9CD9F5FFF8482FFB1B28FD428304C4" author="Greenhill" box="[1175,1409,649,672]" etAl="et al." firstAuthor="Greenhill" journalOrPublisher="Evolutionary Bioinformatics Online" pageId="10" pageNumber="11" pagination="271 - 283" part="4" refId="ref17356" refString="GREENHILL, S. J., R. BLUST, AND R. D. GRAY. 2008. The Austronesian basic vocabulary database: from bioinformatics to lexomics. Evolutionary Bioinformatics Online 4: 271 - 283." title="The Austronesian basic vocabulary database: from bioinformatics to lexomics" type="journal article" year="2008">Greenhill et al. 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF9CD9F5FFF8482FFA1828FD449004A4" author="Robinson" firstAuthor="Robinson" journalOrPublisher="Canberra, Australian Capital Territory, Australia" pageId="10" pageNumber="11" refId="ref19117" refString="ROBINSON, L. C. 2011. Dupaningan Agta: grammar, vocabulary, and texts. Pacific Linguistics. Canberra, Australian Capital Territory, Australia." title="Dupaningan Agta: grammar, vocabulary, and texts. Pacific Linguistics" type="book" year="2011">Robinson 2011</bibRefCitation>
|
||
); we use it as an adjective in the nominative singular neuter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFF8482FFCF4289D44B60524" pageId="10" pageNumber="11" type="distribution">
|
||
<paragraph id="8BB2A404FFF8482FFCF4289D44B60524" blockId="10.[861,1531,201,1856]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9797816FFF8482FFCF4289D431B0564" box="[888,1049,745,768]" italics="true" pageId="10" pageNumber="11">Distribution.—</emphasis>
|
||
Known from three cave sites in the Callao Cave complex, Peñablanca Municipality,
|
||
<collectingRegion id="49C96AE6FFF8482FFABE297D42F80544" box="[1330,1530,777,800]" country="Philippines" name="Cagayan" pageId="10" pageNumber="11">Cagayan Province</collectingRegion>
|
||
(
|
||
<figureCitation id="1336B881FFF8482FFCEA295D44A60524" box="[870,932,809,832]" captionStart="Fig" captionStartId="2.[144,179,2005,2026]" captionTargetBox="[228,1445,202,1991]" captionTargetId="figure-13@2.[226,1448,200,1992]" captionTargetPageId="2" captionText="Fig. 1.—Map of the Callao Cave Complex, showing location of cave sites: Callao (1), Eme (2), Dalan Serkot (3), Minori (4), and Musang (5). Inset shows the location within Luzon Island." figureDoi="http://doi.org/10.5281/zenodo.5613302" httpUri="https://zenodo.org/record/5613302/files/figure.png" pageId="10" pageNumber="11">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFF8482FFCF4293D424702A4" pageId="10" pageNumber="11" type="diagnosis">
|
||
<paragraph id="8BB2A404FFF8482FFCF4293D424702A4" blockId="10.[861,1531,201,1856]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9797816FFF8482FFCF4293D43000504" box="[888,1026,841,864]" italics="true" pageId="10" pageNumber="11">Diagnosis.—</emphasis>
|
||
A member of the genus
|
||
<taxonomicName id="4C0DDF87FFF8482FFB73293D426F0504" box="[1279,1389,841,864]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFF8482FFB73293D426F0504" box="[1279,1389,841,864]" italics="true" pageId="10" pageNumber="11">Carpomys</emphasis>
|
||
</taxonomicName>
|
||
as diagnosed by Musser and Heaney (1992) and as emended above (see Emended Diagnosis and Description and Comparisons); a giant species much larger than the two living species,
|
||
<taxonomicName id="4C0DDF87FFF8482FFAD229DD42F905A4" authorityName="Thomas" authorityYear="1895" box="[1374,1531,937,960]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="melanurus">
|
||
<emphasis id="B9797816FFF8482FFAD229DD42F905A4" box="[1374,1531,937,960]" italics="true" pageId="10" pageNumber="11">Ca. melanurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFF8482FFC0129BD431C0584" authorityName="Thomas" authorityYear="1895" box="[909,1054,969,992]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFF8482FFC0129BD431C0584" box="[909,1054,969,992]" italics="true" pageId="10" pageNumber="11">Ca. phaeurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C68F91BFFFF8482FFBBC29BD43820584" box="[1072,1152,969,992]" captionStart="Table 2" captionStartId="6.[124,180,651,672]" captionTargetPageId="6" captionText="Table 2.—Measurements (in mm) of maxillary and mandibular teeth of Carpomys and Musseromys." httpUri="http://table.plazi.org/id/DF72F48CFFF44823FFF028FF433D04C4" pageId="10" pageNumber="11" tableUuid="DF72F48CFFF44823FFF028FF433D04C4">Table 2</tableCitation>
|
||
). The dentary is robust both absolutely and proportionately, with strongly marked inferior and superior masseteric ridges. Lower incisor is unusually wide,
|
||
<quantity id="4CF509E1FFF8482FFCD12E5D44B30224" box="[861,945,1065,1088]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="11" unit="mm" value="2.3">2.3 mm</quantity>
|
||
at its midpoint. Upper toothrow length is
|
||
<quantity id="4CF509E1FFF8482FFA092E5D42F50224" box="[1413,1527,1065,1088]" metricMagnitude="-2" metricUnit="m" metricValue="1.358" pageId="10" pageNumber="11" unit="mm" value="13.58">13.58 mm</quantity>
|
||
; lower toothrow length ranges from
|
||
<quantity id="4CF509E1FFF8482FFB602E3D42C50204" box="[1260,1479,1097,1120]" metricMagnitude="-2" metricUnit="m" metricValue="1.3355" metricValueMax="1.346" metricValueMin="1.325" pageId="10" pageNumber="11" unit="mm" value="13.355" valueMax="13.46" valueMin="13.25">13.25 to 13.46 mm</quantity>
|
||
. M1 length ranges from
|
||
<quantity id="4CF509E1FFF8482FFBBB2E1D43F602E4" box="[1079,1268,1129,1152]" metricMagnitude="-3" metricUnit="m" metricValue="4.99" metricValueMax="5.49" metricValueMin="4.49" pageId="10" pageNumber="11" unit="mm" value="4.99" valueMax="5.49" valueMin="4.49">4.49 to 5.49 mm</quantity>
|
||
; M1 width ranges from
|
||
<quantity id="4CF509E1FFF8482FFCD12EFD431002C4" box="[861,1042,1161,1184]" metricMagnitude="-3" metricUnit="m" metricValue="3.915" metricValueMax="4.41" metricValueMin="3.42" pageId="10" pageNumber="11" unit="mm" value="3.915" valueMax="4.41" valueMin="3.42">3.42 to 4.41 mm</quantity>
|
||
. m1 length ranges from
|
||
<quantity id="4CF509E1FFF8482FFA942EFD42CF02C4" box="[1304,1485,1161,1184]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.57" metricValueMin="5.93" pageId="10" pageNumber="11" unit="mm" value="6.25" valueMax="6.57" valueMin="5.93">5.93 to 6.57 mm</quantity>
|
||
; m1 width ranges from
|
||
<quantity id="4CF509E1FFF8482FFBA42EDD43D502A4" box="[1064,1239,1193,1216]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.53" metricValueMin="3.37" pageId="10" pageNumber="11" unit="mm" value="3.95" valueMax="4.53" valueMin="3.37">3.37 to 4.53 mm</quantity>
|
||
(
|
||
<tableCitation id="C68F91BFFFF8482FFB6B2EDD423702A4" box="[1255,1333,1193,1216]" captionStart="Table 2" captionStartId="6.[124,180,651,672]" captionTargetPageId="6" captionText="Table 2.—Measurements (in mm) of maxillary and mandibular teeth of Carpomys and Musseromys." httpUri="http://table.plazi.org/id/DF72F48CFFF44823FFF028FF433D04C4" pageId="10" pageNumber="11" tableUuid="DF72F48CFFF44823FFF028FF433D04C4">Table 2</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C317F78FFFF8482EFCF42EBD44DE01A3" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="8BB2A404FFF8482EFCF42EBD46D20163" blockId="10.[861,1531,201,1856]" lastBlockId="11.[101,771,1584,1991]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="B9797816FFF8482FFCF42EBD43DF0284" box="[888,1245,1225,1248]" italics="true" pageId="10" pageNumber="11">Description and comparisons.—</emphasis>
|
||
A giant cloud rat, largest known member of
|
||
<taxonomicName id="4C0DDF87FFF8482FFBBC2E9D439C0364" box="[1072,1182,1257,1280]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFF8482FFBBC2E9D439C0364" box="[1072,1182,1257,1280]" italics="true" pageId="10" pageNumber="11">Carpomys</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="C68F91BFFFF8482FFB3D2E9D42000364" box="[1201,1282,1257,1280]" captionStart="Table 2" captionStartId="6.[124,180,651,672]" captionTargetPageId="6" captionText="Table 2.—Measurements (in mm) of maxillary and mandibular teeth of Carpomys and Musseromys." httpUri="http://table.plazi.org/id/DF72F48CFFF44823FFF028FF433D04C4" pageId="10" pageNumber="11" tableUuid="DF72F48CFFF44823FFF028FF433D04C4">Table 2</tableCitation>
|
||
;
|
||
<figureCitation id="1336B881FFF8482FFA982E9D42570364" box="[1300,1365,1257,1280]" captionStart="Fig" captionStartId="11.[101,136,1461,1482]" captionTargetBox="[112,760,207,1445]" captionTargetId="figure-499@11.[105,767,200,1447]" captionTargetPageId="11" captionText="Fig. 6.—Bivariate plots of Carpomys and Musseromys dental and mandibular measurements (in mm). Data are in Table 2." figureDoi="http://doi.org/10.5281/zenodo.5613321" httpUri="https://zenodo.org/record/5613321/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
|
||
), easily distinguished from the two extant species by its much greater size. The dentary is both larger and proportionately more robust than in extant species, with strongly marked inferior and superior masseteric ridges. The mental foramen is moderately high relative to the apex of the masseteric ridges, situated close to the dorsal surface of the mandible and anterior to m1. The lower incisor, as represented on the
|
||
<typeStatus id="54B61AA6FFF8482FFB1F2FBD43F10384" box="[1171,1267,1481,1504]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
, is robust, orange-yellow on the anterior surface,
|
||
<quantity id="4CF509E1FFF8482FFBD82F9D43A10064" box="[1108,1187,1513,1536]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="11" unit="mm" value="2.3">2.3 mm</quantity>
|
||
wide at its midpoint and
|
||
<quantity id="4CF509E1FFF8482FFA272F9D42F80064" box="[1451,1530,1513,1536]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="10" pageNumber="11" unit="mm" value="8.5">8.5 mm</quantity>
|
||
long from its tip to the point it meets the dentary bone along its medial edge. The dorsal wear surface extends fully to the beginning of the dentary bone slightly below the anteriormost projection of bone along the medial edge, and the tip lies slightly below the occlusal plane of the mandibular molars. The lower incisor of
|
||
<taxonomicName id="4C0DDF87FFF8482FFC422CDD436200A4" authorityName="Thomas" authorityYear="1895" box="[974,1120,1705,1728]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFF8482FFC422CDD436200A4" box="[974,1120,1705,1728]" italics="true" pageId="10" pageNumber="11">Ca. phaeurus</emphasis>
|
||
</taxonomicName>
|
||
also is orange-yellow on the anterior surface, but is both shorter and more slender, 1.0 mm wide at its midpoint and 5.0 mm (
|
||
<emphasis id="B9797816FFF8482FFBFD2C9D437C0164" box="[1137,1150,1769,1792]" italics="true" pageId="10" pageNumber="11">n</emphasis>
|
||
= 2) from its tip to the anteriormost projection of bone along the medial edge. In this species, the dorsal wear surface extends from the tip about 90% of the way to the highest point of the dentary bone; the tip lies slightly below the occlusal plane of the mandibular molars, equivalent to
|
||
<taxonomicName id="4C0DDF87FFF9482EFF082C0447ED00E3" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[132,239,1648,1671]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFF082C0447ED00E3" box="[132,239,1648,1671]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
. The lower incisor of
|
||
<taxonomicName id="4C0DDF87FFF9482EFE602C04458F00E3" authorityName="Thomas" authorityYear="1895" box="[492,653,1648,1671]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="melanurus">
|
||
<emphasis id="B9797816FFF9482EFE602C04458F00E3" box="[492,653,1648,1671]" italics="true" pageId="11" pageNumber="12">Ca. melanurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B9797816FFF9482EFD2C2C0445AF00E3" box="[672,685,1648,1671]" italics="true" pageId="11" pageNumber="12">n</emphasis>
|
||
= 1) is
|
||
<quantity id="4CF509E1FFF9482EFFE92CE447BB00C3" box="[101,185,1680,1703]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="12" unit="mm" value="1.3">1.3 mm</quantity>
|
||
wide at the midpoint, and about 6.0 mm from its tip to the anteriormost projection of bone along the medial edge. Thus, the lower incisor of
|
||
<taxonomicName id="4C0DDF87FFF9482EFE092CA446EC0083" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[389,494,1744,1767]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFE092CA446EC0083" box="[389,494,1744,1767]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
is nearly twice the width of those of the two extant species.
|
||
</paragraph>
|
||
<caption id="DF72F48CFFF8482FFF1C2D3D42B501B6" ID-DOI="http://doi.org/10.5281/zenodo.5613314" ID-Zenodo-Dep="5613314" httpUri="https://zenodo.org/record/5613314/files/figure.png" pageId="10" pageNumber="11" startId="10.[144,178,1865,1886]" targetBox="[148,809,201,1850]" targetPageId="10">
|
||
<paragraph id="8BB2A404FFF8482FFF1C2D3D42B501B6" blockId="10.[144,814,1865,2003]" lastBlockId="10.[861,1532,1923,2002]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9797816FFF8482FFF1C2D3D47D2013A" bold="true" box="[144,208,1865,1886]" pageId="10" pageNumber="11">Fig. 5.</emphasis>
|
||
—Occlusal view of mandibular molars of (A)
|
||
<taxonomicName id="4C0DDF87FFF8482FFD1C2D3D442F013A" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[656,813,1865,1886]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="dakal" status="sp. nov.">
|
||
<emphasis id="B9797816FFF8482FFD1C2D3D442F013A" box="[656,813,1865,1886]" italics="true" pageId="10" pageNumber="11">Carpomys dakal</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFF8482FFF1C2D1247C7011F" box="[144,197,1894,1915]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM II-1977-J11A-16301, holotype), leftside; (B)
|
||
<taxonomicName id="4C0DDF87FFF8482FFD402D1247EA01FC" authorityName="Thomas" authorityYear="1895" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFF8482FFD402D1247EA01FC" italics="true" pageId="10" pageNumber="11">Carpomys phaeurus</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 62291), leftside; (C)
|
||
<taxonomicName id="4C0DDF87FFF8482FFD8C2DF745DF01FC" authorityName="Heaney, Balete, Rickart, Veluz & Jansa" authorityYear="2014" box="[512,733,1923,1944]" class="Mammalia" family="Muridae" genus="Musseromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="inopinatus">
|
||
<emphasis id="B9797816FFF8482FFD8C2DF745DF01FC" box="[512,733,1923,1944]" italics="true" pageId="10" pageNumber="11">Musseromysinopinatus</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 193840), left side; (D)
|
||
<taxonomicName id="4C0DDF87FFF8482FFEE02DD5451501D2" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[364,535,1953,1974]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="ballik" status="sp. nov.">
|
||
<emphasis id="B9797816FFF8482FFEE02DD5451501D2" box="[364,535,1953,1974]" italics="true" pageId="10" pageNumber="11">Crateromys ballik</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFF8482FFD912DD5455301D2" box="[541,593,1953,1974]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM II-1977-I8-402, holotype), right side (mirrored); (E)
|
||
<taxonomicName id="4C0DDF87FFF8482FFE642DCA45D901B7" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[488,731,1982,2003]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFF8482FFE642DCA45D901B7" box="[488,731,1982,2003]" italics="true" pageId="10" pageNumber="11">Crateromys schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 62291), left side; (F)
|
||
<taxonomicName id="4C0DDF87FFF8482FFBA62DF743EB01FC" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1066,1257,1923,1944]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFF8482FFBA62DF743EB01FC" box="[1066,1257,1923,1944]" italics="true" pageId="10" pageNumber="11">Crateromys heaneyi</emphasis>
|
||
</taxonomicName>
|
||
(CMC M626), left side; (G)
|
||
<taxonomicName id="4C0DDF87FFF8482FFCD12DD4433501D1" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[861,1079,1952,1973]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="cagayanensis" status="sp. nov.">
|
||
<emphasis id="B9797816FFF8482FFCD12DD4433501D1" box="[861,1079,1952,1973]" italics="true" pageId="10" pageNumber="11">Batomys cagayanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFF8482FFBB22DD4437101D1" box="[1086,1139,1952,1973]" pageId="10" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM
|
||
<date id="FFB382C4FFF8482FFB322DD4420501D1" box="[1214,1287,1952,1973]" pageId="10" pageNumber="11" value="1977-02">II-1977</date>
|
||
-J3-7554,
|
||
<typeStatus id="54B61AA6FFF8482FFAEB2DD442BC01D1" box="[1383,1470,1952,1973]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
), right side (mirrored); (H)
|
||
<taxonomicName id="4C0DDF87FFF8482FFB932DC943B301B6" box="[1055,1201,1981,2002]" class="Mammalia" family="Muridae" genus="Batomys" kingdom="Animalia" order="Rodentia" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="granti">
|
||
<emphasis id="B9797816FFF8482FFB932DC943B301B6" box="[1055,1201,1981,2002]" italics="true" pageId="10" pageNumber="11">Batomys granti</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 214412), left side.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF72F48CFFF9482EFCBF29F344FA0590" ID-DOI="http://doi.org/10.5281/zenodo.5613316" ID-Zenodo-Dep="5613316" httpUri="https://zenodo.org/record/5613316/files/figure.png" pageId="11" pageNumber="12" startId="11.[819,854,903,924]" targetBox="[822,1484,200,890]" targetPageId="11">
|
||
<paragraph id="8BB2A404FFF9482EFCBF29F344FA0590" blockId="11.[819,1488,903,1012]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9797816FFF9482EFCBF29F3447305F8" bold="true" box="[819,881,903,924]" pageId="11" pageNumber="12">Fig. 7.</emphasis>
|
||
—Lateral lingual (A) and labial (B) view of the partial left maxillary of
|
||
<taxonomicName id="4C0DDF87FFF9482EFC0A29D1432705DE" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[902,1061,933,954]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal" status="sp. nov.">
|
||
<emphasis id="B9797816FFF9482EFC0A29D1432705DE" box="[902,1061,933,954]" italics="true" pageId="11" pageNumber="12">Carpomys dakal</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFF9482EFBA729D1436205DE" box="[1067,1120,933,954]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
|
||
(PNM II-1977-J11A-15997) with M1 to M3, showing the posterior palatine foramen (ppf) and base of the zygomatic arch (za).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF72F48CFFF9482EFFE92FC1457F0383" ID-DOI="http://doi.org/10.5281/zenodo.5613321" ID-Zenodo-Dep="5613321" httpUri="https://zenodo.org/record/5613321/files/figure.png" pageId="11" pageNumber="12" startId="11.[101,136,1461,1482]" targetBox="[112,760,207,1445]" targetPageId="11">
|
||
<paragraph id="8BB2A404FFF9482EFFE92FC1457F0383" blockId="11.[101,771,1461,1511]" pageId="11" pageNumber="12">
|
||
<emphasis id="B9797816FFF9482EFFE92FC147A803AE" bold="true" box="[101,170,1461,1482]" pageId="11" pageNumber="12">Fig. 6.</emphasis>
|
||
—Bivariate plots of
|
||
<taxonomicName id="4C0DDF87FFF9482EFEF02FC146DD03AE" box="[380,479,1461,1482]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFF9482EFEF02FC146DD03AE" box="[380,479,1461,1482]" italics="true" pageId="11" pageNumber="12">Carpomys</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFF9482EFD942FC1458D03AE" box="[536,655,1461,1482]" class="Mammalia" family="Muridae" genus="Musseromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFF9482EFD942FC1458D03AE" box="[536,655,1461,1482]" italics="true" pageId="11" pageNumber="12">Musseromys</emphasis>
|
||
</taxonomicName>
|
||
dental and mandibular measurements (in mm). Data are in Table 2.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB2A404FFF9482EFF0C2D6442C403A3" blockId="11.[101,771,1584,1991]" lastBlockId="11.[819,1489,1104,1991]" pageId="11" pageNumber="12">
|
||
Overall molar morphology follows that shown in
|
||
<figureCitation id="1336B881FFF9482EFDF22D6447860123" captionStart="Fig" captionStartId="4.[861,896,1502,1523]" captionTargetBox="[867,1527,202,1488]" captionTargetId="figure-650@4.[865,1527,200,1489]" captionTargetPageId="4" captionText="Fig. 2.—Maxillary and mandibular left molar toothrows of Carpomys dakal n. sp. (A and B, respectively; PNM II-1977-J11A-15997 and PNM II-1976-M-101) and Crateromys schadenbergi (C and D, respectively; FMNH 62295), showing the location of cusps and conids referred to in the text. pc = posterior cingulum." figureDoi="http://doi.org/10.5281/zenodo.5613304" httpUri="https://zenodo.org/record/5613304/files/figure.png" pageId="11" pageNumber="12">Figs. 2A and 2B</figureCitation>
|
||
. Measurements of individual teeth are provided in
|
||
<tableCitation id="C68F91BFFFF9482EFD212D4445FE0123" box="[685,764,1840,1863]" captionStart="Table 3" captionStartId="7.[124,180,568,589]" captionTargetPageId="7" captionText="Table 3.—Measurements (in mm) of maxillary and mandibular teeth of Crateromys and Batomys. Measurements for Crateromys paulus are summarized as range, mean, and standard deviation (from Reyes et al. 2017). *Estimated value on damaged specimen." httpUri="http://table.plazi.org/id/DF72F48CFFF54822FFF0284C4442040F" pageId="11" pageNumber="12" tableUuid="DF72F48CFFF54822FFF0284C4442040F">Table 3</tableCitation>
|
||
. Molar dimensions of the two living
|
||
<taxonomicName id="4C0DDF87FFF9482EFE512D24459E0103" box="[477,668,1872,1895]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9797816FFF9482EFE512D24454B0103" box="[477,585,1872,1895]" italics="true" pageId="11" pageNumber="12">Carpomys</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
comprise only 40–80% of corresponding measurements in
|
||
<taxonomicName id="4C0DDF87FFF9482EFD1C2D0445FE01E3" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[656,764,1904,1927]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFD1C2D0445FE01E3" box="[656,764,1904,1927]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
. Molar dimensions of
|
||
<taxonomicName id="4C0DDF87FFF9482EFEC12DE4452601C3" box="[333,548,1936,1959]" class="Mammalia" family="Muridae" genus="Musseromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9797816FFF9482EFEC12DE446CD01C3" box="[333,463,1936,1959]" italics="true" pageId="11" pageNumber="12">Musseromys</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are only 20–30% of the corresponding measurements in
|
||
<emphasis id="B9797816FFF9482EFE792DC4456501A3" box="[501,615,1968,1991]" italics="true" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C0DDF87FFF9482EFE792DC4456601A3" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[501,612,1968,1991]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">Ca. dakal</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The variation in molar characters observed in
|
||
<taxonomicName id="4C0DDF87FFF9482EFB1E2E2443FE0203" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1170,1276,1104,1127]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFB1E2E2443FE0203" box="[1170,1276,1104,1127]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
also is seen in
|
||
<taxonomicName id="4C0DDF87FFF9482EFA272E2444A002E3" authorityName="Thomas" authorityYear="1895" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="melanurus">
|
||
<emphasis id="B9797816FFF9482EFA272E2444A002E3" italics="true" pageId="11" pageNumber="12">Ca. melanurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFF9482EFC512E04436D02E3" authorityName="Thomas" authorityYear="1895" box="[989,1135,1136,1159]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFF9482EFC512E04436D02E3" box="[989,1135,1136,1159]" italics="true" pageId="11" pageNumber="12">Ca. phaeurus</emphasis>
|
||
</taxonomicName>
|
||
; size is the main distinguishing character that differentiates the dental morphology of fossil species from the living species. Additional cusplets are often present in M1 and M2 of
|
||
<taxonomicName id="4C0DDF87FFF9482EFBC32EA443B90283" box="[1103,1211,1232,1255]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFF9482EFBC32EA443B90283" box="[1103,1211,1232,1255]" italics="true" pageId="11" pageNumber="12">Carpomys</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1336B881FFF9482EFB412EA442220283" box="[1229,1312,1232,1255]" captionStart="Fig" captionStartId="9.[101,136,1865,1886]" captionTargetBox="[106,767,201,1850]" captionTargetId="figure-521@9.[105,767,200,1851]" captionTargetPageId="9" captionText="Fig. 4.—Occlusal view of maxillary molars of (A) Carpomys dakal n. sp. (PNM II-1977-J11A-15997), left side; (B) Carpomys phaeurus (FMNH62291),leftside;(C) Musseromysinopinatus(FMNH193840), left side; (D) Crateromys ballik n. sp. (PNM II-1977-J3-10389), right side (mirrored); (E) Crateromys schadenbergi (FMNH 62291), right side (mirrored); (F) Crateromys heaneyi (CMC M626), left side; (G) Batomys granti (FMNH 214412), left side." figureDoi="http://doi.org/10.5281/zenodo.5613310" httpUri="https://zenodo.org/record/5613310/files/figure.png" pageId="11" pageNumber="12">Fig. 4A</figureCitation>
|
||
; Supplementary Data SD1B and C): a labial cusplet between t3 and t6 of M1 and a smaller cusplet between t5 and t8 of M2, which are absent in other phloeomyine genera. The M1 cusplet is found in three out of five
|
||
<taxonomicName id="4C0DDF87FFF9482EFC222F24431A0303" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[942,1048,1360,1383]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFC222F24431A0303" box="[942,1048,1360,1383]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
specimens, compared to two out of five
|
||
<taxonomicName id="4C0DDF87FFF9482EFCBF2F0444C003E3" authorityName="Thomas" authorityYear="1895" box="[819,962,1392,1415]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFF9482EFCBF2F0444C003E3" box="[819,962,1392,1415]" italics="true" pageId="11" pageNumber="12">Ca. phaeurus</emphasis>
|
||
</taxonomicName>
|
||
specimens and none in three
|
||
<taxonomicName id="4C0DDF87FFF9482EFA8C2F04429E03E3" authorityName="Thomas" authorityYear="1895" box="[1280,1436,1392,1415]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="melanurus">
|
||
<emphasis id="B9797816FFF9482EFA8C2F04429E03E3" box="[1280,1436,1392,1415]" italics="true" pageId="11" pageNumber="12">Ca. melanurus</emphasis>
|
||
</taxonomicName>
|
||
specimens. The M2 cusplet is present in one of
|
||
<specimenCount id="9D0B6F8DFFF9482EFA992FE442B003C3" box="[1301,1458,1424,1447]" pageId="11" pageNumber="12" type="generic">five specimens</specimenCount>
|
||
of
|
||
<taxonomicName id="4C0DDF87FFF9482EFCBF2FC4449B03A3" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[819,921,1456,1479]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFCBF2FC4449B03A3" box="[819,921,1456,1479]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
and is absent in specimens of the two extant species.
|
||
</paragraph>
|
||
<paragraph id="8BB2A404FFF9482EFCC12FA444DE01A3" blockId="11.[819,1489,1104,1991]" pageId="11" pageNumber="12">
|
||
Molar tooth dimensions of
|
||
<taxonomicName id="4C0DDF87FFF9482EFB092FA443F10383" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1157,1267,1488,1511]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFB092FA443F10383" box="[1157,1267,1488,1511]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
are within the size range of the giant cloud rats
|
||
<taxonomicName id="4C0DDF87FFF9482EFBEF2F84421F0063" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1123,1309,1520,1543]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFF9482EFBEF2F84421F0063" box="[1123,1309,1520,1543]" italics="true" pageId="11" pageNumber="12">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFF9482EFAC22F8442CB0063" authorityName="Gonzales & Kennedy" authorityYear="1996" box="[1358,1481,1520,1543]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="heaneyi">
|
||
<emphasis id="B9797816FFF9482EFAC22F8442CB0063" box="[1358,1481,1520,1543]" italics="true" pageId="11" pageNumber="12">Cr. heaneyi</emphasis>
|
||
</taxonomicName>
|
||
, but compared to them,
|
||
<taxonomicName id="4C0DDF87FFF9482EFBBF2C64439D0043" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[1075,1183,1552,1575]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="dakal">
|
||
<emphasis id="B9797816FFF9482EFBBF2C64439D0043" box="[1075,1183,1552,1575]" italics="true" pageId="11" pageNumber="12">Ca. dakal</emphasis>
|
||
</taxonomicName>
|
||
has a more slender dentary (
|
||
<figureCitation id="1336B881FFF9482EFCB02C4444790023" box="[828,891,1584,1607]" captionStart="Fig" captionStartId="8.[144,179,1893,1914]" captionTargetBox="[271,1405,888,1880]" captionTargetId="figure-359@8.[271,1405,888,1880]" captionTargetPageId="8" captionText="Fig. 3.—Holotype of Carpomys dakal n. sp. (PNM II-1977-J11A-16301), left dentary: (A) labial view of mandible; (B) lingual view of mandible (mirrored)." figureDoi="http://doi.org/10.5281/zenodo.5613308" httpUri="https://zenodo.org/record/5613308/files/figure.png" pageId="11" pageNumber="12">Fig. 3</figureCitation>
|
||
). The maxillary region is robust (
|
||
<figureCitation id="1336B881FFF9482EFB662C44422B0023" box="[1258,1321,1584,1607]" captionStart="Fig" captionStartId="11.[819,854,903,924]" captionTargetBox="[822,1484,200,890]" captionTargetId="figure-458@11.[822,1484,200,890]" captionTargetPageId="11" captionText="Fig. 7.—Lateral lingual (A) and labial (B) view of the partial left maxillary of Carpomys dakal n. sp. (PNM II-1977-J11A-15997) with M1 to M3, showing the posterior palatine foramen (ppf) and base of the zygomatic arch (za)." figureDoi="http://doi.org/10.5281/zenodo.5613316" httpUri="https://zenodo.org/record/5613316/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
|
||
); our fragmentary material implies that the palate is rather smooth with a raised ridge supporting the molar teeth, and a large posterior palatal foramen (ppf) adjacent to M2 (
|
||
<figureCitation id="1336B881FFF9482EFB612CE4424100C3" box="[1261,1347,1680,1703]" captionStart="Fig" captionStartId="11.[819,854,903,924]" captionTargetBox="[822,1484,200,890]" captionTargetId="figure-458@11.[822,1484,200,890]" captionTargetPageId="11" captionText="Fig. 7.—Lateral lingual (A) and labial (B) view of the partial left maxillary of Carpomys dakal n. sp. (PNM II-1977-J11A-15997) with M1 to M3, showing the posterior palatine foramen (ppf) and base of the zygomatic arch (za)." figureDoi="http://doi.org/10.5281/zenodo.5613316" httpUri="https://zenodo.org/record/5613316/files/figure.png" pageId="11" pageNumber="12">Fig. 7A</figureCitation>
|
||
), as seen in the two extant
|
||
<taxonomicName id="4C0DDF87FFF9482EFC572CC443AE00A3" box="[987,1196,1712,1735]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9797816FFF9482EFC572CC4434900A3" box="[987,1099,1712,1735]" italics="true" pageId="11" pageNumber="12">Carpomys</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
No trace of the deep palatal grooves of
|
||
<taxonomicName id="4C0DDF87FFF9482EFC572CA4438D0083" box="[987,1167,1744,1767]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B9797816FFF9482EFC572CA443590083" box="[987,1115,1744,1767]" italics="true" pageId="11" pageNumber="12">Crateromys</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
is evident. It appears that the bony palate terminates about one-third of the way posterior to the anterior edge of M3, as in
|
||
<taxonomicName id="4C0DDF87FFF9482EFB2A2D64424B0143" authorityName="Thomas" authorityYear="1895" box="[1190,1353,1808,1831]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="melanurus">
|
||
<emphasis id="B9797816FFF9482EFB2A2D64424B0143" box="[1190,1353,1808,1831]" italics="true" pageId="11" pageNumber="12">Ca. melanurus</emphasis>
|
||
</taxonomicName>
|
||
, rather than at the anterior edge of M3, as in
|
||
<taxonomicName id="4C0DDF87FFF9482EFB2F2D44423B0123" authorityName="Thomas" authorityYear="1895" box="[1187,1337,1840,1863]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="phaeurus">
|
||
<emphasis id="B9797816FFF9482EFB2F2D44423B0123" box="[1187,1337,1840,1863]" italics="true" pageId="11" pageNumber="12">Ca. phaeurus</emphasis>
|
||
</taxonomicName>
|
||
. The anterior root of the zygomatic arch appears to originate labial to M1 (
|
||
<figureCitation id="1336B881FFF9482EFCB02D04448F01E3" box="[828,909,1904,1927]" captionStart="Fig" captionStartId="11.[819,854,903,924]" captionTargetBox="[822,1484,200,890]" captionTargetId="figure-458@11.[822,1484,200,890]" captionTargetPageId="11" captionText="Fig. 7.—Lateral lingual (A) and labial (B) view of the partial left maxillary of Carpomys dakal n. sp. (PNM II-1977-J11A-15997) with M1 to M3, showing the posterior palatine foramen (ppf) and base of the zygomatic arch (za)." figureDoi="http://doi.org/10.5281/zenodo.5613316" httpUri="https://zenodo.org/record/5613316/files/figure.png" pageId="11" pageNumber="12">Fig. 7B</figureCitation>
|
||
), similar to the extant species of
|
||
<taxonomicName id="4C0DDF87FFF9482EFB742D04426A01E3" box="[1272,1384,1904,1927]" class="Mammalia" family="Muridae" genus="Carpomys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="genus">
|
||
<emphasis id="B9797816FFF9482EFB742D04426A01E3" box="[1272,1384,1904,1927]" italics="true" pageId="11" pageNumber="12">Carpomys</emphasis>
|
||
</taxonomicName>
|
||
, and perhaps a bit further posterior than that of
|
||
<taxonomicName id="4C0DDF87FFF9482EFB6A2DE442A001C3" baseAuthorityName="Meyer" baseAuthorityYear="1895" box="[1254,1442,1936,1959]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="schadenbergi">
|
||
<emphasis id="B9797816FFF9482EFB6A2DE442A001C3" box="[1254,1442,1936,1959]" italics="true" pageId="11" pageNumber="12">Cr. schadenbergi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0DDF87FFF9482EFCBF2DC4449801A3" authority="Ochoa & Mijares & Piper & Reyes & Heaney, 2021" authorityName="Ochoa & Mijares & Piper & Reyes & Heaney" authorityYear="2021" box="[819,922,1968,1991]" class="Mammalia" family="Muridae" genus="Crateromys" kingdom="Animalia" order="Rodentia" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="ballik" status="sp. nov.">
|
||
<emphasis id="B9797816FFF9482EFCBF2DC4449801A3" box="[819,922,1968,1991]" italics="true" pageId="11" pageNumber="12">Cr. ballik</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24AC56DFFF9482EFC2D2DC444DE01A3" box="[929,988,1968,1991]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |