300 lines
35 KiB
XML
300 lines
35 KiB
XML
<document id="E5DC6DEAF266D217A66A724405061A39" ID-DOI="10.11646/zootaxa.4048.3.10" ID-GBIF-Dataset="18d6aadb-aee7-4061-990b-83b89769ff2b" ID-ISSN="1175-5326" ID-Zenodo-Dep="240927" ID-ZooBank="7625A700-4D29-4AE7-9292-E4EE0DD787A8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461347966251" checkinUser="plazi" docAuthor="Yin, Zi-Wei, Coulon, Georges & Bekchiev, Rostislav" docDate="2015" docId="039C925F3453E110FF213D71FB9F7EB1" docLanguage="en" docName="zt04048p450.pdf" docOrigin="Zootaxa 4048 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudophanias spinitarsis Yin, Coulon & Bekchiev, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="449" masterDocId="FFA5EA273452E113FFB63C16FFB87A50" masterDocTitle="A new species of Pseudophanias Raffray from a cave in central Nepal (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="450" masterPageNumber="446" pageNumber="447" updateTime="1698620977355" updateUser="plazi">
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<mods:mods id="08F76742C25B7BA67A400D23CCD39EA1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A12CE2D1E7794FB609F6B1E7B1FEF76E">
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<mods:title id="29F5630D017FC421F1FAF0F24CAEBD89">A new species of Pseudophanias Raffray from a cave in central Nepal (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="C502C1C0C9A8EBBBB9B57090DF4D59B5" type="personal">
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<mods:role id="2A0EBBEB991BCB13CCF6E280A3EB44C5">
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<mods:roleTerm id="AC93D5F891791649F943DB516110740A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E73B1AE192754063D3A3566AE698AFA8">Yin, Zi-Wei</mods:namePart>
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</mods:name>
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<mods:name id="F0502B28CCC9A0E5B2A087A6CC71F081" type="personal">
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<mods:roleTerm id="C0A21F9365390A66F9B1DF8BA8DFE37B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7282DB5968C432B7A3ADEA544D8F073C">Coulon, Georges</mods:namePart>
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</mods:name>
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<mods:name id="B3A4A02C14A7E163BDFDF1AF9C9A39A5" type="personal">
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<mods:role id="B6A88676E1A861C62C8132C0549B8BD1">
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<mods:roleTerm id="AACBCF8534C1E62E9F9CF6C8118FD9B0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A88BF9108E872171970FFD54C59B458E">Bekchiev, Rostislav</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="B58298AED6B1D5471C591D150F8FF2DB">text</mods:typeOfResource>
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<mods:titleInfo id="15BDEEF74FCC035E8A066AF948D23135">
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<mods:title id="C5555BFCF79208447D0E4C694E3C500D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CB7BBB8DA86933D49308C9C395D2D0A7">
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<mods:date id="7A8737C6E42EAD0B81650A3C09EDD572">2015</mods:date>
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<mods:detail id="4C46BA0399BE52C02A1889935D7EC7C4" type="volume">
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<mods:number id="DA428B4D9CAB0BF62E01A8ECF74B6D24">4048</mods:number>
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</mods:detail>
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<mods:detail id="4B99C4C007DAFB792A2EEF67A6B6835F" type="issue">
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<mods:number id="8ACFE3067C9203CF6F081DBB711CED7C">3</mods:number>
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<mods:start id="A1485254BBC7EF13271FC991AACB120B">446</mods:start>
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<mods:classification id="F24A100B5F95092B59C3FEC058221677">journal article</mods:classification>
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<mods:identifier id="A4926AA7D136F23E72EE76E4F8C7FB43" type="DOI">10.11646/zootaxa.4048.3.10</mods:identifier>
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<mods:identifier id="2D157E5FBD43C79DD34CC46DEBCB019A" type="GBIF-Dataset">18d6aadb-aee7-4061-990b-83b89769ff2b</mods:identifier>
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<mods:identifier id="B844D76A08A11A2DB7B57FA07F6B499B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="46243F2B1B4B304F23AA889FC9B339E8" type="Zenodo-Dep">240927</mods:identifier>
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<treatment id="039C925F3453E110FF213D71FB9F7EB1" ID-DOI="http://doi.org/10.5281/zenodo.6100377" ID-GBIF-Taxon="119645443" ID-Zenodo-Dep="6100377" LSID="urn:lsid:plazi:treatment:039C925F3453E110FF213D71FB9F7EB1" httpUri="http://treatment.plazi.org/id/039C925F3453E110FF213D71FB9F7EB1" lastPageId="3" lastPageNumber="449" pageId="1" pageNumber="447">
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<subSubSection id="C32F70C23453E112FF213D71FD427BD2" box="[151,762,359,386]" pageId="1" pageNumber="447" type="nomenclature">
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<paragraph id="8B8A23493453E112FF213D71FD427BD2" blockId="1.[151,762,359,386]" box="[151,762,359,386]" pageId="1" pageNumber="447">
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<heading id="D0C294253453E112FF213D71FD427BD2" bold="true" box="[151,762,359,386]" fontSize="11" level="1" pageId="1" pageNumber="447" reason="1">
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<emphasis id="B941FF5B3453E112FF213D71FD427BD2" bold="true" box="[151,762,359,386]" pageId="1" pageNumber="447">
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<taxonomicName id="4C3558CA3453E112FF213D71FE697BD1" ID-CoL="4P6TG" box="[151,465,359,385]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="spinitarsis" status="sp. nov.">
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<emphasis id="B941FF5B3453E112FF213D71FE697BD1" bold="true" box="[151,465,359,385]" italics="true" pageId="1" pageNumber="447">Pseudophanias spinitarsis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27242203453E112FE563D7EFDC87BD2" box="[480,624,360,386]" pageId="1" pageNumber="447" rank="species">new species</taxonomicNameLabel>
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(
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<figureCitation id="130E3FCC3453E112FD363D7EFD497BD2" box="[640,753,360,386]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,244,1994,2016]" captionStartId-1="2.[151,244,1649,1671]" captionTargetBox-0="[155,1421,718,1966]" captionTargetBox-1="[160,1427,191,1622]" captionTargetId-0="figure@1.[151,1435,714,1970]" captionTargetId-1="figure@2.[151,1435,191,1627]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Dorsal habitus of Pseudophanias spinitarsis. A. Male. B. Female. Scales: 1.0 mm." captionText-1="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri-0="https://zenodo.org/record/240928/files/figure.png" httpUri-1="https://zenodo.org/record/240929/files/figure.png" pageId="1" pageNumber="447">Figs. 1–2</figureCitation>
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||
)
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32F70C23453E112FF213DBBFED77859" pageId="1" pageNumber="447" type="materials_examined">
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||
<paragraph id="8B8A23493453E112FF213DBBFED77859" blockId="1.[151,1436,429,695]" pageId="1" pageNumber="447">
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||
<emphasis id="B941FF5B3453E112FF213DBBFE8E7B94" bold="true" box="[151,310,429,452]" pageId="1" pageNumber="447">
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<typeStatus id="548E9DEB3453E112FF213DBBFF777B94" box="[151,207,429,452]" pageId="1" pageNumber="447">Type</typeStatus>
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material
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</emphasis>
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(5 ♂♂, 6 ♀♀).
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||
<emphasis id="B941FF5B3453E112FE523DBBFD067B94" bold="true" box="[484,702,429,452]" pageId="1" pageNumber="447">
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||
<typeStatus id="548E9DEB3453E112FE523DBBFDE87B94" box="[484,592,429,452]" pageId="1" pageNumber="447" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="F32263D93453E112FDEA3DBBFD017B94" box="[604,697,429,452]" name="Nepal" pageId="1" pageNumber="447">NEPAL</collectingCountry>
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:
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</emphasis>
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♂, labeled ‘
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<collectingCountry id="F32263D93453E112FC893DBBFC3E7B94" box="[831,902,429,452]" name="Nepal" pageId="1" pageNumber="447">Népal</collectingCountry>
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,
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<date id="FF8B05893453E112FC273DBBFC6F7B94" box="[913,983,429,452]" pageId="1" pageNumber="447">29.vii.</date>
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(19)81, Pokhara,
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||
<quantity id="4CCD8EAC3453E112FB393DBBFB6E7B94" box="[1167,1238,429,452]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="447" unit="m" value="900.0">900 m</quantity>
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||
, Mahendra Gupha (cave),
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<geoCoordinate id="EE01458E3453E112FF513DC6FEC17BB7" box="[231,377,464,487]" direction="north" orientation="latitude" pageId="1" pageNumber="447" precision="1" value="28.271389">28.271389°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE01458E3453E112FE323DC6FDAB7BB7" box="[388,531,464,487]" direction="east" orientation="longitude" pageId="1" pageNumber="447" precision="1" value="83.97972">83.979722°E</geoCoordinate>
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, P. Beron’ (
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||
<collectionCode id="ED24BB8C3453E112FD273DC6FD4F7BB7" box="[657,759,464,487]" pageId="1" pageNumber="447">NMNHS</collectionCode>
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||
).
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||
<emphasis id="B941FF5B3453E112FCBF3DC6FC547BB7" bold="true" box="[777,1004,464,487]" pageId="1" pageNumber="447">
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||
<typeStatus id="548E9DEB3453E112FCBF3DC6FCC77BB7" box="[777,895,464,487]" pageId="1" pageNumber="447" type="paratype">Paratypes</typeStatus>
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:
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||
<collectingCountry id="F32263D93453E112FC3D3DC6FC507BB7" box="[907,1000,464,487]" name="Nepal" pageId="1" pageNumber="447">NEPAL</collectingCountry>
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||
:
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||
</emphasis>
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||
4 ♂♂, 6 ♀♀, same data as the
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||
<typeStatus id="548E9DEB3453E112FA883DC6FA237BB7" box="[1342,1435,464,487]" pageId="1" pageNumber="447" type="holotype">holotype</typeStatus>
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||
(
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||
<collectionCode id="ED24BB8C3453E112FF283DE4FEBB7859" box="[158,259,498,521]" pageId="1" pageNumber="447">NMNHS</collectionCode>
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||
,
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||
<collectionCode id="ED24BB8C3453E112FEBB3DE4FEDC7859" box="[269,356,498,521]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="1" pageNumber="447">MHNG</collectionCode>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C32F70C23453E112FF713E03FCE778E7" pageId="1" pageNumber="447" type="diagnosis">
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||
<paragraph id="8B8A23493453E112FF713E03FCE778E7" blockId="1.[151,1436,429,695]" pageId="1" pageNumber="447">
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||
<emphasis id="B941FF5B3453E112FF713E03FE84787C" bold="true" box="[199,316,533,556]" pageId="1" pageNumber="447">Diagnosis.</emphasis>
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Body length over
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||
<quantity id="4CCD8EAC3453E112FDB33E03FDFE787C" box="[517,582,533,556]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="447" unit="mm" value="3.0">3 mm</quantity>
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, with elongate habitus and appendages; pronotal disc bearing a distinct conical spine; elytra trapezoidal, each elytron with a shallow and wide discal stria; metaventrite with two small triangular tubercles; tergite IV more than twice as long as tergite V. Male has antennomeres VIII and IX, protarsomeres II and III, mesotarsomeres II, and base of the profemora modified; symmetric aedeagus with median lobe greatly extended ventrally. Female has simple antennae, tarsomeres, and profemora.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="DF4A73C13453E112FF213BDCFB917D8F" box="[151,1065,1994,2016]" httpUri="https://zenodo.org/record/240928/files/figure.png" pageId="1" pageNumber="447" targetBox="[155,1421,718,1966]" targetPageId="1">
|
||
<paragraph id="8B8A23493453E112FF213BDCFB917D8F" blockId="1.[151,1065,1994,2016]" box="[151,1065,1994,2016]" pageId="1" pageNumber="447">
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||
<emphasis id="B941FF5B3453E112FF213BDCFEB47DB0" bold="true" box="[151,268,1994,2016]" pageId="1" pageNumber="447">FIGURE 1.</emphasis>
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Dorsal habitus of
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<taxonomicName id="4C3558CA3453E112FE0B3BDCFD017D8F" box="[445,697,1994,2015]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="447" phylum="Arthropoda" rank="species" species="spinitarsis">
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||
<emphasis id="B941FF5B3453E112FE0B3BDCFD017D8F" box="[445,697,1994,2015]" italics="true" pageId="1" pageNumber="447">Pseudophanias spinitarsis</emphasis>
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</taxonomicName>
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||
.
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<emphasis id="B941FF5B3453E112FD733BDCFD657DB0" bold="true" box="[709,733,1994,2016]" pageId="1" pageNumber="447">A.</emphasis>
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Male.
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<emphasis id="B941FF5B3453E112FC963BDCFC8E7DB0" bold="true" box="[800,822,1994,2016]" pageId="1" pageNumber="447">B.</emphasis>
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Female. Scales: 1.0 mm.
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</paragraph>
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</caption>
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<caption id="DF4A73C13450E111FF213A67FD857C91" httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="2" pageNumber="448" targetBox="[160,1427,191,1622]" targetPageId="2">
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||
<paragraph id="8B8A23493450E111FF213A67FD857C91" blockId="2.[151,1436,1649,1729]" pageId="2" pageNumber="448">
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||
<emphasis id="B941FF5B3450E111FF213A67FEB47CD7" bold="true" box="[151,268,1649,1671]" pageId="2" pageNumber="448">FIGURE 2.</emphasis>
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Diagnostic features of male
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<taxonomicName id="4C3558CA3450E111FD963A67FCA47CD6" box="[544,796,1649,1670]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="448" phylum="Arthropoda" rank="species" species="spinitarsis">
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||
<emphasis id="B941FF5B3450E111FD963A67FCA47CD6" box="[544,796,1649,1670]" italics="true" pageId="2" pageNumber="448">Pseudophanias spinitarsis</emphasis>
|
||
</taxonomicName>
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||
.
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||
<emphasis id="B941FF5B3450E111FC9E3A67FCE47CD7" bold="true" box="[808,860,1649,1671]" pageId="2" pageNumber="448">A, B.</emphasis>
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||
Antennomeres VIII–XI.
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||
<emphasis id="B941FF5B3450E111FBF83A67FBDD7CD7" bold="true" box="[1102,1125,1649,1671]" pageId="2" pageNumber="448">C.</emphasis>
|
||
Maxillary palpus.
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||
<emphasis id="B941FF5B3450E111FAAC3A67FA8A7CD7" bold="true" box="[1306,1330,1649,1671]" pageId="2" pageNumber="448">D.</emphasis>
|
||
Pronotum, in lateral view.
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||
<emphasis id="B941FF5B3450E111FE9A3A98FEFA7CF4" bold="true" box="[300,322,1678,1700]" pageId="2" pageNumber="448">E.</emphasis>
|
||
Profemur.
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||
<emphasis id="B941FF5B3450E111FE1B3A98FE627CF4" bold="true" box="[429,474,1678,1700]" pageId="2" pageNumber="448">F, G.</emphasis>
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||
Protarsus.
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||
<emphasis id="B941FF5B3450E111FDFE3A98FDD97CF4" bold="true" box="[584,609,1678,1700]" pageId="2" pageNumber="448">H.</emphasis>
|
||
Mesotarsus.
|
||
<emphasis id="B941FF5B3450E111FD573A98FD487CF4" bold="true" box="[737,752,1678,1700]" pageId="2" pageNumber="448">I.</emphasis>
|
||
Aedeagus, in lateral view.
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||
<emphasis id="B941FF5B3450E111FC403A98FBB07CF4" bold="true" box="[1014,1032,1678,1700]" pageId="2" pageNumber="448">J.</emphasis>
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Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F–H = 0.1 mm; C, I, J = 0.2 mm.
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||
</paragraph>
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||
</caption>
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||
<subSubSection id="C32F70C23450E110FF713AFAFA3F799C" lastPageId="3" lastPageNumber="449" pageId="2" pageNumber="448" type="description">
|
||
<paragraph id="8B8A23493450E110FF713AFAFCDE7889" blockId="2.[151,1436,1772,2003]" lastBlockId="3.[151,1436,152,1249]" lastPageId="3" lastPageNumber="449" pageId="2" pageNumber="448">
|
||
<emphasis id="B941FF5B3450E111FF713AFAFEEA7D53" bold="true" box="[199,338,1772,1795]" pageId="2" pageNumber="448">Description.</emphasis>
|
||
Male (
|
||
<figureCitation id="130E3FCC3450E111FE113AFAFE507D53" box="[423,488,1772,1795]" captionStart="FIGURE 1" captionStartId="1.[151,244,1994,2016]" captionTargetBox="[155,1421,718,1966]" captionTargetId="figure@1.[151,1435,714,1970]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Pseudophanias spinitarsis. A. Male. B. Female. Scales: 1.0 mm." httpUri="https://zenodo.org/record/240928/files/figure.png" pageId="2" pageNumber="448">Fig. 1</figureCitation>
|
||
A). Body length
|
||
<quantity id="4CCD8EAC3450E111FD1A3AFAFCE87D53" box="[684,848,1772,1795]" metricMagnitude="-3" metricUnit="m" metricValue="3.16" metricValueMax="3.2" metricValueMin="3.12" pageId="2" pageNumber="448" unit="mm" value="3.16" valueMax="3.2" valueMin="3.12">3.12–3.20 mm</quantity>
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||
. Integuments of dorsal surface without granulations, punctation not ocellate. Punctation of head strong, coarse, confluent at antennal tubercles, punctures well separated from each other, but close, becoming small towards eyes and cheeks; frontal rostrum bearing only few punctures, frons and vertex virtually impunctate. Punctation of pronotum strong laterally and especially at base, confluent, becoming fine and sparse on disk. Punctation of elytra fine and sparse. Pubescence long. Head elongate, HL
|
||
<quantity id="4CCD8EAC3450E111FBE33B61FB4A7DDE" box="[1109,1266,1911,1934]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.4" metricValueMin="6.3" pageId="2" pageNumber="448" unit="mm" value="0.635" valueMax="0.64" valueMin="0.63">0.63–0.64 mm</quantity>
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||
, HW
|
||
<quantity id="4CCD8EAC3450E111FA873B61FF7C7DE0" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.5" metricValueMin="5.3" pageId="2" pageNumber="448" unit="mm" value="0.54" valueMax="0.55" valueMin="0.53">0.53–0.55 mm</quantity>
|
||
; frontal rostrum narrow, setose sub-antennal C-shaped sulci clearly seen in dorsal view; each eye composed of about 50 facets. Temples long, oblique, weakly rounded. Antennae long and slender, antennal club formed by apical four antennomeres; antennomeres each elongate, scape as long as antennomeres II–IV combined, antennomeres VIII angularly expanded laterally, IX triangularly expanded (
|
||
<figureCitation id="130E3FCC3451E110FD4A3CACFC867A81" box="[764,830,186,209]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">Fig. 2</figureCitation>
|
||
A, 2B). Maxillary palpi (
|
||
<figureCitation id="130E3FCC3451E110FBE43CACFB2C7A81" box="[1106,1172,186,209]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">Fig. 2</figureCitation>
|
||
C) simple, palpomeres I minute, II (
|
||
<quantity id="4CCD8EAC3451E110FEA63CCBFECB7AA4" box="[272,371,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="449" unit="mm" value="0.34">0.34 mm</quantity>
|
||
) elongate, slightly pedunculate at basal half, III (
|
||
<quantity id="4CCD8EAC3451E110FC333CCBFC507AA4" box="[901,1000,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="449" unit="mm" value="0.16">0.16 mm</quantity>
|
||
) widest at apex, IV (
|
||
<quantity id="4CCD8EAC3451E110FB7D3CCBFA967AA4" box="[1227,1326,221,244]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="449" unit="mm" value="0.33">0.33 mm</quantity>
|
||
) lengthily fusiform. Pronotum (
|
||
<figureCitation id="130E3FCC3451E110FECA3D16FE067B47" box="[380,446,256,279]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">Fig. 2</figureCitation>
|
||
D) slightly wider than long, PL
|
||
<quantity id="4CCD8EAC3451E110FCA93D16FC077B47" box="[799,959,256,279]" metricMagnitude="-4" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="3" pageNumber="449" unit="mm" value="0.625" valueMax="0.64" valueMin="0.61">0.61–0.64 mm</quantity>
|
||
, PW
|
||
<quantity id="4CCD8EAC3451E110FC4A3D16FB247B47" box="[1020,1180,256,279]" metricMagnitude="-4" metricUnit="m" metricValue="5.7" metricValueMax="5.8" metricValueMin="5.6" pageId="3" pageNumber="449" unit="mm" value="0.57" valueMax="0.58" valueMin="0.56">0.56–0.58 mm</quantity>
|
||
, widest at near middle, constricted at apex, gradually narrowed from middle toward base, disc with strong, conical spine. Elytra trapezoidal, EL
|
||
<quantity id="4CCD8EAC3451E110FF213D53FE8E7B0C" box="[151,310,325,348]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="8.9" metricValueMin="8.8" pageId="3" pageNumber="449" unit="mm" value="0.885" valueMax="0.89" valueMin="0.88">0.88–0.89 mm</quantity>
|
||
, EW
|
||
<quantity id="4CCD8EAC3451E110FEC53D53FDAA7B0C" box="[371,530,325,348]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.2" metricValueMin="1.16" pageId="3" pageNumber="449" unit="mm" value="1.18" valueMax="1.2" valueMin="1.16">1.16–1.20 mm</quantity>
|
||
, widest at posterior third; discal stria shallow and wide, extending to half of elytral length. Metaventrite impunctate, elevated, with two small triangular, dull tubercles surrounding shallow impression. All legs extremely elongate and slender; protibiae straight, slightly broadened at apex; mesotibiae (length:
|
||
<quantity id="4CCD8EAC3451E110FB613D9CFA8F7BF1" box="[1239,1335,394,417]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" pageId="3" pageNumber="449" unit="mm" value="1.04">1.04 mm</quantity>
|
||
) straight, slightly sinuated inwards at apical third; metatibiae (length:
|
||
<quantity id="4CCD8EAC3451E110FCAC3DBBFCC37B94" box="[794,891,429,452]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="3" pageNumber="449" unit="mm" value="1.26">1.26 mm</quantity>
|
||
) straight, slightly broadened at apex; protarsomere II and III (
|
||
<figureCitation id="130E3FCC3451E110FEBF3DC6FEF17BB7" box="[265,329,464,487]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">Fig. 2</figureCitation>
|
||
F, 2G), and mesotarsomere II (
|
||
<figureCitation id="130E3FCC3451E110FD203DC6FD6E7BB7" box="[662,726,464,487]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">Fig. 2</figureCitation>
|
||
H) each spinose; profemora (
|
||
<figureCitation id="130E3FCC3451E110FBA73DC6FBE97BB7" box="[1041,1105,464,487]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">Fig. 2</figureCitation>
|
||
E) concave at basal third, with bunch of thick setae. Abdomen broadest at tergite IV (first visible tergite), AL 1.00–
|
||
<quantity id="4CCD8EAC3451E110FBA33DE4FBCD7859" box="[1045,1141,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="3" pageNumber="449" unit="mm" value="1.03">1.03 mm</quantity>
|
||
, AW
|
||
<quantity id="4CCD8EAC3451E110FB063DE4FAF67859" box="[1200,1358,498,521]" metricMagnitude="-3" metricUnit="m" metricValue="1.165" metricValueMax="1.17" metricValueMin="1.16" pageId="3" pageNumber="449" unit="mm" value="1.165" valueMax="1.17" valueMin="1.16">1.16–1.17 mm</quantity>
|
||
; tergite IV more than twice as long as next one (
|
||
<quantity id="4CCD8EAC3451E110FDF73E03FD18787C" box="[577,672,533,556]" metricMagnitude="-4" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="449" unit="mm" value="0.51">0.51 mm</quantity>
|
||
vs.
|
||
<quantity id="4CCD8EAC3451E110FD7D3E03FC95787C" box="[715,813,533,556]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" pageId="3" pageNumber="449" unit="mm" value="0.24">0.24 mm</quantity>
|
||
), lacking median carina; discal carinae short, but thick and conspicuous. Aedeagus (
|
||
<figureCitation id="130E3FCC3451E110FE113E2EFE53781F" box="[423,491,568,591]" captionStart="FIGURE 2" captionStartId="2.[151,244,1649,1671]" captionTargetBox="[160,1427,191,1622]" captionTargetId="figure@2.[151,1435,191,1627]" captionTargetPageId="2" captionText="FIGURE 2. Diagnostic features of male Pseudophanias spinitarsis. A, B. Antennomeres VIII – XI. C. Maxillary palpus. D. Pronotum, in lateral view. E. Profemur. F, G. Protarsus. H. Mesotarsus. I. Aedeagus, in lateral view. J. Same, in ventral view. Scales: A, B, D, E = 0.3 mm; F – H = 0.1 mm; C, I, J = 0.2 mm." httpUri="https://zenodo.org/record/240929/files/figure.png" pageId="3" pageNumber="449">
|
||
Fig.
|
||
<date id="FF8B05893451E110FE6A3E2EFE53781F" box="[476,491,568,591]" pageId="3" pageNumber="449">2</date>
|
||
</figureCitation>
|
||
I, 2J) moderately sclerotized, symmetric in dorso-ventral view, length
|
||
<quantity id="4CCD8EAC3451E110FB403E2EFA2F781F" box="[1270,1431,568,591]" metricMagnitude="-4" metricUnit="m" metricValue="6.55" metricValueMax="6.6" metricValueMin="6.5" pageId="3" pageNumber="449" unit="mm" value="0.655" valueMax="0.66" valueMin="0.65">0.65–0.66 mm</quantity>
|
||
; basal part strongly curved ventrally, parameres greatly extended basolaterally, each paramere bearing two preapical and two apical setae; median lobe strongly extended ventrally, composed of two elongate, weakly-sclerotized lateral lobes, and broad median plate which is demarcated by two parts, basal part narrowed at base and broadened apically, apical part nearly trapezoidal, broadly and deeply emarginate at apical margin.
|
||
</paragraph>
|
||
<paragraph id="8B8A23493451E110FF713EF3FC047911" blockId="3.[151,1436,152,1249]" pageId="3" pageNumber="449">
|
||
Female (
|
||
<figureCitation id="130E3FCC3451E110FE913EF3FED078AC" box="[295,360,741,764]" captionStart="FIGURE 1" captionStartId="1.[151,244,1994,2016]" captionTargetBox="[155,1421,718,1966]" captionTargetId="figure@1.[151,1435,714,1970]" captionTargetPageId="1" captionText="FIGURE 1. Dorsal habitus of Pseudophanias spinitarsis. A. Male. B. Female. Scales: 1.0 mm." httpUri="https://zenodo.org/record/240928/files/figure.png" pageId="3" pageNumber="449">Fig. 1</figureCitation>
|
||
B). Generally similar to male, antennae and legs simple. Each eye composed of about 40 facets. Measurements: BL
|
||
<quantity id="4CCD8EAC3451E110FEDA3F1EFDB2794F" box="[364,522,776,799]" metricMagnitude="-3" metricUnit="m" metricValue="3.235" metricValueMax="3.36" metricValueMin="3.11" pageId="3" pageNumber="449" unit="mm" value="3.235" valueMax="3.36" valueMin="3.11">3.11–3.36 mm</quantity>
|
||
, HL 0.66–
|
||
<quantity id="4CCD8EAC3451E110FDC93F1EFD59794F" box="[639,737,776,799]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="449" unit="mm" value="0.65">0.65 mm</quantity>
|
||
, HW 0.53–
|
||
<quantity id="4CCD8EAC3451E110FCEB3F1EFC79794F" box="[861,961,776,799]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="3" pageNumber="449" unit="mm" value="0.52">0.52 mm</quantity>
|
||
, PL
|
||
<quantity id="4CCD8EAC3451E110FC423F1EFB2B794F" box="[1012,1171,776,799]" metricMagnitude="-4" metricUnit="m" metricValue="6.35" metricValueMax="6.4" metricValueMin="6.3" pageId="3" pageNumber="449" unit="mm" value="0.635" valueMax="0.64" valueMin="0.63">0.63–0.64 mm</quantity>
|
||
, PW 0.57–
|
||
<quantity id="4CCD8EAC3451E110FABD3F1EFAD6794F" box="[1291,1390,776,799]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="449" unit="mm" value="0.56">0.56 mm</quantity>
|
||
, EL
|
||
<quantity id="4CCD8EAC3451E110FF213F3CFE8C7911" box="[151,308,810,833]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" metricValueMax="8.8" metricValueMin="8.4" pageId="3" pageNumber="449" unit="mm" value="0.86" valueMax="0.88" valueMin="0.84">0.84–0.88 mm</quantity>
|
||
, EW
|
||
<quantity id="4CCD8EAC3451E110FED93F3CFDB47911" box="[367,524,810,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.1549999999999998" metricValueMax="1.18" metricValueMin="1.13" pageId="3" pageNumber="449" unit="mm" value="1.1549999999999998" valueMax="1.18" valueMin="1.13">1.13–1.18 mm</quantity>
|
||
, AL
|
||
<quantity id="4CCD8EAC3451E110FDF73F3CFD667911" box="[577,734,810,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.085" metricValueMax="1.19" metricValueMin="0.98" pageId="3" pageNumber="449" unit="mm" value="1.085" valueMax="1.19" valueMin="0.98">0.98–1.19 mm</quantity>
|
||
, AW 1.20–
|
||
<quantity id="4CCD8EAC3451E110FCE13F3CFC007911" box="[855,952,810,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="3" pageNumber="449" unit="mm" value="1.17">1.17 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8A23493451E110FF713F5BFA3F799C" blockId="3.[151,1436,152,1249]" pageId="3" pageNumber="449">
|
||
<emphasis id="B941FF5B3451E110FF713F5BFE1F7934" bold="true" box="[199,423,845,868]" pageId="3" pageNumber="449">Comparative notes.</emphasis>
|
||
Adults of most known species of
|
||
<taxonomicName id="4C3558CA3451E110FCA93F5BFC797934" box="[799,961,845,868]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B941FF5B3451E110FCA93F5BFC797934" box="[799,961,845,868]" italics="true" pageId="3" pageNumber="449">Pseudophanias</emphasis>
|
||
</taxonomicName>
|
||
have a relatively stout habitus, thus can be readily distinguished from the new species described here.
|
||
<taxonomicName id="4C3558CA3451E110FCA73F66FB9279D7" box="[785,1066,880,903]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="species" species="spinitarsis">
|
||
<emphasis id="B941FF5B3451E110FCA73F66FB9279D7" box="[785,1066,880,903]" italics="true" pageId="3" pageNumber="449">Pseudophanias spinitarsis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to the Sumatran
|
||
<taxonomicName id="4C3558CA3451E110FA3E3F66FE8779F9" authority="Raffray" authorityName="Raffray" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B941FF5B3451E110FA3E3F66FA2D79D7" box="[1416,1429,880,903]" italics="true" pageId="3" pageNumber="449">P</emphasis>
|
||
.
|
||
<emphasis id="B941FF5B3451E110FF213F84FF5079F9" box="[151,232,914,937]" italics="true" pageId="3" pageNumber="449">elegans</emphasis>
|
||
Raffray
|
||
</taxonomicName>
|
||
which also has a more or less elongate general form. These two species can be separated by the distinctly larger body size (>
|
||
<quantity id="4CCD8EAC3451E110FED63FA3FE24799C" box="[352,412,949,972]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="449" unit="mm" value="3.0">3 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C3558CA3451E110FE093FA3FDF0799C" box="[447,584,949,972]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="species" species="spinitarsis">
|
||
<emphasis id="B941FF5B3451E110FE093FA3FE74799C" box="[447,460,949,972]" italics="true" pageId="3" pageNumber="449">P</emphasis>
|
||
.
|
||
<emphasis id="B941FF5B3451E110FE6F3FA3FDF0799C" box="[473,584,949,972]" italics="true" pageId="3" pageNumber="449">spinitarsis</emphasis>
|
||
</taxonomicName>
|
||
vs.
|
||
<quantity id="4CCD8EAC3451E110FDC53FA3FD08799C" box="[627,688,949,972]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="449" unit="mm" value="2.0">2 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C3558CA3451E110FD643FA3FC86799C" box="[722,830,949,972]" class="Insecta" family="Staphylinidae" genus="Pseudophanias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="449" phylum="Arthropoda" rank="species" species="elegans">
|
||
<emphasis id="B941FF5B3451E110FD643FA3FD67799C" box="[722,735,949,972]" italics="true" pageId="3" pageNumber="449">P</emphasis>
|
||
.
|
||
<emphasis id="B941FF5B3451E110FD5B3FA3FC86799C" box="[749,830,949,972]" italics="true" pageId="3" pageNumber="449">elegans</emphasis>
|
||
</taxonomicName>
|
||
), and relatively much longer scape of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F70C23451E110FF713FCEFBE97E64" pageId="3" pageNumber="449" type="distribution">
|
||
<paragraph id="8B8A23493451E110FF713FCEFDD979BF" blockId="3.[151,1436,152,1249]" box="[199,609,984,1007]" pageId="3" pageNumber="449">
|
||
<emphasis id="B941FF5B3451E110FF713FCEFEE179BF" bold="true" box="[199,345,984,1007]" pageId="3" pageNumber="449">Distribution.</emphasis>
|
||
Central
|
||
<collectingCountry id="F32263D93451E110FE023FCEFE4079BF" box="[436,504,984,1007]" name="Nepal" pageId="3" pageNumber="449">Nepal</collectingCountry>
|
||
: Pokhara.
|
||
</paragraph>
|
||
<paragraph id="8B8A23493451E110FF713FECFBE97E64" blockId="3.[151,1436,152,1249]" pageId="3" pageNumber="449">
|
||
<emphasis id="B941FF5B3451E110FF713FECFE9F7E41" bold="true" box="[199,295,1018,1041]" pageId="3" pageNumber="449">Ecology.</emphasis>
|
||
All individuals were found in the deeper part of the cave (P. Beron pers. comm.), but the presence of welldeveloped eyes and hind wings reveals a modest hypogean adaptation in the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32F70C23451E110FF713856FB9F7EB1" pageId="3" pageNumber="449" type="etymology">
|
||
<paragraph id="8B8A23493451E110FF713856FB347E07" blockId="3.[151,1436,152,1249]" box="[199,1164,1088,1111]" pageId="3" pageNumber="449">
|
||
<emphasis id="B941FF5B3451E110FF713856FEFF7E07" bold="true" box="[199,327,1088,1111]" pageId="3" pageNumber="449">Etymology.</emphasis>
|
||
The epithet refers to the modified male pro- and mesotarsi of the new species.
|
||
</paragraph>
|
||
<paragraph id="8B8A23493451E110FF713874FB9F7EB1" blockId="3.[151,1436,152,1249]" pageId="3" pageNumber="449">
|
||
<emphasis id="B941FF5B3451E110FF713874FE267E29" bold="true" box="[199,414,1122,1145]" pageId="3" pageNumber="449">Acknowledgments.</emphasis>
|
||
The senior author thanks Giulio Cuccodoro and Ivan Löbl for their hospitality and logical support when the author visited the MHNG in
|
||
<date id="FF8B05893451E110FD353893FC9C7ECC" box="[643,804,1157,1180]" pageId="3" pageNumber="449" value="2012" valueMax="2015">2012 and 2015</date>
|
||
. An anonymous reviewer is thanked for commenting on the manuscript. The present study was supported by the National Science Foundation of
|
||
<collectingCountry id="F32263D93451E110FBF238BEFB3C7EEF" box="[1092,1156,1192,1215]" name="China" pageId="3" pageNumber="449">China</collectingCountry>
|
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(No. 31501874), and the Science and Technology Commission of Shanghai Municipality (No. 15YF1408700).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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</document> |