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<document id="2138754AAAB20E28A9A6DD5D2ADEF1EC" ID-DOI="10.11646/zootaxa.3915.3.1" ID-GBIF-Dataset="49559d7f-3675-4921-9d62-692c4ff89e88" ID-ISSN="1175-5326" ID-Zenodo-Dep="240898" ID-ZooBank="BBA4F5B8-F240-41F9-9DC5-E64A66E4FA0D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460970807921" checkinUser="plazi" docAuthor="Kits, Joel H. &amp; Marshall, Stephen A." docDate="2015" docId="039C2B7EBD15FFB3FF73FB3DFAE44397" docLanguage="en" docName="zt03915p355.pdf" docOrigin="Zootaxa 3915 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Boreantrops machinator Kits &amp; Marshall, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="341" masterDocId="FFA55306BD33FF9BFFE4FFC9FF84426D" masterDocTitle="A revision of Boreantrops Kits &amp; Marshall (Diptera: Sphaeroceridae: Archiborborinae)" masterLastPageNumber="355" masterPageNumber="301" pageNumber="339" updateTime="1698558812934" updateUser="plazi">
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<mods:title id="42F500369B01166D7672509A5866523B">A revision of Boreantrops Kits &amp; Marshall (Diptera: Sphaeroceridae: Archiborborinae)</mods:title>
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<mods:namePart id="55B7C9014A7AFF716A70887CC1F57A9B">Kits, Joel H.</mods:namePart>
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<mods:namePart id="BA82F4B55B30E332C154E3305838C86A">Marshall, Stephen A.</mods:namePart>
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<mods:date id="762B337BCD3218FE999E030A6D6FED0A">2015</mods:date>
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<treatment id="039C2B7EBD15FFB3FF73FB3DFAE44397" ID-DOI="http://doi.org/10.5281/zenodo.6095929" ID-GBIF-Taxon="119593923" ID-Zenodo-Dep="6095929" LSID="urn:lsid:plazi:treatment:039C2B7EBD15FFB3FF73FB3DFAE44397" httpUri="http://treatment.plazi.org/id/039C2B7EBD15FFB3FF73FB3DFAE44397" lastPageId="40" lastPageNumber="341" pageId="38" pageNumber="339">
<subSubSection id="C32FC9E3BD15FFBDFF73FB3DFE3D4742" pageId="38" pageNumber="339" type="nomenclature">
<paragraph id="8B8A9A68BD15FFBDFF73FB3DFDA14763" blockId="38.[151,549,1268,1327]" box="[151,549,1268,1294]" pageId="38" pageNumber="339">
<heading id="D0C22D04BD15FFBDFF73FB3DFDA14763" bold="true" box="[151,549,1268,1294]" fontSize="11" level="1" pageId="38" pageNumber="339" reason="1">
<emphasis id="B941467ABD15FFBDFF73FB3DFDA14763" bold="true" box="[151,549,1268,1294]" pageId="38" pageNumber="339">
<taxonomicName id="4C35E1EBBD15FFBDFF73FB3DFE444763" ID-CoL="9HYMV" box="[151,448,1268,1294]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="machinator" status="sp. nov.">
<emphasis id="B941467ABD15FFBDFF73FB3DFE444763" bold="true" box="[151,448,1268,1294]" italics="true" pageId="38" pageNumber="339">Boreantrops machinator</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A272FB01BD15FFBDFE23FB3DFDA14763" box="[455,549,1268,1294]" pageId="38" pageNumber="339" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8A9A68BD15FFBDFF73FADFFE3D4742" blockId="38.[151,549,1268,1327]" box="[151,441,1302,1327]" pageId="38" pageNumber="339">(Figs. 19, 86, 87, 88, 125)</paragraph>
</subSubSection>
<subSubSection id="C32FC9E3BD15FFBDFF73FA97FE514788" pageId="38" pageNumber="339" type="diagnosis">
<paragraph id="8B8A9A68BD15FFBDFF73FA97FE514788" blockId="38.[151,1437,1374,2024]" pageId="38" pageNumber="339">
<emphasis id="B941467ABD15FFBDFF73FA97FE95471A" bold="true" box="[151,273,1374,1399]" pageId="38" pageNumber="339">Diagnosis.</emphasis>
Separable from other members of the
<taxonomicName id="4C35E1EBBD15FFBDFD2BFA96FCDE471B" box="[719,858,1375,1398]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="339" phylum="Arthropoda" rank="species" species="emarginatus">
<emphasis id="B941467ABD15FFBDFD2BFA96FCDE471B" box="[719,858,1375,1398]" italics="true" pageId="38" pageNumber="339">emarginatus</emphasis>
</taxonomicName>
group by the following characters: Occiput black, prementum reddish brown. Anterior margin of anepisternum shiny below spiracle. Mid tibia without row of anterodorsal bristles.
<collectingCountry id="F322DAF8BD15FFBDFE6AFA6FFE4247D2" box="[398,454,1446,1471]" name="India" pageId="38" pageNumber="339">Hind</collectingCountry>
tibia with one ventroapical bristle. Wing with white spots on vein R4+5 basal to r-m crossvein and on vein CuA1.
</paragraph>
</subSubSection>
<subSubSection id="C32FC9E3BD15FFBCFF23FA3DFE1445D9" lastPageId="39" lastPageNumber="340" pageId="38" pageNumber="339" type="description">
<paragraph id="8B8A9A68BD15FFBDFF23FA3DFD5744AC" blockId="38.[151,1437,1374,2024]" pageId="38" pageNumber="339">
<emphasis id="B941467ABD15FFBDFF23FA3DFEDD4460" bold="true" box="[199,345,1524,1549]" pageId="38" pageNumber="339">Description.</emphasis>
Occiput black, frons dark brown posteriorly, reddish brown anteriorly, face, gena, prementum, scape, and pedicel reddish brown, first flagellomere orange. Occiput and gena covered with microtomentum, frons with microtomentum on orbital plate extending medially to interfrontal setae and on frontal triangle extending anteriorly to ocellar bristles, with some scattered microtomentum to ptilinal suture, face with microtomentum below lunule and in dense crescent below antenna. Ocellar bristles at level of median ocellus. Subvibrissal and anterior genal bristle about 0.3X length of vibrissa.
</paragraph>
<paragraph id="8B8A9A68BD15FFBDFF23F905FE22453C" blockId="38.[151,1437,1374,2024]" pageId="38" pageNumber="339">Thorax black, mostly covered with microtomentum. Proepisternum shiny, anepisternum mostly shiny with microtomentum covering dorsal quarter and posterior third, katepisternum with a shiny spot behind fore coxa, meron and metapleuron shiny with band of microtomentum between posterior spiracle and hind coxa. Halter white, pale brown below knob.</paragraph>
<paragraph id="8B8A9A68BD15FFBDFF23F894FC3845F4" blockId="38.[151,1437,1374,2024]" pageId="38" pageNumber="339">
Legs black, trochanters and joints dark reddish brown, tarsi yellow. Mid tibia with 1 anteroventral, 6 subapical bristles.
<collectingCountry id="F322DAF8BD15FFBDFF12F849FEAA45F4" box="[246,302,1920,1945]" name="India" pageId="38" pageNumber="339">Hind</collectingCountry>
tibia with 1 long thin anteroventral, 1 ventroapical bristle.
</paragraph>
<paragraph id="8B8A9A68BD15FFBDFF23F86DFAF5458E" blockId="38.[151,1437,1374,2024]" pageId="38" pageNumber="339">Wing brown, veins dark brown. White spots on crossveins r-m, bm-cu, and dm-cu, vein R4+5 with 1 spot basal to crossvein r-m and 2 distal to crossvein r-m, vein CuA1 with 1 medial spot. Vein R2+3 evenly curved (Fig. 19).</paragraph>
<caption id="DF4ACAE0BD14FFBCFF73FA71FF754447" httpUri="https://zenodo.org/record/240912/files/figure.png" pageId="39" pageNumber="340" targetBox="[154,1426,198,1427]" targetPageId="39">
<paragraph id="8B8A9A68BD14FFBCFF73FA71FF754447" blockId="39.[151,1436,1464,1578]" pageId="39" pageNumber="340">
<emphasis id="B941467ABD14FFBCFF73FA71FED247A0" bold="true" box="[151,342,1464,1487]" pageId="39" pageNumber="340">FIGURES 8692.</emphasis>
<emphasis id="B941467ABD14FFBCFEB9FA73FE5E47A2" box="[349,474,1466,1487]" italics="true" pageId="39" pageNumber="340">Boreantrops</emphasis>
male terminalia.
<taxonomicName id="4C35E1EBBD14FFBCFD74FA73FCA647A3" box="[656,802,1464,1487]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="machinator">
<emphasis id="B941467ABD14FFBCFD74FA73FCA647A3" box="[656,802,1464,1487]" italics="true" pageId="39" pageNumber="340">B. machinator</emphasis>
</taxonomicName>
: 86) phallus, postgonite, and phallapodeme, lateral view, 87) sternite 5, 88) right surstylus, interior view;
<taxonomicName id="4C35E1EBBD14FFBCFDBAFA11FD3E4780" box="[606,698,1495,1517]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="pollex">
<emphasis id="B941467ABD14FFBCFDBAFA11FD3E4780" box="[606,698,1495,1517]" italics="true" pageId="39" pageNumber="340">B. pollex</emphasis>
</taxonomicName>
: 89) phallus, postgonite, and phallapodeme, lateral view, 90) sternite 5, 91) right surstylus, interior view;
<taxonomicName id="4C35E1EBBD14FFBCFE1AFA3EFD1B4461" box="[510,671,1527,1548]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="340" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B941467ABD14FFBCFE1AFA3EFD1B4461" box="[510,671,1527,1548]" italics="true" pageId="39" pageNumber="340">B. punctipennis</emphasis>
</taxonomicName>
: 92) phallus, postgonite, and phallapodeme, lateral view. sls: spike
<emphasis id="B941467ABD14FFBCFA8DFA3FFAF14466" bold="true" box="[1385,1397,1526,1547]" pageId="39" pageNumber="340"></emphasis>
like sclerites.
</paragraph>
</caption>
<paragraph id="8B8A9A68BD14FFBCFF23F99EFA9C44F9" blockId="39.[151,1437,1623,2008]" pageId="39" pageNumber="340">Abdomen with tergites and sternites weakly sclerotized. Syntergite 1+2 with posterolateral corners and a thin band between them strongly sclerotized, tergites 34 with a thin posterior band moderately sclerotized.</paragraph>
<paragraph id="8B8A9A68BD14FFBCFF23F956FDD845FD" blockId="39.[151,1437,1623,2008]" pageId="39" pageNumber="340">
Male postabdomen: Sternite 5 with lateral margins strongly rounded, posterior margin deeply notched with a sharp point on either side of notch, anterior apodeme longer than external portion, without keel (
<figureCitation id="130E86EDBD14FFBCFAD1F90AFA0B44B1" box="[1333,1423,1731,1756]" captionStart="FIGURES 86 92" captionStartId="39.[151,264,1464,1487]" captionTargetBox="[154,1426,198,1427]" captionTargetId="figure@39.[151,1436,193,1443]" captionTargetPageId="39" captionText="FIGURES 86 92. Boreantrops male terminalia. B. machinator: 86) phallus, postgonite, and phallapodeme, lateral view, 87) sternite 5, 88) right surstylus, interior view; B. pollex: 89) phallus, postgonite, and phallapodeme, lateral view, 90) sternite 5, 91) right surstylus, interior view; B. punctipennis: 92) phallus, postgonite, and phallapodeme, lateral view. sls: spike like sclerites." httpUri="https://zenodo.org/record/240912/files/figure.png" pageId="39" pageNumber="340">Fig. 87</figureCitation>
). Epandrium fused above cerci. Surstylus with anterior thumb-like projection, expanded basally, ventral portion narrow (
<figureCitation id="130E86EDBD14FFBCFF11F8C2FECD4549" box="[245,329,1803,1828]" captionStart="FIGURES 86 92" captionStartId="39.[151,264,1464,1487]" captionTargetBox="[154,1426,198,1427]" captionTargetId="figure@39.[151,1436,193,1443]" captionTargetPageId="39" captionText="FIGURES 86 92. Boreantrops male terminalia. B. machinator: 86) phallus, postgonite, and phallapodeme, lateral view, 87) sternite 5, 88) right surstylus, interior view; B. pollex: 89) phallus, postgonite, and phallapodeme, lateral view, 90) sternite 5, 91) right surstylus, interior view; B. punctipennis: 92) phallus, postgonite, and phallapodeme, lateral view. sls: spike like sclerites." httpUri="https://zenodo.org/record/240912/files/figure.png" pageId="39" pageNumber="340">Fig. 88</figureCitation>
). Pregonite not fused with postgonite, fairly large. Postgonite very narrow, posterior margin with a rounded projection, lobes not separated, posterior lobe longer. Basiphallus with long, thin epiphallus, preepiphallus very short. Distiphallus with a long dorsal tube with spines near base, flanked by two spike-like sclerites, distal sclerite not projecting dorsally (
<figureCitation id="130E86EDBD14FFBCFE18F8BEFDD445FD" box="[508,592,1911,1936]" captionStart="FIGURES 86 92" captionStartId="39.[151,264,1464,1487]" captionTargetBox="[154,1426,198,1427]" captionTargetId="figure@39.[151,1436,193,1443]" captionTargetPageId="39" captionText="FIGURES 86 92. Boreantrops male terminalia. B. machinator: 86) phallus, postgonite, and phallapodeme, lateral view, 87) sternite 5, 88) right surstylus, interior view; B. pollex: 89) phallus, postgonite, and phallapodeme, lateral view, 90) sternite 5, 91) right surstylus, interior view; B. punctipennis: 92) phallus, postgonite, and phallapodeme, lateral view. sls: spike like sclerites." httpUri="https://zenodo.org/record/240912/files/figure.png" pageId="39" pageNumber="340">Fig. 86</figureCitation>
).
</paragraph>
<paragraph id="8B8A9A68BD14FFBCFF23F852FE1445D9" blockId="39.[151,1437,1623,2008]" box="[199,400,1947,1972]" pageId="39" pageNumber="340">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C32FC9E3BD14FFB3FF23F876FF7042B9" lastPageId="40" lastPageNumber="341" pageId="39" pageNumber="340" type="materials_examined">
<paragraph id="8B8A9A68BD14FFB3FF23F876FF7042B9" blockId="39.[151,1437,1623,2008]" lastBlockId="40.[151,1436,151,506]" lastPageId="40" lastPageNumber="341" pageId="39" pageNumber="340">
<emphasis id="B941467ABD14FFBCFF23F876FEF145B5" bold="true" box="[199,373,1983,2008]" pageId="39" pageNumber="340">
<typeStatus id="548E24CABD14FFBCFF23F876FE8645B5" box="[199,258,1983,2008]" pageId="39" pageNumber="340">Type</typeStatus>
material.
</emphasis>
<typeStatus id="548E24CABD14FFBCFE98F876FE6045B5" box="[380,484,1983,2008]" pageId="39" pageNumber="340" type="holotype">Holotype</typeStatus>
♂:
<emphasis id="B941467ABD14FFBCFDEBF876FD3145B5" bold="true" box="[527,693,1983,2008]" pageId="39" pageNumber="340">
<collectingCountry id="F322DAF8BD14FFBCFDEBF876FDDB45B5" box="[527,607,1983,2008]" name="Peru" pageId="39" pageNumber="340">PERU</collectingCountry>
: Cusco
</emphasis>
:
<taxonomicName id="4C35E1EBBD14FFBCFD20F876FCBC45B5" box="[708,824,1983,2008]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="353" phylum="Arthropoda" rank="species" species="wayqecha">Wayqecha</taxonomicName>
Biol. Stn., ~
<quantity id="4CCD378DBD14FFBCFC23F876FB8445B5" box="[967,1024,1983,2008]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="39" pageNumber="340" unit="km" value="9.0">9 km</quantity>
NE
<taxonomicName id="4C35E1EBBD14FFBCFBD0F876FB4F45B5" box="[1076,1227,1983,2008]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="352" phylum="Arthropoda" rank="species" species="challabamba">Challabamba</taxonomicName>
,
<geoCoordinate id="EE01FCAFBD14FFBCFB32F876FAAA45BA" box="[1238,1326,1983,2007]" direction="south" orientation="latitude" pageId="39" pageNumber="340" precision="925" value="-13.166667">13°10'S</geoCoordinate>
<geoCoordinate id="EE01FCAFBD14FFBCFAD1F876FA1C45BA" box="[1333,1432,1983,2007]" direction="west" orientation="longitude" pageId="39" pageNumber="340" precision="925" value="-71.566666">71°34'W</geoCoordinate>
,
<quantity id="4CCD378DBD1BFFB3FF73FF5EFEBC42C2" box="[151,312,151,176]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="40" pageNumber="341" unit="m" value="2650.0" valueMax="2700.0" valueMin="2600.0">26002700 m</quantity>
,
<date id="FF8BBCA8BD1BFFB3FEACFF5EFE6B42DD" box="[328,495,151,176]" pageId="40" pageNumber="341" value="2011-12-01" valueMax="2011-12-06" valueMin="2011-12-01">16 Dec 2011</date>
, S.A.
<collectingCountry id="F322DAF8BD1BFFB3FDDEFF5EFD2442DD" box="[570,672,151,176]" name="Marshall Islands" pageId="40" pageNumber="341">Marshall</collectingCountry>
, debu00351213 (
<collectionCode id="ED2402ADBD1BFFB3FC8DFF51FC4142DD" box="[873,965,152,176]" pageId="40" pageNumber="341">MUSM</collectionCode>
).
<typeStatus id="548E24CABD1BFFB3FC38FF51FBC642DD" box="[988,1090,152,176]" pageId="40" pageNumber="341" type="paratype">Paratype</typeStatus>
: same data as
<typeStatus id="548E24CABD1BFFB3FB15FF5EFAD742DD" box="[1265,1363,151,176]" pageId="40" pageNumber="341" type="holotype">holotype</typeStatus>
(
<specimenCount id="9D3351E1BD1BFFB3FA8DFF51FA1342C2" box="[1385,1431,152,176]" pageId="40" pageNumber="341" type="generic">1 ♂</specimenCount>
,
<collectionCode id="ED2402ADBD1BFFB3FF73FF75FF6C42B9" box="[151,232,188,212]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="40" pageNumber="341">DEBU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32FC9E3BD1BFFB3FF23FF16FBAC4295" box="[199,1064,223,248]" pageId="40" pageNumber="341" type="distribution">
<paragraph id="8B8A9A68BD1BFFB3FF23FF16FBAC4295" blockId="40.[151,1436,151,506]" box="[199,1064,223,248]" pageId="40" pageNumber="341">
<emphasis id="B941467ABD1BFFB3FF23FF16FEE44295" bold="true" box="[199,352,223,248]" pageId="40" pageNumber="341">Distribution.</emphasis>
Only known from the
<typeStatus id="548E24CABD1BFFB3FD84FF29FD144295" box="[608,656,224,248]" pageId="40" pageNumber="341">type</typeStatus>
locality in southern
<collectingCountry id="F322DAF8BD1BFFB3FC92FF29FC2E429A" box="[886,938,224,247]" name="Peru" pageId="40" pageNumber="341">Peru</collectingCountry>
(Fig. 125).
</paragraph>
</subSubSection>
<subSubSection id="C32FC9E3BD1BFFB3FF23FECDFE18432D" pageId="40" pageNumber="341" type="etymology">
<paragraph id="8B8A9A68BD1BFFB3FF23FECDFE18432D" blockId="40.[151,1436,151,506]" pageId="40" pageNumber="341">
<emphasis id="B941467ABD1BFFB3FF23FECDFECA4370" bold="true" box="[199,334,260,285]" pageId="40" pageNumber="341">Etymology.</emphasis>
The species name is Latin for &quot;a maker of machines&quot; and refers to the wrench-shaped sternite 5; it is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32FC9E3BD1BFFB3FF23FE85FAE44397" pageId="40" pageNumber="341" type="discussion">
<paragraph id="8B8A9A68BD1BFFB3FF23FE85FAE44397" blockId="40.[151,1436,151,506]" pageId="40" pageNumber="341">
<emphasis id="B941467ABD1BFFB3FF23FE85FECB4308" bold="true" box="[199,335,332,357]" pageId="40" pageNumber="341">Comments.</emphasis>
This species appears to be related to
<taxonomicName id="4C35E1EBBD1BFFB3FD1BFE84FCE04309" box="[767,868,332,356]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="341" phylum="Arthropoda" rank="species" species="pollex">
<emphasis id="B941467ABD1BFFB3FD1BFE84FCE04309" box="[767,868,332,356]" italics="true" pageId="40" pageNumber="341">B. pollex</emphasis>
</taxonomicName>
and
<taxonomicName id="4C35E1EBBD1BFFB3FC79FE84FBC94309" box="[925,1101,333,356]" class="Insecta" family="Sphaeroceridae" genus="Boreantrops" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="341" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B941467ABD1BFFB3FC79FE84FBC94309" box="[925,1101,333,356]" italics="true" pageId="40" pageNumber="341">B. punctipennis</emphasis>
</taxonomicName>
, sharing a basal spot on vein R4+5, a lack of anterodorsal bristles on the mid tibia, and a thumb-like projection on the surstylus. It differs in a number of ways, including lacking wing spots on vein tips and having a spot on vein CuA1 (unique in the genus), anal opening not keyhole-shaped, and having a thicker basiphallus. Despite considerable collecting effort, we did not encounter this species at the
<typeStatus id="548E24CABD1BFFB3FDE7FE2BFDB74397" box="[515,563,482,506]" pageId="40" pageNumber="341">type</typeStatus>
locality in
<date id="FF8BBCA8BD1BFFB3FD56FE2BFCAC4397" box="[690,808,482,506]" pageId="40" pageNumber="341" value="2007-05">May 2007</date>
; perhaps adults are not present at that time of year.
</paragraph>
</subSubSection>
</treatment>
</document>