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<document id="8C3AFB3F89A63FECE5D2949E448AAAF2" ID-DOI="10.25221/fee.480.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="10134955" ID-ZooBank="F798770D-B8E3-4EDF-93AB-F9666E087A28" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699901156432" checkinUser="felipe" docAuthor="Lelej, A. S." docDate="2023" docId="0396364DFFC7794BFE00E5718C0BFDAB" docLanguage="en" docName="FarEastEntomol.480.1-22.pdf.imf" docOrigin="Far Eastern Entomologist 480" docSource="http://dx.doi.org/10.25221/fee.480.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.12:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="12" docTitle="Cephalotilla Bischoff 1920" docType="treatment" docVersion="4" lastPageNumber="8" masterDocId="FFAF4E35FFC2794CFFE6E0638842FFDA" masterDocTitle="Review of the tribe Ctenotillini (Hymenoptera: Mutillidae) from Oriental and Palaearctic regions" masterLastPageNumber="22" masterPageNumber="1" pageNumber="6" updateTime="1712597546037" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="63CA2032D5F29AEF257C589CB4AAB8BA">Review of the tribe Ctenotillini (Hymenoptera: Mutillidae) from Oriental and Palaearctic regions</mods:title>
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<mods:date id="21CD38D00327FD2DA91580E49C67D6FC">2023</mods:date>
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<heading id="D0C83037FFC77949FE00E5718B21FAF0" box="[486,867,1298,1322]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="6" reason="2">
<emphasis id="B94B5B49FFC77949FE00E5718B21FAF0" bold="true" box="[486,867,1298,1322]" pageId="5" pageNumber="6">
Genus
<taxonomicName id="4C3FFCD8FFC77949FDD0E5718B21FAF0" authority="Bischoff, 1920" authorityName="Bischoff" authorityYear="1920" box="[566,867,1298,1322]" class="Insecta" genus="Cephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94B5B49FFC77949FDD0E5718AFFFAF0" bold="true" box="[566,701,1298,1322]" italics="true" pageId="5" pageNumber="6">Cephalotilla</emphasis>
<bibRefCitation id="EFAEFAAAFFC77949FD25E5718B21FAF0" author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[707,867,1298,1322]" pageId="5" pageNumber="6" pagination="4 - 5" refId="ref8146" refString="Bischoff, H. 1920 - 1921. Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, 86 A (1 - 3), 1 - 480 [1920]; 86 A (4 - 5), 481 - 830 [1921]." type="journal article" year="1920">Bischoff, 1920</bibRefCitation>
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<treatmentCitation id="0A9EA14AFFC77949FF05E5598A5AFA95" author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[227,536,1338,1360]" page="24" pageId="5" pageNumber="6" year="1920">
<taxonomicName id="4C3FFCD8FFC77949FF05E5598A5AFA95" authority="Bischoff, 1920: 24" authorityName="Bischoff" authorityPageNumber="24" authorityYear="1920" box="[227,536,1338,1360]" class="Insecta" genus="Cephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94B5B49FFC77949FF05E5598919FA95" box="[227,347,1338,1359]" italics="true" pageId="5" pageNumber="6">Cephalotilla</emphasis>
<bibRefCitation id="EFAEFAAAFFC77949FE84E5598A5AFA95" author="Bischoff, H. &amp; Monographie der Mutilliden Afrikas" box="[354,536,1338,1360]" pageId="5" pageNumber="6" pagination="4 - 5" refId="ref8146" refString="Bischoff, H. 1920 - 1921. Monographie der Mutilliden Afrikas. Archiv fur Naturgeschichte, 86 A (1 - 3), 1 - 480 [1920]; 86 A (4 - 5), 481 - 830 [1921]." type="journal article" year="1920">Bischoff, 1920: 24</bibRefCitation>
</taxonomicName>
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(in key); 1921, 86A(4): 509 (description),
<specimenCount id="9D394CD2FFC77949FC5DE5598B8DFA95" box="[955,975,1338,1359]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
,
<specimenCount id="9D394CD2FFC77949FC3CE5598BAFFA95" box="[986,1005,1338,1359]" count="1" pageId="5" pageNumber="6" type="female">♀</specimenCount>
;
<treatmentCitation id="0A9EA14AFFC77949FC1CE5598923FAB0" author="Nonveiller, G." page="81" pageId="5" pageNumber="6" year="1979">
<bibRefCitation id="EFAEFAAAFFC77949FC1CE5598923FAB0" author="Nonveiller, G." pageId="5" pageNumber="6" pagination="7 - 184" refId="ref8994" refString="Nonveiller, G. 1979 [&quot; 1978 &quot;] Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 7 - 184." type="journal article" year="1979">Nonveiller, 1979: 81</bibRefCitation>
</treatmentCitation>
,
<specimenCount id="9D394CD2FFC77949FE8AE53689C2FAB0" box="[364,384,1365,1386]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
,
<specimenCount id="9D394CD2FFC77949FE6DE53689DCFAB0" box="[395,414,1365,1386]" count="1" pageId="5" pageNumber="6" type="female">♀</specimenCount>
;
<treatmentCitation id="0A9EA14AFFC77949FE4DE5368AF2FAB0" author="Lelej, A. S. &amp; Brothers, D. J." box="[427,688,1365,1387]" page="14" pageId="5" pageNumber="6" year="2008">
<bibRefCitation id="EFAEFAAAFFC77949FE4DE5368AF2FAB0" author="Lelej, A. S. &amp; Brothers, D. J." box="[427,688,1365,1387]" pageId="5" pageNumber="6" pagination="1 - 79" refId="ref8650" refString="Lelej, A. S. &amp; Brothers, D. J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889, 1 - 79. DOI: 10.11646 / zootaxa. 1889.1.1" type="journal article" year="2008">Lelej &amp; Brothers, 2008: 14</bibRefCitation>
</treatmentCitation>
,
<specimenCount id="9D394CD2FFC77949FD5DE5368A8CFAB0" box="[699,718,1365,1386]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
,
<specimenCount id="9D394CD2FFC77949FD3CE5368AAEFAB0" box="[730,748,1365,1386]" count="1" pageId="5" pageNumber="6" type="female">♀</specimenCount>
; Brother &amp; Lelej, 2017: 95,
<specimenCount id="9D394CD2FFC77949FC1BE5368C52FAB0" box="[1021,1040,1365,1386]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
,
<specimenCount id="9D394CD2FFC77949FBFAE5368C6CFAB0" box="[1052,1070,1365,1386]" count="1" pageId="5" pageNumber="6" type="female">♀</specimenCount>
;
<treatmentCitation id="0A9EA14AFFC77949FBDDE536899CFA5C" author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Matteini Palmerini, M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." page="157" pageId="5" pageNumber="6" year="2020">
<bibRefCitation id="EFAEFAAAFFC77949FBDDE536899CFA5C" author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Matteini Palmerini, M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." pageId="5" pageNumber="6" pagination="5 - 427" refId="ref9121" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A. &amp; Romano, M. (2020 [2018]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torino, 36 (1 - 2), 5 - 427." type="journal article" year="2020">
Pagliano
<emphasis id="B94B5B49FFC77949FED9E513892BFA5F" box="[319,361,1392,1413]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2020: 157
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B80875BFFC77949FF05E5E88A54FA7B" blockId="5.[227,1127,1338,1441]" box="[227,534,1419,1441]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB2FA075FFC77949FF05E5E88A54FA7B" box="[227,534,1419,1441]" pageId="5" pageNumber="6">
<taxonomicName id="4C3FFCD8FFC77949FF05E5E88901FA7A" authorityName="sensu Bischoff" authorityYear="1920" box="[227,323,1419,1440]" class="Insecta" family="Mutillidae" genus="Ctenotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B94B5B49FFC77949FF05E5E88901FA7A" box="[227,323,1419,1440]" italics="true" pageId="5" pageNumber="6">Ctenotilla</emphasis>
</taxonomicName>
:
<treatmentCitation id="0A9EA14AFFC77949FEB6E5E88A50FA7B" author="Brothers, D. J." box="[336,530,1419,1441]" page="592" pageId="5" pageNumber="6" year="1975">
<bibRefCitation id="EFAEFAAAFFC77949FEB6E5E88A50FA7B" author="Brothers, D. J." box="[336,530,1419,1441]" pageId="5" pageNumber="6" pagination="483 - 648" refId="ref8193" refString="Brothers, D. J. 1975. Phylogeny and classification of the aculeate Hymenoptera, with special reference to Mutillidae. University of Kansas Science Bulletin, 50: 483 - 648." type="journal article" year="1975">Brothers, 1975: 592</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C325D4D0FFC77949FEEFE5D18A83FA3D" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8B80875BFFC77949FEEFE5D18A83FA3D" blockId="5.[227,1127,1457,1511]" pageId="5" pageNumber="6">
<typeStatus id="548439F9FFC77949FEEFE5D18902FA10" box="[265,320,1458,1482]" pageId="5" pageNumber="6">Type</typeStatus>
species:
<taxonomicName id="4C3FFCD8FFC77949FE40E5D28B13FA13" authority="Bischoff, 1921" authorityName="Bischoff" authorityYear="1921" box="[422,849,1457,1481]" class="Insecta" genus="Cephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kamogana">
<emphasis id="B94B5B49FFC77949FE40E5D28AE6FA13" box="[422,676,1457,1481]" italics="true" pageId="5" pageNumber="6">Cephalotilla kamogana</emphasis>
Bischoff, 1921
</taxonomicName>
,
<specimenCount id="9D394CD2FFC77949FCB8E5D18B36FA13" box="[862,884,1458,1481]" count="1" pageId="5" pageNumber="6" type="male">♂</specimenCount>
, by subsequent designation of Bischoff, 1921: 509 [Afrotropical].
</paragraph>
</subSubSection>
<subSubSection id="C325D4D0FFC7794BFEEFE59E896AFDEE" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8B80875BFFC7794AFEEFE59E8C21FEE4" blockId="5.[227,1127,1533,1588]" lastBlockId="6.[227,1126,264,319]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">DIAGNOSIS. MALE. T4–6 with medial longitudinal carina. S8 medially with deep concavity bordered by lateral tuft of setae, without subbasal lateral tubercle. S7 with lateral tubercle. S2 invaginated sublaterally with distinct lateral longitudinal carina and medial longitudinal carina. FEMALE. Inner margin of mandible without</paragraph>
<paragraph id="8B80875BFFC4794AFEEFE6498C10F99A" blockId="6.[227,1106,1578,1627]" box="[265,1106,1578,1600]" pageId="6" pageNumber="7">
Figs 1–6. 1–3 –
<taxonomicName id="4C3FFCD8FFC4794AFE42E6498A00F9E5" baseAuthorityName="Radoszkowski" baseAuthorityYear="1879" box="[420,578,1578,1599]" class="Insecta" family="Mutillidae" genus="Ctenotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caeca">
<emphasis id="B94B5B49FFC4794AFE42E6498A00F9E5" box="[420,578,1578,1599]" italics="true" pageId="6" pageNumber="7">Ctenotilla caeca</emphasis>
</taxonomicName>
,
<specimenCount id="9D394CD2FFC4794AFDA8E6498A23F9E5" box="[590,609,1578,1599]" count="1" pageId="6" pageNumber="7" type="female">♀</specimenCount>
; 4–6 –
<taxonomicName id="4C3FFCD8FFC4794AFD4FE6498B35F9E5" authorityName="Lelej" authorityYear="2023" baseAuthorityName="Turner" baseAuthorityYear="1911" box="[681,887,1578,1599]" class="Insecta" genus="Cephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="porcella">
<emphasis id="B94B5B49FFC4794AFD4FE6498B35F9E5" box="[681,887,1578,1599]" italics="true" pageId="6" pageNumber="7">Cephalotilla porcella</emphasis>
</taxonomicName>
,
<specimenCount id="9D394CD2FFC4794AFC65E6498BD4F9E5" box="[899,918,1578,1599]" count="1" pageId="6" pageNumber="7" type="female">♀</specimenCount>
. 1, 2, 4, 5 – habitus
</paragraph>
<paragraph id="8B80875BFFC4794AFF05E6258B13F981" blockId="6.[227,1106,1578,1627]" box="[227,849,1605,1627]" pageId="6" pageNumber="7">(1, 4 – dorsal view, 2, 5 – lateral view); 3, 6 – T6, pygidial plate.</paragraph>
<paragraph id="8B80875BFFC5794BFF05E16B8AE6FEA6" blockId="7.[227,1127,264,626]" pageId="7" pageNumber="8">subbasal denticle. Pygidial plate granulose. T2 without two subbasal white spots disposed transversely. T2 apically and T3 with non interrupted band of yellowish or white setae, band on T2 widened medially. T1 width much less than propodeum width. Postgenal carina anterad with dent.</paragraph>
<paragraph id="8B80875BFFC5794BFEEFE1E0896AFDEE" blockId="7.[227,1127,264,626]" pageId="7" pageNumber="8">
SPECIES INCLUDED. Oriental
<taxonomicName id="4C3FFCD8FFC5794BFD8BE1E18BBBFE41" authority="(Turner, 1911)" authorityName="Lelej" authorityYear="2023" baseAuthorityName="Turner" baseAuthorityYear="1911" box="[621,1017,386,411]" class="Insecta" genus="Cephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="porcella" status="comb. nov.">
<emphasis id="B94B5B49FFC5794BFD8BE1E18B10FE40" box="[621,850,386,410]" italics="true" pageId="7" pageNumber="8">Cephalotilla porcella</emphasis>
(
<bibRefCitation id="EFAEFAAAFFC5794BFC87E1E08BB3FE41" author="Turner, R. E." box="[865,1009,387,411]" pageId="7" pageNumber="8" pagination="141 - 154" refId="ref9265" refString="Turner, R. E. 1911. New Hymenoptera from Ceylon. Mutillidae and Scoliidae. Spolia Zeylanica, 7 (27): 141 - 154." type="journal article" year="1911">Turner, 1911</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B94B5B49FFC5794BFBE3E1E08C24FE41" bold="true" box="[1029,1126,387,411]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="A278E632FFC5794BFBE3E1E08C24FE41" box="[1029,1126,387,411]" pageId="7" pageNumber="8" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
from nominotypical subgenus and undescribed Indian species with reddish-brown head, black legs and narrow pygidial plate. The subgenera
<taxonomicName id="4C3FFCD8FFC5794BFC81E1A3895AFE2D" authority="Nonveiller, 1979" authorityName="Nonveiller" authorityYear="1979" class="Insecta" genus="Bidentotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94B5B49FFC5794BFC81E1A38BA7FE02" box="[871,997,448,472]" italics="true" pageId="7" pageNumber="8">Bidentotilla</emphasis>
<bibRefCitation id="EFAEFAAAFFC5794BFC08E1A3895AFE2D" author="Nonveiller, G." pageId="7" pageNumber="8" pagination="7 - 184" refId="ref8994" refString="Nonveiller, G. 1979 [&quot; 1978 &quot;] Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 7 - 184." type="journal article" year="1979">Nonveiller, 1979</bibRefCitation>
</taxonomicName>
with one species,
<taxonomicName id="4C3FFCD8FFC5794BFE3AE1BD8B4AFE2D" authority="Nonveiller, 1979" authorityName="Nonveiller" authorityYear="1979" box="[476,776,478,503]" class="Insecta" genus="Taeniotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94B5B49FFC5794BFE3AE1BD8A0DFE2C" box="[476,591,478,502]" italics="true" pageId="7" pageNumber="8">Taeniotilla</emphasis>
<bibRefCitation id="EFAEFAAAFFC5794BFDB3E1BD8B4AFE2D" author="Nonveiller, G." box="[597,776,478,503]" pageId="7" pageNumber="8" pagination="7 - 184" refId="ref8994" refString="Nonveiller, G. 1979 [&quot; 1978 &quot;] Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 7 - 184." type="journal article" year="1979">Nonveiller, 1979</bibRefCitation>
</taxonomicName>
with one species, and 52 species from nominotypical subgenus are distributed in Afrotropical Region (
<bibRefCitation id="EFAEFAAAFFC5794BFC0AE19E895FFDEE" author="Nonveiller, G." pageId="7" pageNumber="8" pagination="7 - 184" refId="ref8994" refString="Nonveiller, G. 1979 [&quot; 1978 &quot;] Recherches sur les Mutillides de l'Afrique (Hymenoptera, Mutillidae). VIII. Revision des genres Ctenotilla, Cephalotilla et Pseudocephalotilla sensu Bischoff. Memoires publies par l'Institut pour la Protection des Plantes, Belgrad, 13, 7 - 184." type="journal article" year="1979">Nonveiller, 1979</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C325D4D0FFC5794BFEEFE2588C0DFD88" box="[265,1103,570,595]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8B80875BFFC5794BFEEFE2588C0DFD88" blockId="7.[227,1127,264,626]" box="[265,1103,570,595]" pageId="7" pageNumber="8">
DISTRIBUTION. Oriental (
<collectingCountry id="F328C7CBFFC5794BFDD0E2598AE1FD88" box="[566,675,570,594]" name="Sri Lanka" pageId="7" pageNumber="8">Sri Lanka</collectingCountry>
, South
<collectingCountry id="F328C7CBFFC5794BFD14E2598B68FD88" box="[754,810,570,594]" name="India" pageId="7" pageNumber="8">India</collectingCountry>
) and Afrotropical Regions.
</paragraph>
</subSubSection>
<subSubSection id="C325D4D0FFC5794BFEEFE23A8C0BFDAB" box="[265,1097,601,626]" pageId="7" pageNumber="8" type="discussion">
<paragraph id="8B80875BFFC5794BFEEFE23A8C0BFDAB" blockId="7.[227,1127,264,626]" box="[265,1097,601,626]" pageId="7" pageNumber="8">
REMARKS. The genus
<taxonomicName id="4C3FFCD8FFC5794BFDEAE23A8AD3FDAB" authorityName="Bischoff" authorityYear="1920" box="[524,657,601,625]" class="Insecta" genus="Cephalotilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B94B5B49FFC5794BFDEAE23A8AD3FDAB" box="[524,657,601,625]" italics="true" pageId="7" pageNumber="8">Cephalotilla</emphasis>
</taxonomicName>
is newly recorded from Oriental Region.
</paragraph>
</subSubSection>
</treatment>
</document>
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