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<document id="BAB41279033EA1A4E8ABC4250822753D" ID-DOI="10.1111/j.1096-3642.2011.00748.x" ID-ISSN="0024-4082" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630453203665" checkinUser="valdenar" docAuthor="Jadin, Robert C., Smith, Eric N. &amp; Campbell, Jonathan A." docDate="2011" docId="03957E16FFB0FFD3FCF7FD661D50D97D" docLanguage="en" docName="j.1096-3642.2011.00748.x.pdf" docOrigin="Zoological Journal of the Linnean Society 163 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00748.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Cerrophidion CAMPBELL &amp; LAMAR 1992" docType="treatment" docVersion="3" lastPageNumber="953" masterDocId="FFAC066EFFBAFFD8FF95FFB81C16DA00" masterDocTitle="Unravelling a tangle of Mexican serpents: a systematic revision of highland pitvipers" masterLastPageNumber="958" masterPageNumber="943" pageNumber="953" updateTime="1699276107582" updateUser="plazi">
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<mods:title id="09E2715A5222DEA5B935459D3038B291">Unravelling a tangle of Mexican serpents: a systematic revision of highland pitvipers</mods:title>
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<mods:namePart id="60E6F859DD173653CC6FFA2AB56AC74F">Jadin, Robert C.</mods:namePart>
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<mods:namePart id="A33ED57D9761C88E6E3C2368373C1812">Campbell, Jonathan A.</mods:namePart>
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<subSubSection id="C3269C8BFFB0FFD2FCF7FD661947D8F2" box="[866,1361,733,755]" pageId="10" pageNumber="953" type="nomenclature">
<paragraph id="8B83CF00FFB0FFD2FCF7FD661947D8F2" blockId="10.[866,1361,733,755]" box="[866,1361,733,755]" pageId="10" pageNumber="953">
<heading id="D0CB786CFFB0FFD2FCF7FD661947D8F2" box="[866,1361,733,755]" centered="true" fontSize="8" level="2" pageId="10" pageNumber="953" reason="8">
<taxonomicName id="4C3CB483FFB0FFD2FCF7FD661947D8F2" authority="CAMPBELL &amp; LAMAR, 1992" authorityName="CAMPBELL &amp; LAMAR" authorityYear="1992" box="[866,1361,733,755]" class="Reptilia" family="Viperidae" genus="Cerrophidion" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="953" phylum="Chordata" rank="genus">
<emphasis id="B9481312FFB0FFD2FCF7FD661818D8F1" box="[866,1038,734,755]" italics="true" pageId="10" pageNumber="953">CERROPHIDION</emphasis>
<bibRefCitation id="EFADB2F1FFB0FFD2FB8DFD651947D8F2" author="Campbell JA &amp; Lamar WW" box="[1048,1361,733,755]" pageId="10" pageNumber="953" pagination="1 - 31" refId="ref10009" refString="Campbell JA, Lamar WW. 1992. Taxonomic status of miscellaneous Neotropical viperids, with the description of a new genus. Occasional Papers of the Museum of Texas Tech University 153: 1 - 31." type="journal article" year="1992">CAMPBELL &amp; LAMAR, 1992</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3269C8BFFB0FFD3FCB3FCBC1D50D97D" lastPageId="11" lastPageNumber="954" pageId="10" pageNumber="953" type="materials_examined">
<paragraph id="8B83CF00FFB0FFD2FCB3FCBC1998D938" blockId="10.[806,1422,772,824]" pageId="10" pageNumber="953">
<emphasis id="B9481312FFB0FFD2FCB3FCBC1FA3D919" box="[806,949,772,793]" italics="true" pageId="10" pageNumber="953">
<typeStatus id="548771A2FFB0FFD2FCB3FCBC1F4CD919" box="[806,858,772,793]" pageId="10" pageNumber="953">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C3CB483FFB0FFD2FC28FCBC1970D91A" authority="Gunther, 1863" authorityName="Gunther" authorityYear="1863" box="[957,1382,772,794]" class="Reptilia" family="Viperidae" genus="Bothriechis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="953" phylum="Chordata" rank="species" species="godmanni">
<emphasis id="B9481312FFB0FFD2FC28FCBC18AED919" box="[957,1208,772,793]" italics="true" pageId="10" pageNumber="953">Bothriechis godmanni</emphasis>
<bibRefCitation id="EFADB2F1FFB0FFD2FB55FCBC1970D91A" author="Gunther ACL" box="[1216,1382,772,794]" pageId="10" pageNumber="953" pagination="348 - 365" refId="ref10617" refString="Gunther ACL. 1863. Hird account of the snakes in the collection of the British Museum. Annals and Magazine of Natural History, 3 rd Series 12: 348 - 365." type="book chapter" year="1863">Günther, 1863</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<bibRefCitation id="EFADB2F1FFB0FFD2FBCCFC9A199FD938" author="Campbell JA &amp; Lamar WW" box="[1113,1417,802,824]" pageId="10" pageNumber="953" pagination="1 - 31" refId="ref10009" refString="Campbell JA, Lamar WW. 1992. Taxonomic status of miscellaneous Neotropical viperids, with the description of a new genus. Occasional Papers of the Museum of Texas Tech University 153: 1 - 31." type="journal article" year="1992">Campbell &amp; Lamar (1992)</bibRefCitation>
.
</paragraph>
<paragraph id="8B83CF00FFB0FFD2FCB3FCD9180FD9D3" blockId="10.[806,1421,865,979]" pageId="10" pageNumber="953">
<emphasis id="B9481312FFB0FFD2FCB3FCD91FB2D976" box="[806,932,865,886]" italics="true" pageId="10" pageNumber="953">Etymology:</emphasis>
The generic name comes from the Spanish
<emphasis id="B9481312FFB0FFD2FCB3FC391F48D996" box="[806,862,897,918]" italics="true" pageId="10" pageNumber="953">cerro</emphasis>
, meaning mountain, an allusion to the habitat, and the Greek
<emphasis id="B9481312FFB0FFD2FC5AFC271825D9B4" box="[975,1075,927,948]" italics="true" pageId="10" pageNumber="953">ophidion</emphasis>
, meaning small snake (
<bibRefCitation id="EFADB2F1FFB0FFD2FAD4FC27181BD9D3" author="Campbell JA &amp; Lamar WW" pageId="10" pageNumber="953" pagination="1 - 31" refId="ref10009" refString="Campbell JA, Lamar WW. 1992. Taxonomic status of miscellaneous Neotropical viperids, with the description of a new genus. Occasional Papers of the Museum of Texas Tech University 153: 1 - 31." type="journal article" year="1992">Campbell &amp; Lamar, 1992</bibRefCitation>
).
</paragraph>
<paragraph id="8B83CF00FFB0FFD2FCB3FC45180BDEE8" blockId="10.[806,1421,1020,1256]" pageId="10" pageNumber="953">
<emphasis id="B9481312FFB0FFD2FCB3FC451F93DE12" box="[806,901,1021,1042]" italics="true" pageId="10" pageNumber="953">Content:</emphasis>
The genus
<taxonomicName id="4C3CB483FFB0FFD2FBA4FC4418DEDE11" authorityName="CAMPBELL &amp; LAMAR" authorityYear="1992" box="[1073,1224,1020,1041]" class="Reptilia" family="Viperidae" genus="Cerrophidion" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="953" phylum="Chordata" rank="genus">
<emphasis id="B9481312FFB0FFD2FBA4FC4418DEDE11" box="[1073,1224,1020,1041]" italics="true" pageId="10" pageNumber="953">Cerrophidion</emphasis>
</taxonomicName>
contains three species:
<taxonomicName id="4C3CB483FFB0FFD2FC1CFBA31886DE30" baseAuthorityName="Gunther" baseAuthorityYear="1863" box="[905,1168,1051,1072]" class="Reptilia" family="Viperidae" genus="Cerrophidion" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="953" phylum="Chordata" rank="species" species="godmani">
<emphasis id="B9481312FFB0FFD2FC1CFBA31886DE30" box="[905,1168,1051,1072]" italics="true" pageId="10" pageNumber="953">Cerrophidion godmani</emphasis>
</taxonomicName>
,
<taxonomicName id="4C3CB483FFB0FFD2FB37FBA31F69DE4E" baseAuthorityName="Lopez-Luna, Vogt &amp; de la Torre-Loranca" baseAuthorityYear="1999" class="Reptilia" family="Viperidae" genus="Cerrophidion" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="953" phylum="Chordata" rank="species" species="petlalcalensis">
<emphasis id="B9481312FFB0FFD2FB37FBA31F69DE4E" italics="true" pageId="10" pageNumber="953">Cerrophidion petlalcalensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C3CB483FFB0FFD2FC57FB8118F4DE4E" baseAuthorityName="Campbell" baseAuthorityYear="1985" box="[962,1250,1081,1102]" class="Reptilia" family="Viperidae" genus="Cerrophidion" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="953" phylum="Chordata" rank="species" species="tzotzilorum">
<emphasis id="B9481312FFB0FFD2FC57FB8118F4DE4E" box="[962,1250,1081,1102]" italics="true" pageId="10" pageNumber="953">Cerrophidion tzotzilorum</emphasis>
</taxonomicName>
. These species occur in pine-oak and cloud forests from
<collectingRegion id="49F801E2FFB0FFD2FABDFBE1199BDE6E" box="[1320,1421,1113,1134]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="10" pageNumber="953">Veracruz</collectingRegion>
(
<collectingCountry id="F32B8F90FFB0FFD2FCB8FBCF1F69DE8D" box="[813,895,1143,1165]" name="Mexico" pageId="10" pageNumber="953">Mexico</collectingCountry>
) southward through the highlands of Central America to
<collectingCountry id="F32B8F90FFB0FFD2FC2CFB2E1801DEAB" box="[953,1047,1174,1195]" name="Panama" pageId="10" pageNumber="953">Panama</collectingCountry>
(
<bibRefCitation id="EFADB2F1FFB0FFD2FBBBFB2D18FCDEAB" author="Campbell JA" box="[1070,1258,1173,1195]" pageId="10" pageNumber="953" pagination="48 - 54" refId="ref9898" refString="Campbell JA. 1985. A new species of highland pitviper of the genus Bothrops from southern Mexico. Journal of Herpetology 19: 48 - 54." type="journal article" year="1985">Campbell, 1985</bibRefCitation>
;
<bibRefCitation id="EFADB2F1FFB0FFD2FB68FB2D1FAFDEC9" author="Campbell JA &amp; Lamar WW" pageId="10" pageNumber="953" refId="ref10047" refString="Campbell JA, Lamar WW. 2004. Venomous reptiles of the Western Hemisphere. 2 vols. Ithaca, NY: Comstock Publishing Associates." type="book" year="2004">Campbell &amp; Lamar, 2004</bibRefCitation>
: maps 79, 80) with a vertical distribution from
<emphasis id="B9481312FFB0FFD2FCF0FB6B1F61DEE8" box="[869,887,1235,1256]" italics="true" pageId="10" pageNumber="953">c.</emphasis>
<quantity id="4CC462E5FFB0FFD2FCEBFB6B180FDEE8" box="[894,1049,1235,1256]" metricMagnitude="3" metricUnit="m" metricValue="2.4455" metricValueMax="3.491" metricValueMin="1.4" pageId="10" pageNumber="953" unit="m" value="2445.5" valueMax="3491.0" valueMin="1400.0">14003491 m</quantity>
.
</paragraph>
<paragraph id="8B83CF00FFB0FFD2FCB3FAAA1998DF27" blockId="10.[806,1422,1297,1319]" box="[806,1422,1297,1319]" pageId="10" pageNumber="953">
<emphasis id="B9481312FFB0FFD2FCB3FAAA1FCCDF27" box="[806,986,1298,1319]" italics="true" pageId="10" pageNumber="953">Common name:</emphasis>
Middle American montane pitvipers.
</paragraph>
<paragraph id="8B83CF00FFB0FFD2FCB3FAE91FAEDD32" blockId="10.[806,1421,1361,1903]" pageId="10" pageNumber="953">
<emphasis id="B9481312FFB0FFD2FCB3FAE91844DF66" box="[806,1106,1361,1382]" italics="true" pageId="10" pageNumber="953">Definition and diagnosis:</emphasis>
Rostral wider than high, front surface flat; three preoculars, upper largest, entire, and squarish, lower forming posterior border of pit and excluded from orbit; single, large, flat, plate-like supraocular above eye; seven to 11 supralabials; eight to 12 infralabials; canthals and internasals relatively large and flat; two to seven intersupraoculars; crown of head covered with variably sized, flat or keeled scales; keeling prominent in parietal area; second supralabial discrete from prelacunal; supralabial and subocular series in contact or separated by single row of scales; 1923 (mode 21) middorsal dorsal scale rows; mid-dorsal scales at midbody moderately slender and pointed; 120150 ventrals; 2236 undivided subcaudals; tail spine straight, moderately long.
</paragraph>
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Lateral edge of nasal broadly expanded, bone roughly quadrangular; frontal bones mostly flat, dorsal surface with slightly elevated margins, longer than wide; postfrontal large, not reaching frontal; transverse distance of postfrontal greater than its distance along parietal bone; posterolateral edges of dorsal surface of parietals forming low to moderately distinct raised ridge continuing posteriorly on parietal as low ridge; junction between parietal and prootic rounded to almost flat; squamosal extending to level posterior to posterior edge of exoccipital; ectopterygoid about same length as expanded, flattened base of pterygoid (posterior to the articulation with ectopterygoid) with flat shaft gradually tapering posteriorly; dorsal surface of parietal roughly triangular to sometimes rounded; three to five palatine teeth; seven to 18 pterygoid teeth; eight to 16 dentary teeth; pterygoid teeth extending just posterior to level of articulation of pterygoid with ectopterygoid in
<taxonomicName id="4C3CB483FFB1FFD3FD2FFD081CC9D8E4" baseAuthorityName="Gunther" baseAuthorityYear="1863" class="Reptilia" family="Viperidae" genus="Cerrophidion" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="954" phylum="Chordata" rank="species" species="godmani">
<emphasis id="B9481312FFB1FFD3FD2FFD081CC9D8E4" italics="true" pageId="11" pageNumber="954">C. godmani</emphasis>
</taxonomicName>
, but not reaching this far back in congeners; maxillary fang relatively short, being about equal in length to height of maxilla; fang at rest extending to level of about middle of supralabial 5 or suture between supralabials 56 (mostly after
<bibRefCitation id="EFADB2F1FFB1FFD3FDEBFCF01D2CD97D" author="Campbell JA &amp; Lamar WW" pageId="11" pageNumber="954" refId="ref10047" refString="Campbell JA, Lamar WW. 2004. Venomous reptiles of the Western Hemisphere. 2 vols. Ithaca, NY: Comstock Publishing Associates." type="book" year="2004">Campbell &amp; Lamar, 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
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