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<mods:title id="222A4A03F71E231C98E330DC4EB7045A">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<subSubSection id="C338FF806445FF9DFCA1EF5BD4F5FD87" box="[813,1220,587,611]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="8B9DAC0B6445FF9DFCA1EF5BD4F5FD87" blockId="44.[813,1220,587,643]" box="[813,1220,587,611]" pageId="44" pageNumber="45">
<heading id="D0D51B676445FF9DFCA1EF5BD4F5FD87" box="[813,1220,587,611]" fontSize="9" level="2" pageId="44" pageNumber="45" reason="8">
<taxonomicName id="4C22D7886445FF9DFCA1EF5BD4F5FD87" authority="(Muller, 1956)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[813,1220,587,611]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Ozarkodinida" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFCA1EF5BD390FD87" box="[813,929,587,611]" italics="true" pageId="44" pageNumber="45">Discretella</emphasis>
aff.
<emphasis id="B95670196445FF9DFC42EF5BD415FD87" box="[974,1060,587,611]" italics="true" pageId="44" pageNumber="45">discreta</emphasis>
(
<bibRefCitation id="EFB3D1FA6445FF9DFBBDEF5BD48AFD87" author="Muller" box="[1073,1211,587,611]" firstAuthor="Muller" pageId="44" pageNumber="45" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C338FF806445FF9DFCA1EF7BD50BF9C1" pageId="44" pageNumber="45" type="description">
<paragraph id="8B9DAC0B6445FF9DFCA1EF7BD3DDFD67" blockId="44.[813,1220,587,643]" box="[813,1004,619,643]" pageId="44" pageNumber="45">
<figureCitation id="1319B08E6445FF9DFCA1EF7BD394FD67" box="[813,933,619,643]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="44" pageNumber="45">Figs. 15B, J</figureCitation>
;
<figureCitation id="1319B08E6445FF9DFC23EF7BD3EDFD67" box="[943,988,619,643]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. GI, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. PR Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="44" pageNumber="45">16C</figureCitation>
?.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCA1EFBBD3AAFD07" blockId="44.[813,1437,683,867]" pageId="44" pageNumber="45">
1989
<taxonomicName id="4C22D7886445FF9DFCE6EFBBD4FEFD27" authority="(Muller)" baseAuthorityName="Muller" baseAuthorityYear="1956" box="[874,1231,683,707]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196445FF9DFCE6EFBBD45BFD27" box="[874,1130,683,707]" italics="true" pageId="44" pageNumber="45">Neospathodus bransoni</emphasis>
(Müller)
</taxonomicName>
; Tang, p. 417, pl. 29, fig. 10.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCA1EFFBD431FCC7" blockId="44.[813,1437,683,867]" pageId="44" pageNumber="45">
2009
<taxonomicName id="4C22D7886445FF9DFCE4EFFBD3CAFCE7" authorityName="Zhang &amp; Yang" authorityYear="1991" box="[872,1019,747,771]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="genus">
<emphasis id="B95670196445FF9DFCE4EFFBD3CAFCE7" box="[872,1019,747,771]" italics="true" pageId="44" pageNumber="45">Guangxidella</emphasis>
</taxonomicName>
? sp. A; Orchard &amp; Zonneveld, p. 780, fig. 14, parts 33, 34.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCA1EE3BD3B3FC87" blockId="44.[813,1437,683,867]" pageId="44" pageNumber="45">
2013
<taxonomicName id="4C22D7886445FF9DFCE3EE3BD489FCA7" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[879,1208,811,835]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFCE3EE3BD47DFCA7" box="[879,1100,811,835]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
(Müller)
</taxonomicName>
; Yan et al., p. 516, fig.
<specimenCount id="9D2467826445FF9DFCDAEE5BD3B3FC87" box="[854,898,843,867]" count="6" pageId="44" pageNumber="45" type="generic">6W.</specimenCount>
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCA1EE9BD463FC47" blockId="44.[813,1106,907,931]" box="[813,1106,907,931]" pageId="44" pageNumber="45">Number of specimens:&gt;15</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCA1EEDDD403FBA0" blockId="44.[813,1437,972,1573]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFCA1EEDDD3ACFC00" box="[813,925,973,996]" italics="true" pageId="44" pageNumber="45">Diagnosis.</emphasis>
Segminate P
<subScript id="17A6AE4E6445FF9DFBA6EEC9D402FC0E" attach="left" box="[1066,1075,985,1002]" fontSize="7" pageId="44" pageNumber="45">1</subScript>
element with rounded, upturned basal cavity at the posteriormost part. Large conspicuous cusp above basal cavity. Discrete, smaller upright denticles in the anterior part.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCCEE95DD4B5FAC0" blockId="44.[813,1437,972,1573]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFCCEE95DD3F7FB80" box="[834,966,1101,1124]" italics="true" pageId="44" pageNumber="45">Description.</emphasis>
Tese P
<subScript id="17A6AE4E6445FF9DFBAFE949D41DFB8E" attach="left" box="[1059,1068,1113,1130]" fontSize="7" pageId="44" pageNumber="45">1</subScript>
elements have markedly discrete denticles and a large cusp. Te lower margin is straight in the anterior and upturned at the posterior part. Te basal cavity is drop-shaped, being rounded at the posterior edge and tapered at the anterior end. Te cusp is relatively large and reclined. A small, accessory, posterior denticle is occasionally present.
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCB3E83DD52DFA00" blockId="44.[813,1437,972,1573]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFCB3E83DD395FAA1" box="[831,932,1325,1349]" italics="true" pageId="44" pageNumber="45">Remarks.</emphasis>
In comparison to
<taxonomicName id="4C22D7886445FF9DFB05E83DD550FAA1" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[1161,1377,1325,1349]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFB05E83DD550FAA1" box="[1161,1377,1325,1349]" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
</taxonomicName>
, this species has a rounded posterior margin of the basal cavity, more discrete denticles and the cusp is relatively larger and often recurved. In lateral view, the lower margin is upturned as in morphotype B of
<taxonomicName id="4C22D7886445FF9DFB78E8BDD352FA00" baseAuthorityName="MUller" baseAuthorityYear="1956" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="discreta">
<emphasis id="B95670196445FF9DFB78E8BDD352FA00" italics="true" pageId="44" pageNumber="45">Discretella discreta</emphasis>
</taxonomicName>
(Maekawa &amp; Igo, in
<bibRefCitation id="EFB3D1FA6445FF9DFBCFE8DDD53CFA01" author="Shigeta" box="[1091,1293,1485,1509]" etAl="et al." firstAuthor="Shigeta" pageId="44" pageNumber="45" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
).
</paragraph>
<paragraph id="8B9DAC0B6445FF9DFCCEE8FDD50BF9C1" blockId="44.[813,1437,972,1573]" pageId="44" pageNumber="45">
<emphasis id="B95670196445FF9DFCCEE8FDD38CF9E0" box="[834,957,1517,1540]" italics="true" pageId="44" pageNumber="45">Occurrence</emphasis>
. Luolou formation,
<taxonomicName id="4C22D7886445FF9DFB1AE8FDD4CBF9E0" authorityName="Hyatt &amp; Smith" authorityYear="1905" box="[1174,1274,1517,1540]" class="Cephalopoda" family="Melagathiceratidae" genus="Owenites" kingdom="Animalia" pageId="44" pageNumber="45" phylum="Mollusca" rank="genus">
<emphasis id="B95670196445FF9DFB1AE8FDD4CBF9E0" box="[1174,1274,1517,1540]" italics="true" pageId="44" pageNumber="45">Owenites</emphasis>
</taxonomicName>
beds Smithian age, Shanggang road cut,
<collectingRegion id="49E662E96445FF9DFBCDEB1DD491F9C1" box="[1089,1184,1549,1573]" country="China" name="Guangxi" pageId="44" pageNumber="45">Guangxi</collectingRegion>
, South
<collectingCountry id="F335EC9B6445FF9DFB7DEB1DD507F9C1" box="[1265,1334,1549,1573]" name="China" pageId="44" pageNumber="45">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
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