treatments-xml/data/03/86/18/03861853FFB4FFE1D9C0FDA2FDEBF884.xml
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<document id="AC203C38308513D97BEB33680B1A3210" ID-DOI="10.2478/if-2018-0001" ID-ISSN="2533-4069" ID-Zenodo-Dep="4586465" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614969889443" checkinUser="felipe" docAuthor="Cleal, Christopher J. &amp; Thomas, Barry A." docDate="2018" docId="03861853FFB4FFE1D9C0FDA2FDEBF884" docLanguage="en" docName="FossilImprint.74.1-2.9-28.pdf" docOrigin="Fossil Imprint 74 (1 - 2)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Filicites (sect. Sphenopteris) BRONGNIART" docType="treatment" docVersion="5" lastPageNumber="18" masterDocId="FFBF602BFFBCFFE8DA70F96EFFD5FF9F" masterDocTitle="Nomenclatural Status Of The Palaeobotanical “ Artificial Taxa ” Established In BrongniartS 1822 “ Classification ” Paper" masterLastPageNumber="28" masterPageNumber="9" pageNumber="17" updateTime="1698881448928" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="72889E3924AEC62CB40E5CC617E29112">Nomenclatural Status Of The Palaeobotanical “ Artificial Taxa ” Established In BrongniartS 1822 “ Classification ” Paper</mods:title>
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<mods:namePart id="D5FDE40E150D1AF56A1968EA74C0C6E3">Cleal, Christopher J.</mods:namePart>
<mods:affiliation id="ABD40DA7883645FAC690AC8D0A32755C">Department of Biodiversity and Systematic Biology, National Museum Wales, Cathays Park, Cardiff CF 10 3 NP, United Kingdom; e-mail: chris. cleal @ museumwales. ac. uk.</mods:affiliation>
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<mods:namePart id="900C8FC17770A2808DE6AC166919DB42">Thomas, Barry A.</mods:namePart>
<mods:affiliation id="1FC3C3552E4C9B6ECB93C4418538117A">Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Penglais, Aberystwyth, Ceredigion, SY 201 NL, United Kingdom; e-mail: bat @ aber. ac. uk.</mods:affiliation>
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<mods:title id="5721A6AC90FEEDDD928DC54379D9FE46">Fossil Imprint</mods:title>
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<mods:date id="70FB4B1E5AFF9799BD1CFADDCCA13DDC">2018</mods:date>
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<subSubSection id="C335FACEFFB4FFE0D9C0FDA2FAB6FB7D" box="[944,1379,1228,1252]" pageId="8" pageNumber="17" type="nomenclature">
<paragraph id="8B90A945FFB4FFE0D9C0FDA2FAB6FB7D" blockId="8.[944,1379,1228,1252]" box="[944,1379,1228,1252]" pageId="8" pageNumber="17">
<heading id="D0D81E29FFB4FFE0D9C0FDA2FAB6FB7D" box="[944,1379,1228,1252]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="17" reason="2">
<taxonomicName id="4C2FD2C6FFB4FFE0D9C0FDA2FAB6FB7D" authority="BRONGNIART" authorityName="BRONGNIART" box="[944,1379,1228,1252]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="section" section="Sphenopteris">
<emphasis id="B95B7557FFB4FFE0D9C0FDA2FAB6FB7D" bold="true" box="[944,1379,1228,1252]" pageId="8" pageNumber="17">
<emphasis id="B95B7557FFB4FFE0D9C0FDA2FBDCFB7B" bold="true" box="[944,1033,1228,1252]" italics="true" pageId="8" pageNumber="17">Filicites</emphasis>
sect.
<emphasis id="B95B7557FFB4FFE0DE38FDA2FB0DFB7B" bold="true" box="[1096,1240,1228,1252]" italics="true" pageId="8" pageNumber="17">Sphenopteris</emphasis>
<emphasis id="B95B7557FFB4FFE0DEAFFDA2FAB6FB7D" bold="true" box="[1247,1379,1228,1252]" pageId="8" pageNumber="17">BRONGNIART</emphasis>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C335FACEFFB4FFE1DE34FD82FDEBF884" lastPageId="9" lastPageNumber="18" pageId="8" pageNumber="17" type="description">
<paragraph id="8B90A945FFB4FFE0DE34FD82FB05FA9B" blockId="8.[1092,1232,1260,1284]" box="[1092,1232,1260,1284]" pageId="8" pageNumber="17">
<figureCitation id="1314B5C0FFB4FFE0DE34FD82FB05FA9B" box="[1092,1232,1260,1284]" captionStart="Text-fig" captionStartId="3.[125,206,1872,1893]" captionText="Text-fig. 1. Illustrations from Brongniart (1822a, b) of specimens that remain legitimate taxonomic types of fossil-species. All same scale as in original plate unless otherwise stated. a) Endongenites echinatus BRONGNIART, Eocene lignite, Vailly, near Soissons, France, × 0.5. b) Culmites nodosus BRONGNIART, Eocene Series, Montrouge near Paris, × 0.5. c) Sigillaria scutellata BRONGNIART, MNHN.F.1072.1, unknown locality. d) Clathraria brardii BRONGNIART (≡ Sigillaria brardii (BRONGNIART) BRONGNIART), Pennsylvanian Subsystem. e) Sagenaria coelata BRONGNIART (≡ Lepidodendron coelatum (BRONGNIART) BRONGNIART; = Lepidodendron aculeatum STERNBERG), Pennsylvanian Subsystem. f, j) Filicites (Sphenopteris) elegans BRONGNIART (≡ Sphenopteris elegans (BRONGNIART) STERNBERG), Wadenburg Formation (Serpukhovian Stage), Upper Silesia. gi) Filicites (Pecopteris) penniformis BRONGNIART (≡ Pecopteris penniformis (BRONGNIART) BRONGNIART), Pennsylvanian Subsystem." pageId="8" pageNumber="17">Text-fig. 1f, j</figureCitation>
</paragraph>
<paragraph id="8B90A945FFB4FFE0D93BFC74FAEAFAB0" blockId="8.[843,1483,1306,1357]" box="[843,1343,1306,1328]" pageId="8" pageNumber="17">
1822a
<taxonomicName id="4C2FD2C6FFB4FFE0D9E6FC74FB25FAB1" authority="BRONGNIART" authorityName="BRONGNIART" box="[918,1264,1306,1328]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="subGenus" subGenus="Sphenopteris">
<emphasis id="B95B7557FFB4FFE0D9E6FC74FC30FAB0" box="[918,997,1306,1327]" italics="true" pageId="8" pageNumber="17">Filicites</emphasis>
(
<emphasis id="B95B7557FFB4FFE0D983FC74FBA5FAB0" box="[1011,1136,1306,1327]" italics="true" pageId="8" pageNumber="17">Sphenopteris</emphasis>
) BRONGNIART
</taxonomicName>
, p. 233.
</paragraph>
<paragraph id="8B90A945FFB4FFE0D93BFC59FA1EFAD2" blockId="8.[843,1483,1306,1357]" box="[843,1483,1335,1357]" pageId="8" pageNumber="17">
1825 “
<taxonomicName id="4C2FD2C6FFB4FFE0D9D1FC56FBFFFAD2" authority="(BRONGNIART) STERNBERG, Tentamen" authorityName="Sternberg" authorityYear="1825" baseAuthorityName="BRONGNIART" box="[929,1066,1336,1357]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphaenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB4FFE0D9D1FC56FBFFFAD2" box="[929,1066,1336,1357]" italics="true" pageId="8" pageNumber="17">Sphaenopteris</emphasis>
</taxonomicName>
” (BRONGNIART) STERNBERG, Tentamen p. xv.
</paragraph>
<paragraph id="8B90A945FFB4FFE0D901FC0BFA70FA43" blockId="8.[843,1482,1381,1501]" pageId="8" pageNumber="17">
Ty p e.
<taxonomicName id="4C2FD2C6FFB4FFE0D9B4FC0BFC32FA02" authority="BRONGNIART, 1822 a, p. 233" authorityName="BRONGNIART" authorityPageNumber="233" authorityYear="1822" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" section="Sphenopteris" species="elegans">
<emphasis id="B95B7557FFB4FFE0D9B4FC0BFBCEFAE2" box="[964,1051,1381,1405]" italics="true" pageId="8" pageNumber="17">Filicites</emphasis>
(sect.
<emphasis id="B95B7557FFB4FFE0DE2AFC0BFB30FAE2" box="[1114,1253,1381,1405]" italics="true" pageId="8" pageNumber="17">Sphenopteris</emphasis>
)
<emphasis id="B95B7557FFB4FFE0DE80FC0BFA94FAE2" box="[1264,1345,1381,1405]" italics="true" pageId="8" pageNumber="17">elegans</emphasis>
<bibRefCitation id="EFBED4B4FFB4FFE0DF34FC0BFC5BFA02" author="Brongniart, A." pageId="8" pageNumber="17" pagination="203 - 240" refId="ref15150" refString="Brongniart, A. (1822 a): Sur la classification et la distribution des vegetaux fossiles en general, et sur ceux des terrains de sediment superieur en particulier. Introduction, Chapitre I. - Memoires du Museum d'Histoire Naturelle, 8: 203 - 240, pls 1 - 4." type="journal article" year="1822">BRONGNIART, 1822a</bibRefCitation>
, p. 233
</taxonomicName>
, pl. 2, fig. 2; Loc.: Waldenburg Formation (Serpukhovian Stage), Upper Silesia; ≡ “
<taxonomicName id="4C2FD2C6FFB4FFE0DF55FCCBFA6BFA22" authorityName="Sternberg" authorityYear="1825" box="[1317,1470,1445,1469]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphaenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB4FFE0DF55FCCBFA6BFA22" box="[1317,1470,1445,1469]" italics="true" pageId="8" pageNumber="17">Sphaenopteris</emphasis>
</taxonomicName>
<taxonomicName id="4C2FD2C6FFB4FFE0D93BFCABFB24FA42" authority="(BRONGNIART) STERNBERG, 1825" authorityName="STERNBERG" authorityYear="1825" baseAuthorityName="BRONGNIART" box="[843,1265,1477,1501]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis id="B95B7557FFB4FFE0D93BFCABFC49FA42" box="[843,924,1477,1501]" italics="true" pageId="8" pageNumber="17">elegans</emphasis>
(BRONGNIART)
<bibRefCitation id="EFBED4B4FFB4FFE0DE4AFCABFB24FA42" author="Sternberg, K. M. von" box="[1082,1265,1477,1501]" pageId="8" pageNumber="17" refId="ref20761" refString="Sternberg, K. M. von (1825): Versuch einer geognostischbotanischen Darstellung der Flora der Vorwelt, Vol. I, 4. - Ernst Brenck's Wittwe, Regensburg, 48 pp., tent. I-XLII." type="book" year="1825">STERNBERG, 1825</bibRefCitation>
</taxonomicName>
, Tentamen p. xv.
</paragraph>
<paragraph id="8B90A945FFB4FFE0D901FC9AFB82F9D3" blockId="8.[843,1482,1524,1612]" pageId="8" pageNumber="17">D i a g n o s i s. “pinnules cunéiformes, arrondies ou lobées à lextrémié, et les nervures palmées ou rayonnantes de la base de la pinnule.”</paragraph>
<paragraph id="8B90A945FFB4FFE1D901FF0AFDA7FEAF" blockId="8.[843,1482,1635,1787]" lastBlockId="9.[106,745,151,1820]" lastPageId="9" lastPageNumber="18" pageId="8" pageNumber="17">
D i s c u s s i o n. It is incorrectly stated in the International Fossil Plant Names
<taxonomicName id="4C2FD2C6FFB4FFE0DE59FFEDFBB0F904" box="[1065,1125,1667,1691]" genus="Index" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="8" pageNumber="22" phylum="Mollusca" rank="genus">Index</taxonomicName>
(http://fossilplants.info/) that the first use of
<taxonomicName id="4C2FD2C6FFB4FFE0D9BBFFCDFB86F924" authorityName="Sternberg" authorityYear="1825" box="[971,1107,1699,1723]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB4FFE0D9BBFFCDFB86F924" box="[971,1107,1699,1723]" italics="true" pageId="8" pageNumber="17">Sphenopteris</emphasis>
</taxonomicName>
as a genus is in
<bibRefCitation id="EFBED4B4FFB4FFE0DF66FFCDFCA5F944" author="Sternberg, K. M. von" pageId="8" pageNumber="17" refId="ref20725" refString="Sternberg, K. M. von (1823): Versuch einer geognostischbotanischen Darstellung der Flora der Vorwelt, Vol. I, 3. - Ernst Brenck's Wittwe, Regensburg, 39 pp." type="book" year="1823">Sternberg (1823: 36)</bibRefCitation>
, but Sternberg clearly stated that he regarded it here as a section (“Abteilung”) and earlier (p. 27) that it was of
<taxonomicName id="4C2FD2C6FFB5FFE1DA1AF9F9FF6AFF30" box="[106,191,151,175]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DA1AF9F9FF6AFF30" box="[106,191,151,175]" italics="true" pageId="9" pageNumber="18">Filicites</emphasis>
</taxonomicName>
. The first use of
<emphasis id="B95B7557FFB5FFE1DB1CF9F9FDC6FF30" box="[364,531,151,175]" italics="true" pageId="9" pageNumber="18">Sph(a)enopteris</emphasis>
as a generic name is in the 1825 Tentamen of Sternberg. Although when first used as a generic name the spelling was altered to
<taxonomicName id="4C2FD2C6FFB5FFE1D823F9B6FD3DFF6F" authorityName="Sternberg" authorityYear="1825" box="[595,744,216,240]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphaenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1D823F9B6FD3DFF6F" box="[595,744,216,240]" italics="true" pageId="9" pageNumber="18">Sphaenopteris</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFBED4B4FFB5FFE1DAFAF996FEE8FE8F" author="Sternberg, K. M. von" box="[138,317,248,272]" pageId="9" pageNumber="18" refId="ref20761" refString="Sternberg, K. M. von (1825): Versuch einer geognostischbotanischen Darstellung der Flora der Vorwelt, Vol. I, 4. - Ernst Brenck's Wittwe, Regensburg, 48 pp., tent. I-XLII." type="book" year="1825">Sternberg (1825)</bibRefCitation>
, the original orthography of
<bibRefCitation id="EFBED4B4FFB5FFE1D816F996FF6EFEAE" author="Brongniart, A." pageId="9" pageNumber="18" pagination="203 - 240" refId="ref15150" refString="Brongniart, A. (1822 a): Sur la classification et la distribution des vegetaux fossiles en general, et sur ceux des terrains de sediment superieur en particulier. Introduction, Chapitre I. - Memoires du Museum d'Histoire Naturelle, 8: 203 - 240, pls 1 - 4." type="journal article" year="1822">Brongniarts (1822a)</bibRefCitation>
section must be retained (ICN, Art. 60.1).
</paragraph>
<caption id="DF50F9CDFFB4FFE0DAE3FE3EFCC2F7AC" pageId="8" pageNumber="17">
<paragraph id="8B90A945FFB4FFE0DAE3FE3EFCC2F7AC" blockId="8.[147,1463,1872,2099]" pageId="8" pageNumber="17">
<emphasis id="B95B7557FFB4FFE0DAE3FE3EFF0FF7AC" bold="true" pageId="8" pageNumber="17">
Text-fig. 2. Illustrations from
<bibRefCitation id="EFBED4B4FFB4FFE0DBCEFE3EFDACF8FA" author="Brongniart, A." box="[446,633,1872,1893]" pageId="8" pageNumber="17" pagination="203 - 240" refId="ref15150" refString="Brongniart, A. (1822 a): Sur la classification et la distribution des vegetaux fossiles en general, et sur ceux des terrains de sediment superieur en particulier. Introduction, Chapitre I. - Memoires du Museum d'Histoire Naturelle, 8: 203 - 240, pls 1 - 4." type="journal article" year="1822">Brongniart (1822a</bibRefCitation>
, b) of specimens that remain legitimate taxonomic types of fossil-species. All same scale as in original plate unless otherwise stated. a, b)
<taxonomicName id="4C2FD2C6FFB4FFE0D8D6FE00FB9DF81D" authority="BRONGNIART" authorityName="BRONGNIART" box="[678,1096,1902,1923]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="brardii" subGenus="Odontopteris">
<emphasis id="B95B7557FFB4FFE0D8D6FE00FD21F81C" bold="true" box="[678,756,1902,1923]" italics="true" pageId="8" pageNumber="17">Filicites</emphasis>
(
<emphasis id="B95B7557FFB4FFE0D970FE00FCA8F81C" bold="true" box="[768,893,1902,1923]" italics="true" pageId="8" pageNumber="17">Odontopteris</emphasis>
)
<emphasis id="B95B7557FFB4FFE0D9FAFE00FC18F81C" bold="true" box="[906,973,1902,1923]" italics="true" pageId="8" pageNumber="17">brardii</emphasis>
BRONGNIART
</taxonomicName>
(≡
<taxonomicName id="4C2FD2C6FFB4FFE0DE17FE00FF28F800" authority="(BRONGNIART) STERNBERG" authorityName="STERNBERG" baseAuthorityName="BRONGNIART" class="Polypodiopsida" family="Lygodiaceae" genus="Odontopteris" higherTaxonomySource="CoL" kingdom="Plantae" order="Schizaeales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="brardii">
<emphasis id="B95B7557FFB4FFE0DE17FE00FAF8F81C" bold="true" box="[1127,1325,1902,1923]" italics="true" pageId="8" pageNumber="17">Odontopteris brardii</emphasis>
(BRONGNIART) STERNBERG
</taxonomicName>
) (a × 0.5). c)
<taxonomicName id="4C2FD2C6FFB4FFE0DB0DFEE5FCF8F83F" authority="BRONGNIART, Miocene" authorityName="BRONGNIART, Miocene" authorityYear="1822" box="[381,813,1931,1952]" class="Magnoliopsida" family="Arecaceae" genus="Palmacites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Arecales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="lamanonis">
<emphasis id="B95B7557FFB4FFE0DB0DFEE5FD87F83F" bold="true" box="[381,594,1931,1952]" italics="true" pageId="8" pageNumber="17">Palmacites lamanonis</emphasis>
BRONGNIART, Miocene
</taxonomicName>
Series, Aix-en-Provence, France, × 0.67. d)
<emphasis id="B95B7557FFB4FFE0DEAEFEE5FF0EF822" bold="true" italics="true" pageId="8" pageNumber="17">Sphenophyllites emarginatus</emphasis>
BRONGNIART (≡
<taxonomicName id="4C2FD2C6FFB4FFE0DB08FEC6FC74F823" authority="(BRONGNIART) BRONGNIART" authorityName="BRONGNIART" baseAuthorityName="BRONGNIART" box="[376,929,1960,1981]" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="emarginatum">
<emphasis id="B95B7557FFB4FFE0DB08FEC6FD4FF822" bold="true" box="[376,666,1960,1981]" italics="true" pageId="8" pageNumber="17">Sphenophyllum emarginatum</emphasis>
(BRONGNIART) BRONGNIART
</taxonomicName>
). e)
<emphasis id="B95B7557FFB4FFE0D9BDFEC6FBF3F822" bold="true" box="[973,1062,1960,1981]" italics="true" pageId="8" pageNumber="17">Fucoides</emphasis>
<taxonomicName id="4C2FD2C6FFB4FFE0DE5CFEC6FA8BF823" authority="AGARDH ex BRONGNIART" authorityName="BRONGNIART" box="[1068,1374,1960,1981]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="strictus">strictus AGARDH ex BRONGNIART</taxonomicName>
, Lile de Aix lignite mine, near La Rochelle, France. f)
<taxonomicName id="4C2FD2C6FFB4FFE0D82EFEA8FC5CF845" authority="BRONGNIART" authorityName="BRONGNIART" box="[606,905,1990,2011]" class="Magnoliopsida" genus="Phyllites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="populina">
<emphasis id="B95B7557FFB4FFE0D82EFEA8FCD8F844" bold="true" box="[606,781,1990,2011]" italics="true" pageId="8" pageNumber="17">Phyllites populina</emphasis>
BRONGNIART
</taxonomicName>
(≡
<taxonomicName id="4C2FD2C6FFB4FFE0D9DAFEA8FA9BF845" authority="(BRONGNIART) KNOBLOCH" authorityName="KNOBLOCH" baseAuthorityName="BRONGNIART" box="[938,1358,1990,2011]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="populina">
<emphasis id="B95B7557FFB4FFE0D9DAFEA8FB83F844" bold="true" box="[938,1110,1990,2011]" italics="true" pageId="8" pageNumber="17">Populus populina</emphasis>
(BRONGNIART) KNOBLOCH
</taxonomicName>
), Miocene Lagerstätte, Öhningen, Switzerland. g)
<taxonomicName id="4C2FD2C6FFB4FFE0D869FE8DFC55F868" authority="BRONGNIART" authorityName="BRONGNIART" authorityYear="1822" box="[537,896,2019,2040]" genus="Carpolithes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" rank="species" species="thalictroides">
<emphasis id="B95B7557FFB4FFE0D869FE8DFCD3F867" bold="true" box="[537,774,2019,2040]" italics="true" pageId="8" pageNumber="17">Carpolithes thalictroides</emphasis>
BRONGNIART
</taxonomicName>
(≡
<taxonomicName id="4C2FD2C6FFB4FFE0D9EFFE8DFABDF868" authority="(BRONGNIART) CHANDLER" authorityName="CHANDLER" baseAuthorityName="BRONGNIART" box="[927,1384,2019,2040]" class="Insecta" family="Carabidae" genus="Stratiotes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="17" phylum="Arthropoda" rank="species" species="thalictroides">
<emphasis id="B95B7557FFB4FFE0D9EFFE8DFBA2F867" bold="true" box="[927,1143,2019,2040]" italics="true" pageId="8" pageNumber="17">Stratiotes thalictroides</emphasis>
(BRONGNIART) CHANDLER
</taxonomicName>
), upper Oligocene Calcaire de Beauce, Paris Basin, France. h)
<taxonomicName id="4C2FD2C6FFB4FFE0D8CBF16EFA9EF78A" authority="BRONGNIART, Eocene Monte Bolca Lagerstatte" authorityName="BRONGNIART, Eocene Monte Bolca Lagerstatte" authorityYear="1822" box="[699,1355,2048,2069]" family="Glossophyllaceae" genus="Antholithes" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" rank="species" species="liliacea">
<emphasis id="B95B7557FFB4FFE0D8CBF16EFCADF78A" bold="true" box="[699,888,2048,2069]" italics="true" pageId="8" pageNumber="17">Antholithes liliacea</emphasis>
BRONGNIART, Eocene Monte Bolca Lagerstätte
</taxonomicName>
in Veneto, Italy. i)
</emphasis>
<taxonomicName id="4C2FD2C6FFB4FFE0DA90F170FCC2F7AC" authority="BRONGNIART, Pennsylvanian Subsystem." box="[224,791,2078,2099]" class="Lycopodiopsida" genus="Poacites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="8" pageNumber="17" phylum="Tracheophyta" rank="species" species="carinata">
<emphasis id="B95B7557FFB4FFE0DA90F170FDD6F7AD" bold="true" box="[224,515,2078,2099]" pageId="8" pageNumber="17">
<emphasis id="B95B7557FFB4FFE0DA90F170FE5DF7AC" bold="true" box="[224,392,2078,2099]" italics="true" pageId="8" pageNumber="17">Poacites carinata</emphasis>
BRONGNIART
</emphasis>
, Pennsylvanian Subsystem.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B90A945FFB5FFE1DAE0F857FDE7FC8B" blockId="9.[106,745,151,1820]" pageId="9" pageNumber="18">
This is one of the most confused of the Brongniart fossil-taxa, and is still regularly used for both fern and seedplant fossil-species. However, the
<typeStatus id="549417E7FFB5FFE1DBACF814FDDFFE0D" box="[476,522,378,402]" pageId="9" pageNumber="18">type</typeStatus>
(
<taxonomicName id="4C2FD2C6FFB5FFE1D86BF814FD52FE0E" authorityName="STERNBERG" authorityYear="1825" baseAuthorityName="BRONGNIART" box="[539,647,377,401]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis id="B95B7557FFB5FFE1D86BF814FD52FE0E" box="[539,647,377,401]" italics="true" pageId="9" pageNumber="18">S. elegans</emphasis>
</taxonomicName>
) is welldocumented as a Mississippian-age early seed-plant (e.g.
<bibRefCitation id="EFBED4B4FFB5FFE1DA1AF8D4FED6FE4D" author="Kidston, R." box="[106,259,442,466]" pageId="9" pageNumber="18" pagination="1 - 670" refId="ref18508" refString="Kidston, R. (1923 - 1925): Fossil plants of the Carboniferous rocks of Great Britain. - Memoirs of the Geological Survey of Great Britain, Palaeontology, 2: 1 - 670." type="journal article" year="1923">Kidston 1923</bibRefCitation>
-1925: 242) and so the name
<taxonomicName id="4C2FD2C6FFB5FFE1D82DF8D4FD3DFE4D" authorityName="Sternberg" authorityYear="1825" box="[605,744,442,466]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1D82DF8D4FD3DFE4D" box="[605,744,442,466]" italics="true" pageId="9" pageNumber="18">Sphenopteris</emphasis>
</taxonomicName>
should only be applied to this group of early seed-plant fronds. It should be noted in passing that Kidston adopted for this species the taxonomic synonym
<emphasis id="B95B7557FFB5FFE1D815FB74FD3CFDAD" box="[613,745,538,562]" italics="true" pageId="9" pageNumber="18">Diplotmema</emphasis>
<taxonomicName id="4C2FD2C6FFB5FFE1DA1AFB55FD17FDCC" authority="(SCHLOTHEIM ex POTONIE) GOTHAN, 1913" authorityName="GOTHAN" authorityYear="1913" baseAuthorityName="SCHLOTHEIM ex POTONIE" box="[106,706,571,595]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="adiantioides">
<emphasis id="B95B7557FFB5FFE1DA1AFB55FF3BFDCC" box="[106,238,571,595]" italics="true" pageId="9" pageNumber="18">adiantioides</emphasis>
(SCHLOTHEIM ex POTONIÉ)
<bibRefCitation id="EFBED4B4FFB5FFE1D853FB55FD17FDCC" author="Gothan, W." box="[547,706,571,595]" pageId="9" pageNumber="18" pagination="1 - 278" refId="ref17867" refString="Gothan, W. (1913): Die oberschlesische Steinkohlenflora. 1 Teil. - Abhandlungen der Koniglich Preussischen Geologischen Landesanstalt, Neue Folge, 75: 1 - 278." type="journal article" year="1913">GOTHAN, 1913</bibRefCitation>
</taxonomicName>
(≡
<emphasis id="B95B7557FFB5FFE1DA1AFB35FED4FDED" box="[106,257,603,626]" italics="true" pageId="9" pageNumber="18">Cuneatopteris</emphasis>
<taxonomicName id="4C2FD2C6FFB5FFE1DB7DFB35FF40FD0C" authority="SCHLOTHEIM ex POTONIE, 1907, p. 2" authorityName="POTONIE" authorityPageNumber="2" authorityYear="1907" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="adiantoides">
<emphasis id="B95B7557FFB5FFE1DB7DFB35FE5CFDEC" box="[269,393,603,627]" italics="true" pageId="9" pageNumber="18">adiantoides</emphasis>
SCHLOTHEIM ex
<bibRefCitation id="EFBED4B4FFB5FFE1D835FB35FD30FDEC" author="Potonie, H." box="[581,741,603,627]" pageId="9" pageNumber="18" refId="ref19941" refString="Potonie, H. (1907): Abbildungen und Beschreibungen fossiler Pflanzen-Reste, Lieferung 5, Nr. 81: Cuneatopteris. - Koniglich Preussische Geologische Landesanstalt, Berlin, 2 pp." type="book" year="1907">POTONIÉ, 1907</bibRefCitation>
, p. 2
</taxonomicName>
) since the first use of the species epithet (as “
<taxonomicName id="4C2FD2C6FFB5FFE1D8E1FB15FE62FD2C" authority="&quot; SCHLOTHEIM, 1820" authorityName="SCHLOTHEIM" authorityYear="1820" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="adiantoides">
<emphasis id="B95B7557FFB5FFE1D8E1FB15FF33FD2C" italics="true" pageId="9" pageNumber="18">Filicites adiantoides</emphasis>
<bibRefCitation id="EFBED4B4FFB5FFE1DA88FBF5FE62FD2C" author="Schlotheim, E. F. von" box="[248,439,667,691]" pageId="9" pageNumber="18" refId="ref20364" refString="Schlotheim, E. F. von (1820): Die Petrefactenkunde auf ihrem jetzigen Standpunkte durch die Beschreibung seiner Sammlung versteinerter und fossiler Uberreste des Thier- und Pflanzenreichs der Vorwelt erlautert. - Becker, Gotha, 437 pp." type="book" year="1820">SCHLOTHEIM, 1820</bibRefCitation>
</taxonomicName>
) pre-dated both Brongniarts and Sternbergs works. However, as “
<taxonomicName id="4C2FD2C6FFB5FFE1D879FBD2FD7FFD4B" authorityName="GOTHAN" authorityYear="1913" baseAuthorityName="SCHLOTHEIM ex POTONIE" box="[521,682,700,724]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="adiantioides">
<emphasis id="B95B7557FFB5FFE1D879FBD2FD7FFD4B" box="[521,682,700,724]" italics="true" pageId="9" pageNumber="18">F. adiantioides</emphasis>
</taxonomicName>
” was published prior to the starting point for publication of fossiltaxa (ICN, Art. 13.1) this must be rejected.
</paragraph>
<paragraph id="8B90A945FFB5FFE1DAE0FA72FDADF904" blockId="9.[106,745,151,1820]" pageId="9" pageNumber="18">
This raises a practical problem with classifying some fossil fern fronds with small lobed pinnules, especially of Palaeozoic age. Although there are numerous welldefined fossil-genera for such ferns (e.g.
<taxonomicName id="4C2FD2C6FFB5FFE1D852FA13FF77FC2A" authority="ZEILLER, 1883" authorityName="ZEILLER" authorityYear="1883" class="Polypodiopsida" family="Marattiaceae" genus="Renaultia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1D852FA13FD5FFC0A" box="[546,650,893,917]" italics="true" pageId="9" pageNumber="18">Renaultia</emphasis>
<bibRefCitation id="EFBED4B4FFB5FFE1D8E1FA13FF77FC2A" author="Zeiller, R." pageId="9" pageNumber="18" pagination="177 - 209" refId="ref21541" refString="Zeiller, R. (1883): Fructifications de fougeres du terrain houiller. - Annales des Sciences Naturelles, 6 e Serie, Botanique, 16: 177 - 209." type="journal article" year="1883">ZEILLER, 1883</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C2FD2C6FFB5FFE1DAC5FAF3FE10FC2A" authority="KIDSTON, 1884" authorityName="KIDSTON" authorityYear="1884" box="[181,453,925,949]" class="Polypodiopsida" family="Marattiaceae" genus="Zeilleria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DAC5FAF3FEC4FC2A" box="[181,273,925,949]" italics="true" pageId="9" pageNumber="18">Zeilleria</emphasis>
<bibRefCitation id="EFBED4B4FFB5FFE1DB51FAF0FE10FC2A" author="Kidston, R." box="[289,453,925,949]" pageId="9" pageNumber="18" pagination="590 - 598" refId="ref18431" refString="Kidston, R. (1884): On the fructification of Zeilleria (Sphenopteris) delicatula, Sternb., sp.; with remarks on Urnatopteris (Sphenopteris) tenella, Brongt., and Hymenophyllites (Sphenopteris) quadridactylites, Gutbier, sp. - Quarterly Journal of the Geological Society, 40: 590 - 598. https: // doi. org / 10.1144 / GSL. JGS. 1884.40.01 - 04.49" type="journal article" year="1884">KIDSTON, 1884</bibRefCitation>
</taxonomicName>
) these are diagnosed by their reproductive structures. Those species for which reproductive structures are unknown have tended to be placed in
<taxonomicName id="4C2FD2C6FFB5FFE1DAA8FA90FEB6FB89" authorityName="Sternberg" authorityYear="1825" box="[216,355,1022,1046]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DAA8FA90FEB6FB89" box="[216,355,1022,1046]" italics="true" pageId="9" pageNumber="18">Sphenopteris</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation id="EFBED4B4FFB5FFE1DBD7FA90FDBCFB89" author="Brousmiche, C." box="[423,617,1022,1046]" pageId="9" pageNumber="18" refId="ref16362" refString="Brousmiche, C. (1983): Les fougeres sphenopteridiennes du bassin houiller Sarro-Lorrain (2 vols) (Publication no. 10). - Societe Geologique du Nord, Lille, 480 pp. + 100 pls." type="book" year="1983">Brousmiche 1983</bibRefCitation>
) but this is not tenable given that the
<typeStatus id="549417E7FFB5FFE1DBF3FD71FE64FBA8" box="[387,433,1055,1079]" pageId="9" pageNumber="18">type</typeStatus>
of that name is a seed-plant. There is in fact no available legitimate name for a fossilgenus of species of such fern fossils for which reproductive structures are unknown. It would be possible to establish an “artificial” genus for such species; this would be analogous to the fossil-genus
<taxonomicName id="4C2FD2C6FFB5FFE1DB34FDAEFD72FB48" authority="BRONGNIART, 1828 a" authorityName="BRONGNIART" authorityYear="1828" box="[324,679,1215,1239]" class="Polypodiopsida" family="Pteridaceae" genus="Taeniopteris" higherTaxonomySource="CoL" kingdom="Plantae" order="Polypodiales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DB34FDAEFE1DFB48" box="[324,456,1216,1239]" italics="true" pageId="9" pageNumber="18">Taeniopteris</emphasis>
<bibRefCitation id="EFBED4B4FFB5FFE1DBA4FDAEFD72FB48" author="Brongniart, A." box="[468,679,1215,1239]" pageId="9" pageNumber="18" pagination="1 - 223" refId="ref15409" refString="Brongniart, A. (1828 a): Prodrome d'une Histoire des vegetaux fossiles. F. G. Levrault, Paris and Strasbourg, 223 pp [also published in Dictionnaire des sciences naturelles, 57: 1 - 223]." type="book chapter" year="1828">BRONGNIART, 1828a</bibRefCitation>
</taxonomicName>
nom. cons., which is now used for entire cycadophyte-like leaves for which neither cuticles nor reproductive structures are known (van Konijnenburg-van Cittert et al. 2017). However, to date no such fossil-genus has been established for these fern fossils.Another option would be to revert to
<bibRefCitation id="EFBED4B4FFB5FFE1D813FC0EFF68FA06" author="Brongniart, A." pageId="9" pageNumber="18" pagination="203 - 240" refId="ref15150" refString="Brongniart, A. (1822 a): Sur la classification et la distribution des vegetaux fossiles en general, et sur ceux des terrains de sediment superieur en particulier. Introduction, Chapitre I. - Memoires du Museum d'Histoire Naturelle, 8: 203 - 240, pls 1 - 4." type="journal article" year="1822">Brongniarts (1822a)</bibRefCitation>
original conception for
<taxonomicName id="4C2FD2C6FFB5FFE1DBC8FCEFFD96FA06" authorityName="Sternberg" authorityYear="1825" box="[440,579,1409,1433]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DBC8FCEFFD96FA06" box="[440,579,1409,1433]" italics="true" pageId="9" pageNumber="18">Sphenopteris</emphasis>
</taxonomicName>
as including all pteridophyllous foliage with lobed pinnules and supressing names such as
<taxonomicName id="4C2FD2C6FFB5FFE1DB7DFCAFFEA0FA46" authorityName="ZEILLER" authorityYear="1883" box="[269,373,1473,1497]" class="Polypodiopsida" family="Marattiaceae" genus="Renaultia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DB7DFCAFFEA0FA46" box="[269,373,1473,1497]" italics="true" pageId="9" pageNumber="18">Renaultia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C2FD2C6FFB5FFE1DBDBFCAFFDD2FA46" authorityName="KIDSTON" authorityYear="1884" box="[427,519,1473,1497]" class="Polypodiopsida" family="Marattiaceae" genus="Zeilleria" higherTaxonomySource="GBIF" kingdom="Plantae" order="Marattiales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B95B7557FFB5FFE1DBDBFCAFFDD2FA46" box="[427,519,1473,1497]" italics="true" pageId="9" pageNumber="18">Zeilleria</emphasis>
</taxonomicName>
but this would seem a retrograde step. The only viable option within the current framework of the ICN is to make a judgement as to the taxonomic position of such species based on similarities in the morphology of the sterile foliage, maybe using an open-nomenclature-like qualifier similar to that used in palaeozoology (
<bibRefCitation id="EFBED4B4FFB5FFE1DB63FFEDFE68F904" author="Matthews, S. C." box="[275,445,1667,1691]" pageId="9" pageNumber="18" pagination="713 - 719" refId="ref19758" refString="Matthews, S. C. (1973): Notes on open nomenclature and on synonymy lists. - Palaeontology, 16: 713 - 719." type="journal article" year="1973">Matthews 1973</bibRefCitation>
,
<bibRefCitation id="EFBED4B4FFB5FFE1DBB7FFEDFDB9F904" author="Bengtson, P." box="[455,620,1667,1691]" pageId="9" pageNumber="18" pagination="223 - 227" refId="ref15058" refString="Bengtson, P. (1988): Open nomenclature. - Palaeontology, 31: 223 - 227." type="journal article" year="1988">Bengtson 1988</bibRefCitation>
).
</paragraph>
<paragraph id="8B90A945FFB5FFE1DAE0FFCDFDEBF884" blockId="9.[106,745,151,1820]" pageId="9" pageNumber="18">
In passing, mention should be made of a later homonym of
<taxonomicName id="4C2FD2C6FFB5FFE1DAF8FFAAFF21F944" authorityName="STERNBERG" authorityYear="1825" baseAuthorityName="BRONGNIART" box="[136,244,1731,1755]" genus="Filicites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis id="B95B7557FFB5FFE1DAF8FFAAFF21F944" box="[136,244,1731,1755]" italics="true" pageId="9" pageNumber="18">S. elegans</emphasis>
</taxonomicName>
,
<taxonomicName id="4C2FD2C6FFB5FFE1DB73FFADFD3DF944" authority="(YOKOYAMA) OISI, 1940" authorityName="OISI" authorityYear="1940" baseAuthorityName="YOKOYAMA" box="[259,744,1731,1758]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis id="B95B7557FFB5FFE1DB73FFADFE32F944" box="[259,487,1731,1755]" italics="true" pageId="9" pageNumber="18">Sphenopteris elegans</emphasis>
(YOKOYAMA) ÔISI, 1940
</taxonomicName>
(≡
<taxonomicName id="4C2FD2C6FFB5FFE1DAFBFF8DFDFEF964" authority="YOKOYAMA, 1895" authorityName="YOKOYAMA" authorityYear="1895" box="[139,555,1763,1787]" class="Polypodiopsida" family="Dicksoniaceae" genus="Onychiopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="9" pageNumber="18" phylum="Tracheophyta" rank="species" species="elegans">
<emphasis id="B95B7557FFB5FFE1DAFBFF8DFEBCF964" box="[139,361,1763,1787]" italics="true" pageId="9" pageNumber="18">Onychiopsis elegans</emphasis>
<bibRefCitation id="EFBED4B4FFB5FFE1DB02FF8AFDFEF964" author="Yokoyama, M." box="[370,555,1763,1787]" pageId="9" pageNumber="18" pagination="201 - 231" refId="ref21502" refString="Yokoyama, M. (1895): Mesozoic plants from Kozuke, Kii, Awa, and Tosa. - Journal of the College of Science, Imperial University, Japan, 7: 201 - 231." type="journal article" year="1895">YOKOYAMA, 1895</bibRefCitation>
</taxonomicName>
) based on a
<typeStatus id="549417E7FFB5FFE1D8CBFF8AFD3CF963" box="[699,745,1764,1788]" pageId="9" pageNumber="18">type</typeStatus>
from the Lower Cretaceous Series of
<collectingCountry id="F338E9D5FFB5FFE1DB8AFE6AFDEFF884" box="[506,570,1796,1819]" name="Japan" pageId="9" pageNumber="18">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>