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<document id="91BB9390D3442A83D13E7F1BAE4CA542" ID-DOI="10.4081/nhs.2020.474" ID-ISSN="2385-0922" ID-Zenodo-Dep="12753933" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1721185017991" checkinUser="felipe" docAuthor="Rosa, Paolo &amp; Pavesi, Maurizio" docDate="2020" docId="0382F67E3D73FFA0E536D5A3781564DB" docLanguage="en" docName="NatHistSci.7.2.39-56.pdf" docOrigin="Natural History Sciences 7 (2)" docSource="http://dx.doi.org/10.4081/nhs.2020.474" docStyle="DocumentStyle:926963A4B76E17DFFD817EF231EB8DA9.3:NatHistSci.2014-.journal_article" docStyleId="926963A4B76E17DFFD817EF231EB8DA9" docStyleName="NatHistSci.2014-.journal_article" docStyleVersion="3" docTitle="Holopyga enslini Linsenmaier 1959" docType="treatment" docVersion="1" lastPageNumber="46" masterDocId="FFBB8E063D74FFA8E676D15C791B6169" masterDocTitle="The case of Holopyga gogorzae Trautmann, 1926 and revision of the H. miranda group (Hymenoptera, Chrysididae)" masterLastPageNumber="56" masterPageNumber="39" pageNumber="46" updateTime="1721982236259" updateUser="tatiana" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="9EFBCECB5CC43A99D5BBE685F8EDA92E">The case of Holopyga gogorzae Trautmann, 1926 and revision of the H. miranda group (Hymenoptera, Chrysididae)</mods:title>
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<mods:namePart id="B0E1368BEF0B2FE6CC39F0EF58281E7E">Rosa, Paolo</mods:namePart>
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<mods:title id="9ADF0397A3624F70EA10A4B1F8B54BAF">Natural History Sciences</mods:title>
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<mods:date id="8C86DFF3B54FEEF4E72789A6F25F8C90">2020</mods:date>
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<treatment id="0382F67E3D73FFA0E536D5A3781564DB" LSID="urn:lsid:plazi:treatment:0382F67E3D73FFA0E536D5A3781564DB" httpUri="http://treatment.plazi.org/id/0382F67E3D73FFA0E536D5A3781564DB" lastPageId="8" lastPageNumber="46" pageId="7" pageNumber="46">
<subSubSection id="C33114E33D73FFAFE536D5A37CBD6470" box="[832,1446,1279,1306]" pageId="7" pageNumber="46" type="nomenclature">
<paragraph id="8B9447683D73FFAFE536D5A37CBD6470" blockId="7.[832,1446,1279,1306]" box="[832,1446,1279,1306]" pageId="7" pageNumber="46">
<heading id="D0DCF0043D73FFAFE536D5A37CBD6470" box="[832,1446,1279,1306]" fontSize="10" level="2" pageId="7" pageNumber="46" reason="6">
<taxonomicName id="4C2B3CEB3D73FFAFE536D5A37DFD6470" authority="Linsenmaier, 1959" authorityName="Linsenmaier" authorityYear="1959" box="[832,1254,1279,1306]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="B95F9B7A3D73FFAFE536D5A37DFD6470" bold="true" box="[832,1254,1279,1306]" pageId="7" pageNumber="46">
<emphasis id="B95F9B7A3D73FFAFE536D5A37D1E6473" bold="true" box="[832,1029,1279,1306]" italics="true" pageId="7" pageNumber="46">Holopyga enslini</emphasis>
<bibRefCitation id="EFBA3A993D73FFAFE27DD5A37DFD6470" author="Linsenmaier W." box="[1035,1254,1279,1305]" pageId="7" pageNumber="46" pagination="1 - 232" refId="ref12179" refString="Linsenmaier W., 1959 - Revision der Familie Chrysi- didae (Hymenoptera) mit besonderer Berucksich- tigung der europaischen Spezies. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 32 (1): 1 - 232." type="journal article" year="1959">Linsenmaier, 1959</bibRefCitation>
</emphasis>
</taxonomicName>
(
<figureCitation id="13105BED3D73FFAFE283D5A37C4D6470" box="[1269,1366,1279,1305]" captionStart="Fig" captionStartId="4.[189,222,1489,1511]" captionTargetBox="[113,1474,879,1456]" captionTargetId="figure-4@4.[113,1474,873,1464]" captionTargetPageId="4" captionText="Fig. 4 - Holopyga species in dorsal view. / Holopyga spp., visione dorsale. A-B) Holopyga calida Linsenmaier, 1951. A) ♀ paratype from Marrakech, Morocco; / ♀ paratypus di Marrakech, Marocco; B) ♂ from Mohammedia, Morocco. / ♂ di Mohammedia, Marocco. C) Holopyga mattheyi Linsenmaier, 1959, ♂ paratype from Marrakech, Morocco. / ♂ paratypus di Marrakech, Marocco. D) Holopyga lucens Rosa, 2018, ♂ paratype from Karabata, Turkmenistan. / ♂ paratypus di Karabata, Turkmenistan. E) Holopyga enslini Linsenmaier, 1959, ♂ paratype from Beersheba, Israel. / ♂ paratypus di Beersheba, Israele. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12753941" httpUri="https://zenodo.org/record/12753941/files/figure.png" pageId="7" pageNumber="46">Figs. 4E</figureCitation>
,
<figureCitation id="13105BED3D73FFAFE317D5A37C766470" box="[1377,1389,1279,1305]" captionStart="Fig" captionStartId="8.[189,222,1104,1126]" captionTargetBox="[113,1474,136,1079]" captionTargetId="figure-334@8.[113,1474,136,1079]" captionTargetPageId="8" captionText="Fig. 8 - Holopyga enslini Linsenmaier, 1959; ♂ paratype / paratypus.A) habitus, lateral view / habitus, visione laterale; B) head, frontal view / capo, visione frontale; C) head, dorsal view / capo, visione dorsale; D) mesosoma, dorsal view / mesosoma, visione dorsale; E) metasoma, postero-lateral view / metasoma, visione postero-laterale; F) metasoma, ventral view / metasoma, visione ventrale." figureDoi="http://doi.org/10.5281/zenodo.12753949" httpUri="https://zenodo.org/record/12753949/files/figure.png" pageId="7" pageNumber="46">8</figureCitation>
A-F)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33114E33D73FFAFE536D4677DB4641D" pageId="7" pageNumber="46" type="reference_group">
<paragraph id="8B9447683D73FFAFE536D4677DB4641D" blockId="7.[832,1474,1339,1396]" pageId="7" pageNumber="46">
<heading id="D0DCF0043D73FFAFE536D4677DB4641D" centered="true" fontSize="10" level="2" pageId="7" pageNumber="46" reason="3">
<treatmentCitationGroup id="AB3B60463D73FFAFE536D4677DB4641D" pageId="7" pageNumber="46">
<treatmentCitation id="0A8A61793D73FFAFE536D4677C05643C" author="Linsenmaier W." box="[832,1310,1339,1366]" page="27" pageId="7" pageNumber="46" year="1959">
<taxonomicName id="4C2B3CEB3D73FFAFE536D4677C05643C" authority="Linsenmaier, 1959: 27" authorityName="Linsenmaier" authorityPageNumber="27" authorityYear="1959" box="[832,1310,1339,1366]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="B95F9B7A3D73FFAFE536D4677D1D643C" box="[832,1030,1339,1365]" italics="true" pageId="7" pageNumber="46">Holopyga enslini</emphasis>
<materialsCitation id="3B434D353D73FFAFE265D4677DB4641D" collectionCode="NMLU" country="Turkey" location="Ulu Kizlar" pageId="7" pageNumber="46" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<bibRefCitation id="EFBA3A993D73FFAFE265D4677C05643C" author="Linsenmaier W." box="[1043,1310,1339,1366]" pageId="7" pageNumber="46" pagination="1 - 232" refId="ref12179" refString="Linsenmaier W., 1959 - Revision der Familie Chrysi- didae (Hymenoptera) mit besonderer Berucksich- tigung der europaischen Spezies. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 32 (1): 1 - 232." type="journal article" year="1959">Linsenmaier, 1959: 27</bibRefCitation>
.
<typeStatus id="5490F9CA3D73FFAFE346D4677C83643C" box="[1328,1432,1339,1365]" pageId="7" pageNumber="46" type="holotype">Holotype</typeStatus>
<specimenCount id="9D2D8CE13D73FFAFE3D3D4607CA6643F" box="[1445,1469,1340,1366]" count="1" pageId="7" pageNumber="46" type="male"></specimenCount>
;
<collectingCountry id="F33C07F83D73FFAFE513D4057AA2641A" box="[869,953,1369,1395]" name="Turkey" pageId="7" pageNumber="46">Turkey</collectingCountry>
:
<location id="8EF411B33D73FFAFE5B2D4057D20641A" LSID="urn:lsid:plazi:treatment:0382F67E3D73FFA0E536D5A3781564DB:8EF411B33D73FFAFE5B2D4057D20641A" box="[964,1083,1369,1395]" country="Turkey" name="Ulu Kizlar" pageId="7" pageNumber="46">Ulu Kizlar</location>
(
<collectionCode id="ED3ADFAD3D73FFAFE23CD4067DB8641D" box="[1098,1187,1370,1396]" pageId="7" pageNumber="46">NMLU</collectionCode>
).
</materialsCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33114E33D73FFAFE513D4C97DF5672E" pageId="7" pageNumber="46" type="materials_examined">
<paragraph id="8B9447683D73FFAFE513D4C97DF5672E" blockId="7.[832,1475,1429,1968]" pageId="7" pageNumber="46">
<emphasis id="B95F9B7A3D73FFAFE513D4C97DC864D9" bold="true" box="[869,1235,1429,1456]" pageId="7" pageNumber="46">
Specimens examined.
<materialsCitation id="3B434D353D73FFAFE201D4CA7CD864A7" collectingDate="1952-07-05" collectorName="Ulu Kizlar &amp; Seidenstucker." country="Turkey" location="Ulukisla" pageId="7" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Nigde" typeStatus="holotype">
<collectingCountry id="F33C07F83D73FFAFE201D4CA7DD564D9" box="[1143,1230,1430,1456]" name="Turkey" pageId="7" pageNumber="46">Turkey</collectingCountry>
.
<emphasis id="B95F9B7A3D73FFAFE2A9D4CA7C3964D9" box="[1247,1314,1430,1456]" italics="true" pageId="7" pageNumber="46">
<collectingRegion id="49EF898A3D73FFAFE2A9D4CA7C3964D9" box="[1247,1314,1430,1456]" country="Turkey" name="Nigde" pageId="7" pageNumber="46">Niğde</collectingRegion>
</emphasis>
:
<specimenCount id="9D2D8CE13D73FFAFE341D4CA7C5764D9" box="[1335,1356,1430,1456]" count="1" pageId="7" pageNumber="46" type="male"></specimenCount>
<typeStatus id="5490F9CA3D73FFAFE32FD4CA7CA564D9" box="[1369,1470,1430,1456]" pageId="7" pageNumber="46" type="holotype">holotype</typeStatus>
,
<collectorName id="26DE22BE3D73FFAFE536D4E87AA064A7" box="[832,955,1460,1486]" pageId="7" pageNumber="46">Ulu Kizlar</collectorName>
[=
<location id="8EF411B33D73FFAFE59AD4EF7D4A64A7" LSID="urn:lsid:plazi:treatment:0382F67E3D73FFA0E536D5A3781564DB:8EF411B33D73FFAFE59AD4EF7D4A64A7" box="[1004,1105,1459,1486]" country="Turkey" name="Ulukisla" pageId="7" pageNumber="46" stateProvince="Nigde">Ulukışla</location>
],
<date id="FF9561A83D73FFAFE210D4E87DC164A6" box="[1126,1242,1460,1487]" pageId="7" pageNumber="46" value="1952-07-05">
<collectingDate id="EFD198403D73FFAFE210D4E87DC164A6" box="[1126,1242,1460,1487]" pageId="7" pageNumber="46" value="1952-07-05">5.vii.1952</collectingDate>
</date>
, leg.
<collectorName id="26DE22BE3D73FFAFE356D4E87CD864A7" box="[1312,1475,1460,1486]" pageId="7" pageNumber="46">Seidenstücker.</collectorName>
</materialsCitation>
</emphasis>
<materialsCitation id="3B434D353D73FFAFE536D48E7AB26485" box="[832,937,1490,1517]" country="Turkey" location="Agri" pageId="7" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Agri" typeStatus="holotype">
<emphasis id="B95F9B7A3D73FFAFE536D48E7A696485" box="[832,882,1490,1516]" italics="true" pageId="7" pageNumber="46">
<collectingRegion id="49EF898A3D73FFAFE536D48E7A696485" box="[832,882,1490,1516]" country="Turkey" name="Agri" pageId="7" pageNumber="46">Ağrı</collectingRegion>
</emphasis>
:
<specimenCount id="9D2D8CE13D73FFAFE5F3D48E7AB26485" box="[901,937,1490,1516]" count="1" pageId="7" pageNumber="46" type="male">1♂</specimenCount>
</materialsCitation>
,
<materialsCitation id="3B434D353D73FFAFE5CFD48E7C776485" box="[953,1388,1490,1517]" collectingDate="1993-06-27" collectorName="K. Denes." country="Turkey" location="Agri" pageId="7" pageNumber="46" specimenCount="1" stateProvince="Agri" typeStatus="holotype">
<collectingRegion id="49EF898A3D73FFAFE5CFD48E7AF76485" box="[953,1004,1490,1516]" country="Turkey" name="Agri" pageId="7" pageNumber="46">Ağrı</collectingRegion>
env.,
<date id="FF9561A83D73FFAFE24ED48E7DA96485" box="[1080,1202,1490,1516]" pageId="7" pageNumber="46" value="1993-06-27">
<collectingDate id="EFD198403D73FFAFE24ED48E7DA96485" box="[1080,1202,1490,1516]" pageId="7" pageNumber="46" value="1993-06-27">27.vi.1993</collectingDate>
</date>
, leg.
<collectorName id="26DE22BE3D73FFAFE281D48F7C776485" box="[1271,1388,1490,1517]" pageId="7" pageNumber="46">K. Deneš.</collectorName>
</materialsCitation>
<materialsCitation id="3B434D353D73FFAFE301D48E7C366740" collectingDate="1959-04-29" collectorName="Beersheba &amp; Nahal Nafha" country="Israel" location="Salz" pageId="7" pageNumber="46" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Southern" typeStatus="paratype">
<emphasis id="B95F9B7A3D73FFAFE301D48E7CD96485" bold="true" box="[1399,1474,1490,1516]" pageId="7" pageNumber="46">
<collectingCountry id="F33C07F83D73FFAFE301D48E7CA66485" box="[1399,1469,1490,1516]" name="Israel" pageId="7" pageNumber="46">Israel</collectingCountry>
.
</emphasis>
<emphasis id="B95F9B7A3D73FFAFE536D4AC7D1C6763" box="[832,1031,1520,1546]" italics="true" pageId="7" pageNumber="46">
<collectingRegion id="49EF898A3D73FFAFE536D4AC7D1C6763" box="[832,1031,1520,1546]" country="Israel" name="Southern" pageId="7" pageNumber="46">Southern District</collectingRegion>
</emphasis>
:
<specimenCount id="9D2D8CE13D73FFAFE26DD4AC7D5B6763" box="[1051,1088,1520,1546]" count="1" pageId="7" pageNumber="46" type="male">1♂</specimenCount>
,
<collectorName id="26DE22BE3D73FFAFE224D4AC7DD16763" box="[1106,1226,1520,1546]" pageId="7" pageNumber="46">Beersheba</collectorName>
, 10.v., leg. Bytinski-
<location id="8EF411B33D73FFAFE536D7527A686741" LSID="urn:lsid:plazi:treatment:0382F67E3D73FFA0E536D5A3781564DB:8EF411B33D73FFAFE536D7527A686741" box="[832,883,1550,1576]" country="Israel" name="Salz" pageId="7" pageNumber="46" stateProvince="Southern">Salz</location>
,
<typeStatus id="5490F9CA3D73FFAFE50BD7537AFA6740" box="[893,993,1551,1577]" pageId="7" pageNumber="46" type="paratype">paratype</typeStatus>
;
<specimenCount id="9D2D8CE13D73FFAFE59DD7527D166741" box="[1003,1037,1550,1576]" count="1" pageId="7" pageNumber="46" type="female">1♀</specimenCount>
,
<collectorName id="26DE22BE3D73FFAFE26CD7527DB16741" box="[1050,1194,1550,1576]" pageId="7" pageNumber="46">Nahal Nafha</collectorName>
,
<date id="FF9561A83D73FFAFE2C2D7527C366740" box="[1204,1325,1550,1577]" pageId="7" pageNumber="46" value="1959-04-29">
<collectingDate id="EFD198403D73FFAFE2C2D7527C366740" box="[1204,1325,1550,1577]" pageId="7" pageNumber="46" value="1959-04-29">29.iv.1959</collectingDate>
</date>
</materialsCitation>
, leg. J. Wahrman. All specimens housed at NMLU.
</paragraph>
</subSubSection>
<subSubSection id="C33114E33D73FFAFE513D7167C88670C" box="[869,1427,1610,1637]" pageId="7" pageNumber="46" type="distribution">
<paragraph id="8B9447683D73FFAFE513D7167C88670C" blockId="7.[832,1475,1429,1968]" box="[869,1427,1610,1637]" pageId="7" pageNumber="46">
<emphasis id="B95F9B7A3D73FFAFE513D7167AE4670D" bold="true" box="[869,1023,1610,1636]" pageId="7" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="F33C07F83D73FFAFE270D7167D5C670D" box="[1030,1095,1610,1636]" name="Israel" pageId="7" pageNumber="46">Israel</collectingCountry>
,
<collectingCountry id="F33C07F83D73FFAFE227D7167DBA670D" box="[1105,1185,1610,1636]" name="Turkey" pageId="7" pageNumber="46">Turkey</collectingCountry>
(
<bibRefCitation id="EFBA3A993D73FFAFE2C6D7167C9D670C" author="Linsenmaier W." box="[1200,1414,1610,1637]" pageId="7" pageNumber="46" pagination="1 - 232" refId="ref12179" refString="Linsenmaier W., 1959 - Revision der Familie Chrysi- didae (Hymenoptera) mit besonderer Berucksich- tigung der europaischen Spezies. Mitteilungen der Schweizerischen Entomologischen Gesellschaft, 32 (1): 1 - 232." type="journal article" year="1959">Linsenmaier, 1959</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33114E33D73FFA0E513D735781564DB" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="discussion">
<paragraph id="8B9447683D73FFA0E513D735781564DB" blockId="7.[832,1475,1429,1968]" lastBlockId="8.[113,756,1280,1458]" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
<emphasis id="B95F9B7A3D73FFAFE513D7357AC267EA" bold="true" box="[869,985,1641,1667]" pageId="7" pageNumber="46">Remarks.</emphasis>
<taxonomicName id="4C2B3CEB3D73FFAFE59ED7357DAB67EA" authorityName="Linsenmaier" authorityYear="1959" box="[1000,1200,1641,1667]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="enslini">
<emphasis id="B95F9B7A3D73FFAFE59ED7357DAB67EA" box="[1000,1200,1641,1667]" italics="true" pageId="7" pageNumber="46">Holopyga enslini</emphasis>
</taxonomicName>
differs from the West Mediterranean species of the
<taxonomicName id="4C2B3CEB3D73FFAFE2EDD7D47C3B67C8" authorityName="Abeille de Perrin. We" authorityYear="1878" box="[1179,1312,1671,1697]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="miranda">
<emphasis id="B95F9B7A3D73FFAFE2EDD7D47C3B67C8" box="[1179,1312,1671,1697]" italics="true" pageId="7" pageNumber="46">H. miranda</emphasis>
</taxonomicName>
group by the ventrally metallic red, instead of black or blackish, mesosoma. It is related to
<taxonomicName id="4C2B3CEB3D73FFAFE255D7987C0A67B4" authority="Rosa, 2018" authorityName="Rosa" authorityYear="2018" box="[1059,1297,1731,1758]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B95F9B7A3D73FFAFE255D7987D9767B4" box="[1059,1164,1731,1757]" italics="true" pageId="7" pageNumber="46">H. lucens</emphasis>
<bibRefCitation id="EFBA3A993D73FFAFE2E7D7987C0A67B4" author="Rosa P." box="[1169,1297,1731,1758]" pageId="7" pageNumber="46" pagination="1 - 27" refId="ref12519" refString="Rosa P., 2018 - New species of Chrysididae (Hymenop- tera) from Central Asia, Russia and Iran. Part 1. Far Eastern Entomologist, 371: 1 - 27." type="journal article" year="2018">Rosa, 2018</bibRefCitation>
</taxonomicName>
, described from Central Asia, yet with blackish, almost without reflections, coxae, trochanters and femora (vs. shining metallic green in
<taxonomicName id="4C2B3CEB3D73FFAFE5D1D6427D08665E" authorityName="Rosa" authorityYear="2018" box="[935,1043,1821,1847]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B95F9B7A3D73FFAFE5D1D6427D08665E" box="[935,1043,1821,1847]" italics="true" pageId="7" pageNumber="46">H. lucens</emphasis>
</taxonomicName>
). Moreover, it differs from
<taxonomicName id="4C2B3CEB3D73FFAFE320D6427CD8665E" authorityName="Rosa" authorityYear="2018" box="[1366,1475,1821,1847]" class="Insecta" family="Chrysididae" genus="Holopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="46" phylum="Arthropoda" rank="species" species="lucens">
<emphasis id="B95F9B7A3D73FFAFE320D6427CD8665E" box="[1366,1475,1821,1847]" italics="true" pageId="7" pageNumber="46">H. lucens</emphasis>
</taxonomicName>
by the shape of the head in dorsal view, larger than pronotum, with posterior edge distinctly concave, and posterolateral angles sharply projecting rearwards (vs. as large as pronotum, with posterior edge almost straight, and posterolateral angles not projecting); head and mesosoma punctation denser to confluent, without shining intervals (vs. scattered, with polished, shining intervals); and unmodified, fully micropunctate and ventrally angulate lower mesopleuron (vs. enlarged, carinate, ventrally polished and rounded).
</paragraph>
</subSubSection>
</treatment>
</document>