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<document id="53846710E071ED0934F2A26381360741" ID-CLB-Dataset="20208" ID-DOI="10.11646/zootaxa.5115.4.1" ID-GBIF-Dataset="b99d2dc5-e0a6-4f73-b2c6-b21b6350c69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6361552" ID-ZooBank="320C2E00-8035-431A-8AD7-D483D81923F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647419732391" checkinUser="plazi" docAuthor="Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid &amp; Garner, Beulah" docDate="2022" docId="291CD3215318DC6CFF6DF9CB87AEFD3B" docLanguage="en" docName="zootaxa.5115.4.1.pdf" docOrigin="Zootaxa 5115 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Epomis) duvaucelii Dej. By H. E. Andrewes" docType="treatment" docVersion="6" lastPageNumber="468" masterDocId="D525AB595317DC7DFFFAFFFA836DFF91" masterDocTitle="Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan" masterLastPageNumber="486" masterPageNumber="451" pageNumber="466" updateTime="1699318106451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B1518B3C30FAA32B38F1FAE2219FB5BC">Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan</mods:title>
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<mods:namePart id="CCB523F1EE584D0ADC7E6C2B2C95EB89">Ullah, Mishkat</mods:namePart>
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<mods:namePart id="96DEB30D8033EA96CEEA9B63400DA6A2">Naeem, Muhammad</mods:namePart>
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<mods:affiliation id="A03F912F2FD53697F3DE5944C359EB3A">muhammad. naeem @ uaar. edu. pk; https: // orcid. org / 0000 - 0002 - 2025 - 2024</mods:affiliation>
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<mods:namePart id="159BE836D1FD18C677747C18CED9CA67">Garner, Beulah</mods:namePart>
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<mods:affiliation id="28ECA14C5066F3D2C184BDF86B4BDB06">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. &amp; b. garner @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 5229 - 2450</mods:affiliation>
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<taxonomicName id="66B519B45318DC72FF6DF9CB81A7F9DA" ID-CoL="5Y7JM" authority="(Dejean, 1831)" authorityName="Dej. By H. E. Andrewes" baseAuthorityName="Dejean" baseAuthorityYear="1831" box="[151,714,1585,1612]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="duvaucelii" subGenus="Epomis">
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<emphasis id="93C1BE255318DC72FF6DF9CB827EF9DD" bold="true" box="[151,275,1585,1612]" italics="true" pageId="15" pageNumber="466">Chlaenius</emphasis>
(
<emphasis id="93C1BE255318DC72FEDEF9CB8212F9DA" bold="true" box="[292,383,1585,1611]" italics="true" pageId="15" pageNumber="466">Epomis</emphasis>
)
<emphasis id="93C1BE255318DC72FE6BF9CB8163F9DD" bold="true" box="[401,526,1585,1612]" italics="true" pageId="15" pageNumber="466">duvaucelii</emphasis>
(
<bibRefCitation id="C5241FC65318DC72FDE7F9CB81AFF9DA" author="Dejean, P. F. M. A." box="[541,706,1585,1611]" pageId="15" pageNumber="466" refId="ref26205" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon- Marvis, Paris, 883 pp." type="book" year="1831">Dejean, 1831</bibRefCitation>
)
</emphasis>
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(
<figureCitation id="398E7EB25318DC72FF5AF9AF839CF9E1" box="[160,241,1621,1648]" captionStart="FIGURE 4" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[193,1395,181,1912]" captionTargetId="figure-18@13.[193,1395,181,1920]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal habitus (a) Chlaenius (Chlaenius) scapularis Chaudoir, 1876 imaged from NHMUK collection; (b) Chlaenius (Chlaeniellus) laeviplaga frater Chaudoir, 1876; (c) Chlaenius (Chlaeniellus) tenuilimbatus Ballion, 1871 imaged from NHM collection; (d) Chlaenius (Chlaeniellus) velocipes Chaudoir, 1876 imaged from NHMUK collection." figureDoi="http://doi.org/10.5281/zenodo.6361567" httpUri="https://zenodo.org/record/6361567/files/figure.png" pageId="15" pageNumber="466">Fig. 4d</figureCitation>
)
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<treatmentCitation id="201444265318DC72FF6DF960817AF923" author="Dejean, P. F. M. A." box="[151,535,1690,1714]" page="668" pageId="15" pageNumber="466" year="1831">
<taxonomicName id="66B519B45318DC72FF6DF960817AF923" ID-CoL="6FX96" authority="Dejean, 1831: 668" authorityName="Dejean" authorityPageNumber="668" authorityYear="1831" box="[151,535,1690,1714]" class="Insecta" family="Carabidae" genus="Epomis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="duvaucelii">
<emphasis id="93C1BE255318DC72FF6DF9608239F923" box="[151,340,1690,1714]" italics="true" pageId="15" pageNumber="466">Epomis duvaucelii</emphasis>
<bibRefCitation id="C5241FC65318DC72FEA3F960817AF923" author="Dejean, P. F. M. A." box="[345,535,1690,1714]" pageId="15" pageNumber="466" refId="ref26205" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon- Marvis, Paris, 883 pp." type="book" year="1831">Dejean, 1831: 668</bibRefCitation>
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.
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<emphasis id="93C1BE255318DC72FF6DF91A82EDF96B" bold="true" box="[151,384,1760,1786]" pageId="15" pageNumber="466">Material examined.</emphasis>
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<collectingCountry id="D9A222A75318DC72FE7DF91A8160F96B" box="[391,525,1760,1786]" name="Pakistan" pageId="15" pageNumber="466">PAKISTAN</collectingCountry>
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;
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,
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D-16;
<date id="D50B44F75318DC72FC32F91A8731F96A" box="[968,1116,1760,1787]" pageId="15" pageNumber="466" value="2015-04-19">
<collectingDate id="C54FBD1F5318DC72FC32F91A8731F96A" box="[968,1116,1760,1787]" pageId="15" pageNumber="466" value="2015-04-19">19 Apr. 2015</collectingDate>
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;
<collectorName id="0C4007E15318DC72FB92F91A87A9F96B" box="[1128,1220,1760,1786]" pageId="15" pageNumber="466">Mishkat</collectorName>
leg.;
<collectionCode id="C7A4FAF25318DC72FAF8F91B863EF96A" box="[1282,1363,1761,1787]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="15" pageNumber="466" type="Museum">PMNH</collectionCode>
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;
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;
<date id="D50B44F75318DC72FE0AF8FE81EFF88E" box="[496,642,1796,1823]" pageId="15" pageNumber="466" value="2015-04-19">
<collectingDate id="C54FBD1F5318DC72FE0AF8FE81EFF88E" box="[496,642,1796,1823]" pageId="15" pageNumber="466" value="2015-04-19">19 Apr. 2015</collectingDate>
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;
<collectorName id="0C4007E15318DC72FD77F8FE804CF88F" box="[653,801,1796,1822]" pageId="15" pageNumber="466">Mishkatullah</collectorName>
leg.;
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;
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,
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;
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<collectingDate id="C54FBD1F5318DC72FA7AF8FE8266F8D3" pageId="15" pageNumber="466" value="2007-09-06">06 Sep. 2007</collectingDate>
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;
<collectorName id="0C4007E15318DC72FEEFF8D2820CF8D3" box="[277,353,1832,1858]" pageId="15" pageNumber="466">Amjad</collectorName>
leg.; NIM
</materialsCitation>
<materialsCitation id="11DD686A5318DC72FE0BF8D287CFF8F7" ID-GBIF-Occurrence="3709719469" collectionCode="NHMUK" collectorName="N. W. India" country="Pakistan" location="Khyber Pakhtunkhwa" pageId="15" pageNumber="466" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Khyber Pakhtunkhwa">
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,
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; “
<collectorName id="0C4007E15318DC72FD94F8D28180F8D3" box="[622,749,1832,1858]" pageId="15" pageNumber="466">N.W. India</collectorName>
” [
<collectingRegion id="6371ACD55318DC72FCFFF8D2876EF8D3" box="[773,1027,1831,1858]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="15" pageNumber="466">Khyber Pakhtunkhwa</collectingRegion>
];
<taxonomicName id="66B519B45318DC72FBEFF8D28264F8F7" authority="Dej. By H. E. Andrewes" authorityName="Dej. By H. E. Andrewes" baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="duvaucelii">
<emphasis id="93C1BE255318DC72FBEFF8D2866EF8D3" box="[1045,1283,1832,1858]" italics="true" pageId="15" pageNumber="466">Chlaenius duvaucelii</emphasis>
Dej. By H.E. Andrewes
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det. [Male specimen with yellow determination label];
<collectionCode id="C7A4FAF25318DC72FC86F8B7808BF8F6" box="[892,998,1869,1895]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="466" type="Museum">NHMUK</collectionCode>
(B.M. 1984-19)
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;
<taxonomicName id="66B519B45318DC72FB54F8B682C5F81B" authority="Dej. By H. E. Andrewes" authorityName="Dej. By H. E. Andrewes" baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="466" phylum="Arthropoda" rank="species" species="duvaucelii">
<emphasis id="93C1BE255318DC72FB54F8B686F0F8F7" box="[1198,1437,1868,1894]" italics="true" pageId="15" pageNumber="466">Chlaenius duvaucelii</emphasis>
Dej. By H.E. Andrewes
</taxonomicName>
(male specimen with yellow determination label);
<collectionCode id="C7A4FAF25318DC72FC0FF88B8732F81A" box="[1013,1119,1905,1931]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="15" pageNumber="466" type="Museum">NHMUK</collectionCode>
(H.E. Andrewes Collection 1945-97).
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5318DC6CFF3DF84281A4FF20" lastPageId="17" lastPageNumber="468" pageId="15" pageNumber="466" type="description">
<paragraph id="A10A62375318DC72FF3DF84280A8F867" blockId="15.[151,1437,1760,2039]" pageId="15" pageNumber="466">
<emphasis id="93C1BE255318DC72FF3DF8428213F843" bold="true" box="[199,382,1976,2002]" pageId="15" pageNumber="466">Measurements.</emphasis>
Male (n=5): TL=
<quantity id="664DCFD25318DC72FDB4F84281AEF842" box="[590,707,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="2.134" pageId="15" pageNumber="466" unit="mm" value="21.34">21.34 mm</quantity>
, HW=
<quantity id="664DCFD25318DC72FCEFF8428011F842" box="[789,892,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="3.73" pageId="15" pageNumber="466" unit="mm" value="3.73">3.73 mm</quantity>
, PL=
<quantity id="664DCFD25318DC72FC3BF8428745F842" box="[961,1064,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="4.27" pageId="15" pageNumber="466" unit="mm" value="4.27">4.27 mm</quantity>
, PW=
<quantity id="664DCFD25318DC72FB8CF84287B0F842" box="[1142,1245,1976,2003]" metricMagnitude="-3" metricUnit="m" metricValue="5.34" pageId="15" pageNumber="466" unit="mm" value="5.34">5.34 mm</quantity>
, EL=
<quantity id="664DCFD25318DC72FAD9F84286F5F842" box="[1315,1432,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="1.335" pageId="15" pageNumber="466" unit="mm" value="13.35">13.35 mm</quantity>
, EW=
<quantity id="664DCFD25318DC72FF20F826822CF866" box="[218,321,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="15" pageNumber="466" unit="mm" value="4.3">4.30 mm</quantity>
. Ratios PL/PW= 0.79, HW/PW= 0.69 and EL/EW= 3.10.
</paragraph>
<caption id="F5CA32BF5307DC6DFF6DF86D827DF869" ID-DOI="http://doi.org/10.5281/zenodo.6361569" ID-Zenodo-Dep="6361569" httpUri="https://zenodo.org/record/6361569/files/figure.png" pageId="16" pageNumber="467" startId="16.[151,250,1943,1967]" targetBox="[216,1371,181,1919]" targetPageId="16">
<paragraph id="A10A62375307DC6DFF6DF86D827DF869" blockId="16.[151,1436,1943,2040]" pageId="16" pageNumber="467">
<emphasis id="93C1BE255307DC6DFF6DF86D827FF821" bold="true" box="[151,274,1943,1968]" pageId="16" pageNumber="467">FIGURE 5.</emphasis>
Dorsal habitus (a)
<taxonomicName id="66B519B45307DC6DFE29F8628714F83E" authority="circumdatus Brulle, 1835" authorityName="circumdatus Brulle" authorityYear="1835" box="[467,1145,1943,1968]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="subSpecies" species="circumdatus" subGenus="Chlaeniostenus" subSpecies="circumdatus">
<emphasis id="93C1BE255307DC6DFE29F8628157F821" box="[467,570,1944,1968]" italics="true" pageId="16" pageNumber="467">Chlaenius</emphasis>
(
<emphasis id="93C1BE255307DC6DFDB2F8628189F821" box="[584,740,1944,1968]" italics="true" pageId="16" pageNumber="467">Chlaeniostenus</emphasis>
)
<emphasis id="93C1BE255307DC6DFD09F862809CF821" box="[755,1009,1944,1968]" italics="true" pageId="16" pageNumber="467">circumdatus circumdatus</emphasis>
<bibRefCitation id="C5241FC65307DC6DFC0DF8628714F83E" author="Brulle, A." box="[1015,1145,1943,1968]" pageId="16" pageNumber="467" pagination="271 - 303" refId="ref25949" refString="Brulle, A. (1835) Observations critiques sur la synonymie des carabiques (suite). Revue Entomologique, 3, 271 - 303." type="journal article" year="1835">Brullé, 1835</bibRefCitation>
</taxonomicName>
; (b)
<taxonomicName id="66B519B45307DC6DFB5CF86282A4F845" authority="Dejean, 1826" authorityName="Dejean" authorityYear="1826" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="nitidicollis" subGenus="Chlaeniostenus">
<emphasis id="93C1BE255307DC6DFB5CF8628660F821" box="[1190,1293,1944,1968]" italics="true" pageId="16" pageNumber="467">Chlaenius</emphasis>
(
<emphasis id="93C1BE255307DC6DFAE1F86283D7F842" italics="true" pageId="16" pageNumber="467">Chlaeniostenus</emphasis>
)
<emphasis id="93C1BE255307DC6DFF30F846825BF845" box="[202,310,1980,2004]" italics="true" pageId="16" pageNumber="467">nitidicollis</emphasis>
<bibRefCitation id="C5241FC65307DC6DFEC7F84682A4F845" author="Dejean, P. F. M. A." box="[317,457,1980,2004]" pageId="16" pageNumber="467" refId="ref26152" refString="Dejean, P. F. M. A. (1826) Species general des coleopteres, de la collection de M. le Comte Dejean. Vol. 2. Crevot, Paris, 501 pp. https: // doi. org / 10.5962 / bhl. title. 8863" type="book" year="1826">Dejean, 1826</bibRefCitation>
</taxonomicName>
; (c)
<taxonomicName id="66B519B45307DC6DFE0DF8468091F842" authority="Nietner, 1857" authorityName="Nietner" authorityYear="1857" box="[503,1020,1979,2004]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="pulcher" subGenus="Chlaeniostenus">
<emphasis id="93C1BE255307DC6DFE0DF8468133F845" box="[503,606,1980,2004]" italics="true" pageId="16" pageNumber="467">Chlaenius</emphasis>
(
<emphasis id="93C1BE255307DC6DFD97F8468064F845" box="[621,777,1980,2004]" italics="true" pageId="16" pageNumber="467">Chlaeniostenus</emphasis>
)
<emphasis id="93C1BE255307DC6DFCE3F846800BF845" box="[793,870,1980,2004]" italics="true" pageId="16" pageNumber="467">pulcher</emphasis>
<bibRefCitation id="C5241FC65307DC6DFC97F8468091F842" author="Nietner, J." box="[877,1020,1979,2004]" pageId="16" pageNumber="467" pagination="241 - 249" refId="ref27780" refString="Nietner, J. (1857) XX. - Descriptions of new Ceylon Coleoptera. Annals and Magazine of Natural History, 19 (111), 241 - 249. https: // doi. org / 10.1080 / 00222935708697726" type="journal article" year="1857">Nietner, 1857</bibRefCitation>
</taxonomicName>
; (d)
<taxonomicName id="66B519B45307DC6DFBD1F8468261F869" authority="(Dejean, 1831)" authorityName="Dej. By H. E. Andrewes" baseAuthorityName="Dejean" baseAuthorityYear="1831" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="467" phylum="Arthropoda" rank="species" species="duvaucelii" subGenus="Epomis">
<emphasis id="93C1BE255307DC6DFBD1F84687FFF845" box="[1067,1170,1980,2004]" italics="true" pageId="16" pageNumber="467">Chlaenius</emphasis>
(
<emphasis id="93C1BE255307DC6DFB58F8468783F842" box="[1186,1262,1980,2003]" italics="true" pageId="16" pageNumber="467">Epomis</emphasis>
)
<emphasis id="93C1BE255307DC6DFB07F846860AF845" box="[1277,1383,1980,2004]" italics="true" pageId="16" pageNumber="467">duvaucelii</emphasis>
(
<bibRefCitation id="C5241FC65307DC6DFA8FF8468269F869" author="Dejean, P. F. M. A." pageId="16" pageNumber="467" refId="ref26205" refString="Dejean, P. F. M. A. (1831) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome cinquieme. Mequignon- Marvis, Paris, 883 pp." type="book" year="1831">Dejean, 1831</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="A10A62375306DC6CFF3DFF6D81A4FF20" blockId="17.[151,1436,151,682]" box="[199,713,151,177]" pageId="17" pageNumber="468">No females were studied / available for study.</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5306DC6CFF3DFF4081B3FE3C" pageId="17" pageNumber="468" type="diagnosis">
<paragraph id="A10A62375306DC6CFF3DFF4081B3FE3C" blockId="17.[151,1436,151,682]" pageId="17" pageNumber="468">
<emphasis id="93C1BE255306DC6CFF3DFF40822FFF45" bold="true" box="[199,322,186,212]" pageId="17" pageNumber="468">Diagnosis.</emphasis>
Head, pronotum and elytra metallic dark cyan green. Antennal articles IIII, legs except tarsus and elytral border interval IX yellowish brown. Palpomeres, antennal articles IVIX, all tarsus reddish brown. Maxillary palpi terminal article truncate. Clypeus smooth, frons sides sparsely punctate towards eyes and smooth at midpoint; vertex and neck area densely covered with deep setiferous punctures. Pronotum, wider than long, apical margin moderately covered with deep setiferous punctures becoming denser towards basal margin, posterior angles obtuse, base slightly concave at midpoint slightly oblique to sides. Elytral striae fine, shallow and distinctly punctate; intervals, slightly raised and densely setiferous punctate.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5306DC6CFF3DFE4C827BFD14" pageId="17" pageNumber="468" type="discussion">
<paragraph id="A10A62375306DC6CFF3DFE4C827BFD14" blockId="17.[151,1436,151,682]" pageId="17" pageNumber="468">
<emphasis id="93C1BE255306DC6CFF3DFE4C8222FE41" bold="true" box="[199,335,438,464]" pageId="17" pageNumber="468">Comments.</emphasis>
This species differs from
<taxonomicName id="66B519B45306DC6CFD89FE4D87FBFE40" authority="Wiedemann, 1821" authorityName="Wiedemann" authorityYear="1821" box="[627,1174,439,465]" class="Insecta" family="Carabidae" genus="Chleanius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="nigricans" subGenus="Epomis">
<emphasis id="93C1BE255306DC6CFD89FE4D8188FE40" box="[627,741,439,465]" italics="true" pageId="17" pageNumber="468">Chleanius</emphasis>
(
<emphasis id="93C1BE255306DC6CFD0FFE4D8024FE41" box="[757,841,439,464]" italics="true" pageId="17" pageNumber="468">Epomis</emphasis>
)
<emphasis id="93C1BE255306DC6CFCA0FE4D80A9FE41" box="[858,964,439,464]" italics="true" pageId="17" pageNumber="468">nigricans</emphasis>
<bibRefCitation id="C5241FC65306DC6CFC30FE4D87FBFE40" author="Wiedemann, C. R. W." box="[970,1174,439,465]" pageId="17" pageNumber="468" pagination="107 - 183" refId="ref28024" refString="Wiedemann, C. R. W. (1821) [New species]. In: Wiedemann, C. R. W. &amp; Germar, E. F., Neue exotische Kafer. Magazin der Entomologie, 4, pp. 107 - 183." type="journal article" year="1821">Wiedemann, 1821</bibRefCitation>
</taxonomicName>
on the following basis, head and pronotum with metallic coppery to green tinge; elytra black with metallic green tinge at base and lateral margin, intervals convex laterally with wide deep setiferous punctures/fovae and middle mostly smooth, interval VIII &amp; IX densely setiferous punctate. In this current study, this species was recorded from Islamabad and Abootabad. This is the first time this species is reported from
<collectingCountry id="D9A222A75306DC6CFC8BFDBD80B9FDF0" box="[881,980,583,609]" name="Pakistan" pageId="17" pageNumber="468">Pakistan</collectingCountry>
. Earlier,
<taxonomicName id="66B519B45306DC6CFBCFFDBD87A7FDF0" box="[1077,1226,583,609]" class="Insecta" family="Carabidae" genus="Chleanius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="468" phylum="Arthropoda" rank="species" species="duvacelii">
<emphasis id="93C1BE255306DC6CFBCFFDBD87A7FDF0" box="[1077,1226,583,609]" italics="true" pageId="17" pageNumber="468">Ch. duvacelii</emphasis>
</taxonomicName>
was recorded only from
<collectingCountry id="D9A222A75306DC6CFF2FFD91827FFD14" box="[213,274,619,645]" name="India" pageId="17" pageNumber="468">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5306DC6CFF3DFD7487AEFD3B" box="[199,1219,654,682]" pageId="17" pageNumber="468" type="distribution">
<paragraph id="A10A62375306DC6CFF3DFD7487AEFD3B" blockId="17.[151,1436,151,682]" box="[199,1219,654,682]" pageId="17" pageNumber="468">
<emphasis id="93C1BE255306DC6CFF3DFD74820CFD39" bold="true" box="[199,353,654,680]" pageId="17" pageNumber="468">Distribution.</emphasis>
<collectingCountry id="D9A222A75306DC6CFE9DFD7582CCFD38" box="[359,417,655,681]" name="India" pageId="17" pageNumber="468">India</collectingCountry>
(Anichtchenko 2017;
<bibRefCitation id="C5241FC65306DC6CFD66FD758041FD38" author="Lorenz, W." box="[668,812,655,681]" pageId="17" pageNumber="468" pagination="2000" refId="ref27320" refString="Lorenz, W. (2018) CarabCat: Global database of ground beetles. Version October 2017. In: Roskov, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P. M., Bourgoin, T., DeWalt, R. E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. &amp; Penev, L. (Eds.), Species 2000 &amp; ITIS Catalogue of Life, 30 th June 2018. Species 2000: Naturalis, Leiden, Available from: https: // www. catalogueoflife. org / col (accessed 20 July 2020)" type="book chapter" year="2018">Lorenz 2018</bibRefCitation>
) and
<collectingCountry id="D9A222A75306DC6CFC90FD7580A4FD38" box="[874,969,655,681]" name="Pakistan" pageId="17" pageNumber="468">Pakistan</collectingCountry>
(first country record).
</paragraph>
</subSubSection>
</treatment>
</document>