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<document id="9A9D531B50A2BE6C74FDF936B22B554D" ID-CLB-Dataset="300033" ID-DOI="10.1590/1678-4766e2020021" ID-GBIF-Dataset="e88f40f8-79ce-4392-ae5b-4c294c8da995" ID-ISSN="1678-4766" ID-Zenodo-Dep="12784936" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1712600270875" checkinUser="luisschmitz" docAuthor="Schmitz, Luís Ricardo, Barcellos, Aline &amp; Eger, Joseph" docDate="2020" docId="C82D477D2A6E7123FCF2454E7A3CC9A3" docLanguage="en" docName="IheringiaSerZool.110.2020021.1-16.pdf" docOrigin="Iheringia, Série Zoologia (e 2020021) 110" docSource="http://dx.doi.org/10.1590/1678-4766e2020021" docStyle="DocumentStyle:CAF80231855201F2281AC57FC1B00651.2:IheringiaSerZool.2016-.journal_article" docStyleId="CAF80231855201F2281AC57FC1B00651" docStyleName="IheringiaSerZool.2016-.journal_article" docStyleVersion="2" docTitle="Coptochilus morrisi Eger &amp; Schmitz 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="15" masterDocId="34143F052A63712DFF98475F7936CE20" masterDocTitle="Revision of Coptochilus (Heteroptera, Scutelleridae, Pachycorinae)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="14" updateTime="1721405175461" updateUser="ExternalLinkService">
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<mods:title id="EAF564669A0BCA2057B03AFEA4C1C55D">Revision of Coptochilus (Heteroptera, Scutelleridae, Pachycorinae)</mods:title>
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<mods:namePart id="59217ABBF42E5ACF13D514FFEA6B9F08">Schmitz, Luís Ricardo</mods:namePart>
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<mods:namePart id="C45565CEE0880AC058846739C761E3BD">Barcellos, Aline</mods:namePart>
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<mods:title id="33137A1E36B7D1AD5DB25757A834BC82">Iheringia, Série Zoologia</mods:title>
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<mods:date id="618D951D50C86066870A91DCACC712D4">2020</mods:date>
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<mods:number id="69EFDDE3EDEADDFBBB00D8AC3C2F54A3">2020-09-18</mods:number>
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<treatment id="C82D477D2A6E7123FCF2454E7A3CC9A3" ID-DOI="http://doi.org/10.5281/zenodo.12784972" ID-Zenodo-Dep="12784972" LSID="urn:lsid:plazi:treatment:C82D477D2A6E7123FCF2454E7A3CC9A3" httpUri="http://treatment.plazi.org/id/C82D477D2A6E7123FCF2454E7A3CC9A3" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="089EA5E02A6E7120FCF2454E7C69CC09" box="[874,1375,529,554]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="403BF66B2A6E7120FCF2454E7C69CC09" blockId="13.[818,1430,529,650]" box="[874,1375,529,554]" pageId="13" pageNumber="14">
<heading id="1B7341072A6E7120FCF2454E7C69CC09" box="[874,1375,529,554]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="14" reason="2">
<emphasis id="72F02A792A6E7120FCF2454E7C69CC09" bold="true" box="[874,1375,529,554]" pageId="13" pageNumber="14">
<taxonomicName id="87848DE82A6E7120FCF2454E7C37CC09" authority="Eger &amp; Schmitz" authorityName="Eger &amp; Schmitz" authorityYear="2020" box="[874,1281,529,554]" class="Insecta" family="Scutelleridae" genus="Coptochilus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="morrisi" status="sp. nov.">
<emphasis id="72F02A792A6E7120FCF2454E7D73CC0A" bold="true" box="[874,1093,529,554]" italics="true" pageId="13" pageNumber="14">Coptochilus morrisi</emphasis>
Eger &amp; Schmitz
</taxonomicName>
<taxonomicNameLabel id="69C397022A6E7120FA90454D7C69CC09" box="[1288,1375,530,554]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="089EA5E02A6E7120FCF4456D7DF4CCAA" pageId="13" pageNumber="14" type="description">
<paragraph id="403BF66B2A6E7120FCF4456D7C6BCC6A" blockId="13.[818,1430,529,650]" box="[876,1373,562,586]" pageId="13" pageNumber="14">
(
<figureCitation id="D8BFEAEE2A6E7120FCEB456D7AD9CC6A" box="[883,1007,562,586]" captionStart="Figs 10–15" captionStartId="4.[151,188,1467,1486]" captionTargetBox="[151,1436,148,1443]" captionTargetId="figure-28@4.[151,1436,148,1443]" captionTargetPageId="4" captionText="Figs 10–15. Coptochilus, habitus, respectively in dorsal, ventral, and lateral view: 10–12, C. morrisi sp. nov.; 13–15, C. neotropicalis Distant, 1899. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12784940" httpUri="https://zenodo.org/record/12784940/files/figure.png" pageId="13" pageNumber="14">Figs 10–12</figureCitation>
,
<figureCitation id="D8BFEAEE2A6E7120FC61456D7D08CC6A" box="[1017,1086,562,586]" captionStart="Figs 16–30" captionStartId="5.[151,188,1924,1943]" captionTargetBox="[204,1383,162,1885]" captionTargetId="figure-28@5.[204,1383,151,1886]" captionTargetPageId="5" captionText="Figs 16–30. Coptochilus, head, respectively in dorsal, lateral, and dorsolateral view: 16–18, C. castaneus (Dallas, 1851), reval.; 19–21, C. ferrugineus Amyot &amp; Serville, 1843; 22–24, C. lentiginosus Berg, 1879; 25–27, C. morrisi sp. nov.; 28–30, C. neotropicalis Distant, 1899. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12784944" httpUri="https://zenodo.org/record/12784944/files/figure.png" pageId="13" pageNumber="14">25–27</figureCitation>
,
<figureCitation id="D8BFEAEE2A6E7120FBD1456D7DB9CC6A" box="[1097,1167,562,586]" captionStart="Figs 31–45" captionStartId="6.[151,188,1899,1918]" captionTargetBox="[150,1436,151,1871]" captionTargetId="figure-28@6.[149,1436,151,1871]" captionTargetPageId="6" captionText="Figs 31–45. Coptochilus, pygophore, respectively in ventral, dorsal, and posterior view: 31–33, C. castaneus (Dallas, 1851), reval.; 34–36, C. ferrugineus Amyot &amp; Serville, 1843; 37–39, C. lentiginosus Berg, 1879; 40–42, C. morrisi sp. nov.; 43–45, C. neotropicalis Distant, 1899 (pr, parameres). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784946" httpUri="https://zenodo.org/record/12784946/files/figure.png" pageId="13" pageNumber="14">40–42</figureCitation>
,
<figureCitation id="D8BFEAEE2A6E7120FB01456D7D80CC6A" box="[1177,1206,562,586]" captionStart="Figs 46–50" captionStartId="7.[151,188,552,571]" captionTargetBox="[151,1436,149,530]" captionTargetId="figure-28@7.[151,1436,149,530]" captionTargetPageId="7" captionText="Figs 46–50. Coptochilus, right parameres, dorsal and ventral view: 46, C. castaneus (Dallas, 1851), reval.; 47, C. ferrugineus Amyot &amp; Serville, 1843; 48, C. lentiginosus Berg, 1879; 49, C. morrisi sp. nov.; 50, C. neotropicalis Distant, 1899. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12784950" httpUri="https://zenodo.org/record/12784950/files/figure.png" pageId="13" pageNumber="14">49</figureCitation>
,
<figureCitation id="D8BFEAEE2A6E7120FB59456D7C30CC6A" box="[1217,1286,562,586]" captionStart="Figs 63–68" captionStartId="9.[151,188,1591,1610]" captionTargetBox="[151,1436,151,1563]" captionTargetId="figure-27@9.[151,1436,151,1563]" captionTargetPageId="9" captionText="Figs 63–68. Coptochilus, phallus, respectively in ventral, dorsal and lateral view: 63–65, C. morrisi sp. nov.; 66–68, C. neotropicalis Distant, 1899. (aed, aedeagus; bp, basal plate; cpI, conjunctival process I; cpII, conjunctival process II; cpIII, conjunctival process III). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784958" httpUri="https://zenodo.org/record/12784958/files/figure.png" pageId="13" pageNumber="14">63–65</figureCitation>
,
<figureCitation id="D8BFEAEE2A6E7120FA89456D7C18CC6A" box="[1297,1326,562,586]" captionStart="Figs 69–73" captionStartId="10.[151,188,1427,1446]" captionTargetBox="[151,1436,151,1365]" captionTargetId="figure-28@10.[151,1436,151,1386]" captionTargetPageId="10" captionText="Figs 69–73. Coptochilus, genital plates: 69, C. castaneus (Dallas, 1851), reval.; 70, C. ferrugineus Amyot &amp; Serville, 1843; 71, C. lentiginosus Berg, 1879; 72, C. morrisi sp. nov.; 73, C. neotropicalis Distant, 1899 (lt8, laterotergites VIII; lt9, laterotergites IX; vf8, valvifers VIII; vf9, valvifer IX). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784962" httpUri="https://zenodo.org/record/12784962/files/figure.png" pageId="13" pageNumber="14">72</figureCitation>
,
<figureCitation id="D8BFEAEE2A6E7120FAA1456D7C63CC6A" box="[1337,1365,562,586]" captionStart="Fig" captionStartId="12.[150,181,956,975]" captionTargetBox="[238,1352,151,932]" captionTargetId="figure-488@12.[238,1352,151,932]" captionTargetPageId="12" captionText="Fig. 80. Distribution of the species of Coptochilus, colored areas based on biogeographical dominions according to MORRONE (2013)." figureDoi="http://doi.org/10.5281/zenodo.12784970" httpUri="https://zenodo.org/record/12784970/files/figure.png" pageId="13" pageNumber="14">80</figureCitation>
)
</paragraph>
<paragraph id="403BF66B2A6E7120FCAA450D7DF4CCAA" blockId="13.[818,1430,529,650]" pageId="13" pageNumber="14">urn:lsid:zoobank.org:act:C78D36CD-3EA5-4092-AFD9- 7D12EA5FD91A</paragraph>
</subSubSection>
<subSubSection id="089EA5E02A6E7120FCB545F67D13CCF5" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="403BF66B2A6E7120FCB545F67D13CCF5" blockId="13.[813,1436,681,726]" pageId="13" pageNumber="14">
<taxonomicName id="87848DE82A6E7120FCB545F67DE7CC9C" authority="Neotropicsis EGER et al., 2015: 761" authorityName="Neotropicsis EGER" authorityPageNumber="761" authorityYear="2015" box="[813,1233,681,700]" class="Insecta" family="Scutelleridae" genus="Coptochilus" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="neotropicsis">
<emphasis id="72F02A792A6E7120FCB545F67D3FCC9B" box="[813,1033,681,700]" italics="true" pageId="13" pageNumber="14">Coptochilus Neotropicsis</emphasis>
<bibRefCitation id="24158B9A2A6E7120FB8845F67DE7CC9C" author="EGER JR., J. E. &amp; BARCELLOS, A. &amp; WEILER, L." box="[1040,1233,681,700]" pageId="13" pageNumber="14" refId="ref10484" refString="EGER JR., J. E.; BARCELLOS, A. &amp; WEILER, L. 2015. Shield bugs (Scutelleridae). In: PANIZZI, A. R. &amp; GRAZIA J. eds. True Bugs (Heteroptera) of the Neotropics. New York, Springer, p. 757 - 788." type="book" year="2015">
EGER
<emphasis id="72F02A792A6E7120FBDE45F67D42CC9C" box="[1094,1140,681,700]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
, 2015:761
</bibRefCitation>
</taxonomicName>
(in part, typographical error, misidentification).
</paragraph>
</subSubSection>
<subSubSection id="089EA5E02A6E7123FCE045A67BC2CDC6" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
<paragraph id="403BF66B2A6E7120FCE045A67D83CD30" blockId="13.[811,1438,760,1970]" box="[888,1205,760,785]" pageId="13" pageNumber="14">Type locality. French Guiana.</paragraph>
<paragraph id="403BF66B2A6E7120FCE044477D96CDD1" blockId="13.[811,1438,760,1970]" pageId="13" pageNumber="14">
Relatively small compared with its congeners, yellowish-brown to orange, with six darker longitudinal stripes distributed on pronotum and scutellum; punctation moderately dense and mostly darker than surrounding surfaces. Venter uniformly light yellowish-brown to red, punctation less dense, unicolorous to clearly darker than the surrounding area (
<figureCitation id="D8BFEAEE2A6E7120FB8F44867DA2CDD1" box="[1047,1172,985,1009]" captionStart="Figs 10–15" captionStartId="4.[151,188,1467,1486]" captionTargetBox="[151,1436,148,1443]" captionTargetId="figure-28@4.[151,1436,148,1443]" captionTargetPageId="4" captionText="Figs 10–15. Coptochilus, habitus, respectively in dorsal, ventral, and lateral view: 10–12, C. morrisi sp. nov.; 13–15, C. neotropicalis Distant, 1899. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12784940" httpUri="https://zenodo.org/record/12784940/files/figure.png" pageId="13" pageNumber="14">Figs 10–12</figureCitation>
).
</paragraph>
<paragraph id="403BF66B2A6E7120FCE044A67C3BCB31" blockId="13.[811,1438,760,1970]" pageId="13" pageNumber="14">
Head (
<figureCitation id="D8BFEAEE2A6E7120FC5244A67D6ECA31" box="[970,1112,1017,1041]" captionStart="Figs 16–30" captionStartId="5.[151,188,1924,1943]" captionTargetBox="[204,1383,162,1885]" captionTargetId="figure-28@5.[204,1383,151,1886]" captionTargetPageId="5" captionText="Figs 16–30. Coptochilus, head, respectively in dorsal, lateral, and dorsolateral view: 16–18, C. castaneus (Dallas, 1851), reval.; 19–21, C. ferrugineus Amyot &amp; Serville, 1843; 22–24, C. lentiginosus Berg, 1879; 25–27, C. morrisi sp. nov.; 28–30, C. neotropicalis Distant, 1899. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12784944" httpUri="https://zenodo.org/record/12784944/files/figure.png" pageId="13" pageNumber="14">Figs 25–27</figureCitation>
). Pale yellow to red, with concolorous to dark brown punctation, clypeus and base of head darker than the remainder of head. Clypeus and mandibular plates strongly elevated anteriorly, mandibular plates converging at apex, their apices separated by about the width of second antennal segment or less (
<figureCitation id="D8BFEAEE2A6E7120FAA443C67CA6CA91" box="[1340,1424,1177,1201]" captionStart="Figs 16–30" captionStartId="5.[151,188,1924,1943]" captionTargetBox="[204,1383,162,1885]" captionTargetId="figure-28@5.[204,1383,151,1886]" captionTargetPageId="5" captionText="Figs 16–30. Coptochilus, head, respectively in dorsal, lateral, and dorsolateral view: 16–18, C. castaneus (Dallas, 1851), reval.; 19–21, C. ferrugineus Amyot &amp; Serville, 1843; 22–24, C. lentiginosus Berg, 1879; 25–27, C. morrisi sp. nov.; 28–30, C. neotropicalis Distant, 1899. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.12784944" httpUri="https://zenodo.org/record/12784944/files/figure.png" pageId="13" pageNumber="14">Fig. 25</figureCitation>
). Antennae pale basally, becoming darker apically, particularly segments IV and V. Rostrum light yellow or red, segment IV and sometimes part of segment III darker.
</paragraph>
<paragraph id="403BF66B2A6E7120FCE042457D52C8D2" blockId="13.[811,1438,760,1970]" pageId="13" pageNumber="14">Pronotum yellow to red with dark to concolorous punctation and six longitudinal stripes. Each lateral stripe extending onto cicatrices anteriorly (not reaching anterior margin of pronotum), reaching posterior margin of pronotum and expanding laterally onto humeri; each intermediate stripe reaching from posterior to anterior margin of pronotum; middle pair of stripes narrower than the other two, also not reaching anterior pronotal margin and separated from each other by pale impunctate line. Anterolateral margins of pronotum straight to shallowly convex. Scutellum similar in color to pronotum, six stripes originating at the anterior margin and extending posteriad for about 2/3 length of scutellum, then coalescing into a broad dark area on about posterior 1/3 of scutellum. Legs immaculate, concolorous with the remainder of venter.</paragraph>
<paragraph id="403BF66B2A6E7123FCE041A579C1CC08" blockId="13.[811,1438,760,1970]" lastBlockId="14.[150,778,151,1923]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Connexiva pale and uniformly colored, with shallow moderately dense punctation laterally, this usually darker than the rest of connexiva. Venter light yellow to red with unicolorous to slightly darker punctation. Sternite VII in females twice as long as sternite VI, almost twice as long in males. Male genitalia. Pygophore (
<figureCitation id="D8BFEAEE2A6D7123FD9D47C87BB6CE90" box="[517,640,151,176]" captionStart="Figs 31–45" captionStartId="6.[151,188,1899,1918]" captionTargetBox="[150,1436,151,1871]" captionTargetId="figure-28@6.[149,1436,151,1871]" captionTargetPageId="6" captionText="Figs 31–45. Coptochilus, pygophore, respectively in ventral, dorsal, and posterior view: 31–33, C. castaneus (Dallas, 1851), reval.; 34–36, C. ferrugineus Amyot &amp; Serville, 1843; 37–39, C. lentiginosus Berg, 1879; 40–42, C. morrisi sp. nov.; 43–45, C. neotropicalis Distant, 1899 (pr, parameres). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784946" httpUri="https://zenodo.org/record/12784946/files/figure.png" pageId="14" pageNumber="15">Figs 40–42</figureCitation>
). Ventral rim evenly concave (
<figureCitation id="D8BFEAEE2A6D7123FED247E578F0CEF2" box="[330,454,186,210]" captionStart="Figs 31–45" captionStartId="6.[151,188,1899,1918]" captionTargetBox="[150,1436,151,1871]" captionTargetId="figure-28@6.[149,1436,151,1871]" captionTargetPageId="6" captionText="Figs 31–45. Coptochilus, pygophore, respectively in ventral, dorsal, and posterior view: 31–33, C. castaneus (Dallas, 1851), reval.; 34–36, C. ferrugineus Amyot &amp; Serville, 1843; 37–39, C. lentiginosus Berg, 1879; 40–42, C. morrisi sp. nov.; 43–45, C. neotropicalis Distant, 1899 (pr, parameres). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784946" httpUri="https://zenodo.org/record/12784946/files/figure.png" pageId="14" pageNumber="15">Figs 40, 41</figureCitation>
), slightly sinuous in posterior view (
<figureCitation id="D8BFEAEE2A6D7123FF4547837806CED4" box="[221,304,220,244]" captionStart="Figs 31–45" captionStartId="6.[151,188,1899,1918]" captionTargetBox="[150,1436,151,1871]" captionTargetId="figure-28@6.[149,1436,151,1871]" captionTargetPageId="6" captionText="Figs 31–45. Coptochilus, pygophore, respectively in ventral, dorsal, and posterior view: 31–33, C. castaneus (Dallas, 1851), reval.; 34–36, C. ferrugineus Amyot &amp; Serville, 1843; 37–39, C. lentiginosus Berg, 1879; 40–42, C. morrisi sp. nov.; 43–45, C. neotropicalis Distant, 1899 (pr, parameres). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784946" httpUri="https://zenodo.org/record/12784946/files/figure.png" pageId="14" pageNumber="15">Fig. 42</figureCitation>
). Head of parameres curved, apex rounded (
<figureCitation id="D8BFEAEE2A6D7123FF3947A179D9CF36" box="[161,239,254,278]" captionStart="Figs 46–50" captionStartId="7.[151,188,552,571]" captionTargetBox="[151,1436,149,530]" captionTargetId="figure-28@7.[151,1436,149,530]" captionTargetPageId="7" captionText="Figs 46–50. Coptochilus, right parameres, dorsal and ventral view: 46, C. castaneus (Dallas, 1851), reval.; 47, C. ferrugineus Amyot &amp; Serville, 1843; 48, C. lentiginosus Berg, 1879; 49, C. morrisi sp. nov.; 50, C. neotropicalis Distant, 1899. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12784950" httpUri="https://zenodo.org/record/12784950/files/figure.png" pageId="14" pageNumber="15">Fig. 49</figureCitation>
). Phallus (unexpanded) (
<figureCitation id="D8BFEAEE2A6D7123FE6B47A17B58CF36" box="[499,622,254,278]" captionStart="Figs 63–68" captionStartId="9.[151,188,1591,1610]" captionTargetBox="[151,1436,151,1563]" captionTargetId="figure-27@9.[151,1436,151,1563]" captionTargetPageId="9" captionText="Figs 63–68. Coptochilus, phallus, respectively in ventral, dorsal and lateral view: 63–65, C. morrisi sp. nov.; 66–68, C. neotropicalis Distant, 1899. (aed, aedeagus; bp, basal plate; cpI, conjunctival process I; cpII, conjunctival process II; cpIII, conjunctival process III). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784958" httpUri="https://zenodo.org/record/12784958/files/figure.png" pageId="14" pageNumber="15">Figs 63–65</figureCitation>
). Conjunctival processes I surpassing conjunctival processes III, broad, appearing somewhat sclerotized apically; conjunctival processes II sickle-shaped, thin, mostly sclerotized, extending apically as far as conjunctival processes I, a little broader than conjunctival processes II; conjunctival processes III right-angled, rounded at apex, half as long as conjunctival processes II. Distal part of aedeagus dilated with margins rounded.
</paragraph>
<paragraph id="403BF66B2A6D7123FF7B456D786DCC91" blockId="14.[150,778,151,1923]" pageId="14" pageNumber="15">
Female genitalia (
<figureCitation id="D8BFEAEE2A6D7123FE36456D7B35CC6B" box="[430,515,562,587]" captionStart="Figs 69–73" captionStartId="10.[151,188,1427,1446]" captionTargetBox="[151,1436,151,1365]" captionTargetId="figure-28@10.[151,1436,151,1386]" captionTargetPageId="10" captionText="Figs 69–73. Coptochilus, genital plates: 69, C. castaneus (Dallas, 1851), reval.; 70, C. ferrugineus Amyot &amp; Serville, 1843; 71, C. lentiginosus Berg, 1879; 72, C. morrisi sp. nov.; 73, C. neotropicalis Distant, 1899 (lt8, laterotergites VIII; lt9, laterotergites IX; vf8, valvifers VIII; vf9, valvifer IX). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784962" httpUri="https://zenodo.org/record/12784962/files/figure.png" pageId="14" pageNumber="15">Fig. 72</figureCitation>
). Laterotergites IX sub trapezoidal, narrowed laterally, anterior margin convex. Valvifers 8 relatively broad, posterior margins shallowly concave (
<figureCitation id="D8BFEAEE2A6D7123FF6745C67878CC91" box="[255,334,665,689]" captionStart="Figs 69–73" captionStartId="10.[151,188,1427,1446]" captionTargetBox="[151,1436,151,1365]" captionTargetId="figure-28@10.[151,1436,151,1386]" captionTargetPageId="10" captionText="Figs 69–73. Coptochilus, genital plates: 69, C. castaneus (Dallas, 1851), reval.; 70, C. ferrugineus Amyot &amp; Serville, 1843; 71, C. lentiginosus Berg, 1879; 72, C. morrisi sp. nov.; 73, C. neotropicalis Distant, 1899 (lt8, laterotergites VIII; lt9, laterotergites IX; vf8, valvifers VIII; vf9, valvifer IX). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12784962" httpUri="https://zenodo.org/record/12784962/files/figure.png" pageId="14" pageNumber="15">Fig. 72</figureCitation>
).
</paragraph>
<paragraph id="403BF66B2A6D7123FF7B45E47B94CD7C" blockId="14.[150,778,151,1923]" pageId="14" pageNumber="15">Male measurements (n=5). Total length 6.60–7.05; head: length 1.50–1.75, width 2.20–2.35; length of anteocular region 0.90–1.00; pronotum: length 1.90–2.30, width 3.85– 4.10; scutellum: length 3.80–4.30, width 3.65–4.00; median length of sternites VI–VII: 0.50–0.55, 0.85–1.00.</paragraph>
<paragraph id="403BF66B2A6D7123FF7B44387BC2CDC6" blockId="14.[150,778,151,1923]" pageId="14" pageNumber="15">Female measurements (n=1). total length 7.20; head: length 1.80, width 2.40; length of anteocular region 1.00; pronotum: length 2.25, width 4.20; scutellum: length 4.50, width 4.05, median length of sternites VI-VII:0.45, 0.90;</paragraph>
</subSubSection>
<subSubSection id="089EA5E02A6D7123FF7B44AF7B0DCB7E" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="403BF66B2A6D7123FF7B44AF7B0DCB7E" blockId="14.[150,778,151,1923]" pageId="14" pageNumber="15">
Type material.
<materialsCitation id="F0ECFC362A6D7123FE1A44AF78D8CA4F" collectingDate="2018-09-01" collectingDateMax="2018-09-12" collectingDateMin="2018-09-01" collectingMethod="MV Light" collectionCode="USNM" collectorName="Morris &amp; Wappes" country="French Guiana" elevation="0" latitude="4.55965" location="Kaw Rd. 6" longLatPrecision="1" longitude="-52.20722" municipality="Amazone Nature Lodge" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="9F3F48C92A6D7123FE1A44AF78D3CA28" box="[386,485,1008,1032]" pageId="14" pageNumber="15">Holotype</typeStatus>
<specimenCount id="56823DE22A6D7123FE7344AF7B34CA27" box="[491,514,1008,1031]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
: a)
<collectingCountry id="3893B6FB2A6D7123FDB044AF7A35CA28" box="[552,771,1008,1032]" name="French Guiana" pageId="14" pageNumber="15">FRENCH GUIANA</collectingCountry>
,
<collectingMunicipality id="A05F6C112A6D7123FF0F434D78AFCA0A" box="[151,409,1042,1066]" pageId="14" pageNumber="15">Amazone Nature Lodge</collectingMunicipality>
,
<location id="455BA0B02A6D7123FE3B434D7B20CA0A" LSID="urn:lsid:plazi:treatment:C82D477D2A6E7123FCF2454E7A3CC9A3:455BA0B02A6D7123FE3B434D7B20CA0A" box="[419,534,1042,1066]" country="French Guiana" latitude="4.55965" longLatPrecision="1" longitude="-52.20722" municipality="Amazone Nature Lodge" name="Kaw Rd. 6" pageId="14" pageNumber="15">Kaw Rd. 6</location>
,
<date id="343AD0AB2A6D7123FDB8434D7B83CA0A" box="[544,693,1042,1066]" pageId="14" pageNumber="15" value="2018-09-01" valueMax="2018-09-12" valueMin="2018-09-01">
<collectingDate id="247E29432A6D7123FDB8434D7B83CA0A" box="[544,693,1042,1066]" pageId="14" pageNumber="15" value="2018-09-01" valueMax="2018-09-12" valueMin="2018-09-01">IX/1-12/2018</collectingDate>
</date>
,
<collectorName id="ED7193BD2A6D7123FD26434D7831CA6C" pageId="14" pageNumber="15">Morris &amp; Wappes</collectorName>
; b)
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<geoCoordinate id="25B090AC2A6D7123FE23436B7B6DCA6C" box="[443,603,1076,1100]" degrees="052" direction="west" minutes="12.433" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-52.20722">W052°12.433ʼ</geoCoordinate>
, Elev. 977ʼ,
<collectingMethod id="99C58E7C2A6D7123FD44436A79E6CA4E" pageId="14" pageNumber="15">MV Light</collectingMethod>
(JEE, deposited in
<collectionCode id="26956EAE2A6D7123FE0E430878D1CA4F" box="[406,487,1111,1135]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="14" pageNumber="15">USNM</collectionCode>
)
</materialsCitation>
<materialsCitation id="F0ECFC362A6D7123FE7643087B95CAB1" collectionCode="USNM" pageId="14" pageNumber="15" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
.
<typeStatus id="9F3F48C92A6D7123FE6E43087B56CA4F" box="[502,608,1111,1135]" pageId="14" pageNumber="15">Paratypes</typeStatus>
:
<specimenCount id="56823DE22A6D7123FDF043087B48CA4E" box="[616,638,1111,1134]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
,
<specimenCount id="56823DE22A6D7123FD1043087BAACA4E" box="[648,668,1111,1134]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
same data as
<typeStatus id="9F3F48C92A6D7123FF2C43267827CAB1" box="[180,273,1145,1169]" pageId="14" pageNumber="15">holotype</typeStatus>
(
<specimenCount id="56823DE22A6D7123FEB943267803CAB1" box="[289,309,1145,1169]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
<collectionCode id="26956EAE2A6D7123FEA44326785CCAB0" box="[316,362,1145,1168]" collectionName="JEE" pageId="14" pageNumber="15">JEE</collectionCode>
,
<specimenCount id="56823DE22A6D7123FEED432678BFCAB0" box="[373,393,1145,1168]" pageId="14" pageNumber="15" type="female">♀</specimenCount>
<collectionCode id="26956EAE2A6D7123FE084326788DCAB0" box="[400,443,1145,1168]" collectionName="JEE" pageId="14" pageNumber="15">JEE</collectionCode>
deposited in
<collectionCode id="26956EAE2A6D7123FDD243267BAACAB1" box="[586,668,1145,1169]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="14" pageNumber="15">USNM</collectionCode>
)
</materialsCitation>
<materialsCitation id="F0ECFC362A6D7123FD3B432679D2CAF5" collectingDate="2019-06-19" collectingDateMax="2019-06-31" collectingDateMin="2019-06-19" collectionCode="FSCA" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
;
<specimenCount id="56823DE22A6D7123FD3643267BF2CAB0" box="[686,708,1145,1168]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
, same data as
<typeStatus id="9F3F48C92A6D7123FF7B43C47809CA93" box="[227,319,1179,1203]" pageId="14" pageNumber="15">holotype</typeStatus>
except for
<date id="343AD0AB2A6D7123FE2843C47B7ACA93" box="[432,588,1179,1203]" pageId="14" pageNumber="15" value="2019-06-19" valueMax="2019-06-31" valueMin="2019-06-19">
<collectingDate id="247E29432A6D7123FE2843C47B7ACA93" box="[432,588,1179,1203]" pageId="14" pageNumber="15" value="2019-06-19" valueMax="2019-06-31" valueMin="2019-06-19">VI/19-31/2019</collectingDate>
</date>
(
<collectionCode id="26956EAE2A6D7123FDC043C47BB2CA93" box="[600,644,1179,1203]" collectionName="JEE" pageId="14" pageNumber="15">JEE</collectionCode>
deposited in
<collectionCode id="26956EAE2A6D7123FF0E43E279EBCAF5" box="[150,221,1213,1237]" collectionName="USA, Florida, Gainesville, Division of Plant Industry, Florida State Collection of Arthropods" pageId="14" pageNumber="15">FSCA</collectionCode>
)
</materialsCitation>
<materialsCitation id="F0ECFC362A6D7123FF7D43E278B2CB3A" collectingDate="1986-06-07" collectingMethod="mercury vapor" collectionCode="DAR" collectorName="E. G. Riley &amp; D. A. Rider" country="French Guiana" location="Hwy ext. N 1, 20 km SE of St. Larent" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1">
;
<specimenCount id="56823DE22A6D7123FF7743E17835CAF5" box="[239,259,1214,1237]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
a)
<collectingCountry id="3893B6FB2A6D7123FEBC43E17888CAF5" box="[292,446,1213,1237]" pageId="14" pageNumber="15">
FR.
<collectingCountry id="3893B6FB2A6D7123FECA43E27888CAF5" box="[338,446,1213,1237]" name="Guyana" pageId="14" pageNumber="15">GUIANA</collectingCountry>
</collectingCountry>
,
<location id="455BA0B02A6D7123FE5043E179E8CAD7" LSID="urn:lsid:plazi:treatment:C82D477D2A6E7123FCF2454E7A3CC9A3:455BA0B02A6D7123FE5043E179E8CAD7" country="FR. GUIANA" name="Hwy ext. N 1, 20 km SE of St. Larent" pageId="14" pageNumber="15">
Hwy ext. N
<quantity id="877C5B8E2A6D7123FDD043E27B96CAF5" box="[584,672,1213,1237]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="14" pageNumber="15" unit="km" value="1.2">1, 20 km</quantity>
SE of St. Larent
</location>
,
<date id="343AD0AB2A6D7123FF7F4380786ECAD7" box="[231,344,1247,1271]" pageId="14" pageNumber="15" value="1986-06-07">
<collectingDate id="247E29432A6D7123FF7F4380786ECAD7" box="[231,344,1247,1271]" pageId="14" pageNumber="15" value="1986-06-07">7-VI-1986</collectingDate>
</date>
,
<collectorName id="ED7193BD2A6D7123FEF943BF7B5BCAD7" box="[353,621,1247,1272]" pageId="14" pageNumber="15">E. G. Riley &amp; D. A. Rider</collectorName>
; b) collected at
<collectingMethod id="99C58E7C2A6D7123FF0E425D7807CB39" box="[150,305,1282,1306]" pageId="14" pageNumber="15">mercury vapor</collectingMethod>
(
<collectionCode id="26956EAE2A6D7123FEA6425D784ACB3A" box="[318,380,1282,1306]" collectionName="DAR" pageId="14" pageNumber="15">DAR</collectionCode>
)
</materialsCitation>
<materialsCitation id="F0ECFC362A6D7123FE1C425D7B0DCB7E" collectingDate="2002-12-03" collectingMethod="MV Light" collectionCode="JEE" collectorName="J. E. Eger" country="French Guiana" elevation="94" latitude="4.2970834" location="17 km W of N 2 on Belizon Rd." longLatPrecision="1" longitude="-52.3703" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1">
;
<specimenCount id="56823DE22A6D7123FE15425D7897CB39" box="[397,417,1282,1305]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
a)
<collectingCountry id="3893B6FB2A6D7123FE59425D7BAACB3A" box="[449,668,1282,1306]" name="French Guiana" pageId="14" pageNumber="15">FRENCH GUIANA</collectingCountry>
:
<location id="455BA0B02A6D7123FD3D425D7847CB1C" LSID="urn:lsid:plazi:treatment:C82D477D2A6E7123FCF2454E7A3CC9A3:455BA0B02A6D7123FD3D425D7847CB1C" country="French Guiana" latitude="4.2970834" longLatPrecision="1" longitude="-52.3703" name="17 km W of N 2 on Belizon Rd." pageId="14" pageNumber="15">
<quantity id="877C5B8E2A6D7123FD3D425D7BDECB3A" box="[677,744,1282,1306]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="15" unit="km" value="17.0">17 km</quantity>
W of N2 on Belizon Rd.
</location>
,
<date id="343AD0AB2A6D7123FEE1427B78C7CB1C" box="[377,497,1316,1340]" pageId="14" pageNumber="15" value="2002-12-03">
<collectingDate id="247E29432A6D7123FEE1427B78C7CB1C" box="[377,497,1316,1340]" pageId="14" pageNumber="15" value="2002-12-03">3-XII-2002</collectingDate>
</date>
,
<collectorName id="ED7193BD2A6D7123FE61427B7B68CB1C" box="[505,606,1316,1340]" pageId="14" pageNumber="15">J. E. Eger</collectorName>
; b)
<geoCoordinate id="25B090AC2A6D7123FD19427B7A31CB1C" box="[641,775,1316,1340]" degrees="04" direction="north" minutes="17.825" orientation="latitude" pageId="14" pageNumber="15" precision="1" value="4.2970834">N04°17.825ʼ</geoCoordinate>
<geoCoordinate id="25B090AC2A6D7123FF0F42197801CB7E" box="[151,311,1350,1374]" degrees="052" direction="west" minutes="22.218" orientation="longitude" pageId="14" pageNumber="15" precision="1" value="-52.3703">W052°22.218ʼ</geoCoordinate>
,
<quantity id="877C5B8E2A6D7123FEDA4219784DCB7E" box="[322,379,1350,1374]" metricMagnitude="1" metricUnit="m" metricValue="9.4" pageId="14" pageNumber="15" unit="m" value="94.0">
<elevation id="CBA911582A6D7123FEDA4219784DCB7E" box="[322,379,1350,1374]" metricMagnitude="1" metricUnit="m" metricValue="9.4" pageId="14" pageNumber="15" unit="m" value="94.0">94 m</elevation>
</quantity>
,
<collectingMethod id="99C58E7C2A6D7123FE1E421878C7CB7E" box="[390,497,1350,1374]" pageId="14" pageNumber="15">MV Light</collectingMethod>
(
<collectionCode id="26956EAE2A6D7123FE6742197B19CB7E" box="[511,559,1350,1374]" pageId="14" pageNumber="15">JEE</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="089EA5E02A6D7123FF7B42377B4CCBA0" box="[227,634,1384,1408]" pageId="14" pageNumber="15" type="distribution">
<paragraph id="403BF66B2A6D7123FF7B42377B4CCBA0" blockId="14.[150,778,151,1923]" box="[227,634,1384,1408]" pageId="14" pageNumber="15">
Distribution. French Guiana (
<figureCitation id="D8BFEAEE2A6D7123FD8542377B5BCBA0" box="[541,621,1384,1408]" captionStart="Fig" captionStartId="12.[150,181,956,975]" captionTargetBox="[238,1352,151,932]" captionTargetId="figure-488@12.[238,1352,151,932]" captionTargetPageId="12" captionText="Fig. 80. Distribution of the species of Coptochilus, colored areas based on biogeographical dominions according to MORRONE (2013)." figureDoi="http://doi.org/10.5281/zenodo.12784970" httpUri="https://zenodo.org/record/12784970/files/figure.png" pageId="14" pageNumber="15">Fig. 80</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="089EA5E02A6D7123FF7B42D4784FCBC8" pageId="14" pageNumber="15" type="etymology">
<paragraph id="403BF66B2A6D7123FF7B42D4784FCBC8" blockId="14.[150,778,151,1923]" pageId="14" pageNumber="15">Etymology. This species is named for Roy Morris, a friend and colleague who collected the holotype and three of the five paratypes.</paragraph>
</subSubSection>
<subSubSection id="089EA5E02A6D7123FF7B42AE7A3CC9A3" pageId="14" pageNumber="15" type="discussion">
<paragraph id="403BF66B2A6D7123FF7B42AE7A3CC9A3" blockId="14.[150,778,151,1923]" pageId="14" pageNumber="15">
Comments. This species resembles
<taxonomicName id="87848DE82A6D7123FDC442AD7A31C829" authorityName="Distant" authorityYear="1899" box="[604,775,1521,1545]" class="Gastropoda" family="Pupinidae" genus="Coptochilus" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="neotropicalis">
<emphasis id="72F02A792A6D7123FDC442AD7B58C829" box="[604,622,1522,1545]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="72F02A792A6D7123FDE342AE7A31C829" box="[635,775,1521,1545]" italics="true" pageId="14" pageNumber="15">neotropicalis</emphasis>
</taxonomicName>
in having the pronotum and scutellum striped. It was misidentified as
<taxonomicName id="87848DE82A6D7123FEC141697B24C86E" authorityName="Distant" authorityYear="1899" box="[345,530,1590,1614]" class="Gastropoda" family="Pupinidae" genus="Coptochilus" kingdom="Animalia" order="Architaenioglossa" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="neotropicalis">
<emphasis id="72F02A792A6D7123FEC14169785DC86D" box="[345,363,1590,1613]" italics="true" pageId="14" pageNumber="15">C</emphasis>
.
<emphasis id="72F02A792A6D7123FEE541697B24C86E" box="[381,530,1590,1614]" italics="true" pageId="14" pageNumber="15">neotropicalis</emphasis>
</taxonomicName>
from French Guiana by
<bibRefCitation id="24158B9A2A6D7123FF23410778BFC850" author="EGER JR., J. E. &amp; BARCELLOS, A. &amp; WEILER, L." box="[187,393,1624,1648]" pageId="14" pageNumber="15" refId="ref10484" refString="EGER JR., J. E.; BARCELLOS, A. &amp; WEILER, L. 2015. Shield bugs (Scutelleridae). In: PANIZZI, A. R. &amp; GRAZIA J. eds. True Bugs (Heteroptera) of the Neotropics. New York, Springer, p. 757 - 788." type="book" year="2015">
EGER
<emphasis id="72F02A792A6D7123FF6541067800C850" box="[253,310,1624,1648]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
(2015)
</bibRefCitation>
. It differs from that species by the mandibular plates converging apically with only a small space between them and by the male internal genitalia. One male specimen has dark punctation ventrally on head and thorax and the abdomen completely black ventrally except for a narrow pale band laterally and posteriorly. This may be a result of discoloration, but it appears to be natural coloration. We were not able to obtain fully expanded male internal genitalia and opted not to dissect the single female paratype.
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