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<document id="E9EC9A9D2CFE43527DEA1E6B7BFB9E6D" ID-DOI="10.11646/zootaxa.4816.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3954419" ID-ZooBank="02EAEA41-160D-41FF-95E5-8F83E4A2BAD7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1595339832409" checkinUser="plazi" docAuthor="Johanson, Kjell Arne, Pham, Thai Hong, Malm, Tobias &amp; Sjöberg, Tin" docDate="2020" docId="03E74143FFE90A01FF4435E47104FE00" docLanguage="en" docName="zootaxa.4816.3.2.pdf" docOrigin="Zootaxa 4816 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oecetis lata Johanson &amp; Pham &amp; Malm &amp; Sjöberg 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="314" masterDocId="FFDE393BFFE80A02FFD336697039FFD3" masterDocTitle="Description of six new species of Oecetis (Trichoptera, Leptoceridae) from Vietnam" masterLastPageNumber="324" masterPageNumber="311" pageNumber="312" updateTime="1698783012615" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="121DDD8274753D4F8E6D4531A5A4BC82">
<mods:title id="E99D0A0D87981D78441D793EBD47A4BB">Description of six new species of Oecetis (Trichoptera, Leptoceridae) from Vietnam</mods:title>
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<mods:name id="F0B94A1D5A4E023FAA164314C3EC11B6" type="personal">
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<mods:namePart id="58D63CFD69ADC9B11865683F3952BA8E">Johanson, Kjell Arne</mods:namePart>
<mods:affiliation id="5557C110CDCB01C96CF8569D85DD771D">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden &amp; kjell. arne. johanson @ nrm. se; https: // orcid. org / 0000 - 0002 - 1893 - 3429</mods:affiliation>
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<mods:namePart id="77404B3CDE9A3751AE08689E40F5DEAB">Pham, Thai Hong</mods:namePart>
<mods:affiliation id="3AE8010E1F87D1C23AF4945AD43AF558">Vietnam National Museum of Nature (VNMN) &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet St, Cau Giay, Hanoi, Vietnam. &amp; Faculty of Biology, Hanoi National University of Education, 136 Xuan Thuy, Cau Giay, Hanoi, Vietnam. &amp; phamthai @ vnmn. vast. vn; https: // orcid. org / 0000 - 0002 - 4763 - 3679</mods:affiliation>
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<mods:namePart id="5E10F42A2E816A1CC5A3CED4DB29B1BF">Malm, Tobias</mods:namePart>
<mods:affiliation id="B713744C0B88FC83CE1FEAF29BEB5A70">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden &amp; tobias. malm @ nrm. se; https: // orcid. org / 0000 - 0003 - 4610 - 4086</mods:affiliation>
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<mods:namePart id="BBC9DD98F50ED741685E5AB8B2B50816">Sjöberg, Tin</mods:namePart>
<mods:affiliation id="B240AE4719B5DD8C9543270EEF6AC3FD">Department of Zoology, Swedish Museum of Natural History, Box 50007, 10405 Stockholm, Sweden &amp; tin. sjoberg @ nrm. se; https: // orcid. org / 0000 - 0002 - 2813 - 9325</mods:affiliation>
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<mods:title id="72551604BBA65C176C1E4F6C37054F40">Zootaxa</mods:title>
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<mods:date id="7504143C675B99EAB1E46F03A4E7ABC6">2020</mods:date>
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<mods:number id="A7AEE16A5D6013735D82A9B6B289FF09">2020-07-17</mods:number>
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<mods:number id="BA9BF70203583587E1E37441C50F49BD">4816</mods:number>
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<subSubSection id="C354A3DEFFE90A03FF4435E471B0FC74" box="[151,393,909,936]" pageId="1" pageNumber="312" type="nomenclature">
<paragraph id="8BF1F055FFE90A03FF4435E471B0FC74" blockId="1.[151,393,909,971]" box="[151,393,909,936]" pageId="1" pageNumber="312">
<heading id="D0B94739FFE90A03FF4435E471B0FC74" bold="true" box="[151,393,909,936]" fontSize="11" level="1" pageId="1" pageNumber="312" reason="1">
<emphasis id="B93A2C47FFE90A03FF4435E471B0FC74" bold="true" box="[151,393,909,936]" pageId="1" pageNumber="312">
<taxonomicName id="4C4E8BD6FFE90A03FF4435E4711DFC7B" authority="Johanson &amp; Pham &amp; Malm &amp; Sjöberg, 2020" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[151,292,909,936]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="lata" status="sp. nov.">
<emphasis id="B93A2C47FFE90A03FF4435E4711DFC7B" bold="true" box="[151,292,909,936]" italics="true" pageId="1" pageNumber="312">Oecetis lata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A209913CFFE90A03FEF835E471B0FC74" box="[299,393,909,935]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C354A3DEFFE90A03FF4435D870C1FC18" box="[151,248,944,971]" pageId="1" pageNumber="312" type="description">
<paragraph id="8BF1F055FFE90A03FF4435D870C1FC18" blockId="1.[151,393,909,971]" box="[151,248,944,971]" pageId="1" pageNumber="312">
<figureCitation id="1375ECD0FFE90A03FF4435D870C1FC18" box="[151,248,944,971]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1945,1970]" captionTargetBox="[208,1377,187,1906]" captionTargetId="figure@2.[193,1395,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Oecetis lata sp. nov., male holotype, genitalia. 1, left lateral; 2, dorsal; 3, ventral; 4, phallus, left lateral; 5, phallus, ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954421" httpUri="https://zenodo.org/record/3954421/files/figure.png" pageId="1" pageNumber="312">Figs 1–5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFE90A03FF4435917523FAE0" pageId="1" pageNumber="312" type="diagnosis">
<paragraph id="8BF1F055FFE90A03FF4435917523FAE0" blockId="1.[151,1437,1016,2016]" pageId="1" pageNumber="312">
<emphasis id="B93A2C47FFE90A03FF443591712BFBC0" bold="true" box="[151,274,1016,1043]" pageId="1" pageNumber="312">Diagnosis.</emphasis>
The male of this species is distinguished from many of the Oriental and Palearctic species by the absence of honeycomb structure on pregenital tergites V–VIII. It resembles
<taxonomicName id="4C4E8BD6FFE90A03FC1E327474D0FBE4" authority="Olah 2013" authorityName="Olah" authorityYear="2013" box="[973,1257,1053,1079]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="varosi">
<emphasis id="B93A2C47FFE90A03FC1E32747454FBE4" box="[973,1133,1053,1079]" italics="true" pageId="1" pageNumber="312">Oecetis varosi</emphasis>
<bibRefCitation id="EFDF8DA4FFE90A03FBA0327474D0FBE4" author="Olah, J." box="[1139,1257,1053,1079]" pageId="1" pageNumber="312" pagination="55 - 134" refId="ref7262" refString="Olah, J. (2013) On the Trichoptera of Vietnam, with description of 52 sp. nov. Annales Historico-Naturales Musei Nationalis Hungarici, 105, 55 - 134." type="journal article" year="2013">
Oláh
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in the genitalia, particularly the shape of the phallus and the coxopodites.
<taxonomicName id="4C4E8BD6FFE90A03FCF432287396FB88" authority="Johanson &amp; Pham &amp; Malm &amp; Sjöberg, 2020" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[807,943,1089,1115]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="lata" status="sp. nov.">
<emphasis id="B93A2C47FFE90A03FCF432287396FB88" box="[807,943,1089,1115]" italics="true" pageId="1" pageNumber="312">Oecetis lata</emphasis>
</taxonomicName>
<emphasis id="B93A2C47FFE90A03FC6532287428FB88" bold="true" box="[950,1041,1089,1115]" pageId="1" pageNumber="312">
<taxonomicNameLabel id="A209913CFFE90A03FC6532287428FB88" box="[950,1041,1089,1115]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from
<taxonomicName id="4C4E8BD6FFE90A03FAC332287541FB88" authorityName="Olah" authorityYear="2013" box="[1296,1400,1089,1115]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="varosi">
<emphasis id="B93A2C47FFE90A03FAC33228751DFB88" box="[1296,1316,1089,1115]" italics="true" pageId="1" pageNumber="312">O</emphasis>
.
<emphasis id="B93A2C47FFE90A03FAE0322B7541FB88" box="[1331,1400,1090,1115]" italics="true" pageId="1" pageNumber="312">varosi</emphasis>
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by the shorter ventral part of segment IX, broader tergum X in lateral and dorsal views, and the larger and more triangularly shaped superior appendages in lateral view. It generally resembles
<taxonomicName id="4C4E8BD6FFE90A03FC3032E07547FB70" authority="(Fabricius 1793)" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[995,1406,1161,1187]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="tripunctata">
<emphasis id="B93A2C47FFE90A03FC3032E07482FB70" box="[995,1211,1161,1187]" italics="true" pageId="1" pageNumber="312">Oecetis tripunctata</emphasis>
(
<bibRefCitation id="EFDF8DA4FFE90A03FB1832E0754FFB70" author="Fabricius, J. C." box="[1227,1398,1161,1187]" pageId="1" pageNumber="312" refId="ref6578" refString="Fabricius, J. C. (1793) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Vol. 2. Proft., Hafniae (Copenhagen), viii + 519 pp. https: // doi. org / 10.5962 / bhl. title. 122153" type="book" year="1793">Fabricius 1793</bibRefCitation>
)
</taxonomicName>
in the shape of tergum X, superior appendages, the coxopodites, and the phallus. It is distinguished from
<taxonomicName id="4C4E8BD6FFE90A03FB2C32C475A5FB14" baseAuthorityName="Fabricius" baseAuthorityYear="1793" box="[1279,1436,1197,1223]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="tripunctata">
<emphasis id="B93A2C47FFE90A03FB2C32C475A5FB14" box="[1279,1436,1197,1223]" italics="true" pageId="1" pageNumber="312">O. tripunctata</emphasis>
</taxonomicName>
by the superior appendages being shorter than tergum X, the broader coxopodites in lateral view, the longer ventral process, and the straight internal spine of the phallus. It also resembles
<taxonomicName id="4C4E8BD6FFE90A03FC61329C746DFADC" authority="Johanson &amp; Pham &amp; Malm &amp; Sjöberg, 2020" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[946,1108,1269,1295]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="312" phylum="Arthropoda" rank="species" species="gretae" status="sp. nov.">
<emphasis id="B93A2C47FFE90A03FC61329C746DFADC" box="[946,1108,1269,1295]" italics="true" pageId="1" pageNumber="312">Oecetis gretae</emphasis>
</taxonomicName>
<emphasis id="B93A2C47FFE90A03FB89329C748AFADC" bold="true" box="[1114,1203,1269,1295]" pageId="1" pageNumber="312">
<taxonomicNameLabel id="A209913CFFE90A03FB89329C748AFADC" box="[1114,1203,1269,1295]" pageId="1" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from which it is distinguished by the much larger superior appendages, taller tergum X, and coxopodites lacking branches.
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFE90A01FF1433557150FE5F" lastPageId="3" lastPageNumber="314" pageId="1" pageNumber="312" type="description">
<paragraph id="8BF1F055FFE90A03FF14335571E2F8A0" blockId="1.[151,1437,1016,2016]" pageId="1" pageNumber="312">
<emphasis id="B93A2C47FFE90A03FF14335571A6FA84" bold="true" box="[199,415,1340,1367]" pageId="1" pageNumber="312">Description, male.</emphasis>
Length of each forewing 6.0–
<quantity id="4CB65DB0FFE90A03FD3C3354737AFA84" box="[751,835,1341,1367]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="1" pageNumber="312" unit="mm" value="6.8">6.8 mm</quantity>
(N = 5). Genitalia (
<figureCitation id="1375ECD0FFE90A03FBC833547444FA84" box="[1051,1149,1340,1367]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1945,1970]" captionTargetBox="[208,1377,187,1906]" captionTargetId="figure@2.[193,1395,181,1919]" captionTargetPageId="2" captionText="FIGURES 1–5. Oecetis lata sp. nov., male holotype, genitalia. 1, left lateral; 2, dorsal; 3, ventral; 4, phallus, left lateral; 5, phallus, ventral. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3954421" httpUri="https://zenodo.org/record/3954421/files/figure.png" pageId="1" pageNumber="312">Figs 1–5</figureCitation>
) slightly higher than long in lateral view. Segment IX oriented almost vertically, ventral part slightly produced anteriorly; dorsal part rectilinear; anterior margin almost straight; acrotergite of segment IX well developed, punctate, undivided, almost as wide as each superior appendage in dorsal view; dorsal process of segment IX minute, forming single semimembranous papilla, wart-like; sternum with large rhomboid membranous area basal of coxopodites; lower posterolateral process of segment IX very shallow; posterior dorsolateral processes of segment IX short and right-angled; longitudinal apodeme absent. Tergum X slightly longer than superior appendages, in dorsal view wide along its length, slightly narrowing posteriorly to truncate apex; dorsal margin almost straight in lateral view; ventral margin membranous and convex in lateral view; apex small and rectangular in lateral view; upper part tergum X absent. Superior appendages wide and long, fused with tergum X, and almost covering tergum X in lateral view; oval in dorsal view. Each coxopodite slender, 1.5X as long as tergum X, unbranched, in lateral view dorsal margin shallowly W-shaped, ventral margin sub-straight; dorsally expanded into subtriangular swelling shortly beyond mid-length, apex almost straight; in ventral view sinuous and slightly tapering, especially beyond convexity at 2/3 length, elongate setae arising at lateral margin of and basal of convexity. Phallus slightly longer than superior appendages, with single, straight internal spine oriented diagonally and angled from left to right posteriorly; phallobase with long apicoventral process directed ventrad.
</paragraph>
<paragraph id="8BF1F055FFE90A03FF1431147118F833" blockId="1.[151,1437,1016,2016]" pageId="1" pageNumber="312">
<emphasis id="B93A2C47FFE90A03FF143114713DF844" bold="true" box="[199,260,1917,1943]" pageId="1" pageNumber="312">Male</emphasis>
<materialsCitation id="3B26FA08FFE90A03FED831147124F833" collectingDate="2011-04-05" collectorName="KA Johanson &amp; Johanson, PH &amp; TT Du" country="Vietnam" latitude="21.394472" location="Ngoc Thanh Village" municipality="Me Linh Distr" pageId="1" pageNumber="312" specimenCount="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<emphasis id="B93A2C47FFE90A03FED8311472FFF844" bold="true" box="[267,710,1916,1943]" pageId="1" pageNumber="312">
<typeStatus id="54F54EF7FFE90A03FED83114714FF844" box="[267,374,1917,1943]" pageId="1" pageNumber="312" type="holotype">holotype</typeStatus>
:
<collectingCountry id="F359B0C5FFE90A03FE51311571D5F844" box="[386,492,1916,1943]" name="Vietnam" pageId="1" pageNumber="312">Vietnam</collectingCountry>
:
<collectingRegion id="498A3EB7FFE90A03FE2B311572F8F844" box="[504,705,1916,1943]" country="Vietnam" name="Vinh Phuc" pageId="1" pageNumber="312">Vinh Phuc Prov.</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6B956A2FFFE90A03FD1E31147349F844" box="[717,880,1917,1943]" pageId="1" pageNumber="312">Me Linh Distr</collectingMunicipality>
, stream near
<location id="8E91A68EFFE90A03FBD5311474D5F844" LSID="urn:lsid:plazi:treatment:03E74143FFE90A01FF4435E47104FE00:8E91A68EFFE90A03FBD5311474D5F844" box="[1030,1260,1917,1943]" country="Vietnam" latitude="21.394472" municipality="Me Linh Distr" name="Ngoc Thanh Village" pageId="1" pageNumber="312" stateProvince="Vinh Phuc">Ngoc Thanh Village</location>
,
<geoCoordinate id="EE7A9692FFE90A03FB24311475A1F844" box="[1271,1432,1917,1943]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="1" pageNumber="312" precision="1" seconds="40.1" value="21.394472">21°23’40.1”N</geoCoordinate>
, 105°42’54.9’E,
<collectingMethod id="520F8842FFE90A03FE9831C87188F868" box="[331,433,1953,1979]" pageId="1" pageNumber="312">light trap</collectingMethod>
<date id="FFF0D695FFE90A03FE6431C97218F868" box="[439,545,1952,1979]" pageId="1" pageNumber="312" value="2011-04-05">
<collectingDate id="EFB42F7DFFE90A03FE6431C97218F868" box="[439,545,1952,1979]" pageId="1" pageNumber="312" value="2011-04-05">5.iv.2011</collectingDate>
</date>
, loc#VN008, leg.
<collectorName id="26BB9583FFE90A03FD3E31C873BCF868" box="[749,901,1953,1979]" pageId="1" pageNumber="312">KA Johanson</collectorName>
, PH Thai &amp;
<collectorName id="26BB9583FFE90A03FBC231C87467F868" box="[1041,1118,1953,1979]" pageId="1" pageNumber="312">TT Du</collectorName>
, DNA voucher JQ8, VNMN (in alcohol)
</materialsCitation>
.
</paragraph>
<caption id="DF31A0DDFFEA0A00FF4431F07234F806" ID-DOI="http://doi.org/10.5281/zenodo.3954421" ID-Zenodo-Dep="3954421" httpUri="https://zenodo.org/record/3954421/files/figure.png" pageId="2" pageNumber="313" startId="2.[151,264,1945,1970]" targetBox="[208,1377,187,1906]" targetPageId="2">
<paragraph id="8BF1F055FFEA0A00FF4431F07234F806" blockId="2.[151,1436,1945,2006]" pageId="2" pageNumber="313">
<emphasis id="B93A2C47FFEA0A00FF4431F0710FF861" bold="true" box="[151,310,1945,1970]" pageId="2" pageNumber="313">FIGURES 1–5</emphasis>
.
<taxonomicName id="4C4E8BD6FFEA0A00FE9631F371F9F861" authority="Johanson &amp; Pham &amp; Malm &amp; Sjöberg, 2020" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[325,448,1946,1970]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="313" phylum="Arthropoda" rank="species" species="lata" status="sp. nov.">
<emphasis id="B93A2C47FFEA0A00FE9631F371F9F861" box="[325,448,1946,1970]" italics="true" pageId="2" pageNumber="313">Oecetis lata</emphasis>
</taxonomicName>
<emphasis id="B93A2C47FFEA0A00FE1B31F37222F861" bold="true" box="[456,539,1946,1970]" pageId="2" pageNumber="313">
<taxonomicNameLabel id="A209913CFFEA0A00FE1B31F37222F861" box="[456,539,1946,1970]" pageId="2" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype, genitalia. 1, left lateral; 2, dorsal; 3, ventral; 4, phallus, left lateral; 5, phallus, ventral. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BF1F055FFEB0A01FF1436F07150FE5F" blockId="3.[151,1437,153,467]" pageId="3" pageNumber="314">
<materialsCitation id="3B26FA08FFEB0A01FF1436F07263FF04" collectingDate="2011-04-04" country="Vietnam" latitude="21.383972" location="Ngoc Thanh Village" longLatPrecision="1" longitude="105.71236" municipality="Me Linh District" pageId="3" pageNumber="314" specimenCount="1" stateProvince="Vinh Phuc" typeStatus="paratype">
<emphasis id="B93A2C47FFEB0A01FF1436F07106FF60" bold="true" box="[199,319,153,179]" pageId="3" pageNumber="314">
<typeStatus id="54F54EF7FFEB0A01FF1436F07106FF60" box="[199,319,153,179]" pageId="3" pageNumber="314" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="F359B0C5FFEB0A01FE8236F0718DFF60" box="[337,436,153,179]" name="Vietnam" pageId="3" pageNumber="314">Vietnam</collectingCountry>
:
<collectingRegion id="498A3EB7FFEB0A01FE1136F072BDFF60" box="[450,644,153,179]" country="Vietnam" name="Vinh Phuc" pageId="3" pageNumber="314">Vinh Phuc Prov.</collectingRegion>
,
<collectingMunicipality id="6B956A2FFFEB0A01FD4036F07362FF60" box="[659,859,153,179]" pageId="3" pageNumber="314">Me Linh District</collectingMunicipality>
, stream near
<location id="8E91A68EFFEB0A01FC2836F074D1FF60" LSID="urn:lsid:plazi:treatment:03E74143FFE90A01FF4435E47104FE00:8E91A68EFFEB0A01FC2836F074D1FF60" box="[1019,1256,153,179]" country="Vietnam" latitude="21.383972" longLatPrecision="1" longitude="105.71236" municipality="Me Linh District" name="Ngoc Thanh Village" pageId="3" pageNumber="314" stateProvince="Vinh Phuc">Ngoc Thanh Village</location>
,
<geoCoordinate id="EE7A9692FFEB0A01FB2436F075A1FF60" box="[1271,1432,153,179]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="314" precision="1" seconds="02.3" value="21.383972">21°23’02.3”N</geoCoordinate>
,
<geoCoordinate id="EE7A9692FFEB0A01FF4436D5717AFF04" box="[151,323,188,215]" degrees="105" direction="east" minutes="42" orientation="longitude" pageId="3" pageNumber="314" precision="1" seconds="44.5" value="105.71236">105°42’44.5”E</geoCoordinate>
,
<collectingMethod id="520F8842FFEB0A01FE8336D47181FF04" box="[336,440,189,215]" pageId="3" pageNumber="314">light trap</collectingMethod>
<date id="FFF0D695FFEB0A01FE1336D47210FF04" box="[448,553,189,215]" pageId="3" pageNumber="314" value="2011-04-04">
<collectingDate id="EFB42F7DFFEB0A01FE1336D47210FF04" box="[448,553,189,215]" pageId="3" pageNumber="314" value="2011-04-04">4.iv.2011</collectingDate>
</date>
, loc
</materialsCitation>
#
<collectingCountry id="F359B0C5FFEB0A01FDBB36D472ABFF04" box="[616,658,189,215]" name="Vietnam" pageId="3" pageNumber="314">VN</collectingCountry>
003, leg.
<collectorName id="26BB9583FFEB0A01FD2836D473ACFF04" box="[763,917,189,215]" pageId="3" pageNumber="314">KA Johanson</collectorName>
,
<materialsCitation id="3B26FA08FFEB0A01FC7136D47394FECC" collectingDate="2011-04-04" collectingDateMax="2011-04-12" collectingDateMin="2011-04-04" collectionCode="NHRS" collectorName="Thai &amp; TT Du" country="Philippines" county="Me Linh District" elevation="233" latitude="21.391167" location="Melinh Station" longLatPrecision="1" longitude="105.71297" municipality="Ngoc Thanh Village" pageId="3" pageNumber="314" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<collectingCountry id="F359B0C5FFEB0A01FC7136D473FFFF04" box="[930,966,189,215]" name="Philippines" pageId="3" pageNumber="314">PH</collectingCountry>
<collectorName id="26BB9583FFEB0A01FC1D36D47438FF04" box="[974,1025,189,215]" pageId="3" pageNumber="314">Thai</collectorName>
&amp;
<collectorName id="26BB9583FFEB0A01FBFB36D4744EFF04" box="[1064,1143,189,215]" pageId="3" pageNumber="314">TT Du</collectorName>
, DNA vouchers JR1, JR5 (
<specimenCount id="9D483BDCFFEB0A01FF73368870C2FF28" box="[160,251,225,251]" pageId="3" pageNumber="314" type="male">2 males</specimenCount>
,
<collectionCode id="ED5F6890FFEB0A01FEDB3688716AFF28" box="[264,339,225,251]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="3" pageNumber="314">NHRS</collectionCode>
in alcohol).
<collectingCounty id="629088D9FFEB0A01FE373688729FFF28" box="[484,678,225,251]" pageId="3" pageNumber="314">Me Linh District</collectingCounty>
, stream near
<collectingMunicipality id="6B956A2FFFEB0A01FCED3688741FFF28" box="[830,1062,225,251]" pageId="3" pageNumber="314">Ngoc Thanh Village</collectingMunicipality>
,
<location id="8E91A68EFFEB0A01FBE1368874E2FF28" LSID="urn:lsid:plazi:treatment:03E74143FFE90A01FF4435E47104FE00:8E91A68EFFEB0A01FBE1368874E2FF28" box="[1074,1243,225,251]" country="Philippines" county="Me Linh District" latitude="21.391167" longLatPrecision="1" longitude="105.71297" municipality="Ngoc Thanh Village" name="Melinh Station" pageId="3" pageNumber="314">Melinh Station</location>
for
<location id="8E91A68EFFEB0A01FADE368875A0FF28" LSID="urn:lsid:plazi:treatment:03E74143FFE90A01FF4435E47104FE00:8E91A68EFFEB0A01FADE368875A0FF28" box="[1293,1433,225,251]" country="Philippines" county="Me Linh District" latitude="21.391167" longLatPrecision="1" longitude="105.71297" municipality="Ngoc Thanh Village" name="Biodiversity" pageId="3" pageNumber="314">Biodiversity</location>
,
<geoCoordinate id="EE7A9692FFEB0A01FF44376C7101FECC" box="[151,312,261,287]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="314" precision="1" seconds="28.2" value="21.391167">21°23’28.2”N</geoCoordinate>
,
<geoCoordinate id="EE7A9692FFEB0A01FE96376D71C9FECC" box="[325,496,260,287]" degrees="105" direction="east" minutes="42" orientation="longitude" pageId="3" pageNumber="314" precision="1" seconds="46.7" value="105.71297">105°42’46.7”E</geoCoordinate>
,
<quantity id="4CB65DB0FFEB0A01FE2E376C7271FECC" box="[509,584,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="3" pageNumber="314" unit="m" value="233.0">
<elevation id="00631766FFEB0A01FE2E376C7271FECC" box="[509,584,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.33" pageId="3" pageNumber="314" unit="m" value="233.0">233 m</elevation>
</quantity>
,
<collectingMethod id="520F8842FFEB0A01FD86376C72DBFECC" box="[597,738,261,287]" pageId="3" pageNumber="314">Malaise trap</collectingMethod>
<date id="FFF0D695FFEB0A01FD39376C7344FECC" box="[746,893,261,287]" pageId="3" pageNumber="314" value="2011-04-04" valueMax="2011-04-12" valueMin="2011-04-04">
<collectingDate id="EFB42F7DFFEB0A01FD39376C7344FECC" box="[746,893,261,287]" pageId="3" pageNumber="314" value="2011-04-04" valueMax="2011-04-12" valueMin="2011-04-04">4–12.iv.2011</collectingDate>
</date>
, loc
</materialsCitation>
#
<collectingCountry id="F359B0C5FFEB0A01FC6F376C73DFFECC" box="[956,998,261,287]" name="Vietnam" pageId="3" pageNumber="314">VN</collectingCountry>
001, leg.
<collectorName id="26BB9583FFEB0A01FB9D376C74D0FECC" box="[1102,1257,261,287]" pageId="3" pageNumber="314">KA Johanson</collectorName>
,
<materialsCitation id="3B26FA08FFEB0A01FB26376C726DFEB4" collectingDate="2011-04-04" collectionCode="NHRS" collectorName="Thai &amp; TT Du" country="Philippines" latitude="21.394138" location="Ngoc Thanh Village" longLatPrecision="1" longitude="105.71408" municipality="Me Linh District" pageId="3" pageNumber="314" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F359B0C5FFEB0A01FB26376C7520FECC" box="[1269,1305,261,287]" name="Philippines" pageId="3" pageNumber="314">PH</collectingCountry>
<collectorName id="26BB9583FFEB0A01FAF2376C756DFECC" box="[1313,1364,261,287]" pageId="3" pageNumber="314">Thai</collectorName>
&amp;
<collectorName id="26BB9583FFEB0A01FAA8376C7085FE90" pageId="3" pageNumber="314">TT Du</collectorName>
, DNA voucher JN5 (
<specimenCount id="9D483BDCFFEB0A01FE74374071CBFE90" box="[423,498,297,323]" pageId="3" pageNumber="314" type="male">1 male</specimenCount>
,
<collectionCode id="ED5F6890FFEB0A01FE2E37407271FE90" box="[509,584,297,323]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="3" pageNumber="314">NHRS</collectionCode>
in alcohol).
<collectingMunicipality id="6B956A2FFFEB0A01FD00374073A8FE90" box="[723,913,297,323]" pageId="3" pageNumber="314">Me Linh District</collectingMunicipality>
, stream nr
<location id="8E91A68EFFEB0A01FBDA374074D4FE90" LSID="urn:lsid:plazi:treatment:03E74143FFE90A01FF4435E47104FE00:8E91A68EFFEB0A01FBDA374074D4FE90" box="[1033,1261,297,323]" country="Philippines" latitude="21.394138" longLatPrecision="1" longitude="105.71408" municipality="Me Linh District" name="Ngoc Thanh Village" pageId="3" pageNumber="314">Ngoc Thanh Village</location>
,
<geoCoordinate id="EE7A9692FFEB0A01FB24374075A1FE90" box="[1271,1432,297,323]" degrees="21" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="314" precision="1" seconds="38.9" value="21.394138">21°23’38.9”N</geoCoordinate>
,
<geoCoordinate id="EE7A9692FFEB0A01FF443725717EFEB4" box="[151,327,332,359]" degrees="105" direction="east" minutes="42" orientation="longitude" pageId="3" pageNumber="314" precision="1" seconds="50.7" value="105.71408">105°42’ 50.7”E</geoCoordinate>
,
<collectingMethod id="520F8842FFEB0A01FE8137247181FEB4" box="[338,440,333,359]" pageId="3" pageNumber="314">light trap</collectingMethod>
<date id="FFF0D695FFEB0A01FE6D3724721EFEB4" box="[446,551,333,359]" pageId="3" pageNumber="314" value="2011-04-04">
<collectingDate id="EFB42F7DFFEB0A01FE6D3724721EFEB4" box="[446,551,333,359]" pageId="3" pageNumber="314" value="2011-04-04">4.iv.2011</collectingDate>
</date>
, loc
</materialsCitation>
#
<collectingCountry id="F359B0C5FFEB0A01FDB0372472B4FEB4" box="[611,653,333,359]" name="Vietnam" pageId="3" pageNumber="314">VN</collectingCountry>
002, leg.
<collectorName id="26BB9583FFEB0A01FD21372473B3FEB4" box="[754,906,333,359]" pageId="3" pageNumber="314">KA Johanson</collectorName>
,
<materialsCitation id="3B26FA08FFEB0A01FC463724715CFE5F" collectionCode="NHRS" collectorName="Thai &amp; TT Du" country="Philippines" location="PH" pageId="3" pageNumber="314" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F359B0C5FFEB0A01FC4637247380FEB4" box="[917,953,333,359]" name="Philippines" pageId="3" pageNumber="314">PH</collectingCountry>
<collectorName id="26BB9583FFEB0A01FC6D372473C8FEB4" box="[958,1009,333,359]" pageId="3" pageNumber="314">Thai</collectorName>
&amp;
<collectorName id="26BB9583FFEB0A01FBC737247458FEB4" box="[1044,1121,333,359]" pageId="3" pageNumber="314">TT Du</collectorName>
, DNA voucher JQ9 (
<specimenCount id="9D483BDCFFEB0A01FA9F372475A1FEB4" box="[1356,1432,333,359]" pageId="3" pageNumber="314" type="male">1 male</specimenCount>
,
<collectionCode id="ED5F6890FFEB0A01FF44371870DBFE58" box="[151,226,369,395]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="3" pageNumber="314">NHRS</collectionCode>
in alcohol)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C354A3DEFFEB0A01FF1437FC7104FE00" pageId="3" pageNumber="314" type="etymology">
<paragraph id="8BF1F055FFEB0A01FF1437FC7104FE00" blockId="3.[151,1437,153,467]" pageId="3" pageNumber="314">
<emphasis id="B93A2C47FFEB0A01FF1437FC7177FE7C" bold="true" box="[199,334,405,431]" pageId="3" pageNumber="314">Etymology.</emphasis>
Latin adjective
<taxonomicName id="4C4E8BD6FFEB0A01FDD137FC7217FE7C" authorityName="Johanson &amp; Pham &amp; Malm &amp; Sjöberg" authorityYear="2020" box="[514,558,405,431]" class="Insecta" family="Leptoceridae" genus="Oecetis" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="314" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B93A2C47FFEB0A01FDD137FC7217FE7C" box="[514,558,405,431]" italics="true" pageId="3" pageNumber="314">lata</emphasis>
</taxonomicName>
, meaning “broad” or “wide,” referring to the shape of the superior appendages in lateral view.
</paragraph>
</subSubSection>
</treatment>
</document>
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