273 lines
37 KiB
XML
273 lines
37 KiB
XML
<document id="3D9407A685777F1DAF68A691E95382AD" ID-CLB-Dataset="45928" ID-DOI="10.11646/zootaxa.3793.2.3" ID-GBIF-Dataset="cec96d21-19fa-4e66-8305-9e70166f04f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="230581" ID-ZooBank="5E6DB00C-E98F-4D9D-958B-95B634E6D9E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460412815762" checkinUser="plazi" docAuthor="Baca, Stephen M., Gustafson, Grey T., Toledo, Mario & Miller, Kelly B." docDate="2014" docId="03D9AD22FFBBFFFB75A8FA05FFA693EB" docLanguage="en" docName="zt03793p246.pdf" docOrigin="Zootaxa 3793 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liocanthydrus armulatus Baca, Gustafson, Toledo & Miller, 2014, sp. n." docType="treatment" docVersion="9" lastPageNumber="241" masterDocId="FFE0D55AFFB2FFF1753FFF9FFF65972F" masterDocTitle="Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species" masterLastPageNumber="246" masterPageNumber="231" pageNumber="240" updateTime="1698301202102" updateUser="ExternalLinkService">
|
||
<mods:mods id="7BAFC1A344372CD6A180237E65D6D329" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="F3039CE502EDC2B8F5E14764D57B930E">
|
||
<mods:title id="3E8658FEAE7ACD07252DF78259822D51">Revision of the Neotropical burrowing water beetle genus Liocanthydrus Guignot (Coleoptera: Noteridae: Noterinae: Noterini) with the description of two new species</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4540C0F440764565D0729C4114C33732" type="personal">
|
||
<mods:role id="006A02230778C1DA5111B61FE195B6B8">
|
||
<mods:roleTerm id="3404CAF964E5401F783C814A036494A8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2163CA2D2262D51B2533F77BAEC5DDF2">Baca, Stephen M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D49EE727DC62E2B146C3041911FF36D2" type="personal">
|
||
<mods:role id="2EEF14C6826AD83731C08AD4FF672830">
|
||
<mods:roleTerm id="3965EFCA9903EC4FF375F51F7A13BDFD">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9C7C7271BC77B0E56C4057492CA1438A">Gustafson, Grey T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="527BE71EDED2DC948BD3E2174130C923" type="personal">
|
||
<mods:role id="CBAFEC22DEC126B6FDE1422728CD846E">
|
||
<mods:roleTerm id="C4DFCAE460479DDFF31CC2729FD30B4A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="018C9D164367728877D383BAA41ADF38">Toledo, Mario</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F4E0BAB7B037CA1FF699DEC820F8BD8A" type="personal">
|
||
<mods:role id="2565B3F172FAD0DD1ABB13A5CC55AEE4">
|
||
<mods:roleTerm id="C0D5AA354E3AEAF54378BC560F6ADEBA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="66A7341AFBFA98161E3815B7DCECD9F0">Miller, Kelly B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="A93117D8D27E5646CB3C157505B5F1BB">text</mods:typeOfResource>
|
||
<mods:relatedItem id="2673106664449393257F704FF427265A" type="host">
|
||
<mods:titleInfo id="62ED81A1FE18882863C2281AB5E12B6A">
|
||
<mods:title id="C21277FA997DA4B54E04F536E28ECDC2">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="113E9084B4BE1B3DAFD4E35C3DCE4139">
|
||
<mods:date id="1B396E7D8567EAC58542C60CE98DA2B5">2014</mods:date>
|
||
<mods:detail id="A3AB339224FDC0659D5DBCF89D504967" type="volume">
|
||
<mods:number id="FB200885AB30F2C0128CDD2C9E405817">3793</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="156723F0A3E1D6E1062137DE22FC5DDD" type="issue">
|
||
<mods:number id="AD71E54FF2B4B00481F9E41C657FBBB4">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="8B0D8A8B010E85CE401464C3E4A6F2B4" unit="page">
|
||
<mods:start id="7009BEB206AF3EE75D62C07F3E14F4F4">231</mods:start>
|
||
<mods:end id="8BACC092584F7812AA44E5B198951155">246</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="948A2862EAEAB22A60738088CE2C18B4">journal article</mods:classification>
|
||
<mods:identifier id="D22263B7A59F9B9E83503DA16470E94A" type="CLB-Dataset">45928</mods:identifier>
|
||
<mods:identifier id="70C8D76CAF576C5614CF8FF9E8F7A33F" type="DOI">10.11646/zootaxa.3793.2.3</mods:identifier>
|
||
<mods:identifier id="73D03732A097851D967B751576D567A7" type="GBIF-Dataset">cec96d21-19fa-4e66-8305-9e70166f04f7</mods:identifier>
|
||
<mods:identifier id="7B000AFD6522C5F9034E5E980182A972" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="59A48A82A46259088C515CB780ACC61C" type="Zenodo-Dep">230581</mods:identifier>
|
||
<mods:identifier id="2EEFEF69BAB601CFE749E1393FBED1FE" type="ZooBank">5E6DB00C-E98F-4D9D-958B-95B634E6D9E4</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D9AD22FFBBFFFB75A8FA05FFA693EB" ID-DOI="http://doi.org/10.5281/zenodo.6131931" ID-GBIF-Taxon="119423981" ID-Zenodo-Dep="6131931" LSID="urn:lsid:plazi:treatment:03D9AD22FFBBFFFB75A8FA05FFA693EB" httpUri="http://treatment.plazi.org/id/03D9AD22FFBBFFFB75A8FA05FFA693EB" lastPageId="10" lastPageNumber="241" pageId="9" pageNumber="240">
|
||
<subSubSection id="C36A4FBFFFBBFFF875A8FA05FD76929A" box="[151,531,1434,1461]" pageId="9" pageNumber="240" type="nomenclature">
|
||
<paragraph id="8BCF1C34FFBBFFF875A8FA05FD76929A" blockId="9.[151,531,1434,1461]" box="[151,531,1434,1461]" pageId="9" pageNumber="240">
|
||
<heading id="D087AB58FFBBFFF875A8FA05FD76929A" bold="true" box="[151,531,1434,1461]" fontSize="11" level="1" pageId="9" pageNumber="240" reason="1">
|
||
<emphasis id="B904C026FFBBFFF875A8FA05FD76929A" bold="true" box="[151,531,1434,1461]" pageId="9" pageNumber="240">
|
||
<taxonomicName id="4C7067B7FFBBFFF875A8FA05FEAF929B" box="[151,458,1434,1460]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="240" phylum="Arthropoda" rank="species" species="armulatus" status="sp. nov.">
|
||
<emphasis id="B904C026FFBBFFF875A8FA05FEAF929B" bold="true" box="[151,458,1434,1460]" italics="true" pageId="9" pageNumber="240">Liocanthydrus armulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377D5DFFBBFFF874EEFA04FD76929A" box="[465,531,1435,1461]" pageId="9" pageNumber="240" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBBFFF875A8FA7EFD5D9035" pageId="9" pageNumber="240" type="materials_examined">
|
||
<paragraph id="8BCF1C34FFBBFFF875A8FA7EFD9892D5" blockId="9.[151,1436,1505,2034]" box="[151,765,1505,1530]" pageId="9" pageNumber="240">
|
||
<emphasis id="B904C026FFBBFFF875A8FA7EFE4892D5" bold="true" box="[151,301,1505,1530]" pageId="9" pageNumber="240">
|
||
<typeStatus id="54CBA296FFBBFFF875A8FA7EFFB792D5" box="[151,210,1505,1530]" pageId="9" pageNumber="240">Type</typeStatus>
|
||
locality
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="F3675CA4FFBBFFF87403FA7EFED792D5" box="[316,434,1505,1530]" name="Venezuela" pageId="9" pageNumber="240">Venezuela</collectingCountry>
|
||
, Bolivar State, Road to Pauji.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFBBFFF875F8F99AFD5D9035" blockId="9.[151,1436,1505,2034]" pageId="9" pageNumber="240">
|
||
<emphasis id="B904C026FFBBFFF875F8F99AFE949131" bold="true" box="[199,497,1541,1566]" pageId="9" pageNumber="240">
|
||
<typeStatus id="54CBA296FFBBFFF875F8F99AFE679131" box="[199,258,1541,1566]" pageId="9" pageNumber="240">Type</typeStatus>
|
||
material examined.
|
||
</emphasis>
|
||
<typeStatus id="54CBA296FFBBFFF874C6F99AFD049131" box="[505,609,1541,1566]" pageId="9" pageNumber="240" type="holotype">Holotype</typeStatus>
|
||
(male): “
|
||
<collectingCountry id="F3675CA4FFBBFFF877F3F999FC1C9131" box="[716,889,1542,1566]" name="Venezuela" pageId="9" pageNumber="240">VENEZUELA</collectingCountry>
|
||
: Bolivar State/
|
||
<geoCoordinate id="EE447AF3FFBBFFF8711AF99AFBC89131" box="[1061,1197,1541,1566]" direction="north" orientation="latitude" pageId="9" pageNumber="240" precision="1" value="4.614222">4°36'51.2"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE447AF3FFBBFFF87185F99AFA369131" box="[1210,1363,1541,1566]" direction="west" orientation="longitude" pageId="9" pageNumber="240" precision="1" value="-61.391724">61°23'30.2"W</geoCoordinate>
|
||
/Road to Pauji;
|
||
<date id="FFCE3AF4FFBBFFF875C3F9B6FE17916D" box="[252,370,1577,1602]" pageId="9" pageNumber="240" value="2008-08-03">3.viii.2008</date>
|
||
/
|
||
<quantity id="4C88B1D1FFBBFFF87444F9B6FEA0916E" box="[379,453,1577,1602]" metricMagnitude="2" metricUnit="m" metricValue="8.95" pageId="9" pageNumber="240" unit="m" value="895.0">895 m</quantity>
|
||
, leg. A. Short/AS-08-070; Stream margin” [printed], “(barcode) SM0828443
|
||
<collectionCode id="ED6184F1FFBBFFF87016F9B5FAF9916E" box="[1321,1436,1578,1601]" pageId="9" pageNumber="240">KUNHM-</collectionCode>
|
||
<collectionCode id="ED6184F1FFBBFFF875A8F9D1FFB59149" LSID="urn:lsid:biocol.org:col:13589" box="[151,208,1614,1638]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="9" pageNumber="240">ENT</collectionCode>
|
||
” [printed], “
|
||
<typeStatus id="54CBA296FFBBFFF87464F9D1FE949149" box="[347,497,1614,1638]" pageId="9" pageNumber="240" type="holotype">HOLOTYPE</typeStatus>
|
||
/
|
||
<emphasis id="B904C026FFBBFFF874C6F9D2FC719149" box="[505,788,1613,1638]" italics="true" pageId="9" pageNumber="240">
|
||
<taxonomicName id="4C7067B7FFBBFFF874C6F9D2FDFD9149" box="[505,664,1613,1638]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="240" phylum="Arthropoda" rank="genus">Liocanthydrus</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBBFFF8779AF9D2FC719149" box="[677,788,1613,1638]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="240" phylum="Arthropoda" rank="species" species="armulatus">armulatus</taxonomicName>
|
||
</emphasis>
|
||
/Baca, Gustafson, Toledo & Miller, 2014” [red with black line border; printed] (
|
||
<collectionCode id="ED6184F1FFBBFFF874B7F9EDFEB391A5" LSID="urn:lsid:biocol.org:col:33834" box="[392,470,1650,1674]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="9" pageNumber="240">MIZA</collectionCode>
|
||
).
|
||
<typeStatus id="54CBA296FFBBFFF874D7F9EDFD3091A5" box="[488,597,1650,1674]" pageId="9" pageNumber="240" type="paratype">Paratypes</typeStatus>
|
||
(males): Same data as
|
||
<typeStatus id="54CBA296FFBBFFF87666F9EEFCA491A5" box="[857,961,1649,1674]" pageId="9" pageNumber="240" type="holotype">Holotype</typeStatus>
|
||
except “SM0828445 KUNHM-ENT” and “SM0828448 KUNHM-ENT” (2:
|
||
<collectionCode id="ED6184F1FFBBFFF87714F909FD199181" box="[555,636,1686,1710]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="240">SEMC</collectionCode>
|
||
).
|
||
<typeStatus id="54CBA296FFBBFFF877AAF909FC679181" box="[661,770,1686,1710]" pageId="9" pageNumber="240" type="paratype">Paratypes</typeStatus>
|
||
(females): Same data as
|
||
<typeStatus id="54CBA296FFBBFFF87106F90AFBC49181" box="[1081,1185,1685,1710]" pageId="9" pageNumber="240" type="holotype">Holotype</typeStatus>
|
||
except “SM0828446 KUNHM-ENT” and “SM0828447 KUNHM-ENT” (2:
|
||
<collectionCode id="ED6184F1FFBBFFF87635F925FC3E91FD" box="[778,859,1722,1746]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="240">SEMC</collectionCode>
|
||
).
|
||
<emphasis id="B904C026FFBBFFF87651F926FCDF91FD" bold="true" box="[878,954,1721,1746]" pageId="9" pageNumber="240">Notes:</emphasis>
|
||
All
|
||
<typeStatus id="54CBA296FFBBFFF876D0F925FB3F91FD" box="[1007,1114,1722,1746]" pageId="9" pageNumber="240" type="paratype">paratypes</typeStatus>
|
||
labeled with: “
|
||
<typeStatus id="54CBA296FFBBFFF87039F925FAF191FD" box="[1286,1428,1722,1746]" pageId="9" pageNumber="240" type="paratype">PARATYPE</typeStatus>
|
||
/
|
||
<emphasis id="B904C026FFBBFFF875A8F942FED791D9" box="[151,434,1757,1782]" italics="true" pageId="9" pageNumber="240">
|
||
<taxonomicName id="4C7067B7FFBBFFF875A8F942FE5291D9" box="[151,311,1757,1782]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="240" phylum="Arthropoda" rank="genus">Liocanthydrus</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C7067B7FFBBFFF8747CF942FED791D9" box="[323,434,1757,1782]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="240" phylum="Arthropoda" rank="species" species="armulatus">armulatus</taxonomicName>
|
||
</emphasis>
|
||
/Baca, Gustafson, Toledo & Miller, 2014” [blue with black line border; printed]. All label descriptions are included in brackets.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFBBFFFB75F8F8BAFBC89633" lastPageId="10" lastPageNumber="241" pageId="9" pageNumber="240" type="diagnosis">
|
||
<paragraph id="8BCF1C34FFBBFFF875F8F8BAFA3190DD" blockId="9.[151,1436,1505,2034]" pageId="9" pageNumber="240">
|
||
<emphasis id="B904C026FFBBFFF875F8F8BAFE249011" bold="true" box="[199,321,1829,1854]" pageId="9" pageNumber="240">Diagnosis.</emphasis>
|
||
Elytra maculate. Distinguishable from other maculate species by the following combination of characters: a) robust, humeral region very broad, TL/GW = 2.00–2.03, PNWpost./PNWant. = 1.64–1.68 (
|
||
<figureCitation id="134B00B1FFBBFFF87014F8D6FA0A904D" box="[1323,1391,1865,1890]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="9" pageNumber="240">Fig. 6</figureCitation>
|
||
); b) yellow bands on elytra broad, not reduced to spots, and all extending perpendicular to elytral suture (
|
||
<figureCitation id="134B00B1FFBBFFF87018F8F2FA0B90A9" box="[1319,1390,1901,1926]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="9" pageNumber="240">Fig. 6</figureCitation>
|
||
); c) aedeagus as in
|
||
<figureCitation id="134B00B1FFBBFFF8747DF80EFEF09085" box="[322,405,1937,1962]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="9" pageNumber="240">Figs 19</figureCitation>
|
||
a–e, with distal region of median lobe curved dorsally, moderately elongate and subparallel in lateral aspect, (
|
||
<figureCitation id="134B00B1FFBBFFF8745DF82AFEDF90E1" box="[354,442,1973,1998]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="9" pageNumber="240">Figs 19</figureCitation>
|
||
a, c); left lateral lobe with setal fringe extended basally to ca. median of overall length along dorsal margin(
|
||
<figureCitation id="134B00B1FFBBFFF874BDF846FEB590DD" box="[386,464,2009,2034]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="9" pageNumber="240">Fig. 19</figureCitation>
|
||
d); right lateral lobe subtriangular, distal angle projected and rounded (
|
||
<figureCitation id="134B00B1FFBBFFF871D8F846FA5C90DD" box="[1255,1337,2009,2034]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="9" pageNumber="240">Fig. 19</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FF08FBC89633" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
Externally
|
||
<taxonomicName id="4C7067B7FFB8FFFB747CFF06FEB6979F" box="[323,467,151,176]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="241" phylum="Arthropoda" rank="species" species="armulatus" status="sp. nov.">
|
||
<emphasis id="B904C026FFB8FFFB747CFF06FEB6979F" box="[323,467,151,176]" italics="true" pageId="10" pageNumber="241">L. armulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2377D5DFFB8FFFB74E4FF07FD729780" box="[475,535,152,175]" pageId="10" pageNumber="241" rank="species">n. sp.</taxonomicNameLabel>
|
||
appears very similar to
|
||
<emphasis id="B904C026FFB8FFFB7617FF06FCFE979F" box="[808,923,151,176]" italics="true" pageId="10" pageNumber="241">
|
||
<taxonomicName id="4C7067B7FFB8FFFB7617FF06FCF2979F" box="[808,919,151,176]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="241" phylum="Arthropoda" rank="species" species="bicolor">L. bicolor</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
but is noticeably more robust.
|
||
<taxonomicName id="4C7067B7FFB8FFFB71C5FF08FE6C97FA" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="241" phylum="Arthropoda" rank="species" species="armulatus">
|
||
<emphasis id="B904C026FFB8FFFB71C5FF08FE6C97FA" italics="true" pageId="10" pageNumber="241">Liocanthydrus armulatus</emphasis>
|
||
</taxonomicName>
|
||
is also distinguished by its distinct aedeagus, with a median lobe that, in lateral aspect, is distally more elongate and subparallel, not blunt. The aedeagus of
|
||
<taxonomicName id="4C7067B7FFB8FFFB77DBFF7EFC1697D7" box="[740,883,223,248]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="241" phylum="Arthropoda" rank="species" species="armulatus">
|
||
<emphasis id="B904C026FFB8FFFB77DBFF7EFC1697D7" box="[740,883,223,248]" italics="true" pageId="10" pageNumber="241">L. armulatus</emphasis>
|
||
</taxonomicName>
|
||
also differs by its subtriangular right lateral lobe, with a distinctly projected and rounded distal angle, not triangular with the distal angle acute.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFB8FFFB75F8FEB8FE0794C2" pageId="10" pageNumber="241" type="description">
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FEB8FEF496A7" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FEB8FE3C966F" bold="true" box="[199,345,295,320]" pageId="10" pageNumber="241">Description.</emphasis>
|
||
<emphasis id="B904C026FFB8FFFB745EFEB6FD6E966F" box="[353,523,297,320]" italics="true" pageId="10" pageNumber="241">Measurements.</emphasis>
|
||
TL =
|
||
<quantity id="4C88B1D1FFB8FFFB7767FEB8FD979610" box="[600,754,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="3.125" metricValueMax="3.15" metricValueMin="3.1" pageId="10" pageNumber="241" unit="mm" value="3.125" valueMax="3.15" valueMin="3.1">3.1–3.15 mm</quantity>
|
||
, GW =
|
||
<quantity id="4C88B1D1FFB8FFFB7678FEB8FC859610" box="[839,992,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.55" metricValueMin="1.5" pageId="10" pageNumber="241" unit="mm" value="1.525" valueMax="1.55" valueMin="1.5">1.5–1.55 mm</quantity>
|
||
, TL/GW = 2.00–2.03, HW =
|
||
<quantity id="4C88B1D1FFB8FFFB7005FEB8FFA0964B" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="10" pageNumber="241" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.9–0.95 mm</quantity>
|
||
, EW =
|
||
<quantity id="4C88B1D1FFB8FFFB7423FED3FED1964B" box="[284,436,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="10" pageNumber="241" unit="mm" value="0.575" valueMax="0.6" valueMin="0.55">0.55–0.6 mm</quantity>
|
||
, HW/EW = 1.58–1.64, PNWant. =
|
||
<quantity id="4C88B1D1FFB8FFFB767AFED3FCB8964B" box="[837,989,332,357]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="10" pageNumber="241" unit="mm" value="0.925" valueMax="0.95" valueMin="0.9">0.9–0.95 mm</quantity>
|
||
, PNWpost. =
|
||
<quantity id="4C88B1D1FFB8FFFB7143FED3FA71964B" box="[1148,1300,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.55" metricValueMin="1.5" pageId="10" pageNumber="241" unit="mm" value="1.525" valueMax="1.55" valueMin="1.5">1.5–1.55 mm</quantity>
|
||
, PNWpost./ PNWant. = 1.64–1.68.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FE0BFBEE9537" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FE0BFE2F9682" box="[199,330,404,429]" italics="true" pageId="10" pageNumber="241">Coloration.</emphasis>
|
||
Elytra maculate, with transverse bands; color of elytra brown to dark brown; color of head, pronotum and elytral bands ranging from yellow to brownish yellow; all examined specimens with small darkened maculation situated on anteromedial region of pronotum; elytral bands as in
|
||
<figureCitation id="134B00B1FFB8FFFB76C4FE43FB2596DA" box="[1019,1088,476,501]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="10" pageNumber="241">Fig. 6</figureCitation>
|
||
, broad, not obliquely oriented, and discontinuous across elytral suture; color of venter yellowish brown to reddish brown.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FDBBFE28954F" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FDBBFE629512" box="[199,263,548,573]" italics="true" pageId="10" pageNumber="241">Body.</emphasis>
|
||
Body outline as in
|
||
<figureCitation id="134B00B1FFB8FFFB74D5FDBBFD4A9512" box="[490,559,548,573]" captionStart="FIGURES 3 – 9" captionStartId="5.[151,264,1206,1228]" captionTargetBox="[192,1397,212,1170]" captionTargetId="figure@5.[172,1415,193,1185]" captionTargetPageId="5" captionText="FIGURES 3 – 9. Liocanthydrus species, dorsal habitus: 3) L. angustus; 4) L. uniformis; 5) L. bicolor; 6) L. armulatus sp. n.; 7) L. clayae; 8) L. nanops sp. n.; 9) L. octoguttatus (illustration). Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230583/files/figure.png" pageId="10" pageNumber="241">Fig. 6</figureCitation>
|
||
; oval, robust, very broad, attenuate posteriorly; moderately convex in lateral aspect (
|
||
<figureCitation id="134B00B1FFB8FFFB75D3FDD8FE5A954F" box="[236,319,583,608]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="10" pageNumber="241">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FDF3FCE195AA" blockId="10.[151,1437,151,1220]" box="[199,900,620,645]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FDF3FE6E95AA" box="[199,267,620,645]" italics="true" pageId="10" pageNumber="241">Head.</emphasis>
|
||
Eye size relative to head capsule: HW/EW = 1.58–1.64.
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FD10FBEB9587" blockId="10.[151,1437,151,1220]" box="[199,1166,655,680]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FD10FE459587" box="[199,288,655,680]" italics="true" pageId="10" pageNumber="241">Thorax.</emphasis>
|
||
Elytra with margins moderately ascendant medially in lateral aspect (
|
||
<figureCitation id="134B00B1FFB8FFFB7112FD10FBE59587" box="[1069,1152,655,680]" captionStart="FIGURES 10 – 15" captionStartId="5.[151,264,1970,1992]" captionTargetBox="[164,1430,1309,1937]" captionTargetId="figure@5.[151,1436,1309,1949]" captionTargetPageId="5" captionText="FIGURES 10 – 15. Liocanthydrus species, lateral aspect: 10) L. angustus; 11) L. uniformis; 12) L. bicolor; 13) L. armulatus sp. n.; 14) L. clayae; 15) L. nanops sp. n. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/230584/files/figure.png" pageId="10" pageNumber="241">Fig. 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FD2BFE3C9473" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FD2BFE5D95E2" box="[199,312,692,717]" italics="true" pageId="10" pageNumber="241">Abdomen.</emphasis>
|
||
Aedeagus as in
|
||
<figureCitation id="134B00B1FFB8FFFB74CBFD2BFD2D95E3" box="[500,584,692,717]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="10" pageNumber="241">Figs 19</figureCitation>
|
||
a–e; distal portion of median lobe subparallel in lateral aspect (
|
||
<figureCitation id="134B00B1FFB8FFFB702AFD2BFA0995E3" box="[1301,1388,692,717]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="10" pageNumber="241">Figs 19</figureCitation>
|
||
a, c) and rounding at apex, lateral margins sinuate (
|
||
<figureCitation id="134B00B1FFB8FFFB7794FD48FD9B95DF" box="[683,766,727,752]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="10" pageNumber="241">Fig. 19</figureCitation>
|
||
b), ventral groove nearly extending to apex (
|
||
<figureCitation id="134B00B1FFB8FFFB71C1FD48FA3795DF" box="[1278,1362,727,752]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="10" pageNumber="241">Fig. 19</figureCitation>
|
||
c); left lateral lobe weakly curved; setal fringe extending ca. to, or just short of, median of inner dorsal margin (
|
||
<figureCitation id="134B00B1FFB8FFFB7013FD63FA1B943A" box="[1324,1406,764,789]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="10" pageNumber="241">Fig. 19</figureCitation>
|
||
d); right lateral lobe subtriangular, distal angle strongly projected and dorsal margin broadly emarginated along its length (
|
||
<figureCitation id="134B00B1FFB8FFFB75D3FCDBFE5B9473" box="[236,318,836,861]" captionStart="FIGURES 16 – 22" captionStartId="6.[151,264,965,987]" captionTargetBox="[163,1436,193,935]" captionTargetId="figure@6.[151,1436,193,944]" captionTargetPageId="6" captionText="FIGURES 16 – 22. Aedeagi of Liocanthydrus species. a) median lobe, right lateral aspect; b) median lobe, dorsal aspect; c) median lobe, left lateral aspect; d) left lateral lobe, right lateral aspect; e) right lateral lobe, left lateral aspect: 16) L. angustus; 17) L. uniformis; 18) L. bicolor; 19) L. armulatus sp. n.; 20) L. clayae; 21) L. nanops sp. n.; 22) L. octoguttatus. Scale bars = 0.2 mm. Notes: Terminology follows K. B. Miller & Nilsson (2003). Illustrations of lateral lobes depict medial (inner) surfaces." httpUri="https://zenodo.org/record/230585/files/figure.png" pageId="10" pageNumber="241">Fig. 19</figureCitation>
|
||
e).
|
||
</paragraph>
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FCF8FE0794C2" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FCF8FE5E94AF" bold="true" box="[199,315,871,896]" pageId="10" pageNumber="241">Variation.</emphasis>
|
||
The variation exhibited by this species was difficult to assess due to the limited number of specimens available for examination. One specimen appeared to be teneral and was relatively pale in overall coloration. All other specimens were otherwise nearly identical, including the
|
||
<typeStatus id="54CBA296FFB8FFFB7138FC2FFB5F94E7" box="[1031,1082,944,968]" pageId="10" pageNumber="241">type</typeStatus>
|
||
, with some very slightly darker or lighter in color.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFB8FFFB75F8FC68FE199377" pageId="10" pageNumber="241" type="etymology">
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FC68FE199377" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FC68FE2B933F" bold="true" box="[199,334,1015,1040]" pageId="10" pageNumber="241">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="4C7067B7FFB8FFFB7704FC68FDC8933F" box="[571,685,1015,1040]" class="Insecta" family="Noteridae" genus="Liocanthydrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="241" phylum="Arthropoda" rank="species" species="armulatus">
|
||
<emphasis id="B904C026FFB8FFFB7704FC68FDC8933F" box="[571,685,1015,1040]" italics="true" pageId="10" pageNumber="241">armulatus</emphasis>
|
||
</taxonomicName>
|
||
is a combination of the Latin words
|
||
<emphasis id="B904C026FFB8FFFB7171FC66FBF1933F" box="[1102,1172,1017,1040]" italics="true" pageId="10" pageNumber="241">armus</emphasis>
|
||
, meaning shoulder and
|
||
<emphasis id="B904C026FFB8FFFB75A8FB83FFA8931A" box="[151,205,1052,1077]" italics="true" pageId="10" pageNumber="241">latus</emphasis>
|
||
, meaning broad, in reference to the species’ broad humeral region. It is treated as an adjective in the nominative singular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFB8FFFB75F8FBFBFA3D9352" box="[199,1368,1124,1149]" pageId="10" pageNumber="241" type="biology_ecology">
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FBFBFA3D9352" blockId="10.[151,1437,151,1220]" box="[199,1368,1124,1149]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FBFBFE429352" bold="true" box="[199,295,1124,1149]" pageId="10" pageNumber="241">Biology.</emphasis>
|
||
All five specimens of this species were taken from a single collection event at a stream margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A4FBFFFB8FFFB75F8FB18FFA693EB" pageId="10" pageNumber="241" type="distribution">
|
||
<paragraph id="8BCF1C34FFB8FFFB75F8FB18FFA693EB" blockId="10.[151,1437,151,1220]" pageId="10" pageNumber="241">
|
||
<emphasis id="B904C026FFB8FFFB75F8FB18FE05938F" bold="true" box="[199,352,1159,1184]" pageId="10" pageNumber="241">Distribution.</emphasis>
|
||
Known from the
|
||
<typeStatus id="54CBA296FFB8FFFB7714FB16FD3E938E" box="[555,603,1161,1185]" pageId="10" pageNumber="241">type</typeStatus>
|
||
locality in the extreme southeast region of Bolivar State,
|
||
<collectingCountry id="F3675CA4FFB8FFFB71D4FB18FA3B938F" box="[1259,1374,1159,1184]" name="Venezuela" pageId="10" pageNumber="241">Venezuela</collectingCountry>
|
||
(
|
||
<figureCitation id="134B00B1FFB8FFFB7050FB18FFD293EB" captionStart="FIGURE 23" captionStartId="7.[151,250,1491,1513]" captionText="FIGURE 23. Distribution map of Liocanthydrus species." pageId="10" pageNumber="241">Fig. 23</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |