141 lines
15 KiB
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141 lines
15 KiB
XML
<document id="FA2B53BBD97A0B29C07C71CB1C83D973" ID-CLB-Dataset="6630" ID-DOI="10.5281/zenodo.4648829" ID-GBIF-Dataset="ef7c103d-43ea-4929-8ceb-401a3053d901" ID-ISSN="1536-9307" ID-Zenodo-Dep="4648829" ID-ZooBank="634A674E-E08C-42DB-96E0-F6E5416D71CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1617136517793" checkinUser="carolina" docAuthor="Kovařík, František, Lowe, Graeme & Elmi, Hassan Sh Abdirahman" docDate="2020" docId="03C087E9450BCD3AFFE9FBC1FAEFFB43" docLanguage="en" docName="Euscorpius.322.1-23.pdf" docOrigin="Euscorpius 322" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Pandinurus intermedius" docType="treatment" docVersion="6" lastPageNumber="18" masterDocId="FFF9FF914518CD29FF98FF89FF99FFA2" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXV. Description of Pandinurus awalei sp. n. and the male of Pandiborellius somalilandus (Kovařík, 2012), with remarks on recent synonymies (Scorpionidae: Pandininae)" masterLastPageNumber="21" masterPageNumber="1" pageNumber="18" updateTime="1698935510268" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="C00CD9CB70766E71B781303925406CBB">
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<mods:title id="7E3A030C04A1E65129530407E0F67625">Scorpions of the Horn of Africa (Arachnida: Scorpiones). Part XXV. Description of Pandinurus awalei sp. n. and the male of Pandiborellius somalilandus (Kovařík, 2012), with remarks on recent synonymies (Scorpionidae: Pandininae)</mods:title>
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<mods:name id="74B150CFD7C64212557DE872196DACCF" type="personal">
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<mods:roleTerm id="C604D82C4BD6F29BA69FCEE3F8E20086">Author</mods:roleTerm>
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<mods:namePart id="2D81DA3D01821EB7FFD14AF0672E4528">Kovařík, František</mods:namePart>
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<mods:namePart id="72628167A0EE9E8364E2E20A0F85F7E4">Lowe, Graeme</mods:namePart>
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<mods:name id="0B32896D6C1EF63BC399E385A1C53D5A" type="personal">
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<mods:roleTerm id="6EEE9D6F90241E36126BC6C8DF83FBE1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B1EAAC761758E2FE5C462EA03C49A15B">Elmi, Hassan Sh Abdirahman</mods:namePart>
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<mods:typeOfResource id="3FA9D61CED132A504E87440339E6819A">text</mods:typeOfResource>
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<mods:title id="325B8306B84F2BE0209C2BACEF5A2069">Euscorpius</mods:title>
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<mods:part id="ADA784F6C541073ABF39BBA746917A96">
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<mods:date id="B72EE32F8BA0D1CE7AFD322F9BDA4637">2020</mods:date>
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<mods:number id="98FCAB627250132073C7E2FBE99D22B7">322</mods:number>
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<treatment id="03C087E9450BCD3AFFE9FBC1FAEFFB43" ID-DOI="http://doi.org/10.5281/zenodo.4773468" ID-GBIF-Taxon="182237302" ID-Zenodo-Dep="4773468" LSID="urn:lsid:plazi:treatment:03C087E9450BCD3AFFE9FBC1FAEFFB43" httpUri="http://treatment.plazi.org/id/03C087E9450BCD3AFFE9FBC1FAEFFB43" lastPageNumber="18" pageId="19" pageNumber="18">
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<subSubSection id="C3736574450BCD3AFFE9FBC1FEBBFB27" pageId="19" pageNumber="18" type="nomenclature">
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<paragraph id="8BD636FF450BCD3AFFE9FBC1FEBBFB27" blockId="19.[113,691,1095,1158]" pageId="19" pageNumber="18">
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<emphasis id="B91DEAED450BCD3AFFE9FBC1FE1CFBC0" bold="true" box="[113,389,1096,1122]" pageId="19" pageNumber="18">Taxonomic position of</emphasis>
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<taxonomicName id="4C694D7C450BCD3AFE15FBCEFEBBFB27" ID-CoL="4CBTF" authority="(Borelli, 1919)" baseAuthorityName="Borelli" baseAuthorityYear="1919" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B91DEAED450BCD3AFE15FBCEFF56FB27" bold="true" pageId="19" pageNumber="18">
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<emphasis id="B91DEAED450BCD3AFE15FBCEFD2AFBC0" bold="true" box="[397,691,1095,1122]" italics="true" pageId="19" pageNumber="18">Pandinurus intermedius</emphasis>
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(Borelli
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</emphasis>
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,
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<emphasis id="B91DEAED450BCD3AFF46FBE2FEBBFB27" bold="true" box="[222,290,1131,1157]" pageId="19" pageNumber="18">1919)</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3736574450BCD3AFFE9FB24FAEFFB43" pageId="19" pageNumber="18" type="discussion">
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<paragraph id="8BD636FF450BCD3AFFE9FB24FAEFFB43" blockId="19.[113,772,1197,2053]" lastBlockId="19.[816,1474,1096,1249]" pageId="19" pageNumber="18">
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Prendini & Loria (2020: 442) formally synonymized
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<taxonomicName id="4C694D7C450BCD3AFFE9FB44FE6EFB47" authority="Borelli, 1919" authorityName="Borelli" authorityYear="1919" box="[113,503,1229,1253]" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B91DEAED450BCD3AFFE9FB44FEC7FB47" box="[113,350,1229,1253]" italics="true" pageId="19" pageNumber="18">Pandinus intermedius</emphasis>
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Borelli, 1919
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</taxonomicName>
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under
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<taxonomicName id="4C694D7C450BCD3AFDD0FB44FE9FFAA7" authority="Borelli, 1919" authorityName="Borelli" authorityYear="1919" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="citernii">
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<emphasis id="B91DEAED450BCD3AFDD0FB44FC9AFB47" box="[584,771,1229,1253]" italics="true" pageId="19" pageNumber="18">Pandinus citernii</emphasis>
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Borelli, 1919
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</taxonomicName>
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(both now in the genus
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<taxonomicName id="4C694D7C450BCD3AFDAEFB64FD2BFAA7" authorityName="Fet" authorityYear="1997" box="[566,690,1261,1285]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
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<emphasis id="B91DEAED450BCD3AFDAEFB64FD2BFAA7" box="[566,690,1261,1285]" italics="true" pageId="19" pageNumber="18">Pandinurus</emphasis>
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</taxonomicName>
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), “
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<emphasis id="B91DEAED450BCD3AFD45FB67FECFFA87" italics="true" pageId="19" pageNumber="18">two Ethiopian species of</emphasis>
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<taxonomicName id="4C694D7C450BCD3AFEFAFA84FE43FA87" authorityName="Fet" authorityYear="1997" box="[354,474,1293,1317]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">Pandinurus</taxonomicName>
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,
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<emphasis id="B91DEAED450BCD3AFE75FA84FF63FAE7" italics="true" pageId="19" pageNumber="18">
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described from the same
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<typeStatus id="54D2885D450BCD3AFFE9FAA7FF04FAE7" box="[113,157,1326,1349]" pageId="19" pageNumber="18">type</typeStatus>
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locality
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</emphasis>
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”, without explanation of their taxonomic act. In their paragraph, the authors cited only “
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<emphasis id="B91DEAED450BCD3AFD19FAC4FF39FA27" italics="true" pageId="19" pageNumber="18">unpublished data</emphasis>
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”. They did not address previously proposed diagnostic characters, in particular chelal trichobothrium
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<emphasis id="B91DEAED450BCD3AFDE1FA07FD0BFA07" box="[633,658,1422,1445]" italics="true" pageId="19" pageNumber="18">ea</emphasis>
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, which is present in
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<taxonomicName id="4C694D7C450BCD3AFF78FA27FEE1FA67" authorityName="Borelli" authorityYear="1919" box="[224,376,1453,1477]" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B91DEAED450BCD3AFF78FA27FEE1FA67" box="[224,376,1453,1477]" italics="true" pageId="19" pageNumber="18">P. intermedius</emphasis>
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</taxonomicName>
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and absent in
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<taxonomicName id="4C694D7C450BCD3AFD89FA27FDE1FA67" authorityName="Borelli" authorityYear="1919" box="[529,632,1454,1477]" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="citernii">
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<emphasis id="B91DEAED450BCD3AFD89FA27FDE1FA67" box="[529,632,1454,1477]" italics="true" pageId="19" pageNumber="18">P. citernii</emphasis>
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</taxonomicName>
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, and ignored other morphological characters published in KovařÍk et al. (2017c). They characterized these as “Ethiopian” species, although the originally cited
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<typeStatus id="54D2885D450BCD3AFE56F984FE65F987" box="[462,508,1549,1573]" pageId="19" pageNumber="18">type</typeStatus>
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localities are actually located in
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<collectingCountry id="F37E766F450BCD3AFF7BF9A4FEA7F9E7" box="[227,318,1581,1605]" name="Somalia" pageId="19" pageNumber="18">Somalia</collectingCountry>
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. In fact, two separate areas of distribution in
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<collectingCountry id="F37E766F450BCD3AFF16F9C4FF71F9C7" box="[142,232,1613,1637]" name="Ethiopia" pageId="19" pageNumber="18">Ethiopia</collectingCountry>
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were indeed confirmed by KovařÍk et al. (2017c). However, the co-occurrence of both species at “the same
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<typeStatus id="54D2885D450BCD3AFD4DF9E4FC9AF927" box="[725,771,1645,1669]" pageId="19" pageNumber="18">type</typeStatus>
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locality” in
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<collectingCountry id="F37E766F450BCD3AFF6DF904FED5F907" box="[245,332,1677,1701]" name="Somalia" pageId="19" pageNumber="18">Somalia</collectingCountry>
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had not been confirmed. Borelli (1919) studied specimens collected by Carlo Citerni, and there is a significant possibility that Citerni did not personally collect these specimens and assign locality labels himself. It has been repeatedly documented that old specimens in Italian museums are labelled with imprecise localities, and their true geographic distributions need to be independently confirmed (see also KovařÍk et al, 2019: 61). The confirmed distributions of both the species in
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<collectingCountry id="F37E766F450BCD3AFEFDF804FE5BF807" box="[357,450,1933,1957]" name="Ethiopia" pageId="19" pageNumber="18">Ethiopia</collectingCountry>
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, as well as their unconfirmed
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<typeStatus id="54D2885D450BCD3AFFE9F824FF06F867" box="[113,159,1965,1989]" pageId="19" pageNumber="18">type</typeStatus>
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locality in
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<collectingCountry id="F37E766F450BCD3AFE86F824FEE1F867" box="[286,376,1965,1989]" name="Somalia" pageId="19" pageNumber="18">Somalia</collectingCountry>
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, are mapped in
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<figureCitation id="13522A7A450BCD3AFDB1F824FDE3F867" box="[553,634,1965,1989]" captionStart="Figure 83" captionStartId="20.[113,181,2018,2040]" captionTargetBox="[113,1474,944,2005]" captionTargetId="figure-31@20.[113,1474,941,2005]" captionTargetPageId="20" captionText="Figure 83. Map showing confirmed distribution of Pandinurus spp." figureDoi="http://doi.org/10.5281/zenodo.4648868" httpUri="https://zenodo.org/record/4648868/files/figure.png" pageId="19" pageNumber="18">Fig. 83</figureCitation>
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. In addition, recent DNA and chromosomal analysis further supports the view that both are separate and valid species. For example,
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<taxonomicName id="4C694D7C450BCD3AFCA8FBC0FC56FBC2" authorityName="Borelli" authorityYear="1919" box="[816,975,1096,1120]" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B91DEAED450BCD3AFCA8FBC0FC56FBC2" box="[816,975,1096,1120]" italics="true" pageId="19" pageNumber="18">P. intermedius</emphasis>
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</taxonomicName>
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has a constant number of chromosomes of 2n=110, whereas
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<taxonomicName id="4C694D7C450BCD3AFC65FBE0FBF4FB22" authorityName="Borelli" authorityYear="1919" box="[1021,1133,1129,1152]" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="citernii">
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<emphasis id="B91DEAED450BCD3AFC65FBE0FBF4FB22" box="[1021,1133,1129,1152]" italics="true" pageId="19" pageNumber="18">P. citernii</emphasis>
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</taxonomicName>
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has a different and constant number of chromosomes of 2n=120 (
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<bibRefCitation id="EFF84B0E450BCD3AFB49FB00FA13FB03" author="STUNDLOVA" box="[1233,1418,1161,1187]" pageId="19" pageNumber="18" refId="ref6739" refString="STUNDLOVA, J. 2019. Mechanisms of the karyotype evolution in scorpions (Arachnida: Scorpiones). Ph. D. Dissertation, Department of Zoology, Charles University of Prague, Czech Republic." type="book" year="2019">ŠtundlovÁ, 2019</bibRefCitation>
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). We therefore reinstate
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<taxonomicName id="4C694D7C450BCD3AFC63FB21FA3DFB63" authority="(Borelli, 1919)" baseAuthorityName="Borelli" baseAuthorityYear="1919" box="[1019,1444,1192,1217]" class="Arachnida" family="Scorpionidae" genus="Pandinurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="intermedius">
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<emphasis id="B91DEAED450BCD3AFC63FB21FB64FB62" box="[1019,1277,1192,1216]" italics="true" pageId="19" pageNumber="18">Pandinurus intermedius</emphasis>
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(Borelli, 1919)
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</taxonomicName>
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as a valid species distinct from
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<taxonomicName id="4C694D7C450BCD3AFBFAFB40FAEBFB43" authority="(Borelli, 1919)" baseAuthorityName="Borelli" baseAuthorityYear="1919" box="[1122,1394,1224,1249]" class="Arachnida" family="Scorpionidae" genus="Pandinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="citernii">
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<emphasis id="B91DEAED450BCD3AFBFAFB40FB50FB42" box="[1122,1225,1225,1248]" italics="true" pageId="19" pageNumber="18">P. citernii</emphasis>
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(Borelli, 1919)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |