167 lines
18 KiB
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167 lines
18 KiB
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<document id="6546D1C8CE8B3BB3A37DA63715E26E73" ID-CLB-Dataset="6398" ID-DOI="10.5252/g2014n3a1" ID-GBIF-Dataset="d46af7eb-bf62-4022-8dbe-5c6f8c918820" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538455" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150288044" checkinUser="felipe" docAuthor="Newman, Michael J., Burrow, Carole J., Den Blaauwen, Jan L. & Davidson, Robert G." docDate="2014" docId="03AE87E08022C11B2098FCEC38E2FED9" docLanguage="en" docName="Geodiversitas.36.3.321-348.pdf" docOrigin="Geodiversitas 36 (3)" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Euthacanthidae Berg 1940" docType="treatment" docVersion="7" lastPageNumber="325" masterDocId="FF97FF988026C11F2028FFFE3B56FFA9" masterDocTitle="The Early Devonian acanthodian Euthacanthus macnicoli Powrie, 1864 from the Midland Valley of Scotland" masterLastPageNumber="348" masterPageNumber="321" pageNumber="325" updateTime="1698929422071" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="A2B08A1EFF687BD904886FEA42DFCDDB">
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<mods:title id="9BE968F39DA4BAD0E1010CE18743136E">The Early Devonian acanthodian Euthacanthus macnicoli Powrie, 1864 from the Midland Valley of Scotland</mods:title>
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<mods:namePart id="173057966FCE34158C961D3405510ACC">Newman, Michael J.</mods:namePart>
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<mods:namePart id="798B839A1232D0B31A6B42C9F85D2890">Burrow, Carole J.</mods:namePart>
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<mods:namePart id="449AEB5E04CBC2F87ADB33F0A3329BF4">Den Blaauwen, Jan L.</mods:namePart>
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<mods:namePart id="0C822CB7213ECF16AE3EBBFE1711158A">Davidson, Robert G.</mods:namePart>
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<mods:typeOfResource id="DD61FD52F37891C96ABD65E710A4130F">text</mods:typeOfResource>
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<mods:title id="A0382873053B6F70E7CED4F6A9A59EB7">Geodiversitas</mods:title>
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<mods:date id="4D1445C42F3E7A5A64FC25F802A9AA23">2014</mods:date>
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<mods:number id="1ECF8EDDB1D089ED6D913A29F003121C">2014-09-26</mods:number>
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<mods:number id="5B4E9219F7928ADB701A4DAE435DBE75">36</mods:number>
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<treatment id="03AE87E08022C11B2098FCEC38E2FED9" ID-DOI="http://doi.org/10.5281/zenodo.4538779" ID-GBIF-Taxon="182401039" ID-Zenodo-Dep="4538779" LSID="urn:lsid:plazi:treatment:03AE87E08022C11B2098FCEC38E2FED9" httpUri="http://treatment.plazi.org/id/03AE87E08022C11B2098FCEC38E2FED9" lastPageNumber="325" pageId="4" pageNumber="325">
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<subSubSection id="C31D657D8022C11B2098FCEC391EFC85" box="[176,584,786,813]" pageId="4" pageNumber="325" type="nomenclature">
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<paragraph id="8BB836F68022C11B2098FCEC391EFC85" blockId="4.[176,584,786,813]" box="[176,584,786,813]" pageId="4" pageNumber="325">
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<heading id="D0F0819A8022C11B2098FCEC391EFC85" box="[176,584,786,813]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="325" reason="3">
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Family
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<taxonomicName id="4C074D758022C11B212AFCED391EFC85" authority="Berg, 1940" authorityName="Berg" authorityYear="1940" box="[258,584,786,813]" class="Acanthodii" family="Euthacanthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="325" phylum="Chordata" rank="family">
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EUTHACANTHIDAE
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<bibRefCitation id="EF964B078022C11B21F8FCED391EFC85" author="BERG L. S." box="[464,584,786,813]" pageId="4" pageNumber="325" pagination="85 - 517" refId="ref13131" refString="BERG L. S. 1940. - Classification of fishes both recent and fossil. Transactions of the Zoological Institute, Leningrad, USSR 5: 85 - 517. [85 - 345 in Russian]" type="journal article" year="1940">Berg, 1940</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D657D8022C11B2059FCB239D1FCCA" box="[113,647,843,867]" pageId="4" pageNumber="325" type="materials_examined">
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<paragraph id="8BB836F68022C11B2059FCB239D1FCCA" blockId="4.[113,647,843,867]" box="[113,647,843,867]" pageId="4" pageNumber="325">
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<collectionCode id="ED16AE338022C11B2059FCB23A19FCCB" box="[113,335,844,867]" pageId="4" pageNumber="325">
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<typeStatus id="54BC88548022C11B2059FCB23BF4FCC8" box="[113,162,844,867]" pageId="4" pageNumber="325">TYPE</typeStatus>
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AND ONLY GENUS
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</collectionCode>
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. —
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<taxonomicName id="4C074D758022C11B2151FCB539D5FCCA" authority="Powrie, 1864" authorityName="Powrie" authorityYear="1864" box="[377,643,843,867]" family="Euthacanthidae" genus="Euthacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B2151FCB53AAFFCCA" box="[377,505,843,867]" italics="true" pageId="4" pageNumber="325">Euthacanthus</emphasis>
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<bibRefCitation id="EF964B078022C11B21D7FCB539D5FCCA" author="POWRIE J." box="[511,643,843,867]" pageId="4" pageNumber="325" pagination="413 - 429" refId="ref14777" refString="POWRIE J. 1864. - On the fossiliferous rocks of Forfarshire and their contents. Quarterly Journal of the Geological Society of London 47: 413 - 429." type="journal article" year="1864">Powrie, 1864</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D657D8022C11B2059FC853A32FB1C" pageId="4" pageNumber="325" type="diagnosis">
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<paragraph id="8BB836F68022C11B2059FC853A32FB1C" blockId="4.[112,648,891,1206]" pageId="4" pageNumber="325">DIAGNOSIS. — Acanthodians lacking an oral dentition; ring of five sclerotic bones; closeset branchiostegal plates over part of the branchial region; dermal pectoral girdle comprises a single pair of triangular postbranchial plates, each with one spine, that are not contiguous with the scapulocoracoid; simple scapulocoracoid with a columnar shaft; scapulocoracoid is covered laterally only by normal body scales, not dermal tesserae; anterior dorsal, posterior dorsal, anal, pectoral and pelvic fin spines are robust with thick walls and a wide pulp canal, and longitudinal ornament ridges; four to six pairs of prepelvic spines; polygonal tectal tesserae.</paragraph>
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</subSubSection>
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<subSubSection id="C31D657D8022C11B2059FB3038E2FED9" pageId="4" pageNumber="325" type="discussion">
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<paragraph id="8BB836F68022C11B2059FB303B83FB4F" blockId="4.[113,649,1230,1700]" box="[113,213,1230,1256]" pageId="4" pageNumber="325">REMARKS</paragraph>
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<paragraph id="8BB836F68022C11B2059FB1338E2FED9" blockId="4.[113,649,1230,1700]" lastBlockId="4.[677,1211,216,369]" pageId="4" pageNumber="325">
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Cladistic analyses (e.g.,
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<bibRefCitation id="EF964B078022C11B2153FB1339D7FAAE" author="HANKE G. F. & WILSON M. V. H." box="[379,641,1261,1287]" pageId="4" pageNumber="325" pagination="189 - 216" refId="ref13931" refString="HANKE G. F. & WILSON M. V. H. 2004. - New teleostome fishes and acanthodian systematic, in AR- RATIA G., WILSON M. V. H. & CLOUTIER R. (eds), Recent Advances in the Origin and Early Radiation of Vertebrates. Verlag Dr Friedrich Pfeil, Munchen: 189 - 216." type="book chapter" year="2004">Hanke & Wilson 2004</bibRefCitation>
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;
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<bibRefCitation id="EF964B078022C11B205BFAF33A43FA8E" author="BRAZEAU M. D." box="[115,277,1293,1319]" pageId="4" pageNumber="325" pagination="305 - 308" refId="ref13169" refString="BRAZEAU M. D. 2009. - The braincase and jaws of a Devonian " acanthodian " and modern gnathostome origins. Nature 457: 305 - 308." type="journal article" year="2009">Brazeau 2009</bibRefCitation>
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;
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<bibRefCitation id="EF964B078022C11B210BFAF33961FA8E" author="BURROW C. J. & TURNER S." box="[291,567,1293,1319]" pageId="4" pageNumber="325" pagination="123 - 144" refId="ref13272" refString="BURROW C. J. & TURNER S. 2010. - Reassessment of " Protodus " scoticus from the Early Devonian of Scotland, in ELLIOTT D. K., MAISEY J. G., YU X & MIAO D. (eds), Morphology, Phylogeny and Paleobiogeography of Fossil Fishes. Verlag Dr. Friedrich Pfeil, Munchen: 123 - 144." type="book chapter" year="2010">Burrow & Turner 2010</bibRefCitation>
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;
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<bibRefCitation id="EF964B078022C11B226EFAF33BBEFAEE" author="DAVIS S. P. & FINARELLI J. A. & COATES M. I." pageId="4" pageNumber="325" pagination="247 - 250" refId="ref13534" refString="DAVIS S. P., FINARELLI J. A. & COATES M. I. 2012. - Acanthodes and shark-like conditions in the last common ancestor of modern gnathostomes. Nature 486: 247 - 250." type="journal article" year="2012">
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Davis
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<emphasis id="B973EAE48022C11B205CFAD33BF0FAEE" box="[116,166,1325,1351]" italics="true" pageId="4" pageNumber="325">et al.</emphasis>
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2012
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</bibRefCitation>
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;
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<bibRefCitation id="EF964B078022C11B20DBFAD33ACBFAEE" author="ZHU M. & YU X. & AHLBERG P. E. & CHOO B. & LU J. & QIAO T. & QU Q. & ZHAO W. & JIA L. & BLOM H. & ZHU Y." box="[243,413,1325,1351]" pageId="4" pageNumber="325" pagination="188 - 193" refId="ref15427" refString="ZHU M., YU X., AHLBERG P. E., CHOO B., LU J., QIAO T., QU Q., ZHAO W., JIA L., BLOM H. & ZHU Y. 2013. - A Silurian placoderm with osteichthyan-like marginal jaw bones. Nature 502: 188 - 193." type="journal article" year="2013">
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Zhu
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<emphasis id="B973EAE48022C11B2101FAD33A0BFAEE" box="[297,349,1325,1351]" italics="true" pageId="4" pageNumber="325">et al.</emphasis>
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2013
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</bibRefCitation>
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) have shown that the order
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<taxonomicName id="4C074D758022C11B2090FAB2393DFACF" authority="sensu Denison (1979)" authorityName="sensu Denison" authorityYear="1979" box="[184,619,1356,1383]" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="325" phylum="Chordata" rank="order">
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Climatiiformes sensu
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<bibRefCitation id="EF964B078022C11B219CFAB2393DFACF" author="DENISON R." box="[436,619,1356,1382]" pageId="4" pageNumber="325" refId="ref13595" refString="DENISON R. 1979. - Acanthodii, in SCHULTZE H. - P. (ed.), Handbook of palaeoichthyology, 5. Gustav Fischer Verlag, Stuttgart, 62 p." type="book" year="1979">Denison (1979)</bibRefCitation>
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</taxonomicName>
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is paraphyletic, and
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<taxonomicName id="4C074D758022C11B2110FA923A93FA2F" authorityName="Powrie" authorityYear="1864" box="[312,453,1388,1414]" family="Euthacanthidae" genus="Euthacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B2110FA923A93FA2F" box="[312,453,1388,1414]" italics="true" pageId="4" pageNumber="325">Euthacanthus</emphasis>
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</taxonomicName>
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does not fall into other recognized orders.
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<bibRefCitation id="EF964B078022C11B21A2FA723944FA0F" author="BERG L. S." box="[394,530,1420,1446]" pageId="4" pageNumber="325" pagination="85 - 517" refId="ref13131" refString="BERG L. S. 1940. - Classification of fishes both recent and fossil. Transactions of the Zoological Institute, Leningrad, USSR 5: 85 - 517. [85 - 345 in Russian]" type="journal article" year="1940">Berg (1940)</bibRefCitation>
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raised the family
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<taxonomicName id="4C074D758022C11B2093FA553A36FA6C" authorityName="Berg" authorityYear="1940" box="[187,352,1451,1477]" class="Acanthodii" family="Euthacanthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="325" phylum="Chordata" rank="family">Euthacanthidae</taxonomicName>
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to incorporate
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<taxonomicName id="4C074D758022C11B21D6FA5539DFFA6C" authorityName="Powrie" authorityYear="1864" box="[510,649,1451,1477]" family="Euthacanthidae" genus="Euthacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B21D6FA5539DFFA6C" box="[510,649,1451,1477]" italics="true" pageId="4" pageNumber="325">Euthacanthus</emphasis>
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</taxonomicName>
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and possibly
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<taxonomicName id="4C074D758022C11B212BFA353906FA4C" authority="Egerton, 1860" authorityName="Egerton" authorityYear="1860" box="[259,592,1483,1510]" family="Climatiidae" genus="Brachyacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B212BFA353AF0FA4C" box="[259,422,1483,1509]" italics="true" pageId="4" pageNumber="325">Brachyacanthus</emphasis>
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Egerton, 1860
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</taxonomicName>
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, and
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<bibRefCitation id="EF964B078022C11B205BFA153BADF9AF" author="MILES R. S." box="[115,251,1515,1542]" pageId="4" pageNumber="325" pagination="147 - 194" refId="ref14361" refString="MILES R. S. 1966. - The acanthodian fishes of the Devonian Plattenkalk of the Paffrath trough in the Rhineland. Arkiv fur Zoologi 18: 147 - 194." type="journal article" year="1966">Miles (1966)</bibRefCitation>
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also considered that
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<taxonomicName id="4C074D758022C11B21F8FA15390CF9AC" authorityName="Powrie" authorityYear="1864" box="[464,602,1515,1541]" family="Euthacanthidae" genus="Euthacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B21F8FA15390CF9AC" box="[464,602,1515,1541]" italics="true" pageId="4" pageNumber="325">Euthacanthus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C074D758022C11B2059F9F43A44F98D" authorityName="Egerton" authorityYear="1860" box="[113,274,1546,1572]" family="Climatiidae" genus="Brachyacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B2059F9F43A44F98D" box="[113,274,1546,1572]" italics="true" pageId="4" pageNumber="325">Brachyacanthus</emphasis>
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</taxonomicName>
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were members of the same family as both forms have no teeth and a principle gill cover that did not cover the whole gill chamber.
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<bibRefCitation id="EF964B078022C11B205BF9943A3DF92D" author="NEWMAN M. J. & DAVIDSON R. G. & DEN BLAAUWEN J. L. & BURROW C. J." box="[115,363,1641,1668]" pageId="4" pageNumber="325" pagination="101 - 111" refId="ref14554" refString="NEWMAN M. J., DAVIDSON R. G., DEN BLAAUWEN J. L. & BURROW C. J. 2011. - The Early Devonian acanthodian Euthacanthus gracilis from the Midland Valley of Scotland. Scottish Journal of Geology 47: 101 - 111." type="journal article" year="2011">
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Newman
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<emphasis id="B973EAE48022C11B20C8F9943A41F92A" box="[224,279,1641,1667]" italics="true" pageId="4" pageNumber="325">et al.</emphasis>
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(2011)
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</bibRefCitation>
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excluded
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<taxonomicName id="4C074D758022C11B21C8F99739D0F92A" authorityName="Egerton" authorityYear="1860" box="[480,646,1641,1667]" family="Climatiidae" genus="Brachyacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B21C8F99739D0F92A" box="[480,646,1641,1667]" italics="true" pageId="4" pageNumber="325">Brachyacanthus</emphasis>
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</taxonomicName>
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from the
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<taxonomicName id="4C074D758022C11B20F4F9773AD1F90A" authorityName="Berg" authorityYear="1940" box="[220,391,1673,1699]" class="Acanthodii" family="Euthacanthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="325" phylum="Chordata" rank="family">Euthacanthidae</taxonomicName>
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based on the fin spine ridge ornamentation, and as none of the recent cladistic analyses support a sister relationship between
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<taxonomicName id="4C074D758022C11B22C7FEE93829FE98" authorityName="Powrie" authorityYear="1864" box="[751,895,279,305]" family="Euthacanthidae" genus="Euthacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B22C7FEE93829FE98" box="[751,895,279,305]" italics="true" pageId="4" pageNumber="325">Euthacanthus</emphasis>
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</taxonomicName>
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|
and
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<taxonomicName id="4C074D758022C11B2393FEE93F36FE98" authorityName="Egerton" authorityYear="1860" box="[955,1120,279,305]" family="Climatiidae" genus="Brachyacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="325" phylum="Chordata" rank="genus">
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<emphasis id="B973EAE48022C11B2393FEE93F36FE98" box="[955,1120,279,305]" italics="true" pageId="4" pageNumber="325">Brachyacanthus</emphasis>
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|
</taxonomicName>
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|
(or any other genera) we consider the
|
|
<taxonomicName id="4C074D758022C11B23DCFEC93FCBFEF8" authorityName="Berg" authorityYear="1940" box="[1012,1181,311,337]" class="Acanthodii" family="Euthacanthidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Climatiiformes" pageId="4" pageNumber="325" phylum="Chordata" rank="family">Euthacanthidae</taxonomicName>
|
|
to be a monogeneric family.
|
|
</paragraph>
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|
</subSubSection>
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|
</treatment>
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|
</document> |