247 lines
29 KiB
XML
247 lines
29 KiB
XML
<document id="B621244A0ADBB371A97650DC28AF661E" ID-DOI="10.11646/zootaxa.5424.4.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821574" ID-ZooBank="C8692FDA-9AB4-4C5C-B235-E131BBC30A7A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710501336329" checkinUser="plazi" docAuthor="Sendi, Hemen" docDate="2024" docId="03AE87CDFFD85A1DF2ABFE35FBB3764C" docLanguage="en" docName="zootaxa.5424.4.6.pdf" docOrigin="Zootaxa 5424 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nigropterix cummingi Sendi 2024, sp. n." docType="treatment" docUuid="B0D98387-7252-4293-A972-C991AD673082" docUuidSource="ZooBank" docVersion="2" lastPageNumber="480" masterDocId="FF97FFB5FFDA5A19F23CFFC1FF8E770D" masterDocTitle="Nigropterix cummingi sp. n., an umenocoleid (Insecta: Dictyoptera) from mid-Cretaceous northern Myanmar amber" masterLastPageNumber="482" masterPageNumber="476" pageNumber="478" updateTime="1710502004080" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="81FA21C6283ACFE4531CF82762E46E82">Nigropterix cummingi sp. n., an umenocoleid (Insecta: Dictyoptera) from mid-Cretaceous northern Myanmar amber</mods:title>
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<mods:namePart id="F97F986EC74099735207CAED73A93FBB">Sendi, Hemen</mods:namePart>
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<mods:typeOfResource id="BD3A745709A1E9CEB812D918EA1189F7">text</mods:typeOfResource>
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<mods:title id="218646751F286D28589E18F2514E8453">Zootaxa</mods:title>
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<mods:part id="2544C68908236BBB2C6D1AB9FBCC0EED">
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<mods:date id="278225973B573BBFFE452F98D11D58F9">2024</mods:date>
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<mods:detail id="10D137127187609EFBB76D97F89BE4BB" type="pubDate">
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<mods:number id="CCCEA3A11A5309AC457B14ADFCFB5954">2024-03-15</mods:number>
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<mods:number id="5B6901874D83930CFD0E910949D18105">5424</mods:number>
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<mods:url id="F8636DE7099F60E65729B8F5E85F6585">http://dx.doi.org/10.11646/zootaxa.5424.4.6</mods:url>
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<mods:classification id="E635B6C29366E6BF856B59F6DB8562FE">journal article</mods:classification>
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<mods:identifier id="16DB029128AE86E92DE9E952A715636F" type="DOI">10.11646/zootaxa.5424.4.6</mods:identifier>
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<mods:identifier id="9A2FF925BBF792788380F70D1D8ED158" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03AE87CDFFD85A1DF2ABFE35FBB3764C" LSID="urn:lsid:zoobank.org:act:B0D98387-7252-4293-A972-C991AD673082" httpUri="http://treatment.plazi.org/id/03AE87CDFFD85A1DF2ABFE35FBB3764C" lastPageId="4" lastPageNumber="480" pageId="2" pageNumber="478">
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<subSubSection id="C31D6550FFD85A1BF2ABFE35FBFF753E" pageId="2" pageNumber="478" type="nomenclature">
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<paragraph id="8BB836DBFFD85A1BF2ABFE35FDC17503" blockId="2.[151,1137,500,563]" box="[151,591,500,527]" pageId="2" pageNumber="478">
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<heading id="D0F081B7FFD85A1BF2ABFE35FDC17503" bold="true" box="[151,591,500,527]" fontSize="11" level="1" pageId="2" pageNumber="478" reason="1">
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<emphasis id="B973EAC9FFD85A1BF2ABFE35FDC17503" bold="true" box="[151,591,500,527]" pageId="2" pageNumber="478">
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<taxonomicName id="4C074D58FFD85A1BF2ABFE35FE2B7503" authority="Sendi, 2024" authorityName="Sendi" authorityYear="2024" box="[151,421,500,526]" class="Insecta" family="Umenocoleidae" genus="Nigropterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="478" phylum="Arthropoda" rank="species" species="cummingi" status="sp. nov.">
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<emphasis id="B973EAC9FFD85A1BF2ABFE35FE2B7503" bold="true" box="[151,421,500,526]" italics="true" pageId="2" pageNumber="478">Nigropterix cummingi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24057B2FFD85A1BF390FE34FE607502" box="[428,494,501,527]" pageId="2" pageNumber="478" rank="species">sp. n.</taxonomicNameLabel>
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(
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<figureCitation id="133C2A5EFFD85A1BF3C2FE35FDCB7503" box="[510,581,500,526]" captionStart="FIGURE 1" captionStartId="3.[151,250,1870,1894]" captionTargetBox="[177,1410,181,1847]" captionTargetId="figure-31@3.[177,1410,181,1847]" captionTargetPageId="3" captionText="FIGURE 1 Nigropterix cummingi sp. n. from Burmese amber of Hukawng Valley, North Myanmar, holotype (SNM Z 40731). A: Illustration of dorsal view. B: Photograph of the dorsal view. C: Photograph of the ventral view. D: Right forewing of the specimen. E: Detail of the forewing. F: Detail of terminalia with characteristic chaetotaxy. G: Detail of the ovipositor. Scale bar is 1mm." figureDoi="http://doi.org/10.5281/zenodo.10821576" httpUri="https://zenodo.org/record/10821576/files/figure.png" pageId="2" pageNumber="478">Fig. 1</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB836DBFFD85A1BF2ABFDD8FBFF753E" blockId="2.[151,1137,500,563]" box="[151,1137,536,563]" pageId="2" pageNumber="478">
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<emphasis id="B973EAC9FFD85A1BF2ABFDD8FEC4753E" bold="true" box="[151,330,537,563]" pageId="2" pageNumber="478">ZooBank code.</emphasis>
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<uri id="FF963AD9FFD85A1BF36DFDD9FBFF753E" box="[337,1137,536,563]" pageId="2" pageNumber="478">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA10C0EFFD85A1BF05DFDD9FBFF753E" box="[609,1137,536,563]" pageId="2" pageNumber="478">B0D98387-7252-4293-A972-C991AD673082</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D6550FFD85A1BF2ABFDA1FA8F714B" pageId="2" pageNumber="478" type="description">
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<paragraph id="8BB836DBFFD85A1BF2ABFDA1FC117577" blockId="2.[151,1437,608,2039]" box="[151,927,608,635]" pageId="2" pageNumber="478">
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<emphasis id="B973EAC9FFD85A1BF2ABFDA1FEE97577" bold="true" box="[151,359,608,634]" pageId="2" pageNumber="478">Autapomorphies.</emphasis>
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Punctured ovipositor with setae, wide paranotalia.
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</paragraph>
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<paragraph id="8BB836DBFFD85A1BF2FBFD45FC287592" blockId="2.[151,1437,608,2039]" box="[199,934,644,671]" pageId="2" pageNumber="478">
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<emphasis id="B973EAC9FFD85A1BF2FBFD45FD477593" bold="true" box="[199,713,644,671]" pageId="2" pageNumber="478">
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||
Symplesiomorhies with standard
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<taxonomicName id="4C074D58FFD85A1BF06AFD45FD4D7593" box="[598,707,644,670]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattaria" pageId="2" pageNumber="478" phylum="Arthropoda" rank="order">Blattaria</taxonomicName>
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.
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</emphasis>
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Paranotalia present
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</paragraph>
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<paragraph id="8BB836DBFFD85A1BF2FBFD69FD777423" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
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<emphasis id="B973EAC9FFD85A1BF2FBFD69FDC275CF" bold="true" box="[199,588,680,707]" pageId="2" pageNumber="478">
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Description (based on
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<typeStatus id="54BC8879FFD85A1BF3E9FD69FDB175CF" box="[469,575,680,706]" pageId="2" pageNumber="478" type="holotype">holotype</typeStatus>
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).
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</emphasis>
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Adult female,
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<quantity id="4CFF9B3EFFD85A1BF0CBFD69FCD275CE" box="[759,860,680,707]" metricMagnitude="-3" metricUnit="m" metricValue="8.86" pageId="2" pageNumber="478" unit="mm" value="8.86">8.86 mm</quantity>
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from head to end of abdomen, and
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||
<quantity id="4CFF9B3EFFD85A1BF6CEFD68FAD975CE" box="[1266,1367,681,707]" metricMagnitude="-3" metricUnit="m" metricValue="3.21" pageId="2" pageNumber="478" unit="mm" value="3.21">3.21 mm</quantity>
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||
wide. Head transverse in dorsal view (1.78/
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||
<quantity id="4CFF9B3EFFD85A1BF072FD0DFD3B75EA" box="[590,693,716,743]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="478" unit="mm" value="0.52">0.52 mm</quantity>
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||
) with large bulging compound eyes (0.33/
|
||
<quantity id="4CFF9B3EFFD85A1BF69BFD0CFA9F75EA" box="[1191,1297,717,743]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="2" pageNumber="478" unit="mm" value="0.37">0.37 mm</quantity>
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||
), protruding from pronotum. Antenna with 34 visible segments (0.06/ 0.12–0.18/
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||
<quantity id="4CFF9B3EFFD85A1BF190FD30FB987406" box="[940,1046,753,779]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="478" unit="mm" value="0.04">0.04 mm</quantity>
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||
). Distal segments are thinner than proximal one. It is only about half as long as the body.
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||
</paragraph>
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||
<paragraph id="8BB836DBFFD85A1BF2FBFCF8FD15747B" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
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||
Pronotum (1.46/
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||
<quantity id="4CFF9B3EFFD85A1BF3BBFCF9FE62745E" box="[391,492,824,851]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="478" unit="mm" value="2.6">2.60 mm</quantity>
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||
) flattened, broadest posteriorly (broader than head) with oblique corners and darkly coloured, except translucent wide paranotalia.
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||
</paragraph>
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||
<paragraph id="8BB836DBFFD85A1BF2FBFC41FD637343" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
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||
Forewings elongate (7.31/
|
||
<quantity id="4CFF9B3EFFD85A1BF3C5FC40FDED7496" box="[505,611,897,923]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" pageId="2" pageNumber="478" unit="mm" value="1.87">1.87 mm</quantity>
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||
), reaching (overlapping in repose) to the end of ovipositor, and strongly sclerotized. Posterior margin almost straight, anterior margin convex with sparse minute setae (
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<figureCitation id="133C2A5EFFD85A1BF6CDFC65FAC974B3" box="[1265,1351,932,958]" captionStart="FIGURE 1" captionStartId="3.[151,250,1870,1894]" captionTargetBox="[177,1410,181,1847]" captionTargetId="figure-31@3.[177,1410,181,1847]" captionTargetPageId="3" captionText="FIGURE 1 Nigropterix cummingi sp. n. from Burmese amber of Hukawng Valley, North Myanmar, holotype (SNM Z 40731). A: Illustration of dorsal view. B: Photograph of the dorsal view. C: Photograph of the ventral view. D: Right forewing of the specimen. E: Detail of the forewing. F: Detail of terminalia with characteristic chaetotaxy. G: Detail of the ovipositor. Scale bar is 1mm." figureDoi="http://doi.org/10.5281/zenodo.10821576" httpUri="https://zenodo.org/record/10821576/files/figure.png" pageId="2" pageNumber="478">Fig. 1d</figureCitation>
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||
). Apex slightly rounded. Colouration dark, except in subcostal area. It consists of minute setae (
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||
<quantity id="4CFF9B3EFFD85A1BF644FC08FB5074EE" box="[1144,1246,969,995]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="478" unit="mm" value="0.03">0.03 mm</quantity>
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||
long), which are formed in horizontal, somewhat irregular rows of bulges, which have small pits where the setae grow from (
|
||
<figureCitation id="133C2A5EFFD85A1BF702FC2DFA03730A" box="[1342,1421,1004,1031]" captionStart="FIGURE 1" captionStartId="3.[151,250,1870,1894]" captionTargetBox="[177,1410,181,1847]" captionTargetId="figure-31@3.[177,1410,181,1847]" captionTargetPageId="3" captionText="FIGURE 1 Nigropterix cummingi sp. n. from Burmese amber of Hukawng Valley, North Myanmar, holotype (SNM Z 40731). A: Illustration of dorsal view. B: Photograph of the dorsal view. C: Photograph of the ventral view. D: Right forewing of the specimen. E: Detail of the forewing. F: Detail of terminalia with characteristic chaetotaxy. G: Detail of the ovipositor. Scale bar is 1mm." figureDoi="http://doi.org/10.5281/zenodo.10821576" httpUri="https://zenodo.org/record/10821576/files/figure.png" pageId="2" pageNumber="478">Fig. 1e</figureCitation>
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). Venation somewhat traceable with one R vein almost reaching wing apex, presumably two M veins dichotomised near the apex, and two CuA veins branched centrally.
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</paragraph>
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<paragraph id="8BB836DBFFD85A1BF2FBFB99FD8F720F" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
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Hindwings membranous with minute seta (
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||
<quantity id="4CFF9B3EFFD85A1BF098FB98FC87737E" box="[676,777,1113,1139]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="478" unit="mm" value="0.02">0.02 mm</quantity>
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||
long), and overlapping forewings and abdomen (7.29/
|
||
<quantity id="4CFF9B3EFFD85A1BF757FB98FF49739A" metricMagnitude="-3" metricUnit="m" metricValue="3.03" pageId="2" pageNumber="478" unit="mm" value="3.03">3.03 mm</quantity>
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||
). Sc reaching ½ of anterior margin. R veins reduced and divided into R1 and Rs. R1 with one visible branch, Rs with one detectable vein. M partially preserved with only one visible straight vein. Pterostigma not visible. CuA branched with at least 3 visible veins. A branched with at least 4 visible veins, two of which are part of the vannus and folded under the hindwings.
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||
</paragraph>
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<paragraph id="8BB836DBFFD85A1BF2FBFACCFED8729F" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
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||
Legs standardly cursorial and robust. Midfemur partially preserved (
|
||
<quantity id="4CFF9B3EFFD85A1BF1E8FACCFBB2722A" box="[980,1084,1293,1319]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="478" unit="mm" value="0.38">0.38 mm</quantity>
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||
wide). Tibiae (2.46/
|
||
<quantity id="4CFF9B3EFFD85A1BF712FACCFA1A722A" box="[1326,1428,1293,1319]" metricMagnitude="-4" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="478" unit="mm" value="0.31">0.31 mm</quantity>
|
||
) with abundant spurs (
|
||
<quantity id="4CFF9B3EFFD85A1BF3B6FAF0FE7F7246" box="[394,497,1329,1355]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="478" unit="mm" value="0.34">0.34 mm</quantity>
|
||
long). Tarsi divided into 5 segments (0.83/ 0.11; 0.28/ 0.11; 0.12/ 0.11; 0.09/ 0.06; 0.24/
|
||
<quantity id="4CFF9B3EFFD85A1BF2EAFA94FEB37262" box="[214,317,1365,1391]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="478" unit="mm" value="0.12">0.12 mm</quantity>
|
||
). Tarsal claw partially preserved (
|
||
<quantity id="4CFF9B3EFFD85A1BF093FA95FC9A7262" box="[687,788,1364,1391]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="2" pageNumber="478" unit="mm" value="0.16">0.16 mm</quantity>
|
||
long) with arolium (
|
||
<quantity id="4CFF9B3EFFD85A1BF1CAFA94FBD57262" box="[1014,1115,1365,1391]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="478" unit="mm" value="0.14">0.14 mm</quantity>
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||
in diameter), last 3 segments are recognisable.
|
||
</paragraph>
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||
<paragraph id="8BB836DBFFD85A1BF2FBFA5DFA8F714B" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
|
||
Abdomen somewhat narrow (
|
||
<quantity id="4CFF9B3EFFD85A1BF02EFA5CFDF872BA" box="[530,630,1437,1463]" metricMagnitude="-3" metricUnit="m" metricValue="2.11" pageId="2" pageNumber="478" unit="mm" value="2.11">2.11 mm</quantity>
|
||
wide) with a row of setae between the cerci (
|
||
<quantity id="4CFF9B3EFFD85A1BF653FA5CFB5A72BA" box="[1135,1236,1437,1463]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="2" pageNumber="478" unit="mm" value="0.07">0.07 mm</quantity>
|
||
long). Cerci (
|
||
<figureCitation id="133C2A5EFFD85A1BF753FA5DFF2072D7" captionStart="FIGURE 1" captionStartId="3.[151,250,1870,1894]" captionTargetBox="[177,1410,181,1847]" captionTargetId="figure-31@3.[177,1410,181,1847]" captionTargetPageId="3" captionText="FIGURE 1 Nigropterix cummingi sp. n. from Burmese amber of Hukawng Valley, North Myanmar, holotype (SNM Z 40731). A: Illustration of dorsal view. B: Photograph of the dorsal view. C: Photograph of the ventral view. D: Right forewing of the specimen. E: Detail of the forewing. F: Detail of terminalia with characteristic chaetotaxy. G: Detail of the ovipositor. Scale bar is 1mm." figureDoi="http://doi.org/10.5281/zenodo.10821576" httpUri="https://zenodo.org/record/10821576/files/figure.png" pageId="2" pageNumber="478">Fig. 1f</figureCitation>
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) striated with at least 8 segments (0.11/ 0.12; 0.17/ 0.17; 0.19/ 0.22; 0.12/ 0.17; 0.12/ 0.12; 0.12/ 0.08; 0.12/ 0.06; 0.12/
|
||
<quantity id="4CFF9B3EFFD85A1BF2EAFA25FEB372F2" box="[214,317,1508,1535]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="478" unit="mm" value="0.05">0.05 mm</quantity>
|
||
), with abundant setae (
|
||
<quantity id="4CFF9B3EFFD85A1BF006FA24FD1172F2" box="[570,671,1509,1535]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="478" unit="mm" value="0.1">0.10 mm</quantity>
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||
long). Ovipostor robust, protruding (0.56/
|
||
<quantity id="4CFF9B3EFFD85A1BF647FA25FB6C72F2" box="[1147,1250,1508,1535]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="478" unit="mm" value="0.25">0.25 mm</quantity>
|
||
), possibly
|
||
<typeStatus id="54BC8879FFD85A1BF764FA24FA0672F2" box="[1368,1416,1509,1535]" pageId="2" pageNumber="478">type</typeStatus>
|
||
2 (
|
||
<bibRefCitation id="EF964B2AFFD85A1BF2A3F9C9FEFF712E" author="Hornig, M. K. & Haug, C. & Schneider, J. W. & Haug, J. T." box="[159,369,1544,1571]" pageId="2" pageNumber="478" pagination="1 - 24" refId="ref3237" refString="Hornig, M. K., Haug, C., Schneider, J. W. & Haug, J. T. (2018) Evolution of reproductive strategies in dictyopteran insects-clues from ovipositor morphology of extinct roachoids. Acta Palaeontologica Polonica, 63 (1), 1 - 24. https: // doi. org / 10.4202 / app. 00324.2016" type="journal article" year="2018">
|
||
Hörnig
|
||
<emphasis id="B973EAC9FFD85A1BF2CAF9C8FEA0712F" box="[246,302,1544,1570]" italics="true" pageId="2" pageNumber="478">et al.</emphasis>
|
||
2018
|
||
</bibRefCitation>
|
||
). It is punctured on the surface with small minute setae (
|
||
<quantity id="4CFF9B3EFFD85A1BF1DEF9C9FBC9712E" box="[994,1095,1544,1571]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="478" unit="mm" value="0.05">0.05 mm</quantity>
|
||
long) (
|
||
<figureCitation id="133C2A5EFFD85A1BF6ABF9C9FB69712E" box="[1175,1255,1544,1571]" captionStart="FIGURE 1" captionStartId="3.[151,250,1870,1894]" captionTargetBox="[177,1410,181,1847]" captionTargetId="figure-31@3.[177,1410,181,1847]" captionTargetPageId="3" captionText="FIGURE 1 Nigropterix cummingi sp. n. from Burmese amber of Hukawng Valley, North Myanmar, holotype (SNM Z 40731). A: Illustration of dorsal view. B: Photograph of the dorsal view. C: Photograph of the ventral view. D: Right forewing of the specimen. E: Detail of the forewing. F: Detail of terminalia with characteristic chaetotaxy. G: Detail of the ovipositor. Scale bar is 1mm." figureDoi="http://doi.org/10.5281/zenodo.10821576" httpUri="https://zenodo.org/record/10821576/files/figure.png" pageId="2" pageNumber="478">
|
||
Fig.
|
||
<quantity id="4CFF9B3EFFD85A1BF6F7F9C8FB69712E" box="[1227,1255,1545,1571]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="478" unit="g" value="1.0">1g</quantity>
|
||
</figureCitation>
|
||
) and with a row of long setae laterally (
|
||
<quantity id="4CFF9B3EFFD85A1BF3A4F9ECFE61714A" box="[408,495,1581,1607]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="478" unit="mm" value="0.8">0.8 mm</quantity>
|
||
long). Valves are tapering towards its apex, and posteriorly separated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D6550FFD85A1BF2FBF990FB1C71BF" pageId="2" pageNumber="478" type="materials_examined">
|
||
<paragraph id="8BB836DBFFD85A1BF2FBF990FDDA7166" blockId="2.[151,1437,608,2039]" box="[199,596,1616,1643]" pageId="2" pageNumber="478">
|
||
<materialsCitation id="3B6F3C86FFD85A1BF2FBF990FDDA7166" box="[199,596,1616,1643]" collectionCode="SNM" pageId="2" pageNumber="478" specimenCode="Z 40731" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<emphasis id="B973EAC9FFD85A1BF2FBF990FEB57166" bold="true" box="[199,315,1617,1643]" pageId="2" pageNumber="478">
|
||
<typeStatus id="54BC8879FFD85A1BF2FBF990FEB87166" box="[199,310,1617,1643]" pageId="2" pageNumber="478" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult
|
||
<specimenCount id="9D01FD52FFD85A1BF3BBF990FE127166" box="[391,412,1617,1643]" count="1" pageId="2" pageNumber="478" type="female">♀</specimenCount>
|
||
,
|
||
<collectionCode id="ED16AE1EFFD85A1BF396F990FE697166" box="[426,487,1617,1643]" country="Slovakia" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="2" pageNumber="478">SNM</collectionCode>
|
||
<specimenCode id="DBA19EA0FFD85A1BF3D2F990FDDE7166" box="[494,592,1617,1643]" collectionCode="SNM" country="Slovakia" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="2" pageNumber="478">Z 40731</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB836DBFFD85A1BF2FBF9B5FB1C71BF" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
|
||
<emphasis id="B973EAC9FFD85A1BF2FBF9B5FE377182" bold="true" box="[199,441,1652,1679]" pageId="2" pageNumber="478">Derivation of name.</emphasis>
|
||
The species is named after the entomologist Royce Cumming, who contributed to the study of
|
||
<taxonomicName id="4C074D58FFD85A1BF2C5F959FE2771BF" authorityName="Chen et Tan" authorityYear="1973" box="[249,425,1688,1714]" class="Insecta" family="Umenocoleidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="478" phylum="Arthropoda" rank="family">Umenocoleidae</taxonomicName>
|
||
by providing several spectacular specimens from Burmese amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D6550FFD85A1BF2FBF97DFE1C71F7" pageId="2" pageNumber="478" type="diagnosis">
|
||
<paragraph id="8BB836DBFFD85A1BF2FBF97DFE1C71F7" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
|
||
<emphasis id="B973EAC9FFD85A1BF2FBF97DFE5E71DB" bold="true" box="[199,464,1724,1750]" pageId="2" pageNumber="478">Differential diagnosis.</emphasis>
|
||
The specimen differs from
|
||
<taxonomicName id="4C074D58FFD85A1BF12CF97CFB8671DB" box="[784,1032,1725,1750]" class="Insecta" family="Umenocoleidae" genus="Nigropterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="478" phylum="Arthropoda" rank="species" species="angustata">
|
||
<emphasis id="B973EAC9FFD85A1BF12CF97CFB8671DB" box="[784,1032,1725,1750]" italics="true" pageId="2" pageNumber="478">Nigropterix angustata</emphasis>
|
||
</taxonomicName>
|
||
by the parallel M, and R area with numerous cross-veins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D6550FFD85A1DF2FBF8C4FBB3764C" lastPageId="4" lastPageNumber="480" pageId="2" pageNumber="478" type="discussion">
|
||
<paragraph id="8BB836DBFFD85A1BF2FBF8C4FE2C70FB" blockId="2.[151,1437,608,2039]" pageId="2" pageNumber="478">
|
||
<emphasis id="B973EAC9FFD85A1BF2FBF8C4FEB57012" bold="true" box="[199,315,1797,1823]" pageId="2" pageNumber="478">Remarks.</emphasis>
|
||
The specimen is classified within the genus
|
||
<taxonomicName id="4C074D58FFD85A1BF177F8C4FC437013" box="[843,973,1797,1822]" class="Insecta" family="Umenocoleidae" genus="Nigropterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B973EAC9FFD85A1BF177F8C4FC437013" box="[843,973,1797,1822]" italics="true" pageId="2" pageNumber="478">Nigropterix</emphasis>
|
||
</taxonomicName>
|
||
because it has dark coloured forewings with translucent subcostal area, and hindwing with narrow CuA. In addition, the texture of the forewings (formed by irregular rows of bulges) is similar to
|
||
<taxonomicName id="4C074D58FFD85A1BF054F88DFB55706B" authority=", Archaeospinapteryx and Poikiloprosopon." authorityName="Archaeospinapteryx and Poikiloprosopon." box="[616,1243,1868,1894]" class="Insecta" family="Umenocoleidae" genus="Alienopterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B973EAC9FFD85A1BF054F88DFC6F706B" box="[616,993,1868,1894]" italics="true" pageId="2" pageNumber="478">Alienopterix, Archaeospinapteryx</emphasis>
|
||
and
|
||
<emphasis id="B973EAC9FFD85A1BF625F88DFB55706B" box="[1049,1243,1868,1894]" italics="true" pageId="2" pageNumber="478">Poikiloprosopon.</emphasis>
|
||
</taxonomicName>
|
||
The pronotum is widened and shield-like, somewhat similar to
|
||
<taxonomicName id="4C074D58FFD85A1BF09DF8B0FC937087" authorityName="Koubova" authorityYear="2020" box="[673,797,1905,1930]" class="Insecta" family="Cratovitismidae" genus="Perspicuus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B973EAC9FFD85A1BF09DF8B0FC937087" box="[673,797,1905,1930]" italics="true" pageId="2" pageNumber="478">Perspicuus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C074D58FFD85A1BF16AF8B0FC637087" authorityName="Vrsansky et Grimaldi" authorityYear="2003" box="[854,1005,1905,1930]" class="Insecta" family="Umenocoleidae" genus="Jantaropterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="478" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B973EAC9FFD85A1BF16AF8B0FC637087" box="[854,1005,1905,1930]" italics="true" pageId="2" pageNumber="478">Jantaropterix</emphasis>
|
||
</taxonomicName>
|
||
, but more pronounced. The triangular shaped pronotum without reduced paranotalia, and shortened punctured ovipositor with small minute setae on the surface are probably apomorphic, and not observed in other umenocoleids. Legs are cursorial and robust, standard as in most cockroaches.
|
||
</paragraph>
|
||
<caption id="DF786653FFD95A1AF2ABF88FFF6770DE" ID-DOI="http://doi.org/10.5281/zenodo.10821576" ID-Zenodo-Dep="10821576" httpUri="https://zenodo.org/record/10821576/files/figure.png" pageId="3" pageNumber="479" startId="3.[151,250,1870,1894]" targetBox="[177,1410,181,1847]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BB836DBFFD95A1AF2ABF88FFF6770DE" blockId="3.[151,1437,1870,2003]" pageId="3" pageNumber="479">
|
||
<emphasis id="B973EAC9FFD95A1AF2ABF88FFE83706B" bold="true" box="[151,269,1870,1894]" pageId="3" pageNumber="479">FIGURE 1</emphasis>
|
||
<taxonomicName id="4C074D58FFD95A1AF32FF88EFE7C706B" authority="Sendi, 2024" authorityName="Sendi" authorityYear="2024" box="[275,498,1871,1894]" class="Insecta" family="Umenocoleidae" genus="Nigropterix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="479" phylum="Arthropoda" rank="species" species="cummingi" status="sp. nov.">
|
||
<emphasis id="B973EAC9FFD95A1AF32FF88EFE7C706B" box="[275,498,1871,1894]" italics="true" pageId="3" pageNumber="479">Nigropterix cummingi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B973EAC9FFD95A1AF3C5F88EFDBC706A" bold="true" box="[505,562,1871,1895]" pageId="3" pageNumber="479">
|
||
<taxonomicNameLabel id="A24057B2FFD95A1AF3C5F88EFDBC706A" box="[505,562,1871,1895]" pageId="3" pageNumber="479" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from Burmese amber of Hukawng Valley, North Myanmar, holotype (SNM Z 40731). A: Illustration of dorsal view. B: Photograph of the dorsal view. C: Photograph of the ventral view. D: Right forewing of the specimen. E: Detail of the forewing. F: Detail of terminalia with characteristic chaetotaxy. G: Detail of the ovipositor. Scale bar is 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB836DBFFDE5A1DF2FBFF57FBB3764C" blockId="4.[151,1436,150,321]" pageId="4" pageNumber="480">
|
||
<emphasis id="B973EAC9FFDE5A1DF2FBFF57FD8A77BD" bold="true" box="[199,516,150,176]" pageId="4" pageNumber="480">Character of preservation.</emphasis>
|
||
The amber sample has a honey-yellow colour with numerous small syninclusions, probably detritus and possibly
|
||
<taxonomicName id="4C074D58FFDE5A1DF038FF7AFCEA77D8" authorityName="Vršanská & Hinkelman" authorityYear="2020" box="[516,868,187,213]" class="Chlorophyceae" family="Chlamydomonadaceae" genus="Chlamydomonas" kingdom="Plantae" order="Chlamydomonadales" pageId="4" pageNumber="480" phylum="Chlorophyta" rank="species" species="hanublicanus">
|
||
<emphasis id="B973EAC9FFDE5A1DF038FF7AFCEA77D8" box="[516,868,187,213]" italics="true" pageId="4" pageNumber="480">Chlamydomonas hanublicanus</emphasis>
|
||
</taxonomicName>
|
||
cysts (
|
||
<bibRefCitation id="EF964B2AFFDE5A1DF187FF7AFA9877D8" author="Hinkelman, J. & Vrsanska, L." box="[955,1302,187,213]" pageId="4" pageNumber="480" pagination="13" refId="ref3185" refString="Hinkelman, J. & Vrsanska, L. (2020) A Myanmar amber cockroach with protruding feces contains pollen and a rich microcenosis. The Science of Nature, 107, 13. https: // doi. org / 10.1007 / s 00114 - 020 - 1669 - y" type="journal article" year="2020">Hinkelman & Vršanská 2020</bibRefCitation>
|
||
). The adult umenocoleid is almost completely preserved, except parts of the legs, distal cercal segments, and antennae are missing. Details of the forewing venation are barely visible due to strong sclerotization of the wings. Details of the head (e.g. ocelli, palpomeres) cannot be recognised due to the state of preservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |