142 lines
18 KiB
XML
142 lines
18 KiB
XML
<document id="0D1F0B9AC7B37CF6B43BFEF926951B74" ID-CLB-Dataset="39152" ID-DOI="10.11646/zootaxa.4107.3.14" ID-GBIF-Dataset="1b00d70d-1952-4628-a1da-ad47cd02628a" ID-ISSN="1175-5326" ID-Zenodo-Dep="262394" ID-ZooBank="C1A137C8-888E-4303-B2B4-397ABB9D5D58" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1462258958507" checkinUser="plazi" docAuthor="Leister, Matthew & Miller, Kelly B." docDate="2016" docId="03AE8797FFAAFFBB69F08FDD0358B0FA" docLanguage="en" docName="zootaxa.4107.3.14.pdf.imf" docOrigin="Zootaxa 4107 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hoedillus Simon 1898" docType="treatment" docVersion="10" lastPageNumber="448" masterDocId="FF97FFEFFFAAFFBA696789400124B11D" masterDocTitle="First description of the male Hoedillus sexpunctatus Simon, 1898; a redescription of the female, and transfer of Hoedillus to Zoropsidae Bertkau, 1882 (Araneae)" masterLastPageNumber="450" masterPageNumber="447" pageNumber="447" updateTime="1698662222870" updateUser="ExternalLinkService">
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<mods:title id="B67CE4A0B240847DAD51AD22859AA919">First description of the male Hoedillus sexpunctatus Simon, 1898; a redescription of the female, and transfer of Hoedillus to Zoropsidae Bertkau, 1882 (Araneae)</mods:title>
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<mods:namePart id="81DAE84091784C4F31575F27B192F1CF">Leister, Matthew</mods:namePart>
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<mods:namePart id="D7AC42F1CF334AC2A55EB00D63685779">Miller, Kelly B.</mods:namePart>
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<mods:title id="177355A5CEBCA4FCF00AA28FC7D76A3F">Zootaxa</mods:title>
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<mods:date id="C0FCA2C15F9F97BFF6947F7037797FAF">2016</mods:date>
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<mods:number id="1F079525AD2A9F9EA8174F2EA55AF2F4">4107</mods:number>
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<treatment id="03AE8797FFAAFFBB69F08FDD0358B0FA" ID-DOI="http://doi.org/10.5281/zenodo.6057504" ID-GBIF-Taxon="120620748" ID-Zenodo-Dep="6057504" LSID="urn:lsid:plazi:treatment:03AE8797FFAAFFBB69F08FDD0358B0FA" httpUri="http://treatment.plazi.org/id/03AE8797FFAAFFBB69F08FDD0358B0FA" lastPageId="1" lastPageNumber="448" pageId="0" pageNumber="447">
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<subSubSection id="C31D650AFFAAFFBA69F08FDD0089B7AA" box="[151,429,1693,1719]" pageId="0" pageNumber="447" type="nomenclature">
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<paragraph id="8BB83681FFAAFFBA69F08FDD0089B7AA" blockId="0.[151,1149,1624,1748]" box="[151,429,1693,1719]" pageId="0" pageNumber="447">
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<heading id="D0F081EDFFAAFFBA69F08FDD0089B7AA" bold="true" box="[151,429,1693,1719]" fontSize="11" level="1" pageId="0" pageNumber="447" reason="1">
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<taxonomicName id="4C074D02FFAAFFBA69F08FDD0089B7AA" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[151,429,1693,1719]" class="Arachnida" family="Zoridae" genus="Hoedillus" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="447" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA93FFAAFFBA69F08FDD0089B7AA" bold="true" box="[151,429,1693,1719]" pageId="0" pageNumber="447">
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<emphasis id="B973EA93FFAAFFBA69F08FDD0028B7AA" bold="true" box="[151,268,1693,1719]" italics="true" pageId="0" pageNumber="447">Hoedillus</emphasis>
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<bibRefCitation id="EF964B70FFAAFFBA68748FDD0089B7AA" author="Simon" box="[275,429,1693,1719]" pageId="0" pageNumber="447" refString="Simon, E. (1898) Histoire naturelle des araignees. Tome 2. Librairie Encyclopedique de Roret, Paris, pp. 193 - 380." type="journal article" year="1898">Simon, 1898</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D650AFFAAFFBA69F08FFF0559B7C9" box="[151,1149,1727,1748]" pageId="0" pageNumber="447" type="reference_group">
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<paragraph id="8BB83681FFAAFFBA69F08FFF0559B7C9" blockId="0.[151,1149,1624,1748]" box="[151,1149,1727,1748]" pageId="0" pageNumber="447">
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<treatmentCitationGroup id="AB1711AFFFAAFFBA69F08FFF0559B7C9" box="[151,1149,1727,1748]" pageId="0" pageNumber="447">
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<taxonomicName id="4C074D02FFAAFFBA69F08FFF0053B7C9" authority="Simon, 1898" authorityName="Simon" authorityYear="1898" box="[151,375,1727,1748]" class="Arachnida" family="Zoridae" genus="Hoedillus" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="447" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA93FFAAFFBA69F08FFF01D0B7C9" box="[151,244,1727,1748]" italics="true" pageId="0" pageNumber="447">Hoedillus</emphasis>
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<bibRefCitation id="EF964B70FFAAFFBA699D8FFF0053B7C9" author="Simon" box="[250,375,1727,1748]" pageId="0" pageNumber="447" refString="Simon, E. (1898) Histoire naturelle des araignees. Tome 2. Librairie Encyclopedique de Roret, Paris, pp. 193 - 380." type="journal article" year="1898">Simon, 1898</bibRefCitation>
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</taxonomicName>
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;
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<typeStatus id="54BC8823FFAAFFBA68E68FFF008EB7C9" box="[385,426,1727,1748]" pageId="0" pageNumber="447">type</typeStatus>
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species by original designation:
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<taxonomicName id="4C074D02FFAAFFBA6B828FFF055DB7C9" authority="Simon, 1898: 202" authorityName="Simon" authorityPageNumber="202" authorityYear="1898" box="[741,1145,1727,1748]" class="Arachnida" family="Zoridae" genus="Hoedillus" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="447" phylum="Arthropoda" rank="species" species="sexpunctatus">
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<emphasis id="B973EA93FFAAFFBA6B828FFF02E0B7C9" box="[741,964,1727,1748]" italics="true" pageId="0" pageNumber="447">Hoedillus sexpunctatus</emphasis>
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<treatmentCitation id="0AA61090FFAAFFBA6AAD8FFF055DB7C9" author="Simon" box="[970,1145,1727,1748]" page="202" pageId="0" pageNumber="447" year="1898">
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<bibRefCitation id="EF964B70FFAAFFBA6AAD8FFF056DB7C9" author="Simon" box="[970,1097,1727,1748]" pageId="0" pageNumber="447" refString="Simon, E. (1898) Histoire naturelle des araignees. Tome 2. Librairie Encyclopedique de Roret, Paris, pp. 193 - 380." type="journal article" year="1898">Simon, 1898</bibRefCitation>
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: 202
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D650AFFAAFFBB69F08E400358B0FA" lastPageId="1" lastPageNumber="448" pageId="0" pageNumber="447" type="diagnosis">
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<paragraph id="8BB83681FFAAFFBA69F08E400239B662" blockId="0.[151,1436,1791,2023]" pageId="0" pageNumber="447">
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<emphasis id="B973EA93FFAAFFBA69F08E400028B60A" bold="true" box="[151,268,1792,1815]" pageId="0" pageNumber="447">Diagnosis.</emphasis>
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The genus
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<taxonomicName id="4C074D02FFAAFFBA68E08FBF00CBB60A" box="[391,495,1791,1815]" class="Arachnida" family="Zoridae" genus="Hoedillus" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="447" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA93FFAAFFBA68E08FBF00CBB60A" box="[391,495,1791,1815]" italics="true" pageId="0" pageNumber="447">Hoedillus</emphasis>
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</taxonomicName>
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is distinguished by the following combination of characters: (1) eight eyes forming two strongly recurved rows, (2) lateral eyes of both rows 1.5 times the diameter of median eyes, (
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<docIssue id="CF64496EFFAAFFBA6DC58E62058BB624" box="[1186,1199,1826,1849]" pageId="0" pageNumber="447">3</docIssue>
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) four to five pairs of ventral spines in raised sockets on tibia I and II, and two pairs of ventral spines in raised sockets on metatarsi I and II, and (4) three promarginal and two retromarginal cheliceral teeth.
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</paragraph>
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<paragraph id="8BB83681FFAAFFBB69A08EC9008FB024" blockId="0.[151,1436,1791,2023]" lastBlockId="1.[151,1436,152,487]" lastPageId="1" lastPageNumber="448" pageId="0" pageNumber="447">
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<emphasis id="B973EA93FFAAFFBA69A08EC900F5B6BC" box="[199,465,1929,1953]" italics="true" pageId="0" pageNumber="447">Justification of transfer.</emphasis>
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An examination of the male pedipalpus fails to reveal synapomorphic characters for
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<taxonomicName id="4C074D02FFAAFFBA69F08EED0029B6D9" box="[151,269,1965,1988]" class="Arachnida" family="Miturgidae" kingdom="Animalia" order="Araneae" pageId="0" pageNumber="447" phylum="Arthropoda" rank="family">Miturgidae</taxonomicName>
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(
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<emphasis id="B973EA93FFAAFFBA68798EED007DB6D9" box="[286,345,1965,1988]" italics="true" pageId="0" pageNumber="447">sensu</emphasis>
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<bibRefCitation id="EF964B70FFAAFFBA68058EED0324B6D9" author="Ramirez" box="[354,512,1965,1988]" pageId="0" pageNumber="447" refString="Ramirez, M. J. (2014) The morphology and phylogeny of the dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014</bibRefCitation>
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): a retrolateral cymbial groove (
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<bibRefCitation id="EF964B70FFAAFFBA6A398EED01E6B6FA" author="Ramirez" pageId="0" pageNumber="447" refString="Ramirez, M. J. (2014) The morphology and phylogeny of the dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014: figs. 145F, 148D, 162A, B; character 331</bibRefCitation>
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), retrolateral tibial apophysis (RTA) with membranous area (
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<bibRefCitation id="EF964B70FFAAFFBA6A298E900455B6FA" author="Ramirez" box="[846,1393,2000,2023]" pageId="0" pageNumber="447" refString="Ramirez, M. J. (2014) The morphology and phylogeny of the dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014: figs. 145A, 163A, B; character 315</bibRefCitation>
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), or a canal on the RTA (
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<bibRefCitation id="EF964B70FFABFFBB681089D802C1B1B2" author="Ramirez" box="[375,997,152,175]" pageId="1" pageNumber="448" refString="Ramirez, M. J. (2014) The morphology and phylogeny of the dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014: figs. 142E, 146B, 160E, H; character 316</bibRefCitation>
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). Furthermore, the shape of the embolus and median apophysis is inconsistent with this group;
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<taxonomicName id="4C074D02FFABFFBB6B8E89F90275B1CC" box="[745,849,185,209]" class="Arachnida" family="Zoridae" genus="Hoedillus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA93FFABFFBB6B8E89F90275B1CC" box="[745,849,185,209]" italics="true" pageId="1" pageNumber="448">Hoedillus</emphasis>
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</taxonomicName>
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lacks the long circular embolus and a forward facing median apophysis seen in the
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<taxonomicName id="4C074D02FFABFFBB68BC899D0375B1E9" box="[475,593,221,244]" class="Arachnida" family="Miturgidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="448" phylum="Arthropoda" rank="family">Miturgidae</taxonomicName>
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(
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<bibRefCitation id="EF964B70FFABFFBB6B07899D0284B1E9" author="Ramirez" box="[608,928,221,244]" pageId="1" pageNumber="448" refString="Ramirez, M. J. (2014) The morphology and phylogeny of the dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014: figs. 145, 146</bibRefCitation>
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). The examination also fails to reveal a tegular division (
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<bibRefCitation id="EF964B70FFABFFBB699C884000B3B00A" author="Ramirez" box="[251,407,256,279]" pageId="1" pageNumber="448" refString="Ramirez, M. J. (2014) The morphology and phylogeny of the dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 - 374. http: // dx. doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014</bibRefCitation>
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: figs. 150A–C, E; character 343 and
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<bibRefCitation id="EF964B70FFABFFBB6A42884002B8B00A" author="Silva" box="[805,924,256,279]" pageId="1" pageNumber="448" refString="Silva, D. (2003) Higher-level relationships of the spider family Ctenidae (Araneae: Ctenoidea). Bulletin of the American Museum of Natural History, 274, 1 - 86. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2003) 274 <0001: HLROTS> 2.0. CO; 2" type="journal article" year="2003">Silva 2003</bibRefCitation>
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: fig. 18; character 38), which is synapomorphic for the
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<taxonomicName id="4C074D02FFABFFBB69838862007BB024" box="[228,351,290,313]" class="Arachnida" family="Zoridae" genus="Xenoctenus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA93FFABFFBB69838862007BB024" box="[228,351,290,313]" italics="true" pageId="1" pageNumber="448">Xenoctenus</emphasis>
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</taxonomicName>
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group.
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</paragraph>
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<paragraph id="8BB83681FFABFFBB69A088050358B0FA" blockId="1.[151,1436,152,487]" pageId="1" pageNumber="448">
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Both males and females of
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<taxonomicName id="4C074D02FFABFFBB688988040371B041" box="[494,597,324,348]" class="Arachnida" family="Zoridae" genus="Hoedillus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="448" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA93FFABFFBB688988040371B041" box="[494,597,324,348]" italics="true" pageId="1" pageNumber="448">Hoedillus</emphasis>
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</taxonomicName>
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show characters previously considered as synapomorphies of
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<taxonomicName id="4C074D02FFABFFBB6D9388050454B041" box="[1268,1392,325,348]" class="Arachnida" family="Zoropsidae" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="448" phylum="Arthropoda" rank="family">Zoropsidae</taxonomicName>
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: the lateral eyes are larger than the median eyes (
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<figureCitation id="133C2A04FFABFFBB6B1688280389B062" box="[625,685,360,383]" captionStart="FIGURE 1" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[223,1348,199,1872]" captionTargetId="figure@2.[212,1360,194,1882]" captionTargetPageId="2" captionText="FIGURE 1. Hoedillus sexpunctatus Simon, 1898, male. A. Habitus. B – F. Right pedipalpus; B. Retrolateral view; C. Ventral view; D. Prolateral view; E. Tibia ventral view; F. Pedipalpus, dorsal view, highlighting dorsal cymbial scopula. Con—conductor, Em—embolus, MA—median apophysis, TA—tegular apophysis." httpUri="https://zenodo.org/record/262395/files/figure.png" pageId="1" pageNumber="448">Fig. 1</figureCitation>
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A,
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<bibRefCitation id="EF964B70FFABFFBB6BB6882802F4B062" author="Raven" box="[721,976,360,383]" pageId="1" pageNumber="448" refString="Raven, R. J. & Stumkat, K. S. (2005) Revisions of Australian ground-hunting spiders: II. Zoropsidae (Lycosoidea: Araneae). Memoirs of the Queensland Museum, 50, 347 - 423." type="journal article" year="2005">Raven & Stumkat 2005</bibRefCitation>
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: character 42). The male pedipalpus shows dense dorsal cymbial scopula with flattened surface (
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<figureCitation id="133C2A04FFABFFBB6BA888CA0220B0BC" box="[719,772,394,417]" captionStart="FIGURE 1" captionStartId="2.[151,244,1944,1966]" captionTargetBox="[223,1348,199,1872]" captionTargetId="figure@2.[212,1360,194,1882]" captionTargetPageId="2" captionText="FIGURE 1. Hoedillus sexpunctatus Simon, 1898, male. A. Habitus. B – F. Right pedipalpus; B. Retrolateral view; C. Ventral view; D. Prolateral view; E. Tibia ventral view; F. Pedipalpus, dorsal view, highlighting dorsal cymbial scopula. Con—conductor, Em—embolus, MA—median apophysis, TA—tegular apophysis." httpUri="https://zenodo.org/record/262395/files/figure.png" pageId="1" pageNumber="448">Fig 1</figureCitation>
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B–D, F;
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<bibRefCitation id="EF964B70FFABFFBB6A0188CA0542B0BC" author="Raven" box="[870,1126,394,417]" pageId="1" pageNumber="448" refString="Raven, R. J. & Stumkat, K. S. (2005) Revisions of Australian ground-hunting spiders: II. Zoropsidae (Lycosoidea: Araneae). Memoirs of the Queensland Museum, 50, 347 - 423." type="journal article" year="2005">Raven & Stumkat 2005</bibRefCitation>
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: character 1), and the female is found to have dorsal spigots on the posterior median spinnerets (
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<bibRefCitation id="EF964B70FFABFFBB6AFE88ED0460B0D9" author="Raven" box="[921,1348,429,452]" pageId="1" pageNumber="448" refString="Raven, R. J. & Stumkat, K. S. (2005) Revisions of Australian ground-hunting spiders: II. Zoropsidae (Lycosoidea: Araneae). Memoirs of the Queensland Museum, 50, 347 - 423." type="journal article" year="2005">Raven & Stumkat 2005: character 63</bibRefCitation>
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). These characters justify its placement in this family.
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</paragraph>
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</subSubSection>
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</treatment>
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||
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