210 lines
24 KiB
XML
210 lines
24 KiB
XML
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<mods:title id="3125DCEC84C262231461109A653E53FA">New species, occurrence records and observations of predation by deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA ship Okeanos Explorer</mods:title>
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<mods:namePart id="5A666756EDAACDF13EEB7B67D2EC1B4B">Mah, Christopher L.</mods:namePart>
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<mods:title id="9B4BD38FAC01DBEB83E261F21988F4B5">Zootaxa</mods:title>
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<mods:date id="881B51DF35D44C66C7D6EC096F3FBD6B">2020</mods:date>
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<treatment id="03AE8786FFB3D54FFF4024BEFD24581C" ID-DOI="http://doi.org/10.5281/zenodo.3803745" ID-GBIF-Taxon="163737300" ID-Zenodo-Dep="3803745" LSID="urn:lsid:plazi:treatment:03AE8786FFB3D54FFF4024BEFD24581C" httpUri="http://treatment.plazi.org/id/03AE8786FFB3D54FFF4024BEFD24581C" lastPageId="15" lastPageNumber="216" pageId="14" pageNumber="215">
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<subSubSection id="C31D651BFFB3D54EFF4024BEFE4E58EB" box="[151,400,1449,1475]" pageId="14" pageNumber="215" type="nomenclature">
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<paragraph id="8BB83690FFB3D54EFF4024BEFE4E58EB" blockId="14.[151,400,1449,1475]" box="[151,400,1449,1475]" pageId="14" pageNumber="215">
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<taxonomicName id="4C074D13FFB3D54EFF4024BEFE4E58EB" ID-CoL="8RYP7" box="[151,400,1449,1475]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="family">
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<emphasis id="B973EA82FFB3D54EFF4024BEFE4E58EB" bold="true" box="[151,400,1449,1475]" pageId="14" pageNumber="215">GONIASTERIDAE</emphasis>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFB3D54EFF4024E6FD115A5B" pageId="14" pageNumber="215" type="discussion">
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<paragraph id="8BB83690FFB3D54EFF4024E6FC4C5B07" blockId="14.[151,1437,1521,1907]" pageId="14" pageNumber="215">
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The
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<taxonomicName id="4C074D13FFB3D54EFF1D24E6FEB85B23" box="[202,358,1521,1547]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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is the most taxon-rich group within the
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<taxonomicName id="4C074D13FFB3D54EFCF324E6FC7E5B23" authorityName="Database (Mah" authorityYear="2020" box="[804,928,1521,1547]" class="Asteroidea" kingdom="Animalia" pageId="14" pageNumber="215" phylum="Echinodermata" rank="class">Asteroidea</taxonomicName>
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, including over 270 species in approximately 70 genera with many new species being described (e.g., Mah 2019).
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</paragraph>
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<paragraph id="8BB83690FFB3D54EFF40272EFD2F5B5F" blockId="14.[151,1437,1521,1907]" pageId="14" pageNumber="215">
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Although an overview of the group is presented in
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<bibRefCitation id="EF964B61FFB3D54EFD1F272EFC335B7B" author="Clark, A. M. & Downey, M. E." box="[712,1005,1593,1620]" pageId="14" pageNumber="215" refId="ref28216" refString="Clark, A. M. & Downey, M. E. (1992) Starfishes of the Atlantic. Chapman and Hall, London, 794 pp." type="book" year="1992">Clark and Downey (1992)</bibRefCitation>
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,
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<bibRefCitation id="EF964B61FFB3D54EFC2F272EFB7C5B7B" author="Halpern, J. A." box="[1016,1186,1593,1619]" pageId="14" pageNumber="215" pagination="193 - 286" refId="ref29158" refString="Halpern, J. A. (1970) Biological investigations of the deep sea. 51. Goniasteridae (Echinodermata: Asteroidea) of the Straits of Florida. Bulletin of Marine Science, 20 (1), 193 - 286." type="journal article" year="1970">Halpern (1970)</bibRefCitation>
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provided an extensive treatment on
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<taxonomicName id="4C074D13FFB3D54EFEFC274AFE195B5F" box="[299,455,1629,1655]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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from the tropical Atlantic.
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</paragraph>
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<paragraph id="8BB83690FFB3D54EFF102796FAD45A2F" blockId="14.[151,1437,1521,1907]" pageId="14" pageNumber="215">
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Although several subfamilies have been recognized, the most frequently encountered group on the
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<taxonomicName id="4C074D13FFB3D54EFAED2796FF245B97" authority="Explorer" authorityName="Explorer" class="Insecta" family="Pentatomidae" genus="Okeanos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="215" phylum="Arthropoda" rank="genus">
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<emphasis id="B973EA82FFB3D54EFAED2796FF245B97" italics="true" pageId="14" pageNumber="215">Okeanos Explorer</emphasis>
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</taxonomicName>
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surveys is the
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<taxonomicName id="4C074D13FFB3D54EFE7427B2FD9F5B97" authorityName="Verrill" authorityYear="1899" box="[419,577,1701,1727]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="subFamily" subFamily="Hippasterinae">Hippasterinae</taxonomicName>
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, which includes multiple genera that are now all known to feed on deep-sea octocorals and scleractinians (e.g.
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<bibRefCitation id="EF964B61FFB3D54EFDC927DEFD7C5BCC" author="Mah, C. L." box="[542,674,1737,1764]" pageId="14" pageNumber="215" pagination="1 - 8" refId="ref29743" refString="Mah, C. L. (2015 b) A new Atlantic species of Evoplosoma with taxonomic summary and in situ observations of Atlantic deep-sea corallivorous Goniasteridae (Valvatida; Asteroidea). Marine Biodiversity Records, 8 (e 5), 1 - 8. https: // doi. org / 10.1017 / S 1755267214001407" type="journal article" year="2015">Mah 2015b</bibRefCitation>
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,
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<bibRefCitation id="EF964B61FFB3D54EFD7827DEFCB75BCB" author="Mah, C. L. & Nizinski, M. & Lundsten, L." box="[687,873,1737,1764]" pageId="14" pageNumber="215" pagination="266 - 301" refId="ref30122" refString="Mah, C. L., Nizinski, M. & Lundsten, L. (2010) Phylogenetic Revision of the Hippasterinae (Goniasteridae; Asteroidea): Systematics of Deep Sea Corallivores, including one new genus and three new species. Zoological Journal of the Linnean Society, 160, 266 - 301, 11 figs. https: // doi. org / 10.1111 / j. 1096 - 3642.2010.00638. x" type="book chapter" year="2010">
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Mah
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<emphasis id="B973EA82FFB3D54EFD3D27DDFCC35BCB" box="[746,797,1737,1763]" italics="true" pageId="14" pageNumber="215">et al</emphasis>
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. 2010
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</bibRefCitation>
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). All of the genera within the
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<taxonomicName id="4C074D13FFB3D54EFB6927DEFA825BCB" authorityName="Verrill" authorityYear="1899" box="[1214,1372,1737,1763]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="subFamily" subFamily="Hippasterinae">Hippasterinae</taxonomicName>
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,
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<taxonomicName id="4C074D13FFB3D54EFABE27DDFF2D5A2F" class="Asteroidea" family="Goniasteridae" genus="Hippasteria" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="genus">
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<emphasis id="B973EA82FFB3D54EFABE27DDFF2D5A2F" italics="true" pageId="14" pageNumber="215">Hippasteria</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C074D13FFB3D54EFED627FAFE555A2F" box="[257,395,1773,1799]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="genus">
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<emphasis id="B973EA82FFB3D54EFED627FAFE555A2F" box="[257,395,1773,1799]" italics="true" pageId="14" pageNumber="215">Evoplosoma</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C074D13FFB3D54EFE4E27FAFDFC5A2F" box="[409,546,1773,1799]" class="Asteroidea" family="Goniasteridae" genus="Gilbertaster" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="genus">
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<emphasis id="B973EA82FFB3D54EFE4E27FAFDFC5A2F" box="[409,546,1773,1799]" italics="true" pageId="14" pageNumber="215">Gilbertaster</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C074D13FFB3D54EFD8E27FAFD115A2F" authorityName="Mah, Nizinski & Lundsten" authorityYear="2010" box="[601,719,1773,1799]" class="Asteroidea" family="Goniasteridae" genus="Sthenaster" kingdom="Animalia" order="Valvatida" pageId="14" pageNumber="215" phylum="Echinodermata" rank="genus">
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<emphasis id="B973EA82FFB3D54EFD8E27FAFD115A2F" box="[601,719,1773,1799]" italics="true" pageId="14" pageNumber="215">Sthenaster</emphasis>
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</taxonomicName>
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were observed during the surveys included herein.
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</paragraph>
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<paragraph id="8BB83690FFB3D54EFF102606FD115A5B" blockId="14.[151,1437,1521,1907]" pageId="14" pageNumber="215">
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Corallivory has been reported among many Pacific goniasterid taxa (
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<bibRefCitation id="EF964B61FFB3D54EFC0B2606FB885A03" author="Mah, C. L." box="[988,1110,1809,1835]" pageId="14" pageNumber="215" pagination="917 - 954" refId="ref29603" refString="Mah, C. L. (2006) Phylogeny and biogeography of the deep-sea goniasterid, Circeaster (Echinodermata: Asteroidea) including descriptions of six new species. Zoosystema, 28 (4), 917 - 954." type="journal article" year="2006">Mah 2006</bibRefCitation>
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,
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<bibRefCitation id="EF964B61FFB3D54EFBB42606FB3B5A04" author="Mah, C. L." box="[1123,1253,1809,1836]" pageId="14" pageNumber="215" pagination="211 - 228" refId="ref29686" refString="Mah, C. L. (2015 a) New species, corallivory, in situ video observations and overview of the Goniasteridae (Valvatida, Asteroidea) in the Hawaiian Region. Zootaxa, 3926 (2), 211 - 228. https: // doi. org / 10.11646 / zootaxa. 3926.2.3" type="journal article" year="2015">Mah 2015a</bibRefCitation>
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) whose biology and feeding habits had been previously unknown suggesting that the group, as a whole, is likely significant to understanding the ecology of deep-sea coral habitats.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFB2D54FFF40218EFDF35D9B" box="[151,557,153,180]" pageId="15" pageNumber="216" type="discussion">
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<paragraph id="8BB83690FFB2D54FFF40218EFDF35D9B" blockId="15.[151,557,153,215]" box="[151,557,153,180]" pageId="15" pageNumber="216">
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<emphasis id="B973EA82FFB2D54FFF40218EFDF35D9B" bold="true" box="[151,557,153,180]" pageId="15" pageNumber="216">
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Undescribed
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<taxonomicName id="4C074D13FFB2D54FFEEA218EFE335D9B" box="[317,493,153,179]" class="Asteroidea" family="Goniasteridae" kingdom="Animalia" order="Valvatida" pageId="15" pageNumber="216" phylum="Echinodermata" rank="family">Goniasteridae</taxonomicName>
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sp. 1
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFB2D54FFF4021AAFED75DFF" box="[151,265,189,215]" pageId="15" pageNumber="216" type="description">
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<paragraph id="8BB83690FFB2D54FFF4021AAFED75DFF" blockId="15.[151,557,153,215]" box="[151,265,189,215]" pageId="15" pageNumber="216">
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<figureCitation id="133C2A15FFB2D54FFF4021AAFED75DFF" box="[151,265,189,215]" captionStart="FIGURE 6" captionStartId="16.[151,250,1363,1388]" captionTargetBox="[163,1426,193,1333]" captionTargetId="figure@16.[151,1436,181,1337]" captionTargetPageId="16" captionText="FIGURE 6. Unidentified species and Ceramaster grenadensis. A. Unidentified genus and species 1, NW Florida Escarpment, Gulf of Mexico, 2178 m. B. C. grenadensis in situ feeding. Unidentified stalks emerging from below (indicated with arrows). Richardson Scarp, North Atlantic. 897 m C. C. grenadensis feeding on coral rubble. Richardson Scarp, North Atlantic. 954 m D. C. grenadensis with yellow scale worm associate (indicated by arrow). Okeanos Ridge, 734 m." figureDoi="http://doi.org/10.5281/zenodo.3764030" httpUri="https://zenodo.org/record/3764030/files/figure.png" pageId="15" pageNumber="216">Figure 6A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFB2D54FFF402012FBD25FF8" pageId="15" pageNumber="216" type="discussion">
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<paragraph id="8BB83690FFB2D54FFF402012FD425CDF" blockId="15.[151,1437,261,720]" pageId="15" pageNumber="216">This species was characterized by its strongly stellate body shape (R/r=~4.0) with elongate, tapering arms and weakly curved to straight interradial arcs. Abactinal plates were abutted and were part of a relatively even surface with no spines or other primary accessories (i.e. tubercles, etc.). Fine, widely spaced granules were observed on the surface of the abactinal and superomarginal plates. Arm plates appeared similar in size to those on disk. Superomarginal plates appeared strongly convex, wide and were dark in color relative to disk and peripheral coloration. Distalmost inferomarginal plates possessed sharp tubercles or small spines. Color in life was white with brown highlights on disk and central surface of marginal plates.</paragraph>
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<paragraph id="8BB83690FFB2D54FFF102316FBD25FF8" blockId="15.[151,1437,261,720]" pageId="15" pageNumber="216">
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This species could not be fully reconciled with known taxa, suggesting a likely new genus and species. No specimen was collected. This species’ abactinal plate and arm shape suggests comparison
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<taxonomicName id="4C074D13FFB2D54FFB422331FA485F17" baseAuthorityName="A. H. Clark" baseAuthorityYear="1916" box="[1173,1430,550,576]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="15" pageNumber="216" phylum="Echinodermata" rank="species" species="americanus">
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<emphasis id="B973EA82FFB2D54FFB422331FA485F17" box="[1173,1430,550,576]" italics="true" pageId="15" pageNumber="216">Circeaster americanus</emphasis>
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</taxonomicName>
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, but abactinal arm plates appeared relatively similar in size and arms were much more elongate than known specimens. The superomarginal plates and apparently convex superomarginal plates are also different from
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<emphasis id="B973EA82FFB2D54FFAF42379FA435FA0" box="[1315,1437,622,648]" italics="true" pageId="15" pageNumber="216">
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<taxonomicName id="4C074D13FFB2D54FFAF42379FA475FA0" authorityName="Koehler" authorityYear="1909" box="[1315,1433,622,648]" class="Asteroidea" family="Goniasteridae" genus="Circeaster" kingdom="Animalia" order="Valvatida" pageId="15" pageNumber="216" phylum="Echinodermata" rank="genus">Circeaster</taxonomicName>
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.
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</emphasis>
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The elongate arms and marginal plates also suggest similarity with
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<taxonomicName id="4C074D13FFB2D54FFC422386FBC15F83" box="[917,1055,657,683]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="15" pageNumber="216" phylum="Echinodermata" rank="genus">
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<emphasis id="B973EA82FFB2D54FFC422386FBC15F83" box="[917,1055,657,683]" italics="true" pageId="15" pageNumber="216">Evoplosoma</emphasis>
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</taxonomicName>
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, although the extensive spination on the abactinal or marginal plates and the pulpy body surface was not present.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFB2D54FFF4023EAFDFB5EE3" pageId="15" pageNumber="216" type="biology_ecology">
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<paragraph id="8BB83690FFB2D54FFF4023EAFE515E3F" blockId="15.[151,1437,765,971]" box="[151,399,765,791]" pageId="15" pageNumber="216">
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<emphasis id="B973EA82FFB2D54FFF4023EAFE515E3F" box="[151,399,765,791]" italics="true" pageId="15" pageNumber="216">Feeding Observations</emphasis>
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</paragraph>
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<paragraph id="8BB83690FFB2D54FFF102236FDFB5EE3" blockId="15.[151,1437,765,971]" pageId="15" pageNumber="216">
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The three images of this species all showed it on hard, rocky vertical substratum. Two of the images show one of the arms extended off the substrate into the area above the animal. One of the
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<collectingRegion id="49C3F872FFB2D54FFBFD2252FBA55E77" box="[1066,1147,837,863]" country="United States of America" name="Florida" pageId="15" pageNumber="216">Florida</collectingRegion>
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Escarpment observations (at
|
||
<quantity id="4CFF9B75FFB2D54FFF6A227EFECA5EAC" box="[189,276,873,900]" metricMagnitude="3" metricUnit="m" metricValue="2.006" pageId="15" pageNumber="216" unit="m" value="2006.0">2006 m</quantity>
|
||
) shows a stalk of possibly coral, bryozoan or some other flocculent material emerging from below the oral surface. Possible food was not observed on the other image with the extended arm but the behavior might have been indicative of possible feeding.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFB2D54FFF4022EDFA95593C" box="[151,1355,1017,1044]" pageId="15" pageNumber="216" type="distribution">
|
||
<paragraph id="8BB83690FFB2D54FFF4022EDFA95593C" blockId="15.[151,1355,1017,1044]" box="[151,1355,1017,1044]" pageId="15" pageNumber="216">
|
||
<emphasis id="B973EA82FFB2D54FFF4022EDFEFD593C" box="[151,291,1018,1044]" italics="true" pageId="15" pageNumber="216">Occurrence:</emphasis>
|
||
Incised Escarpment Ridge and Northwest
|
||
<collectingRegion id="49C3F872FFB2D54FFCD422EEFC8A593B" box="[771,852,1017,1043]" country="United States of America" name="Florida" pageId="15" pageNumber="216">Florida</collectingRegion>
|
||
Escarpment, Gulf of
|
||
<collectingRegion id="49C3F872FFB2D54FFB9222EEFB43593B" box="[1093,1181,1017,1043]" country="Mexico" name="Mexico" pageId="15" pageNumber="216">Mexico</collectingRegion>
|
||
,
|
||
<quantity id="4CFF9B75FFB2D54FFB7E22EEFA99593C" box="[1193,1351,1017,1044]" metricMagnitude="3" metricUnit="m" metricValue="2.119" metricValueMax="2.178" metricValueMin="2.06" pageId="15" pageNumber="216" unit="m" value="2119.0" valueMax="2178.0" valueMin="2060.0">2060–2178 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFB2D54FFF402555FD24581C" pageId="15" pageNumber="216" type="discussion">
|
||
<paragraph id="8BB83690FFB2D54FFF402555FE815973" blockId="15.[151,1028,1089,1332]" box="[151,351,1089,1115]" pageId="15" pageNumber="216">
|
||
<emphasis id="B973EA82FFB2D54FFF402555FE815973" box="[151,351,1089,1115]" italics="true" pageId="15" pageNumber="216">Images Examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFB2D54FFF102572FD27598C" blockId="15.[151,1028,1089,1332]" pageId="15" pageNumber="216">
|
||
Incised Escarpment Ridge, Gulf of
|
||
<collectingRegion id="49C3F872FFB2D54FFD832572FD725957" box="[596,684,1125,1151]" country="Mexico" name="Mexico" pageId="15" pageNumber="216">Mexico</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE335057FFB2D54FFD602572FCFD59A8" box="[695,803,1125,1152]" degrees="27.35241" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="1" value="27.35241">27.35241</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFB2D54FFCF92572FC7A59A8" box="[814,932,1125,1152]" degrees="85.43421" direction="west" orientation="longitude" pageId="15" pageNumber="216" precision="1" value="-85.43421">-85.43421</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFB2D54FFC782572FBDA59A8" box="[943,1028,1125,1152]" metricMagnitude="3" metricUnit="m" metricValue="2.06" pageId="15" pageNumber="216" unit="m" value="2060.0">2060 m</quantity>
|
||
EX1711_IMG_20171204T172534Z_ROVHD.jpg
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFB2D54FFF1025B9FD2459C3" blockId="15.[151,1028,1089,1332]" pageId="15" pageNumber="216">
|
||
NW
|
||
<collectingRegion id="49C3F872FFB2D54FFF2B25BAFE9359EF" box="[252,333,1197,1223]" country="United States of America" name="Florida" pageId="15" pageNumber="216">Florida</collectingRegion>
|
||
Escarpment, Gulf of
|
||
<collectingRegion id="49C3F872FFB2D54FFDE825BAFD4959EF" box="[575,663,1197,1223]" country="Mexico" name="Mexico" pageId="15" pageNumber="216">Mexico</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE335057FFB2D54FFD7425BAFCD159EF" box="[675,783,1197,1223]" degrees="28.12419" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="1" value="28.12419">28.12419</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFB2D54FFCCD25BAFC5F59E0" box="[794,897,1197,1224]" degrees="86.6575" direction="west" orientation="longitude" pageId="15" pageNumber="216" precision="5" value="-86.6575">-86.6575</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFB2D54FFC5B25BAFC3F59E0" box="[908,993,1197,1224]" metricMagnitude="3" metricUnit="m" metricValue="2.178" pageId="15" pageNumber="216" unit="m" value="2178.0">2178 m</quantity>
|
||
EX1803_IMG_20180426T163320Z_ROVHD.jpg
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFB2D54FFF1025E1FD24581C" blockId="15.[151,1028,1089,1332]" pageId="15" pageNumber="216">
|
||
NW
|
||
<collectingRegion id="49C3F872FFB2D54FFF2B25E2FE935827" box="[252,333,1269,1295]" country="United States of America" name="Florida" pageId="15" pageNumber="216">Florida</collectingRegion>
|
||
Escarpment, Gulf of
|
||
<collectingRegion id="49C3F872FFB2D54FFDE825E2FD495827" box="[575,663,1269,1295]" country="Mexico" name="Mexico" pageId="15" pageNumber="216">Mexico</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE335057FFB2D54FFD7425E2FCD15838" box="[675,783,1269,1296]" degrees="28.12527" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="1" value="28.12527">28.12527</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFB2D54FFCCD25E2FC515838" box="[794,911,1269,1296]" degrees="86.65618" direction="west" orientation="longitude" pageId="15" pageNumber="216" precision="1" value="-86.65618">-86.65618</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFB2D54FFC4D25E2FC315838" box="[922,1007,1269,1296]" metricMagnitude="3" metricUnit="m" metricValue="2.006" pageId="15" pageNumber="216" unit="m" value="2006.0">2006 m</quantity>
|
||
EX1803_IMG_20180426T181615Z_ROVHD.jpg
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |