424 lines
64 KiB
XML
424 lines
64 KiB
XML
<document id="26A0E8B2440A441F468FEA045C5D5275" ID-CLB-Dataset="22829" ID-DOI="10.11646/zootaxa.4766.2.1" ID-GBIF-Dataset="7fa4a4ce-5db2-4d06-adeb-db21b0574cef" ID-ISSN="1175-5334" ID-Zenodo-Dep="3764018" ID-ZooBank="urn:lsid:zoobank.org:pub:B47DC09C-181A-4DFE-B415-770AFFC11BD3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587671180438" checkinUser="carolina" docAuthor="Mah, Christopher L." docDate="2020" docId="03AE8786FFAED557FF402793FC1259F5" docLanguage="en" docName="zootaxa.4766.2.1.pdf.imf" docOrigin="Zootaxa 4766 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Evoplosoma nizinskiae Mah 2020, n. sp." docType="treatment" docVersion="8" lastPageNumber="224" masterDocId="FF97FFFEFFBDD540FFD72117FFDE5D28" masterDocTitle="New species, occurrence records and observations of predation by deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA ship Okeanos Explorer" masterLastPageNumber="260" masterPageNumber="201" pageNumber="220" updateTime="1702001939344" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="91A594D65D53FBE9C85D3B03C42E8494">New species, occurrence records and observations of predation by deep-sea Asteroidea (Echinodermata) from the North Atlantic by NOAA ship Okeanos Explorer</mods:title>
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<mods:name id="62F0D3156733808ACD74AF5791328EA7" type="personal">
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<mods:namePart id="178978AF370875677864082CDD8BA4AA">Mah, Christopher L.</mods:namePart>
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<mods:title id="24BD3F18AF93A1EF2BC4778887FE5C61">Zootaxa</mods:title>
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<mods:date id="E8532FBD1BDC869B1139A702FD27D2F1">2020</mods:date>
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<mods:number id="A90899331CA6B45F2029429B79CAD2A5">2020-04-20</mods:number>
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<mods:number id="9BDDF0F12EE35022F565BF7A85520E25">4766</mods:number>
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<treatment id="03AE8786FFAED557FF402793FC1259F5" ID-DOI="http://doi.org/10.5281/zenodo.3803733" ID-GBIF-Taxon="163737321" ID-Zenodo-Dep="3803733" LSID="urn:lsid:plazi:treatment:03AE8786FFAED557FF402793FC1259F5" httpUri="http://treatment.plazi.org/id/03AE8786FFAED557FF402793FC1259F5" lastPageId="23" lastPageNumber="224" pageId="19" pageNumber="220">
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<subSubSection id="C31D651BFFAED553FF402793FE295BB7" box="[151,503,1668,1695]" pageId="19" pageNumber="220" type="nomenclature">
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<paragraph id="8BB83690FFAED553FF402793FE295BB7" blockId="19.[151,503,1668,1731]" box="[151,503,1668,1695]" pageId="19" pageNumber="220">
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<heading id="D0F081FCFFAED553FF402793FE295BB7" bold="true" box="[151,503,1668,1695]" fontSize="10" level="2" pageId="19" pageNumber="220" reason="2">
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<emphasis id="B973EA82FFAED553FF402793FE295BB7" bold="true" box="[151,503,1668,1695]" pageId="19" pageNumber="220">
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<taxonomicName id="4C074D13FFAED553FF402793FE735BB7" ID-CoL="848S2" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[151,429,1668,1695]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="19" pageNumber="220" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
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<emphasis id="B973EA82FFAED553FF402793FE735BB7" bold="true" box="[151,429,1668,1695]" italics="true" pageId="19" pageNumber="220">Evoplosoma nizinskiae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24057F9FFAED553FE632792FE295BB7" box="[436,503,1669,1695]" pageId="19" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFAED553FF4027BEFE585BEB" box="[151,390,1705,1731]" pageId="19" pageNumber="220" type="nomenclature">
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<paragraph id="8BB83690FFAED553FF4027BEFE585BEB" blockId="19.[151,503,1668,1731]" box="[151,390,1705,1731]" pageId="19" pageNumber="220">
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<figureCitation id="133C2A15FFAED553FF4027BEFF2A5BEB" box="[151,244,1705,1731]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="19" pageNumber="220">Figure 9</figureCitation>
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A–F, 10A–D
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFAED553FF4027E6FD355A7B" pageId="19" pageNumber="220" type="etymology">
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<paragraph id="8BB83690FFAED553FF4027E6FED15A23" blockId="19.[151,1436,1777,1875]" box="[151,271,1777,1803]" pageId="19" pageNumber="220">
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<emphasis id="B973EA82FFAED553FF4027E6FED15A23" box="[151,271,1777,1803]" italics="true" pageId="19" pageNumber="220">Etymology</emphasis>
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</paragraph>
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<paragraph id="8BB83690FFAED553FF102602FD355A7B" blockId="19.[151,1436,1777,1875]" pageId="19" pageNumber="220">This species is named for my colleague Dr. Martha Nizinski in honor of her tireless contributions to deep-sea coral ecology and NOAA’s Ocean Explorer program.</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFAED554FF402696FA535DFF" lastPageId="20" lastPageNumber="221" pageId="19" pageNumber="220" type="diagnosis">
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<paragraph id="8BB83690FFAED553FF402696FED75AB2" blockId="19.[151,1437,1921,2019]" box="[151,265,1921,1946]" pageId="19" pageNumber="220">
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<emphasis id="B973EA82FFAED553FF402696FED75AB2" box="[151,265,1921,1946]" italics="true" pageId="19" pageNumber="220">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BB83690FFAED553FF1026B2FA435ACB" blockId="19.[151,1437,1921,2019]" pageId="19" pageNumber="220">This species is characterized by the lack of primary spines on the abactinal plates, the evenly distributed granules, the absence of primary spines from interradial superomarginal plates-but present on distal superomarginal</paragraph>
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<paragraph id="8BB83690FFA9D554FF40218EFA535DFF" blockId="20.[151,1437,153,215]" pageId="20" pageNumber="221">plates, mostly bare superomarginal plates save for coarse, widely spaced granules, prominent elongate subambulacral pedicellariae with three to four teeth per valve on each adambulacral plate and 7 to 10 leaflike furrow spines.</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFA9D554FF402012FC065C37" box="[151,984,260,288]" pageId="20" pageNumber="221" type="distribution">
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<paragraph id="8BB83690FFA9D554FF402012FC065C37" blockId="20.[151,984,260,288]" box="[151,984,260,288]" pageId="20" pageNumber="221">
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<emphasis id="B973EA82FFA9D554FF402012FEFD5C37" box="[151,291,261,287]" italics="true" pageId="20" pageNumber="221">Occurrence:</emphasis>
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Gulf of Mexico, Caribbean Sea (
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<collectingCountry id="F3107600FFA9D554FD4F2012FCFD5C08" box="[664,803,261,288]" name="Puerto Rico" pageId="20" pageNumber="221">Puerto Rico</collectingCountry>
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),
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<quantity id="4CFF9B75FFA9D554FCE12013FC0A5C37" box="[822,980,260,287]" metricMagnitude="3" metricUnit="m" metricValue="2.293" metricValueMax="2.549" metricValueMin="2.037" pageId="20" pageNumber="221" unit="m" value="2293.0" valueMax="2549.0" valueMin="2037.0">2037–2549 m</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31D651BFFA9D555FF402059FCA85DD3" lastPageId="21" lastPageNumber="222" pageId="20" pageNumber="221" type="description">
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<paragraph id="8BB83690FFA9D554FF402059FEC55C4F" blockId="20.[151,1437,334,2015]" box="[151,283,334,359]" pageId="20" pageNumber="221">
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<emphasis id="B973EA82FFA9D554FF402059FEC55C4F" box="[151,283,334,359]" italics="true" pageId="20" pageNumber="221">Description</emphasis>
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</paragraph>
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<paragraph id="8BB83690FFA9D554FF102066FCCE5C87" blockId="20.[151,1437,334,2015]" pageId="20" pageNumber="221">
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Body strongly stellate (R/r=4.42), arms elongate with pointed arm tip, interradial arcs weakly curved to straight. Four arms (five on other observed specimens) (
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<figureCitation id="133C2A15FFA9D554FD7E2082FCDD5C87" box="[681,771,405,431]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig. 9A</figureCitation>
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).
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</paragraph>
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<paragraph id="8BB83690FFA9D554FF1020AEFD655E13" blockId="20.[151,1437,334,2015]" pageId="20" pageNumber="221">
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Abactinal surface pulpy, shows strongly convex surface but with some folded and sagged regions, suggesting inflation when alive (
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<figureCitation id="133C2A15FFA9D554FE5120CAFE2D5CDF" box="[390,499,477,503]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig 9A,B</figureCitation>
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). Abactinal plates extend from disk to arm tip. Abactinal plates round to irregular in shape, closely abutted continuous surface. Plate boundaries weakly defined. Surface covered by moderately coarse, widely spaced but evenly distributed granules ranging from round surface to pointed tips, approximately one to four, mostly one to three along a 1.0 mm line. Granules smaller distally becoming larger proximally on disk with some enlarged granules
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<quantity id="4CFF9B75FFA9D554FE73237BFE275FAF" box="[420,505,620,647]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="221" unit="mm" value="0.5">0.5 mm</quantity>
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width. Individual plate surfaces vary from completely 0 to 25 per plate. Plates lacking granules bare and smooth. Where granules are present, they form a periphery, approximately three to five per side with one to eight present centrally. The more granules present, the smaller they are. Pedicellariae present on disk, relatively few with approximately one per interradial region (four on this individual), approximately 1.0 mm in length, elongate, jaw-like with three to five teeth on each valve. Madreporite round, weakly pentagonal, weakly convex with a shallow channel present around it.
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</paragraph>
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<paragraph id="8BB83690FFA9D554FF102252FC5D58EB" blockId="20.[151,1437,334,2015]" pageId="20" pageNumber="221">
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Superomarginals and inferomarginals 54–55 per interradius, with a lateral facing (
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<figureCitation id="133C2A15FFA9D554FBA02252FB295E77" box="[1143,1271,837,863]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig. 9A, C</figureCitation>
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). All marginal plates with approximately 40–60 pointed granules forming the plate periphery, widely spaced, approximately 10–12 per side. Some superomarginals irregular in shape between fully formed superomarginal plates. Superomarginals larger and elongate interradially, (approximately ten plates) becoming wider (i.e. narrower) and more strongly convex distally along the arms. Interradial superomarginals, approximately ten to twelve plates lacking primary spines, each of these superomarginals with 20–30, mostly about 25 coarse, bullet-like, widely spaced granules. Central plate granules larger, smaller granules approximately 20% of the larger granules and present peripherally around the central granules. Superomarginals along the arms (
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<figureCitation id="133C2A15FFA9D554FD232556FC965973" box="[756,840,1089,1115]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig. 9E</figureCitation>
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) mostly with a single prominent, bullet-shaped spine, (approximately 1–2.0 mm in length) remainder of superomarginal surface mostly bare, smooth, forming a continuous series along the arm (
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<figureCitation id="133C2A15FFA9D554FE68259EFDC7598B" box="[447,537,1161,1187]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig. 9C</figureCitation>
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). Most distalmost superomarginals with zero to eight small, round granules on plate surface around base of each spine. Inferomarginals with bald upper lateral plate surface, but with oral surface covered by 20 to 30 coarse, round to cylindrical granules with some appearing as incipient spines. Inferomarginal plates mostly with similar spines to those on superomarginals, with spines absent on more inferomarginals than superomarginals, especially distally. Inferomarginals present basal own arm. Where spines absent, coarse, widely spaced granules, 10–30 per inferomarginal plate surface. Shallow fasciolar grooves between superomarginal and inferomarginal plates. Both superomarginal and inferomarginal interradial plates with a relatively large (1.0–2.0 mm) elongate pedicellariae, similar to those on abactinal surface, on the lateral surface. Terminal plates triangular in shape, widely spaced granules present on terminal plate surface.
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</paragraph>
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<paragraph id="8BB83690FFA9D554FF1024DAFBB15BB3" blockId="20.[151,1437,334,2015]" pageId="20" pageNumber="221">
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Actinal surface with only two to three series, mostly limited to disk and basal arm regions (
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<figureCitation id="133C2A15FFA9D554FB0C24DAFAE858CF" box="[1243,1334,1485,1511]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig. 9D</figureCitation>
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). Actinal plates with quadrate to irregularly shaped plates. Each plate surface with 2 to 20 pointed, evenly spaced granules. Most plates with 4 to 14 peripheral granules with 2 to 6 identical to large sized granules on each central actinal plate surface. Large, prominent pedicellariae, 6 to 9 per interradius, with some actinal plates bearing a single large pedicellariae with a bare smooth region around it. Pedicellariae identical to those on other surfaces. Peripheral granules around these “pedicellariae actinal plates” more widely spaced than other actinal plates.
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</paragraph>
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<paragraph id="8BB83690FFA9D555FF1027B2FED75DFF" blockId="20.[151,1437,334,2015]" lastBlockId="21.[151,1437,153,251]" lastPageId="21" lastPageNumber="222" pageId="20" pageNumber="221">
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Furrow spines 8 to 10 per plate arranged straight to weakly curved (
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<figureCitation id="133C2A15FFA9D554FC1927B2FBFA5B97" box="[974,1060,1701,1727]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="20" pageNumber="221">Fig. 9F</figureCitation>
|
||
). Eight furrow spines proximally becoming ten distally. Furrow spines flattened, leaf-like, angular in cross section. Furrow spines interdigitate along the tube foot groove. Adambulacral plates on disk with large elongate pedicellariae, especially on proximal adambulacrals. Along arm and more distally adambulacral plates have both a prominent pedicellariae but also a single, blunt spine similar in size to the pedicellariae. Adambulacral spines becoming larger and more prominent than the pedicellariae on distalmost plates. Widely space pointed granules present on adambulacral plate adjacent to actinal plates side. Some adambulacral plates lacking spines and/or granules, surface covered by pointed to round granules present. Oral plate furrow spines, eight with central spine projecting into mouth, projecting into mouth. Spines narrow and leaf-like, triangular in cross section. Approximately seven or eight paired multi-spined granules present along either side of fossae on oral plate. Five to eight multi-spined granules present around periphery of each oral plate side.
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF1021F6FCA85DD3" blockId="21.[151,1437,153,251]" box="[199,886,225,251]" pageId="21" pageNumber="222">
|
||
In life, disk is white, arms a light orange to tan (
|
||
<figureCitation id="133C2A15FFA8D555FD3621F6FCEA5DD3" box="[737,820,225,251]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="21" pageNumber="222">Figs 10</figureCitation>
|
||
C–F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFA8D555FF40203EFA985E8F" pageId="21" pageNumber="222" type="discussion">
|
||
<paragraph id="8BB83690FFA8D555FF40203EFD4F5C6B" blockId="21.[151,1437,297,935]" box="[151,657,297,323]" pageId="21" pageNumber="222">
|
||
<emphasis id="B973EA82FFA8D555FF40203EFE785C6B" box="[151,422,297,323]" italics="true" pageId="21" pageNumber="222">Comparisons with other</emphasis>
|
||
<taxonomicName id="4C074D13FFA8D555FE7A203EFDE75C6B" box="[429,569,297,323]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="genus">Evoplosoma</taxonomicName>
|
||
<emphasis id="B973EA82FFA8D555FD97203DFD4F5C6B" box="[576,657,298,323]" italics="true" pageId="21" pageNumber="222">species</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF10205AFD175F4B" blockId="21.[151,1437,297,935]" pageId="21" pageNumber="222">
|
||
This species invites immediate comparison with
|
||
<taxonomicName id="4C074D13FFA8D555FD2E205AFBAC5C4F" authority="Downey 1982" authorityName="Downey" authorityYear="1982" box="[761,1138,333,359]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="virgo">
|
||
<emphasis id="B973EA82FFA8D555FD2E205AFC195C4F" box="[761,967,333,359]" italics="true" pageId="21" pageNumber="222">Evoplosoma virgo</emphasis>
|
||
Downey 1982
|
||
</taxonomicName>
|
||
which also lacks primary spines on the abactinal and the interradial superomarginal plate surface. However
|
||
<taxonomicName id="4C074D13FFA8D555FB942066FACE5CA3" authorityName="Downey" authorityYear="1982" box="[1091,1296,369,395]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="virgo">
|
||
<emphasis id="B973EA82FFA8D555FB942066FACE5CA3" box="[1091,1296,369,395]" italics="true" pageId="21" pageNumber="222">Evoplosoma virgo</emphasis>
|
||
</taxonomicName>
|
||
has a dense covering of pointed granules covering the marginal plate surfaces, whereas
|
||
<taxonomicName id="4C074D13FFA8D555FC222081FB585C87" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[1013,1158,405,431]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFA8D555FC222081FB585C87" box="[1013,1158,405,431]" italics="true" pageId="21" pageNumber="222">E. nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFA8D555FB592082FB125C87" box="[1166,1228,405,431]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
|
||
has a mostly bare and smooth superomarginal surface, save for widely spaced granules on the lateral edge. Then dense abactinal granulation in
|
||
<taxonomicName id="4C074D13FFA8D555FEEA20C9FE495CDF" authorityName="Downey" authorityYear="1982" box="[317,407,478,503]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="virgo">
|
||
<emphasis id="B973EA82FFA8D555FEEA20C9FE495CDF" box="[317,407,478,503]" italics="true" pageId="21" pageNumber="222">E. virgo</emphasis>
|
||
</taxonomicName>
|
||
occurs in discrete, round to polygonal patches versus the more evenly distributed granulation observed in
|
||
<taxonomicName id="4C074D13FFA8D555FE832315FE3D5F33" authorityName="Mah" authorityYear="2020" box="[340,483,513,539]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskiae">
|
||
<emphasis id="B973EA82FFA8D555FE832315FE3D5F33" box="[340,483,513,539]" italics="true" pageId="21" pageNumber="222">E. nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
. Furrow spines in
|
||
<taxonomicName id="4C074D13FFA8D555FD612315FCD15F33" authorityName="Downey" authorityYear="1982" box="[694,783,514,539]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="virgo">
|
||
<emphasis id="B973EA82FFA8D555FD612315FCD15F33" box="[694,783,514,539]" italics="true" pageId="21" pageNumber="222">E. virgo</emphasis>
|
||
</taxonomicName>
|
||
are three to five times as thick as those in
|
||
<taxonomicName id="4C074D13FFA8D555FB272315FAAA5F33" box="[1264,1396,513,539]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskia">
|
||
<emphasis id="B973EA82FFA8D555FB272315FAAA5F33" box="[1264,1396,513,539]" italics="true" pageId="21" pageNumber="222">E. nizinskia</emphasis>
|
||
</taxonomicName>
|
||
e n. sp.
|
||
<taxonomicName id="4C074D13FFA8D555FF692332FE615F17" authorityName="Mah" authorityYear="2020" box="[190,447,549,575]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskiae">
|
||
<emphasis id="B973EA82FFA8D555FF692332FE615F17" box="[190,447,549,575]" italics="true" pageId="21" pageNumber="222">Evoplosoma nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
is also more strongly stellate (R/r=4.42) with elongate arms versus
|
||
<taxonomicName id="4C074D13FFA8D555FB612331FAD15F17" authorityName="Downey" authorityYear="1982" box="[1206,1295,550,575]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="virgo">
|
||
<emphasis id="B973EA82FFA8D555FB612331FAD15F17" box="[1206,1295,550,575]" italics="true" pageId="21" pageNumber="222">E. virgo</emphasis>
|
||
</taxonomicName>
|
||
with a more weakly stellate form (R/r=3.5) and a broader disk.
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF10237AFB7F5FE7" blockId="21.[151,1437,297,935]" pageId="21" pageNumber="222">However, the two species share similar pedicellariae morphology with both species displaying these pedicellariae on their adambulacral plates adjacent to the furrow spination. Both species also lack superomarginal spination interradially but possess a primary spine on more distalmost superomarginals along the arm.</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF1023CEFBFC5E77" blockId="21.[151,1437,297,935]" pageId="21" pageNumber="222">
|
||
<taxonomicName id="4C074D13FFA8D555FF1023CEFE195FDB" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[199,455,729,755]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFA8D555FF1023CEFE195FDB" box="[199,455,729,755]" italics="true" pageId="21" pageNumber="222">Evoplosoma nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFA8D555FE1A23CEFDD65FDB" box="[461,520,729,755]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
|
||
is distinguished from the other Atlantic
|
||
<taxonomicName id="4C074D13FFA8D555FC1423CEFB935FDB" box="[963,1101,729,755]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B973EA82FFA8D555FC1423CEFB935FDB" box="[963,1101,729,755]" italics="true" pageId="21" pageNumber="222">Evoplosoma</emphasis>
|
||
</taxonomicName>
|
||
spp. by the absence of numerous, thick primary spines present on the abactinal and actinal plates (
|
||
<figureCitation id="133C2A15FFA8D555FC7D23EAFBF95E3F" box="[938,1063,765,791]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="21" pageNumber="222">Fig. 9A, B</figureCitation>
|
||
). Both
|
||
<taxonomicName id="4C074D13FFA8D555FBAA23EAFAB55E3F" authorityName="Mah" authorityYear="2015" box="[1149,1387,765,791]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="watlingi">
|
||
<emphasis id="B973EA82FFA8D555FBAA23EAFAB55E3F" box="[1149,1387,765,791]" italics="true" pageId="21" pageNumber="222">Evoplosoma watlingi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C074D13FFA8D555FF402236FEA25E13" authorityName="Downey" authorityYear="1981" box="[151,380,801,827]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="scorpio">
|
||
<emphasis id="B973EA82FFA8D555FF402236FEA25E13" box="[151,380,801,827]" italics="true" pageId="21" pageNumber="222">Evoplosoma scorpio</emphasis>
|
||
</taxonomicName>
|
||
also possess a full series of prominent superomarginal spines from arm tip to arm tip, whereas
|
||
<taxonomicName id="4C074D13FFA8D555FF402252FE475E77" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[151,409,837,863]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFA8D555FF402252FE475E77" box="[151,409,837,863]" italics="true" pageId="21" pageNumber="222">Evoplosoma nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFA8D555FE772252FE025E77" box="[416,476,837,863]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
|
||
lacks superomarginal spines interradially (
|
||
<figureCitation id="133C2A15FFA8D555FC6A2252FBCB5E77" box="[957,1045,837,863]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="21" pageNumber="222">Fig. 9C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF10227EFA985E8F" blockId="21.[151,1437,297,935]" pageId="21" pageNumber="222">
|
||
The unidentified
|
||
<taxonomicName id="4C074D13FFA8D555FE5B227EFDC85EAB" box="[396,534,873,899]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="B973EA82FFA8D555FE5B227EFDC85EAB" box="[396,534,873,899]" italics="true" pageId="21" pageNumber="222">Evoplosoma</emphasis>
|
||
</taxonomicName>
|
||
sp. 1 listed herein, also shows strong similarity with
|
||
<taxonomicName id="4C074D13FFA8D555FBAC227EFAA05EAB" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[1147,1406,873,899]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="21" pageNumber="222" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFA8D555FBAC227EFAA05EAB" box="[1147,1406,873,899]" italics="true" pageId="21" pageNumber="222">Evoplosoma nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFA8D555FA50227EFF675E8F" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
|
||
, in showing a similar body shape, and color, but differs in having a full series of superomarginal spines.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFA8D555FF4022C2FC8F5AF7" pageId="21" pageNumber="222" type="description">
|
||
<paragraph id="8BB83690FFA8D555FF4022C2FE5C5EC7" blockId="21.[151,1437,981,2015]" box="[151,386,981,1007]" pageId="21" pageNumber="222">
|
||
<emphasis id="B973EA82FFA8D555FF4022C2FE5C5EC7" box="[151,386,981,1007]" italics="true" pageId="21" pageNumber="222">Juvenile morphology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF1022EEFDB4591F" blockId="21.[151,1437,981,2015]" pageId="21" pageNumber="222">Body form stellate (R/r=2.25) but more weakly than the larger individual. Arms triangular but less elongate. Interradial arcs weakly curved to straight.</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF102556FC3059C3" blockId="21.[151,1437,981,2015]" pageId="21" pageNumber="222">
|
||
Abactinal surface composed of primarily round plates (
|
||
<figureCitation id="133C2A15FFA8D555FCE32556FC435973" box="[820,925,1089,1115]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="21" pageNumber="222">Fig. 10A</figureCitation>
|
||
), extending to arm tip. Plates becoming more irregular in shape distally, especially near arm tip. Each plate with four to 20 small round granules forming periphery of each plate. Central region devoid of granules, surface bare and smooth. Plates weakly convex becoming more flat distally. Anus surrounded by five triangular, blunt spines. Madreporite weakly convex, with four sides, rounded surrounded by five to six plates. Pedicellariae absent from abactinal surface.
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF1025E2FDFD5B7B" blockId="21.[151,1437,981,2015]" pageId="21" pageNumber="222">At R=1.8, 19–21 superomarginal plates per interradius (arm tip to arm tip), 22–24 inferomarginal plates per interradius. Marginal plates blocky, with distinct angular lateral surface. Interradial superomarginals with no spines, but two proximal superomarginals on arm with distinct, blunt, bullet-shaped spines. on two arms. Remaining superomarginals without larger primary spine, most with 1 to 10 (mostly 6 to 8) widely spaced spiny tipped coarse granules present on lateral and abactinolateral surface. Central granules coarse and approximately twice as large as others on plate surface. Remainder of plate smooth and bare. Inferomarginal plate surface covered by 10–20 pointed granules, widely spaced in linear patterns. Round, bald region on ventral inferomarginal plate surface. Periphery of each marginal with 40–60 widely spaced, irregular to round granules with spiny tips, approximately 10–12 per side. Terminal plate triangular with one to three pointed but blunt spines projecting from each arm tip. Pedicellariae absent from marginal plate surface.</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF10274AFEF05BCB" blockId="21.[151,1437,981,2015]" pageId="21" pageNumber="222">Actinal region with two series, limited to disk. Actinal plates quadrate to irregular in shape. Pedicellariae absent from actinal surface. Actinal plate surface covered by 2 to 16 pointed granules, widely spaced. A single elongate pedicellariae with three to four teeth per valve present on two inter radii. The remaining two actinal regions lacking pedicellariae.</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF1027FAFA4C5A93" blockId="21.[151,1437,981,2015]" pageId="21" pageNumber="222">
|
||
Furrow spines six to seven (
|
||
<figureCitation id="133C2A15FFA8D555FDC627FAFDA45A2F" box="[529,634,1773,1799]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="21" pageNumber="222">Fig. 10B</figureCitation>
|
||
), flattened, with shortest spines farthest from center. Furrow spine in straight to weakly curved series. Proximalmost three or four adambulacral plates with prominent elongate single pedicellariae on surface. Distalmost remaining adambulacral plate surface with prominent, single blunt spines. Adambulacral plate surface with four to seven pointed granules. Oral plates with eight furrow spines identical to other furrow spines but with a single angular spine projecting into mouth. Oral plate surface each with five pairs triangular granules flanking each side of central fossae. Other short thorny granules irregularly present on oral plate surface.
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFA8D555FF1026D2FC8F5AF7" blockId="21.[151,1437,981,2015]" box="[199,849,1989,2015]" pageId="21" pageNumber="222">Color in life almost white but with light orange arm color.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF786618FFABD556FF40267AFE1F5AD9" ID-DOI="http://doi.org/10.5281/zenodo.3764036" ID-Zenodo-Dep="3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="22" pageNumber="223" startId="22.[151,250,1901,1926]" targetBox="[194,1397,192,1868]" targetPageId="22">
|
||
<paragraph id="8BB83690FFABD556FF40267AFE1F5AD9" blockId="22.[151,1437,1901,2034]" pageId="22" pageNumber="223">
|
||
<emphasis id="B973EA82FFABD556FF40267AFD875AAE" bold="true" box="[151,601,1901,1926]" pageId="22" pageNumber="223">
|
||
FIGURE. 9.
|
||
<taxonomicName id="4C074D13FFABD556FEF4267AFDC85AAE" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[291,534,1901,1926]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="22" pageNumber="223" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFABD556FEF4267AFDC85AAE" bold="true" box="[291,534,1901,1926]" italics="true" pageId="22" pageNumber="223">Evoplosoma nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFABD556FDCA2679FD875AAE" box="[541,601,1902,1926]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<typeStatus id="54BC8832FFABD556FDB6267AFD1C5AAE" box="[609,706,1901,1926]" pageId="22" pageNumber="223" type="holotype">Holotype</typeStatus>
|
||
, USNM 1507289, scale bar=1.0 cm.
|
||
<emphasis id="B973EA82FFABD556FB92267AFB835AAE" bold="true" box="[1093,1117,1901,1926]" pageId="22" pageNumber="223">A.</emphasis>
|
||
Abactinal surface
|
||
<emphasis id="B973EA82FFABD556FAF52679FAE75AAE" bold="true" box="[1314,1337,1902,1926]" pageId="22" pageNumber="223">B.</emphasis>
|
||
Close up showing surface detail. Scale bar=3.0 mm.
|
||
<emphasis id="B973EA82FFABD556FDB62686FDA75A82" bold="true" box="[609,633,1937,1962]" pageId="22" pageNumber="223">C.</emphasis>
|
||
Lateral surface showing marginal plates surface. Scale bar=3.0 mm.
|
||
<emphasis id="B973EA82FFABD556FA9A2685FAB85A82" bold="true" box="[1357,1382,1938,1962]" pageId="22" pageNumber="223">D.</emphasis>
|
||
Oral surface. scale bar=1.0 cm.
|
||
<emphasis id="B973EA82FFABD556FE7926A1FE1B5AE6" bold="true" box="[430,453,1974,1998]" pageId="22" pageNumber="223">E.</emphasis>
|
||
Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm.
|
||
<emphasis id="B973EA82FFABD556FAE126A1FA975AE6" bold="true" box="[1334,1353,1974,1998]" pageId="22" pageNumber="223">F.</emphasis>
|
||
Furrow spination. Scale bar=3.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31D651BFFAAD557FF40218EFE3D5CDB" pageId="23" pageNumber="224" type="discussion">
|
||
<paragraph id="8BB83690FFAAD557FF40218EFE0F5D9B" blockId="23.[151,1436,153,500]" box="[151,465,153,179]" pageId="23" pageNumber="224">
|
||
<emphasis id="B973EA82FFAAD557FF40218EFE0F5D9B" box="[151,465,153,179]" italics="true" pageId="23" pageNumber="224">Comments on Juvenile form</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF1021AAFE3D5CDB" blockId="23.[151,1436,153,500]" pageId="23" pageNumber="224">
|
||
The
|
||
<typeStatus id="54BC8832FFAAD557FF2921AAFEBF5DFF" box="[254,353,189,215]" pageId="23" pageNumber="224" type="paratype">paratype</typeStatus>
|
||
, USNM 1550630 is smaller and shows several differences from the
|
||
<typeStatus id="54BC8832FFAAD557FB5F21AAFB335DFF" box="[1160,1261,189,215]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
|
||
, but characters strongly suggest similarity and it is regarded herein as a juvenile of
|
||
<taxonomicName id="4C074D13FFAAD557FC7021F7FB745DD2" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[935,1194,224,250]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="23" pageNumber="224" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFAAD557FC7021F7FB745DD2" box="[935,1194,224,250]" italics="true" pageId="23" pageNumber="224">Evoplosoma nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFAAD557FB6421F6FB2F5DD3" box="[1203,1265,225,251]" pageId="23" pageNumber="224" rank="species">n. sp.</taxonomicNameLabel>
|
||
If correct, this provides a rare opportunity to examine the growth in a deep-sea goniasterid species. It showed a significantly less stellate body from (R/r=2.25) versus the more stellate body form (R/r=4.42) at R=6.2 as well as triangular versus the elongate arms in the
|
||
<typeStatus id="54BC8832FFAAD557FEAA205CFE3D5C4D" box="[381,483,331,357]" pageId="23" pageNumber="224" type="holotype">holotype</typeStatus>
|
||
. The abactinal plates are round to polygonal in outline (
|
||
<figureCitation id="133C2A15FFAAD557FB9F205CFB715C4D" box="[1096,1199,331,357]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="23" pageNumber="224">Fig. 10A</figureCitation>
|
||
), and although many, if not most, of the plates are bare, several plates show single to few granules suggesting incipient development of the granulation that is seen in the larger specimen (
|
||
<figureCitation id="133C2A15FFAAD557FD0B2085FCE95C84" box="[732,823,402,428]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="23" pageNumber="224">Fig. 9A</figureCitation>
|
||
). Furrow spine number also shows some overlap with the
|
||
<typeStatus id="54BC8832FFAAD557FF1620A1FEFF5CF8" box="[193,289,438,464]" pageId="23" pageNumber="224" type="paratype">paratype</typeStatus>
|
||
showing primarily six or seven furrow spines (
|
||
<figureCitation id="133C2A15FFAAD557FCEE20A2FC405CE7" box="[825,926,437,463]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="23" pageNumber="224">Fig. 10B</figureCitation>
|
||
) and the larger specimen displaying seven to ten, primarily eight (
|
||
<figureCitation id="133C2A15FFAAD557FEA920CEFE095CDB" box="[382,471,473,499]" captionStart="FIGURE" captionStartId="22.[151,250,1901,1926]" captionTargetBox="[194,1397,192,1868]" captionTargetId="figure@22.[185,1403,181,1876]" captionTargetPageId="22" captionText="FIGURE. 9. Evoplosoma nizinskiae n. sp. Holotype, USNM 1507289, scale bar=1.0 cm. A. Abactinal surface B. Close up showing surface detail. Scale bar=3.0 mm. C.Lateral surface showing marginal plates surface. Scale bar=3.0 mm. D. Oral surface. scale bar=1.0 cm. E. Close up on abactinal and lateral surface distalmost arm region. Scale bar=3.0 mm. F. Furrow spination. Scale bar=3.0 mm ." figureDoi="http://doi.org/10.5281/zenodo.3764036" httpUri="https://zenodo.org/record/3764036/files/figure.png" pageId="23" pageNumber="224">Fig. 9B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFAAD557FF40230BFCE95F8A" pageId="23" pageNumber="224" type="biology_ecology">
|
||
<paragraph id="8BB83690FFAAD557FF40230BFE515F1E" blockId="23.[151,1436,540,675]" box="[151,399,540,566]" pageId="23" pageNumber="224">
|
||
<emphasis id="B973EA82FFAAD557FF40230BFE515F1E" box="[151,399,540,566]" italics="true" pageId="23" pageNumber="224">Feeding Observations</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF102357FCE95F8A" blockId="23.[151,1436,540,675]" pageId="23" pageNumber="224">
|
||
Of the four
|
||
<emphasis id="B973EA82FFAAD557FE9B2356FE4C5F72" box="[332,402,577,602]" italics="true" pageId="23" pageNumber="224">in situ</emphasis>
|
||
observations of this species (
|
||
<figureCitation id="133C2A15FFAAD557FD092357FCF05F72" box="[734,814,576,602]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="23" pageNumber="224">Fig. 10</figureCitation>
|
||
C–E), three showed
|
||
<taxonomicName id="4C074D13FFAAD557FBC12356FB7B5F72" authority="Mah, 2020" authorityName="Mah" authorityYear="2020" box="[1046,1189,576,602]" class="Asteroidea" family="Goniasteridae" genus="Evoplosoma" kingdom="Animalia" order="Valvatida" pageId="23" pageNumber="224" phylum="Echinodermata" rank="species" species="nizinskiae" status="sp. nov.">
|
||
<emphasis id="B973EA82FFAAD557FBC12356FB7B5F72" box="[1046,1189,576,602]" italics="true" pageId="23" pageNumber="224">E. nizinskiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24057F9FFAAD557FB7A2357FB375F72" box="[1197,1257,576,602]" pageId="23" pageNumber="224" rank="species">n. sp.</taxonomicNameLabel>
|
||
feeding. In two instances, (
|
||
<figureCitation id="133C2A15FFAAD557FEC32373FE495F56" box="[276,407,612,638]" captionStart="FIGURE 10" captionStartId="24.[151,250,1842,1867]" captionTargetBox="[159,1430,189,1812]" captionTargetId="figure@24.[151,1436,181,1816]" captionTargetPageId="24" captionText="FIGURE 10. Evoplosoma nizinskiae & Evoplosoma spp. Small specimen, USNM 1550630 unless otherwise noted. A. Abactinal plates. Scale=1.0 mm. B. Furrow spination differences. Scale=1.0 mm. C. In situ feeding observation. R=1.8 cm. D. E. nizinskiae feeding observation West Florida Escarpment, Gulf of Mexico. 2181 m. E. Evoplosoma sp. 1, Blake Ridge, North Atlantic, 3388 m. F. Evoplosoma sp. 2, Mona Canyon East, Puerto Rico, 2393 m." figureDoi="http://doi.org/10.5281/zenodo.3764038" httpUri="https://zenodo.org/record/3764038/files/figure.png" pageId="23" pageNumber="224">Fig. 10C, E</figureCitation>
|
||
) individuals had climbed up onto higher branches and extended their pyloric stomachs onto the polyps of their isidid octocoral (aka bamboo coral) prey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFAAD557FF4023C6FCB25EEA" pageId="23" pageNumber="224" type="discussion">
|
||
<paragraph id="8BB83690FFAAD557FF4023C6FE815FC2" blockId="23.[151,1034,720,962]" box="[151,351,720,746]" pageId="23" pageNumber="224">
|
||
<emphasis id="B973EA82FFAAD557FF4023C6FE815FC2" box="[151,351,720,746]" italics="true" pageId="23" pageNumber="224">Images Examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF1023E3FD275E1A" blockId="23.[151,1034,720,962]" pageId="23" pageNumber="224">
|
||
Smooth Escarpment Ridge, Gulf of
|
||
<collectingRegion id="49C3F872FFAAD557FD8D23E3FD6D5E26" box="[602,691,756,782]" country="Mexico" name="Mexico" pageId="23" pageNumber="224">Mexico</collectingRegion>
|
||
.
|
||
<geoCoordinate id="EE335057FFAAD557FD6923E3FCF45E26" box="[702,810,756,782]" degrees="28.00404" direction="north" orientation="latitude" pageId="23" pageNumber="224" precision="1" value="28.00404">28.00404</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFAAD557FCE223E3FC745E26" box="[821,938,756,782]" degrees="86.43931" direction="west" orientation="longitude" pageId="23" pageNumber="224" precision="1" value="-86.43931">-86.43931</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFAAD557FC6223E4FBD45E26" box="[949,1034,755,782]" metricMagnitude="3" metricUnit="m" metricValue="2.054" pageId="23" pageNumber="224" unit="m" value="2054.0">2054 m</quantity>
|
||
EX1711_IMG_20171205T154320Z_ROVHD.jpg
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF10222CFC8B5E7E" blockId="23.[151,1034,720,962]" box="[199,853,827,854]" pageId="23" pageNumber="224">
|
||
KC560 site, Gulf of
|
||
<collectingRegion id="49C3F872FFAAD557FE7A222BFDDB5E7E" box="[429,517,828,854]" country="Mexico" name="Mexico" pageId="23" pageNumber="224">Mexico</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE335057FFAAD557FDC7222CFDA25E7E" box="[528,636,827,854]" degrees="26.42891" direction="north" orientation="latitude" pageId="23" pageNumber="224" precision="1" value="26.42891">26.42891</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFAAD557FD57222CFD285E7E" box="[640,758,827,854]" degrees="92.35167" direction="west" orientation="longitude" pageId="23" pageNumber="224" precision="1" value="-92.35167">-92.35167</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFAAD557FCD7222BFC8B5E7E" box="[768,853,828,854]" metricMagnitude="3" metricUnit="m" metricValue="1.918" pageId="23" pageNumber="224" unit="m" value="1918.0">1918 m</quantity>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF102277FD265E52" blockId="23.[151,1034,720,962]" box="[199,760,864,890]" pageId="23" pageNumber="224">EX1711_IMG_20171213T194231Z_ROVHD.jpg</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF102293FCB25EEA" blockId="23.[151,1034,720,962]" pageId="23" pageNumber="224">
|
||
West
|
||
<collectingRegion id="49C3F872FFAAD557FED22293FE885EB6" box="[261,342,900,926]" country="United States of America" name="Florida" pageId="23" pageNumber="224">Florida</collectingRegion>
|
||
Escarpment, Gulf of
|
||
<collectingRegion id="49C3F872FFAAD557FD9F2293FD7F5EB6" box="[584,673,900,926]" country="Mexico" name="Mexico" pageId="23" pageNumber="224">Mexico</collectingRegion>
|
||
.
|
||
<geoCoordinate id="EE335057FFAAD557FD7B2293FCC65EB6" box="[684,792,900,926]" degrees="24.91667" direction="north" orientation="latitude" pageId="23" pageNumber="224" precision="1" value="24.91667">24.91667</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFAAD557FCF42293FC465EB6" box="[803,920,900,926]" degrees="84.49046" direction="west" orientation="longitude" pageId="23" pageNumber="224" precision="1" value="-84.49046">-84.49046</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFAAD557FC742293FC255EB6" box="[931,1019,900,926]" metricMagnitude="3" metricUnit="m" metricValue="2.181" pageId="23" pageNumber="224" unit="m" value="2181.0">2181 m</quantity>
|
||
. EX1803_VID_20180430T164500Z_ROVHD_Low.mp4.tiff
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D651BFFAAD557FF4022FCFC1259F5" pageId="23" pageNumber="224" type="materials_examined">
|
||
<paragraph id="8BB83690FFAAD557FF4022FCFEAE592D" blockId="23.[151,1437,1003,1245]" box="[151,368,1003,1029]" pageId="23" pageNumber="224">
|
||
<emphasis id="B973EA82FFAAD557FF4022FCFEAE592D" box="[151,368,1003,1029]" italics="true" pageId="23" pageNumber="224">Material Examined</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF102518FD805959" blockId="23.[151,1437,1003,1245]" pageId="23" pageNumber="224">
|
||
<materialsCitation id="3B6F3CCDFFAAD557FF102518FD805959" ID-GBIF-Occurrence="2608527144" collectingDate="2018-04-25" collectionCode="USNM" collectorName="Coll. D. Wagner & Deep Discoverer & Ship" elevation="2549" location="Gulf" municipality="DeSoto Canyon" pageId="23" pageNumber="224" specimenCode="USNM 1507289" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
|
||
<emphasis id="B973EA82FFAAD557FF102518FEE55901" bold="true" box="[199,315,1039,1065]" pageId="23" pageNumber="224">
|
||
<typeStatus id="54BC8832FFAAD557FF102518FEE85901" box="[199,310,1039,1065]" pageId="23" pageNumber="224" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCode id="DBA19EEBFFAAD557FE932518FE215901" box="[324,511,1038,1065]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="224" type="Museum">USNM 1507289</specimenCode>
|
||
<collectingMunicipality id="6BDCACEAFFAAD557FDDF2518FD615901" box="[520,703,1039,1065]" pageId="23" pageNumber="224">DeSoto Canyon</collectingMunicipality>
|
||
,
|
||
<location id="8ED8604BFFAAD557FD1C2519FD205900" LSID="urn:lsid:plazi:treatment:03AE8786FFAED557FF402793FC1259F5:8ED8604BFFAAD557FD1C2519FD205900" box="[715,766,1038,1064]" municipality="DeSoto Canyon" name="Gulf" pageId="23" pageNumber="224" stateProvince="Mexico">Gulf</location>
|
||
of
|
||
<collectingRegion id="49C3F872FFAAD557FCF02519FCA15900" box="[807,895,1038,1064]" country="Mexico" name="Mexico" pageId="23" pageNumber="224">Mexico</collectingRegion>
|
||
, 28.28, -87.22,
|
||
<quantity id="4CFF9B75FFAAD557FBE42519FB525901" box="[1075,1164,1038,1065]" metricMagnitude="3" metricUnit="m" metricValue="2.549" pageId="23" pageNumber="224" unit="m" value="2549.0">
|
||
<elevation id="002AD1A3FFAAD557FBE42519FB525901" box="[1075,1164,1038,1065]" metricMagnitude="3" metricUnit="m" metricValue="2.549" pageId="23" pageNumber="224" unit="m" value="2549.0">2549 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="26F25346FFAAD557FB4E2519FA8B5901" box="[1177,1365,1038,1065]" pageId="23" pageNumber="224">Coll. D. Wagner</collectorName>
|
||
, ROV
|
||
<emphasis id="B973EA82FFAAD557FF402524FE8B5964" box="[151,341,1075,1100]" italics="true" pageId="23" pageNumber="224">
|
||
<collectorName id="26F25346FFAAD557FF402524FE8B5964" box="[151,341,1075,1100]" pageId="23" pageNumber="224">Deep Discoverer</collectorName>
|
||
</emphasis>
|
||
from NOAA
|
||
<collectorName id="26F25346FFAAD557FE252525FDFB5964" box="[498,549,1074,1100]" pageId="23" pageNumber="224">Ship</collectorName>
|
||
<taxonomicName id="4C074D13FFAAD557FDF92524FD245965" authority="Explorer" authorityName="Explorer" box="[558,762,1075,1101]" class="Insecta" family="Pentatomidae" genus="Okeanos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B973EA82FFAAD557FDF92524FD245965" box="[558,762,1075,1101]" italics="true" pageId="23" pageNumber="224">Okeanos Explorer</emphasis>
|
||
</taxonomicName>
|
||
EX
|
||
<date id="FFB91050FFAAD557FCFF2524FC1A5964" box="[808,964,1074,1101]" pageId="23" pageNumber="224" value="1803-04-25">
|
||
1803
|
||
<collectingDate id="EFFDE9B8FFAAD557FCB02525FC1A5964" box="[871,964,1074,1101]" pageId="23" pageNumber="224" value="2018-04-25">25 April</collectingDate>
|
||
</date>
|
||
2018. 1 wet spec. R=6.2 r=1.4. EX1803_ IMG_20180425T163351Z_ROVHD.jpg
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BB83690FFAAD557FF10256CFC1259F5" blockId="23.[151,1437,1003,1245]" pageId="23" pageNumber="224">
|
||
<emphasis id="B973EA82FFAAD557FF10256CFEEA59BD" bold="true" box="[199,308,1147,1173]" pageId="23" pageNumber="224">
|
||
<typeStatus id="54BC8832FFAAD557FF10256CFEEA59BD" box="[199,308,1147,1173]" pageId="23" pageNumber="224" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode id="ED16AE55FFAAD557FE93256CFE4B59BD" box="[324,405,1147,1173]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="224" type="Museum">USNM</collectionCode>
|
||
1550630
|
||
<collectingCountry id="F3107600FFAAD557FDDE256DFD7659BC" box="[521,680,1146,1172]" name="Virgin Islands" pageId="23" pageNumber="224">Virgin Islands</collectingCountry>
|
||
Trough,
|
||
<collectingCountry id="F3107600FFAAD557FCC4256CFC4359BC" box="[787,925,1146,1173]" name="Puerto Rico" pageId="23" pageNumber="224">Puerto Rico</collectingCountry>
|
||
, Caribbean Sea.
|
||
<geoCoordinate id="EE335057FFAAD557FB88256CFB6359BD" box="[1119,1213,1147,1173]" degrees="17.7732" direction="north" orientation="latitude" pageId="23" pageNumber="224" precision="5" value="17.7732">17.7732</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE335057FFAAD557FB1D256DFAEF59BD" box="[1226,1329,1146,1173]" degrees="65.4269" direction="west" orientation="longitude" pageId="23" pageNumber="224" precision="5" value="-65.4269">-65.4269</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9B75FFAAD557FAE9256CFA4659BD" box="[1342,1432,1147,1173]" metricMagnitude="3" metricUnit="m" metricValue="2.037" pageId="23" pageNumber="224" unit="m" value="2037.0">2037 m</quantity>
|
||
, Coll. D. Wagner with
|
||
<emphasis id="B973EA82FFAAD557FE592588FD955990" box="[398,587,1183,1208]" italics="true" pageId="23" pageNumber="224">Deep Discoverer</emphasis>
|
||
<collectionCode id="ED16AE55FFAAD557FD852588FD535991" box="[594,653,1183,1209]" country="Italy" lsid="urn:lsid:biocol.org:col:15413" name="Museo Civico di Rovereto" pageId="23" pageNumber="224" type="Herbarium">ROV</collectionCode>
|
||
,
|
||
<collectionCode id="ED16AE55FFAAD557FD4F2588FD395991" box="[664,743,1183,1209]" country="USA" httpUri="http://grbio.org/cool/qzhp-igg5" name="National Oceanic and Atmospeheric Administration" pageId="23" pageNumber="224">NOAA</collectionCode>
|
||
Ship
|
||
<taxonomicName id="4C074D13FFAAD557FCFE2588FC2A5991" authority="Explorer" authorityName="Explorer" box="[809,1012,1183,1209]" class="Insecta" family="Pentatomidae" genus="Okeanos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="224" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B973EA82FFAAD557FCFE2588FC2A5991" box="[809,1012,1183,1209]" italics="true" pageId="23" pageNumber="224">Okeanos Explorer</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<date id="FFB91050FFAAD557FBD42589FB4E5991" box="[1027,1168,1182,1209]" pageId="23" pageNumber="224" value="2018-11-05">5 Nov. 2018</date>
|
||
. 1 wet spec. 1 wet spec.
|
||
<collectionCode id="ED16AE55FFAAD557FF4025D4FF7559F5" box="[151,171,1219,1245]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="224">R</collectionCode>
|
||
=1.8 r=0.8.
|
||
<collectionCode id="ED16AE55FFAAD557FEF925D5FE8A59F5" box="[302,340,1218,1245]" country="Slovenia" name="The Culture Collection of Extremophilic Fungi" pageId="23" pageNumber="224">EX</collectionCode>
|
||
1811_IMG_20181105
|
||
<collectionCode id="ED16AE55FFAAD557FD9A25D5FD8059F5" box="[589,606,1218,1245]" name="Tavera, Department of Geology and Geophysics" pageId="23" pageNumber="224">T</collectionCode>
|
||
170423Z_
|
||
<collectionCode id="ED16AE55FFAAD557FD0725D5FCEB59F5" box="[720,821,1218,1245]" pageId="23" pageNumber="224">ROVHD</collectionCode>
|
||
.jpg (feeding)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |