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<document id="CDC17EC5503A0EB918DA0F68D4366EAB" ID-DOI="10.5281/zenodo.4525415" ID-ISSN="1638-9387" ID-RefBankID="E114FFC7FFB8FFD32C2E585BCB79FFF4" ID-Zenodo-Dep="4525415" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612878832874" checkinUser="felipe" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques &amp; Mauriès, Jean-Paul" docDate="2006" docId="03AE8781FF97FFEBF27DFA609448FD0B" docLanguage="en" docName="Zoosystema.28.4.887-915.pdf" docOrigin="Zoosystema 28 (4)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Hyleoglomeris sulcostriata Golovatch &amp; Geoffroy &amp; Mauriès 2006, n. sp." docType="treatment" docVersion="3" lastPageNumber="904" masterDocId="FF97FFF9FF98FFFAF17FFFAC9009FF85" masterDocTitle="Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia" masterLastPageNumber="915" masterPageNumber="887" pageNumber="902" updateTime="1698868940497" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9F6A1A6A7EAA3AB277FDC6FC1DBCD348">Review of the millipede genus Hyleoglomeris Verhoeff, 1910 (Diplopoda, Glomerida, Glomeridae), with descriptions of new species from caves in Southeast Asia</mods:title>
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<mods:name id="F60B7746F0FC5F635B6ED3CEBB48CCA4" type="personal">
<mods:role id="99F7ADFE164B37805017BC0F6328AAC6">
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<mods:namePart id="32A060D26799B795D70B3692C1710B9F">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="997E6614E17EC67E0EE1F22F0BB0C89F">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
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<mods:namePart id="B35B5FB09525A04FCB2EFA93D7BE3961">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="798AB3A5B086A558EC5DFAB1FFE32AEB">Muséum national dHistoire naturelle, Département Écologie et Gestion de la Biodiversité, EA 3778 - USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
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<mods:namePart id="447268235828668063434C509647973C">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="3612CBB3EBD3A324C2C80D36E73C75E1">Muséum national dHistoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
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<mods:title id="04419D33148EDC0E08D3BA29489185EA">Zoosystema</mods:title>
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<mods:date id="A52C41201AC023008C8F23820011F5FC">2006</mods:date>
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<taxonomicName id="4C074D14FF97FFF5F27DFA609413FA63" ID-CoL="3NF2Q" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[770,1050,1484,1510]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="15" pageNumber="902" phylum="Arthropoda" rank="species" species="sulcostriata" status="sp. nov.">
<emphasis id="B973EA85FF97FFF5F27DFA609413FA63" bold="true" box="[770,1050,1484,1510]" italics="true" pageId="15" pageNumber="902">Hyleoglomeris sulcostriata</emphasis>
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(
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)
</paragraph>
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<collectionCode id="ED16AE52FF97FFF5F3DAF98E934DF9BD" box="[677,836,1570,1593]" pageId="15" pageNumber="902">TYPE MATERIAL</collectionCode>
. —
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.
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,
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(
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), cave,
<date id="FFB91057FF97FFF5F2C2F9909419F9D6" box="[957,1040,1596,1619]" pageId="15" pageNumber="902" value="2000-01-01">
<collectingDate id="EFFDE9BFFF97FFF5F2C2F9909419F9D6" box="[957,1040,1596,1619]" pageId="15" pageNumber="902" value="2000-01-01">1.I.2000</collectingDate>
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, leg.
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&amp;
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(
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;
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;
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(
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). —
<collectingRegion id="49C3F875FF97FFF5F545F9DD92F7F926" country="Laos" name="Khammouan" pageId="15" pageNumber="902">Khammouan Province</collectingRegion>
,
<collectingCounty id="62D94E1BFF97FFF5F274F9209372F926" box="[779,891,1676,1699]" pageId="15" pageNumber="902">Ban Vieng</collectingCounty>
,
<collectingMunicipality id="6BDCACEDFF97FFF5F2F8F920943DF926" box="[903,1076,1676,1699]" pageId="15" pageNumber="902">Tham Houai Sai</collectingMunicipality>
(
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,
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), sink cave,
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<collectingDate id="EFFDE9BFFF88FFEAF044FAA191A8FAA1" box="[315,417,1293,1316]" pageId="16" pageNumber="903" value="2001-02-13">13.II.2001</collectingDate>
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, leg.
<collectorName id="26F25341FF88FFEAF0B2FAA19239FAA1" box="[461,560,1293,1316]" pageId="16" pageNumber="903">X. Noguès</collectorName>
,
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(
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). —
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, Ban Nam Non, Nam Non (
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), cave,
<date id="FFB91057FF88FFEAF08DFAEE9252FADC" box="[498,603,1346,1369]" pageId="16" pageNumber="903" value="1999-02-15">
<collectingDate id="EFFDE9BFFF88FFEAF08DFAEE9252FADC" box="[498,603,1346,1369]" pageId="16" pageNumber="903" value="1999-02-15">15.II.1999</collectingDate>
</date>
, leg.
<collectorName id="26F25341FF88FFEAF10FFAF090D8FAF6" box="[112,209,1372,1395]" pageId="16" pageNumber="903">J. Lordon</collectorName>
,
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,
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(
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.
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<caption id="DF78661FFF88FFEAF10EFB079136FB56" ID-DOI="http://doi.org/10.5281/zenodo.4525439" ID-Zenodo-Dep="4525439" httpUri="https://zenodo.org/record/4525439/files/figure.png" pageId="16" pageNumber="903" startId="16.[113,124,1195,1212]" targetBox="[125,1192,218,1150]" targetPageId="16">
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FIG. 9. —
<taxonomicName id="4C074D14FF88FFEAF1BAFB009198FB38" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[197,401,1196,1213]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="16" pageNumber="903" phylum="Arthropoda" rank="species" species="speophila" status="sp. nov.">
<emphasis id="B973EA85FF88FFEAF1BAFB009198FB38" box="[197,401,1196,1213]" italics="true" pageId="16" pageNumber="903">Hyleoglomeris speophila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF88FFEAF0E8FB0791C2FB39" box="[407,459,1195,1212]" pageId="16" pageNumber="903" rank="species">n. sp.</taxonomicNameLabel>
, ♂ holotype:
<emphasis id="B973EA85FF88FFEAF344FB009241FB38" bold="true" box="[571,584,1196,1213]" pageId="16" pageNumber="903">A</emphasis>
, leg 17;
<emphasis id="B973EA85FF88FFEAF3ECFB0092A9FB38" bold="true" box="[659,672,1196,1213]" pageId="16" pageNumber="903">B</emphasis>
, leg 18;
<emphasis id="B973EA85FF88FFEAF394FB0092F0FB38" bold="true" box="[747,761,1196,1213]" pageId="16" pageNumber="903">C</emphasis>
, telopod, front view;
<emphasis id="B973EA85FF88FFEAF2D2FB0093B2FB38" bold="true" box="[941,955,1196,1213]" pageId="16" pageNumber="903">D</emphasis>
, distal part of telopod, caudal view. Scale bar: 0.4 mm.
</paragraph>
</caption>
<subSubSection id="C31D651CFF88FFEAF10EFA259276FA3F" pageId="16" pageNumber="903" type="etymology">
<paragraph id="8BB83697FF88FFEAF10EFA259276FA3F" blockId="16.[111,646,1417,1466]" pageId="16" pageNumber="903">ETYMOLOGY. — To emphasize a conspicuous lateral sulcus on the thoracic shield turning into a stria middorsally.</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF88FFEAF10EFA7C9256F999" pageId="16" pageNumber="903" type="diagnosis">
<paragraph id="8BB83697FF88FFEAF10EFA7C9256F999" blockId="16.[112,646,1488,1564]" pageId="16" pageNumber="903">DIAGNOSIS. — Differs from congeners in some troglomorphic traits, combined with a conspicuous lateral sulcus on the thoracic shield turning into a stria middorsally.</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF88FFEBF10EF9E691C7FC2E" lastPageId="17" lastPageNumber="904" pageId="16" pageNumber="903" type="description">
<paragraph id="8BB83697FF88FFEAF10EF9E6910BF9E7" blockId="16.[113,646,1610,1700]" box="[113,258,1610,1636]" pageId="16" pageNumber="903">DESCRIPTION</paragraph>
<paragraph id="8BB83697FF88FFEAF10EF9C6928FF921" blockId="16.[113,646,1610,1700]" pageId="16" pageNumber="903">
Length 5.0-6.0 (
<specimenCount id="9D01FD1EFF88FFEAF04FF9C7914FF900" box="[304,326,1643,1669]" pageId="16" pageNumber="903" type="male"></specimenCount>
) or 5.5-7.0 mm (
<specimenCount id="9D01FD1EFF88FFEAF368F9C79222F900" box="[535,555,1643,1669]" pageId="16" pageNumber="903" type="female"></specimenCount>
), width 2.5-3.0 (
<specimenCount id="9D01FD1EFF88FFEAF1B0F92690ECF921" box="[207,229,1674,1700]" pageId="16" pageNumber="903" type="male"></specimenCount>
) or
<quantity id="4CFF9B72FF88FFEAF070F9269186F921" box="[271,399,1674,1700]" metricMagnitude="-3" metricUnit="m" metricValue="3.15" metricValueMax="3.5" metricValueMin="2.8" pageId="16" pageNumber="903" unit="mm" value="3.15" valueMax="3.5" valueMin="2.8">2.8-3.5 mm</quantity>
(
<specimenCount id="9D01FD1EFF88FFEAF0E1F92691B8F921" box="[414,433,1674,1700]" pageId="16" pageNumber="903" type="female"></specimenCount>
).
<typeStatus id="54BC8835FF88FFEAF0B9F9269225F921" box="[454,556,1674,1700]" pageId="16" pageNumber="903" type="holotype">Holotype</typeStatus>
6.0 mm
</paragraph>
<paragraph id="8BB83697FF88FFEAF3DAFAA293ABFAAD" blockId="16.[675,1211,1294,1700]" box="[677,930,1294,1320]" pageId="16" pageNumber="903">
long and
<quantity id="4CFF9B72FF88FFEAF274FAA29368FAAD" box="[779,865,1294,1320]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="903" unit="mm" value="2.8">2.8 mm</quantity>
wide.
</paragraph>
<paragraph id="8BB83697FF88FFEAF3C4FA81947DFAC2" blockId="16.[675,1211,1294,1700]" box="[699,1140,1325,1351]" pageId="16" pageNumber="903">Coloration entirely pallid, unpigmented.</paragraph>
<paragraph id="8BB83697FF88FFEAF3C4FAE19385FA63" blockId="16.[675,1211,1294,1700]" pageId="16" pageNumber="903">Antennomere 6 about 1.6-1.7 times longer than wide, antennomere 7 very short; 5 or 6 poorly visible, unpigmented but convex ocelli on each side of head; Tömösvárys organ transverse-oval, about twice as wide as long.</paragraph>
<paragraph id="8BB83697FF88FFEAF3C4FA479427F980" blockId="16.[675,1211,1294,1700]" box="[699,1070,1515,1541]" pageId="16" pageNumber="903">Collum with two transverse striae.</paragraph>
<paragraph id="8BB83697FF88FFEBF3C4F9A791C6FE14" blockId="16.[675,1211,1294,1700]" lastBlockId="17.[111,649,216,939]" lastPageId="17" lastPageNumber="904" pageId="16" pageNumber="903">Thoracic shield with a medium-sized hyposchism slightly surpassing the caudal tergal contour; 10 or 11 transverse striae, of which 8 or 9 start above the schism and 7 or 8 cross the dorsum; most of the striae lying on an elevation just in front of a conspicuous, rather strongly and regularly curved (like the other striae), midway, lateral sulcus delimiting a slight impression in posterolateral portion of the shield; the sulcus deep basally, starting just at schism end, gradually turning into the last crossing stria in apical third of the shield.</paragraph>
<paragraph id="8BB83697FF89FFEBF1F7FE3B911AFE55" blockId="17.[111,649,216,939]" pageId="17" pageNumber="904">Male pygidium virtually not sinuate medially at caudal margin.</paragraph>
<paragraph id="8BB83697FF89FFEBF1F7FE7A90C2FDCA" blockId="17.[111,649,216,939]" pageId="17" pageNumber="904">
Male leg 17 (
<figureCitation id="133C2A12FF89FFEBF05AFE7A91DEFE75" box="[293,471,470,496]" captionStart="FIG" captionStartId="18.[113,124,1634,1651]" captionTargetBox="[122,1200,218,1524]" captionTargetId="figure-13@18.[112,1211,214,1527]" captionTargetPageId="18" captionText="FIG. 10. — Hyleoglomeris sulcostriata n. sp., ♂ holotype (A-D), ♂ paratype from Nam Non (E-G) and ♂ paratype from Tham Houai Sai (H-J); A, E, H, leg 17; B, F, I, leg 18; C, G, J, telopods, front view; D, distal part of telopod, caudal view. Scale bars: A-D, 0.3 mm; E-J, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525443" httpUri="https://zenodo.org/record/4525443/files/figure.png" pageId="17" pageNumber="904">Fig. 10A, E, H</figureCitation>
) with a low to medium-sized, more or less rounded, sometimes micropapillate outer coxal lobe; telopodite 4-segmented.
</paragraph>
<paragraph id="8BB83697FF89FFEBF1F7FDF9910CFD2B" blockId="17.[111,649,216,939]" pageId="17" pageNumber="904">
Male leg 18 (
<figureCitation id="133C2A12FF89FFEBF058FDF991CEFDEA" box="[295,455,597,623]" captionStart="FIG" captionStartId="18.[113,124,1634,1651]" captionTargetBox="[122,1200,218,1524]" captionTargetId="figure-13@18.[112,1211,214,1527]" captionTargetPageId="18" captionText="FIG. 10. — Hyleoglomeris sulcostriata n. sp., ♂ holotype (A-D), ♂ paratype from Nam Non (E-G) and ♂ paratype from Tham Houai Sai (H-J); A, E, H, leg 17; B, F, I, leg 18; C, G, J, telopods, front view; D, distal part of telopod, caudal view. Scale bars: A-D, 0.3 mm; E-J, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525443" httpUri="https://zenodo.org/record/4525443/files/figure.png" pageId="17" pageNumber="904">Fig. 10B, F, I</figureCitation>
) with a broadly rounded to ogival syncoxital notch; telopodite 4-segmented.
</paragraph>
<paragraph id="8BB83697FF89FFEBF1F7FD1891C7FC2E" blockId="17.[111,649,216,939]" pageId="17" pageNumber="904">
Telopods (
<figureCitation id="133C2A12FF89FFEBF181FD1891C3FD4B" box="[254,458,692,718]" captionStart="FIG" captionStartId="18.[113,124,1634,1651]" captionTargetBox="[122,1200,218,1524]" captionTargetId="figure-13@18.[112,1211,214,1527]" captionTargetPageId="18" captionText="FIG. 10. — Hyleoglomeris sulcostriata n. sp., ♂ holotype (A-D), ♂ paratype from Nam Non (E-G) and ♂ paratype from Tham Houai Sai (H-J); A, E, H, leg 17; B, F, I, leg 18; C, G, J, telopods, front view; D, distal part of telopod, caudal view. Scale bars: A-D, 0.3 mm; E-J, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525443" httpUri="https://zenodo.org/record/4525443/files/figure.png" pageId="17" pageNumber="904">Fig. 10C, D, G, J</figureCitation>
) somewhat variable, with a roundly subtrapeziform to large and emarginate syncoxital lobe flanked by two setose horns crowned with a subapical setoid. Prefemur micropapillate laterally. Caudomedial outgrowth of femur rather narrow, that of tibia with a distinct to very prominent tubercle at base. Tarsus narrowly rounded apically, subacuminate.
</paragraph>
</subSubSection>
<subSubSection id="C31D651CFF89FFEBF10EFC7D9448FD0B" pageId="17" pageNumber="904" type="discussion">
<paragraph id="8BB83697FF89FFEBF10EFC7D90DCFC6C" blockId="17.[111,649,977,1700]" box="[113,213,977,1003]" pageId="17" pageNumber="904">REMARKS</paragraph>
<paragraph id="8BB83697FF89FFEBF10EFC5C9105F9A0" blockId="17.[111,649,977,1700]" pageId="17" pageNumber="904">
As the material comes from three different caves in
<collectingCountry id="F3107607FF89FFEBF10FFBBC90AFFBAF" box="[112,166,1040,1066]" name="Laos" pageId="17" pageNumber="904">Laos</collectingCountry>
, with a distance between both
<collectingRegion id="49C3F875FF89FFEBF08CFBBC928DFBAF" box="[499,644,1040,1066]" country="Laos" name="Khammouan" pageId="17" pageNumber="904">Khammouan</collectingRegion>
caves of about
<quantity id="4CFF9B72FF89FFEBF063FB9C916EFBCF" box="[284,359,1072,1098]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="904" unit="km" value="60.0">60 km</quantity>
(although located in the same karst area), which in turn are separated from Tham Non by a distance of about
<quantity id="4CFF9B72FF89FFEBF356FBC3928BFB0C" box="[553,642,1135,1161]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="904" unit="km" value="300.0">300 km</quantity>
, variation is certainly something to expect. The specimens from the
<collectingRegion id="49C3F875FF89FFEBF033FB0391D4FB4C" box="[332,477,1199,1225]" country="Laos" name="Khammouan" pageId="17" pageNumber="904">Khammouan</collectingRegion>
caves appear to be particularly similar. They are slightly larger in size (3.0-
<quantity id="4CFF9B72FF89FFEBF1A8FB429139FA8D" box="[215,304,1262,1288]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="904" unit="mm" value="3.5">3.5 mm</quantity>
wide), there are usually seven striae crossing the thoracic shield while the respective male legs 17-19 are nearly identical, with an unusually high tubercle at base of the telopod tibial outgrowth (
<figureCitation id="133C2A12FF89FFEBF03EFAC1919DFA02" box="[321,404,1389,1415]" captionStart="FIG" captionStartId="18.[113,124,1634,1651]" captionTargetBox="[122,1200,218,1524]" captionTargetId="figure-13@18.[112,1211,214,1527]" captionTargetPageId="18" captionText="FIG. 10. — Hyleoglomeris sulcostriata n. sp., ♂ holotype (A-D), ♂ paratype from Nam Non (E-G) and ♂ paratype from Tham Houai Sai (H-J); A, E, H, leg 17; B, F, I, leg 18; C, G, J, telopods, front view; D, distal part of telopod, caudal view. Scale bars: A-D, 0.3 mm; E-J, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525443" httpUri="https://zenodo.org/record/4525443/files/figure.png" pageId="17" pageNumber="904">Fig. 10</figureCitation>
E-J). In contrast, the sample from Tham Non is distinct in that the animals are a little smaller (width
<quantity id="4CFF9B72FF89FFEBF08CFA009272FA43" box="[499,635,1452,1478]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="17" pageNumber="904" unit="mm" value="2.65" valueMax="2.8" valueMin="2.5">2.5-2.8 mm</quantity>
), usually showing eight striae crossing the thoracic shield, while male legs 17-19 are slightly different (
<figureCitation id="133C2A12FF89FFEBF105F9A790C0F9A0" box="[122,201,1547,1573]" captionStart="FIG" captionStartId="18.[113,124,1634,1651]" captionTargetBox="[122,1200,218,1524]" captionTargetId="figure-13@18.[112,1211,214,1527]" captionTargetPageId="18" captionText="FIG. 10. — Hyleoglomeris sulcostriata n. sp., ♂ holotype (A-D), ♂ paratype from Nam Non (E-G) and ♂ paratype from Tham Houai Sai (H-J); A, E, H, leg 17; B, F, I, leg 18; C, G, J, telopods, front view; D, distal part of telopod, caudal view. Scale bars: A-D, 0.3 mm; E-J, 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4525443" httpUri="https://zenodo.org/record/4525443/files/figure.png" pageId="17" pageNumber="904">Fig. 10</figureCitation>
A-D).
</paragraph>
<paragraph id="8BB83697FF89FFEBF1F7F98793ADFED4" blockId="17.[111,649,977,1700]" lastBlockId="17.[675,1212,216,654]" pageId="17" pageNumber="904">
Nevertheless, there can be no doubt that the samples are conspecific because of the striking midway sulcus present on the thoracic shield.A similar sulcus is only known to occur in
<taxonomicName id="4C074D14FF89FFEBF0F9F926920FF921" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[390,518,1674,1700]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="17" pageNumber="904" phylum="Arthropoda" rank="species" species="eusulcata" status="sp. nov.">
<emphasis id="B973EA85FF89FFEBF0F9F926920FF921" box="[390,518,1674,1700]" italics="true" pageId="17" pageNumber="904">H. eusulcata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF89FFEBF374F926924EF921" box="[523,583,1674,1700]" pageId="17" pageNumber="904" rank="species">n. sp.</taxonomicNameLabel>
, from
<collectingRegion id="49C3F875FF89FFEBF3DAFF74930DFF77" box="[677,772,216,242]" country="China" name="Guizhou" pageId="17" pageNumber="904">Guizhou</collectingRegion>
(see above), attesting to their particularly close relations. However, in
<taxonomicName id="4C074D14FF89FFEBF2A3FF54947DFE97" authority="Golovatch &amp; Geoffroy &amp; Mauriès, 2006" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès" authorityYear="2006" box="[988,1140,248,274]" class="Diplopoda" family="Glomeridae" genus="Hyleoglomeris" kingdom="Animalia" order="Glomerida" pageId="17" pageNumber="904" phylum="Arthropoda" rank="species" species="sulcostriata" status="sp. nov.">
<emphasis id="B973EA85FF89FFEBF2A3FF54947DFE97" box="[988,1140,248,274]" italics="true" pageId="17" pageNumber="904">H. sulcostriata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24057FEFF89FFEBF502FF5494B3FE97" box="[1149,1210,248,274]" pageId="17" pageNumber="904" rank="species">n. sp.</taxonomicNameLabel>
this sulcus gradually turns middorsally into the last stria crossing the shield.
</paragraph>
<paragraph id="8BB83697FF89FFEBF3C4FEFB933EFDAA" blockId="17.[675,1212,216,654]" pageId="17" pageNumber="904">This species also shows such troglomorphic features as complete depigmentation and an elongate Tömösvárys organ, but the ocelli are still discernible and neither the antennae nor the legs are particularly elongate. To prove that we face a troglobite, additional observations and material are necessary.</paragraph>
<paragraph id="8BB83697FF89FFEBF3C4FD999448FD0B" blockId="17.[675,1212,216,654]" pageId="17" pageNumber="904">
In general,
<collectingCountry id="F3107607FF89FFEBF24FFD999368FDCA" box="[816,865,565,591]" name="Laos" pageId="17" pageNumber="904">Laos</collectingCountry>
is rich in karst caves that support numerous troglobites, presumably several among diplopods as well (
<bibRefCitation id="EF964B66FF89FFEBF214FDD8943CFD0B" author="BESSON J. - P. &amp; DEHARVENG L. &amp; BREHIER F." box="[875,1077,628,654]" pageId="17" pageNumber="904" pagination="1883 - 1889" refId="ref16541" refString="BESSON J. - P., DEHARVENG L. &amp; BREHIER F. 2001. - Laos, in JUBERTHIE C. &amp; DECU V. (eds), Encyclopaedia Biospeologica 3. Societe internationale de Biospeologie, Moulis: 1883 - 1889." type="book chapter" year="2001">
Besson
<emphasis id="B973EA85FF89FFEBF2C0FDD893FBFD0B" box="[959,1010,628,654]" italics="true" pageId="17" pageNumber="904">et al.</emphasis>
2001
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>