395 lines
64 KiB
XML
395 lines
64 KiB
XML
<document id="56E76DDF16F57E5D2AC90B6DE17784A4" ID-DOI="10.1111/j.1096-3642.2012.00850.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5408795" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630349485666" checkinUser="marcus" docAuthor="Capa, Maria, Parapar, Julio & Hutchings, Pat" docDate="2012" docId="03AE8729AB14D45BFEBFF9F5A26CFE1F" docLanguage="en" docName="j.1096-3642.2012.00850.x.pdf" docOrigin="Zoological Journal of the Linnean Society 166 (2)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2012.00850.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Galathowenia quelis Capa & Parapar & Hutchings 2012, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="248" masterDocId="FF97FF51AB1ED457FFACFFA2A67DFFF0" masterDocTitle="Phylogeny of Oweniidae (Polychaeta) based on morphological data and taxonomic revision of Australian fauna" masterLastPageNumber="278" masterPageNumber="236" pageNumber="246" updateTime="1699241006827" updateUser="plazi">
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<mods:title id="B12CBC4FB44E394E043E9D24F970678C">Phylogeny of Oweniidae (Polychaeta) based on morphological data and taxonomic revision of Australian fauna</mods:title>
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<mods:namePart id="9539D27B44C48D8F567F5452A5BFCE1E">Capa, Maria</mods:namePart>
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<mods:namePart id="E5123AAD5857518F8596D10452CACC6A">Parapar, Julio</mods:namePart>
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<mods:namePart id="C07DD79E7558825C5CC2B1B4C0566315">Hutchings, Pat</mods:namePart>
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<mods:title id="2EE0B53600CF81C4ABABB5A9424FAB80">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="73789C6CD6CDB00CC62506B80BF68DC1">2012</mods:date>
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<mods:number id="BE8823AE248F37F657CB2FB9B6032A0E">2012-09-28</mods:number>
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<mods:number id="53A9ADB5CAF625ED85C785996F9C882B">166</mods:number>
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<subSubSection id="C31D65B4AB14D45DFEBFF9F5A4E1F99C" box="[275,668,1623,1644]" pageId="10" pageNumber="246" type="nomenclature">
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<heading id="D0F08153AB14D45DFEBFF9F5A4E1F99C" box="[275,668,1623,1644]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="246" reason="4">
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<taxonomicName id="4C074DBCAB14D45DFEBFF9F5A450F99B" ID-CoL="6K849" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[275,557,1623,1644]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="10" pageNumber="246" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">GALATHOWENIA QUELIS</taxonomicName>
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<emphasis id="B973EA2DAB14D45DFD9BF9F8A4E1F99C" bold="true" box="[567,668,1623,1644]" pageId="10" pageNumber="246">
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<taxonomicNameLabel id="A2405756AB14D45DFD9BF9F8A4E1F99C" box="[567,668,1623,1644]" pageId="10" pageNumber="246" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="C31D65B4AB14D45BFE88F9D5A26CFE1F" lastPageId="12" lastPageNumber="248" pageId="10" pageNumber="246" type="description">
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<paragraph id="8BB8363FAB14D45DFE88F9D5A4F6F97C" blockId="10.[275,668,1623,1676]" box="[292,651,1655,1676]" pageId="10" pageNumber="246">
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<figureCitation id="133C2ABAAB14D45DFE88F9D5A7AEF97C" box="[292,467,1655,1676]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">FIGURES 2E–H</figureCitation>
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,
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<figureCitation id="133C2ABAAB14D45DFE4DF9D5A78CF97C" box="[481,497,1655,1676]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">4</figureCitation>
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,
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<tableCitation id="C6850384AB14D45DFDADF9D5A411F97C" box="[513,620,1655,1676]" captionStart="Table 1" captionStartId="8.[136,200,599,620]" captionText="Table 1. Character matrix of morphological features for resolving oweniid relationships" pageId="10" pageNumber="246">TABLES 1</tableCitation>
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,
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<tableCitation id="C6850384AB14D45DFDD1F9D5A4F6F97C" box="[637,651,1655,1676]" captionStart="Table 2" captionStartId="9.[136,200,525,546]" captionText="Table 2. Comparison of three Australian species of Galathowenia and Galathowenia oculata from different geographical areas" pageId="10" pageNumber="246">2</tableCitation>
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</paragraph>
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<paragraph id="8BB8363FAB14D45DFF08F93FA7C6F900" blockId="10.[164,779,1693,1776]" pageId="10" pageNumber="246">
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<materialsCitation id="3B6F3C62AB14D45DFF08F93FA7C6F900" accessionNumber="W37821" collectingDate="2004-12-02" collectionCode="AM" latitude="-33.595554" location="Sand Point" longLatPrecision="19" longitude="151.30556" municipality="Pittwater" pageId="10" pageNumber="246" specimenCount="1" stateProvince="New South Wales" typeStatus="holotype">
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<emphasis id="B973EA2DAB14D45DFF08F93FA773F942" box="[164,270,1693,1714]" italics="true" pageId="10" pageNumber="246">
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<typeStatus id="54BC889DAB14D45DFF08F93FA775F942" box="[164,264,1693,1714]" pageId="10" pageNumber="246" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectingRegion id="49C3F8DDAB14D45DFEB4F93CA797F943" box="[280,490,1693,1715]" country="Australia" name="New South Wales" pageId="10" pageNumber="246">New South Wales</collectingRegion>
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,
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<collectingMunicipality id="6BDCAC45AB14D45DFE55F93FA41AF943" box="[505,615,1693,1715]" pageId="10" pageNumber="246">Pittwater</collectingMunicipality>
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,
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<collectionCode id="ED16AEFAAB14D45DFDD8F93FA4E0F942" box="[628,669,1693,1714]" country="Australia" name="Australian Museum" pageId="10" pageNumber="246" type="Museum">AM</collectionCode>
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W37821, west of
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,
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,
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<geoCoordinate id="EE3350F8AB14D45DFD85F91EA4CAF921" box="[553,695,1723,1746]" degrees="151" direction="east" minutes="18" orientation="longitude" pageId="10" pageNumber="246" precision="15" seconds="20" value="151.30556">151°18′20″E</geoCoordinate>
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, sandy mud, 15.6 m,
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<collectingDate id="EFFDE917AB14D45DFEE9F978A7CAF900" box="[325,439,1754,1776]" pageId="10" pageNumber="246" value="2004-12-02">2.xii.2004</collectingDate>
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</date>
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.
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</paragraph>
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<paragraph id="8BB8363FAB14D45DFF08F8BEA36AFE84" blockId="10.[164,779,1820,1903]" lastBlockId="10.[826,1441,198,373]" pageId="10" pageNumber="246">
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<materialsCitation id="3B6F3C62AB14D45DFF08F8BEA31EFF2B" accessionNumber="W37822, W37823" collectingDate="2004-09-15" collectionCode="AM, R" latitude="-33.975834" location="Botany Bay" longLatPrecision="19" longitude="151.195" municipality="Pittwater" pageId="10" pageNumber="246" specimenCount="1" stateProvince="New South Wales" typeStatus="paratype">
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<emphasis id="B973EA2DAB14D45DFF08F8BEA766F8C1" box="[164,283,1820,1841]" italics="true" pageId="10" pageNumber="246">
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<typeStatus id="54BC889DAB14D45DFF08F8BEA768F8C1" box="[164,277,1820,1841]" pageId="10" pageNumber="246" type="paratype">Paratypes</typeStatus>
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:
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<collectingRegion id="49C3F8DDAB14D45DFE8FF8BEA78CF8C2" box="[291,497,1820,1842]" country="Australia" name="New South Wales" pageId="10" pageNumber="246">New South Wales</collectingRegion>
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,
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<collectingMunicipality id="6BDCAC45AB14D45DFE51F8BEA416F8C2" box="[509,619,1820,1842]" pageId="10" pageNumber="246">Pittwater</collectingMunicipality>
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,
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<collectionCode id="ED16AEFAAB14D45DFDDAF8BEA4E2F8C1" box="[630,671,1820,1841]" country="Australia" name="Australian Museum" pageId="10" pageNumber="246" type="Museum">AM</collectionCode>
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W37822, same sample (six specs);
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<location id="8ED860E4AB14D45DFE7BF899A41FF8A0" LSID="urn:lsid:plazi:treatment:03AE8729AB14D45BFEBFF9F5A26CFE1F:8ED860E4AB14D45DFE7BF899A41FF8A0" box="[471,610,1851,1872]" latitude="-33.975834" longLatPrecision="19" longitude="151.195" municipality="Pittwater" name="Botany Bay" pageId="10" pageNumber="246" stateProvince="New South Wales">Botany Bay</location>
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,
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<collectionCode id="ED16AEFAAB14D45DFDDDF898A4E7F8BF" box="[625,666,1850,1871]" country="Australia" name="Australian Museum" pageId="10" pageNumber="246" type="Museum">AM</collectionCode>
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W37823, east end of runway 34
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<collectionCode id="ED16AEFAAB14D45DFE6DF8FBA7A8F89E" box="[449,469,1881,1902]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="246">R</collectionCode>
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,
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,
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, sandy mud, 16.5 m,
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<date id="FFB910FFAB14D45DFBEDFF64A2CBFF2C" box="[1089,1206,198,220]" pageId="10" pageNumber="246" value="2004-09-15">
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<collectingDate id="EFFDE917AB14D45DFBEDFF64A2CBFF2C" box="[1089,1206,198,220]" pageId="10" pageNumber="246" value="2004-09-15">15.ix.2004</collectingDate>
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</date>
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(eight specs)
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,
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<materialsCitation id="3B6F3C62AB14D45DFAD4FF64A36EFE84" accessionNumber="W37824, W37825, W37826" collectingDate="1992-07-27" collectingDateMax="1992-07-28" collectingDateMin="1992-07-27" collectionCode="AM" elevation="1000" latitude="-33.97111" location="Port Botany" longLatPrecision="19" longitude="151.19945" municipality="Port Botany" pageId="10" pageNumber="246" specimenCount="3" stateProvince="New South Wales" typeStatus="paratype">
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<collectionCode id="ED16AEFAAB14D45DFAD4FF64A3DCFF2B" box="[1400,1441,198,219]" country="Australia" name="Australian Museum" pageId="10" pageNumber="246" type="Museum">AM</collectionCode>
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W37824, 800–
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<elevation id="002AD10CAB14D45DFC4AFF47A244FF0A" box="[998,1081,229,250]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="246" unit="m" value="1000.0">1000 m</elevation>
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</quantity>
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off
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<collectingMunicipality id="6BDCAC45AB14D45DFBDAFF47A377FF0A" box="[1142,1290,229,250]" pageId="10" pageNumber="246">Port Botany</collectingMunicipality>
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,
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<geoCoordinate id="EE3350F8AB14D45DFAB2FF46A3E1FF0A" box="[1310,1436,227,250]" degrees="33" direction="south" minutes="58" orientation="latitude" pageId="10" pageNumber="246" precision="15" seconds="45" value="-33.979168">33°58′45″S</geoCoordinate>
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,
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<geoCoordinate id="EE3350F8AB14D45DFC96FEA1A5C4FEE9" box="[826,953,258,281]" degrees="151" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="246" precision="15" seconds="1" value="151.18361">151°11′1″E</geoCoordinate>
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,
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</quantity>
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,
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<date id="FFB910FFAB14D45DFBACFEA1A200FEE9" box="[1024,1149,259,281]" pageId="10" pageNumber="246" value="1992-07-28">
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<collectingDate id="EFFDE917AB14D45DFBACFEA1A200FEE9" box="[1024,1149,259,281]" pageId="10" pageNumber="246" value="1992-07-28">28.vii.1992</collectingDate>
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</date>
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(
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<specimenCount id="9D01FDB6AB14D45DFB21FEA1A285FEE9" box="[1165,1272,259,281]" pageId="10" pageNumber="246" type="generic">one spec.</specimenCount>
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),
|
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<collectionCode id="ED16AEFAAB14D45DFAA0FEA1A348FEE8" box="[1292,1333,259,280]" country="Australia" name="Australian Museum" pageId="10" pageNumber="246" type="Museum">AM</collectionCode>
|
||
W37825, 800–
|
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<elevation id="002AD10CAB14D45DFCDEFE80A5B9FEC7" box="[882,964,290,311]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="246" unit="m" value="1000.0">1000 m</elevation>
|
||
</quantity>
|
||
off
|
||
<location id="8ED860E4AB14D45DFC59FE80A2FEFEC7" LSID="urn:lsid:plazi:treatment:03AE8729AB14D45BFEBFF9F5A26CFE1F:8ED860E4AB14D45DFC59FE80A2FEFEC7" box="[1013,1155,290,311]" latitude="-33.97111" longLatPrecision="19" longitude="151.19945" municipality="Port Botany" name="Port Botany" pageId="10" pageNumber="246" stateProvince="New South Wales">Port Botany</location>
|
||
,
|
||
<geoCoordinate id="EE3350F8AB14D45DFB3DFE83A373FEC7" box="[1169,1294,288,311]" degrees="33" direction="south" minutes="58" orientation="latitude" pageId="10" pageNumber="246" precision="15" seconds="45" value="-33.979168">33°58′45″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3350F8AB14D45DFAB2FE83A3E1FEC7" box="[1310,1436,288,311]" degrees="151" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="246" precision="15" seconds="1" value="151.18361">151°11′1″E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9BDAAB14D45DFC96FEE3A51AFEA6" box="[826,871,321,342]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="246" unit="m" value="7.0">
|
||
<elevation id="002AD10CAB14D45DFC96FEE3A51AFEA6" box="[826,871,321,342]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="246" unit="m" value="7.0">7 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB910FFAB14D45DFCD8FEE2A58CFEA6" box="[884,1009,320,342]" pageId="10" pageNumber="246" value="1992-07-28">
|
||
<collectingDate id="EFFDE917AB14D45DFCD8FEE2A58CFEA6" box="[884,1009,320,342]" pageId="10" pageNumber="246" value="1992-07-28">28.vii.1992</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D01FDB6AB14D45DFBADFEE2A210FEA6" box="[1025,1133,320,342]" pageId="10" pageNumber="246" type="generic">one spec.</specimenCount>
|
||
),
|
||
<collectionCode id="ED16AEFAAB14D45DFB2CFEE2A2D4FEA5" box="[1152,1193,320,341]" country="Australia" name="Australian Museum" pageId="10" pageNumber="246" type="Museum">AM</collectionCode>
|
||
W37826,
|
||
<geoCoordinate id="EE3350F8AB14D45DFAB2FEE2A3E1FEA5" box="[1310,1436,319,342]" degrees="33" direction="south" minutes="58" orientation="latitude" pageId="10" pageNumber="246" precision="15" seconds="16" value="-33.97111">33°58′16″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3350F8AB14D45DFC96FEFDA5BAFE84" box="[826,967,350,373]" degrees="151" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="246" precision="15" seconds="58" value="151.19945">151°11′58″E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFF9BDAAB14D45DFC79FEFDA27FFE84" box="[981,1026,351,372]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="246" unit="m" value="7.0">
|
||
<elevation id="002AD10CAB14D45DFC79FEFDA27FFE84" box="[981,1026,351,372]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="246" unit="m" value="7.0">7 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB910FFAB14D45DFBBCFEFDA2F0FE85" box="[1040,1165,351,373]" pageId="10" pageNumber="246" value="1992-07-27">
|
||
<collectingDate id="EFFDE917AB14D45DFBBCFEFDA2F0FE85" box="[1040,1165,351,373]" pageId="10" pageNumber="246" value="1992-07-27">27.vii.1992</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="9D01FDB6AB14D45DFB32FEFDA376FE84" box="[1182,1291,351,372]" pageId="10" pageNumber="246" type="generic">one spec.</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB8363FAB14D45DFC96FE3EA516FDDC" blockId="10.[826,1441,412,556]" pageId="10" pageNumber="246">
|
||
<emphasis id="B973EA2DAB14D45DFC96FE3EA371FE41" box="[826,1292,412,433]" italics="true" pageId="10" pageNumber="246">Other material examined (Appendix 1):</emphasis>
|
||
<collectingRegion id="49C3F8DDAB14D45DFAB7FE3FA500FE21" country="Australia" name="New South Wales" pageId="10" pageNumber="246">New South Wales</collectingRegion>
|
||
(256 specs): Botany Bay, Malabar, Port Jackson, Pittwater;
|
||
<collectingRegion id="49C3F8DDAB14D45DFB88FE7BA2D1FE1F" box="[1060,1196,473,495]" country="Australia" name="Queensland" pageId="10" pageNumber="246">Queensland</collectingRegion>
|
||
(12 specs): Shoalwater Bay;
|
||
<collectingRegion id="49C3F8DDAB14D45DFC1FFE5AA2E8FDFE" box="[947,1173,504,526]" country="Australia" name="Northern Territory" pageId="10" pageNumber="246">Northern Territory</collectingRegion>
|
||
(two specs): Arafura Sea.
|
||
</paragraph>
|
||
<paragraph id="8BB8363FAB14D45DFC96FDF6A5B8F904" blockId="10.[826,1442,596,1780]" pageId="10" pageNumber="246">
|
||
<emphasis id="B973EA2DAB14D45DFC96FDF6A225FD99" box="[826,1112,596,617]" italics="true" pageId="10" pageNumber="246">
|
||
Description of
|
||
<typeStatus id="54BC889DAB14D45DFC5FFDF6A22FFD99" box="[1011,1106,596,617]" pageId="10" pageNumber="246" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Slender, thread-like body, measuring 13.0 mm long, 0.15 mm wide, with 20 chaetigers; cylindrical in cross section (
|
||
<figureCitation id="133C2ABAAB14D45DFB61FD33A334FD56" box="[1229,1353,657,679]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">Figs 2E–G</figureCitation>
|
||
,
|
||
<figureCitation id="133C2ABAAB14D45DFAFFFD33A3EBFD57" box="[1363,1430,657,679]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">4A–C</figureCitation>
|
||
). Head elongated, cylindrical, of same width as anterior segments; anterior end truncated, with terminal ciliated mouth opening (
|
||
<figureCitation id="133C2ABAAB14D45DFB8AFD4FA2E4FCF2" box="[1062,1177,749,771]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">Figs 2E, F</figureCitation>
|
||
,
|
||
<figureCitation id="133C2ABAAB14D45DFB0FFD4FA362FCF2" box="[1187,1311,749,770]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">4A, B, D, E</figureCitation>
|
||
), extending midventrally as an elongated slit. Head smooth, with no folds or grooves, continuing to first segment with no apparent external division, with a pair of ventrolateral brownish eyespots. Anterior region with three short uniramous segments, each with notochaetae only. Second segment twice as long as first and third; RLUS = 1:2:1 (
|
||
<figureCitation id="133C2ABAAB14D45DFC48FC61A220FC28" box="[996,1117,963,985]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">Figs 2E, F</figureCitation>
|
||
,
|
||
<figureCitation id="133C2ABAAB14D45DFBC0FC61A2CDFC29" box="[1132,1200,963,985]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">4B, C</figureCitation>
|
||
). Deep groove encircling the body, between first and second chaetiger, except for a short dorsal portion (
|
||
<figureCitation id="133C2ABAAB14D45DFB17FBA2A34FFBE6" box="[1211,1330,1024,1046]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">Fig. 4A–C</figureCitation>
|
||
). Oesophageal commissures with Y shape in anterior segments. Biramous chaetigers six to ten times longer than wide, becoming shorter and compact in last three posterior segments; chaetiger 6 longest. Notochaetae of all segments similar, capillaries, decreasing in length ventrally within each fascicle, with proximal part smooth or slightly striated and mid and distal end with sculpture resembling scales (
|
||
<figureCitation id="133C2ABAAB14D45DFB0BFB57A358FAFB" box="[1191,1317,1269,1291]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">Fig. 4F, G</figureCitation>
|
||
). Acicular chaetae absent. Posterior chaetigers with fewer but longer chaetae than anterior segments (
|
||
<figureCitation id="133C2ABAAB14D45DFAB9FA90A3E9FAB7" box="[1301,1428,1330,1352]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">Fig. 4 K, L</figureCitation>
|
||
). Neurochaetae, as uncini, present from chaetiger
|
||
<quantity id="4CFF9BDAAB14D45DFADFFAF3A3DCFA97" box="[1395,1441,1361,1383]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="10" pageNumber="246" unit="in" value="4.0">4 in</quantity>
|
||
long and broad lateroventral tori, decreasing posteriorly in size and number of uncini. Anterior uncinal fields with around six to eight transverse rows of uncini (
|
||
<figureCitation id="133C2ABAAB14D45DFC38FA69A591FA11" box="[916,1004,1483,1505]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">Fig. 4H</figureCitation>
|
||
); tori of posterior part much shorter. Each uncinus bifid, provided with two nearly equally convex teeth, one offset slightly higher than the other and arranged in an oblique row (
|
||
<figureCitation id="133C2ABAAB14D45DFB70F985A337F9CC" box="[1244,1354,1575,1597]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">Fig. 4I, J</figureCitation>
|
||
). Anus terminal surrounded by a ciliated rim (
|
||
<figureCitation id="133C2ABAAB14D45DFABEF9E7A3EBF9AB" box="[1298,1430,1605,1627]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="10" pageNumber="246">Fig. 4 K–M</figureCitation>
|
||
), with two low blunted lobes on each side (
|
||
<figureCitation id="133C2ABAAB14D45DFAADF9C6A325F98A" box="[1281,1368,1636,1658]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">Fig. 2G</figureCitation>
|
||
). Tube three to four times longer than animal, with thin secreted layer encrusted with sand grains (
|
||
<figureCitation id="133C2ABAAB14D45DFA91F903A3E9F947" box="[1341,1428,1697,1719]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">Fig. 2H</figureCitation>
|
||
). Colour in alcohol pale yellow, with no pigment pattern (
|
||
<figureCitation id="133C2ABAAB14D45DFCEEF97CA5C7F904" box="[834,954,1758,1780]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="10" pageNumber="246">Fig. 2E–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8363FAB14D45BFC96F8BEA77DFD78" blockId="10.[826,1441,1820,1903]" lastBlockId="12.[164,779,198,648]" lastPageId="12" lastPageNumber="248" pageId="10" pageNumber="246">
|
||
<emphasis id="B973EA2DAB14D45DFC96F8BEA5D6F8C1" box="[826,939,1820,1841]" italics="true" pageId="10" pageNumber="246">Variation:</emphasis>
|
||
The specimens examined vary between 15 to 20 segments and
|
||
<quantity id="4CFF9BDAAB14D45DFBCBF898A2ADF8A0" box="[1127,1232,1850,1872]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.6" metricValueMin="0.6" pageId="10" pageNumber="246" unit="mm" value="11.0" valueMax="16.0" valueMin="6.0">6–16 mm</quantity>
|
||
in length. Some
|
||
<typeStatus id="54BC889DAB14D45DFC96F8FBA5D6F89E" box="[826,939,1881,1902]" pageId="10" pageNumber="246" type="paratype">paratypes</typeStatus>
|
||
and additional material show the anterior end weakly rounded instead of truncated. Eyespots have faded in some specimens, probably related to the time of storage in alcohol. Shape of pygidium varies slightly amongst specimens and probably as a result of methods of fixation and/or manipulation for observation; specimens studied under the dissecting microscope showed a well-marked bilobed pygidium (
|
||
<figureCitation id="133C2ABAAB12D45BFF00FE3EA77DFE42" box="[172,256,412,434]" captionStart="Figure 2" captionStartId="6.[164,243,1366,1385]" captionTargetBox="[164,1444,196,1337]" captionTargetId="figure-179@6.[164,1444,196,1337]" captionTargetPageId="6" captionText="Figure 2. A–D, Galathowenia annae sp. nov. Paratype, AM W37233. A, anterior end, ventrolateral view. B, anterior end, lateral view. C, posterior end, ventral view. D, tube. E–H, Galathowenia quelis sp. nov. AM W37821-37822. E, anterior end, ventral view. F, anterior end, dorsal view. G, posterior end, ventral view. H, tube. Abbreviations: b1, biramous segment; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Scale bars: A–D = 50 Mm; E–H = 75 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408801" httpUri="https://zenodo.org/record/5408801/files/figure.png" pageId="12" pageNumber="248">Fig. 2G</figureCitation>
|
||
) whereas in specimens observed under SEM, the pygidium resembled a short rim with a slightly enlarged dorsal border (
|
||
<figureCitation id="133C2ABAAB12D45BFE73FE7BA41EFE1F" box="[479,611,473,495]" captionStart="Figure 4" captionStartId="11.[144,223,1672,1691]" captionTargetBox="[182,1386,194,1643]" captionTargetId="figure-0@11.[182,1386,194,1643]" captionTargetPageId="11" captionText="Figure 4. Galathowenia quelis sp. nov. A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm." figureDoi="http://doi.org/10.5281/zenodo.5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="12" pageNumber="248">Fig. 4 K–M</figureCitation>
|
||
). We believe that this difference may be because of the critical point drying method. Some specimens are slightly brownish after preservation but none of the material examined exhibited any distinguishable colour pattern.
|
||
</paragraph>
|
||
<caption id="DF7866B7AB15D45CFF3CF92AA7CCF899" ID-DOI="http://doi.org/10.5281/zenodo.5408805" ID-Zenodo-Dep="5408805" httpUri="https://zenodo.org/record/5408805/files/figure.png" pageId="11" pageNumber="247" startId="11.[144,223,1672,1691]" targetBox="[182,1386,194,1643]" targetPageId="11">
|
||
<paragraph id="8BB8363FAB15D45CFF3CF92AA7CCF899" blockId="11.[144,1422,1672,1897]" pageId="11" pageNumber="247">
|
||
<emphasis id="B973EA2DAB15D45CFF3CF92AA42AF96C" bold="true" box="[144,599,1672,1692]" pageId="11" pageNumber="247">
|
||
Figure 4.
|
||
<taxonomicName id="4C074DBCAB15D45CFEA8F92AA78AF96B" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[260,503,1672,1691]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="247" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB15D45CFEA8F92AA78AF96B" bold="true" box="[260,503,1672,1691]" italics="true" pageId="11" pageNumber="247">Galathowenia quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB15D45CFE53F92BA42AF96C" box="[511,599,1673,1692]" pageId="11" pageNumber="247" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, anterior end, ventral view. B, anterior end, ventrolateral view. C, anterior end, dorsal view. D, detail of small spheres (diatoms?) located between the buccal ciliature. E, mouth and ventral slit, ventral view. F, anterior capillary chaetae, proximal end. G, detail of scale covering in capillary chaetae, segment 3. H, neuropodial uncinal field, segment 4 (first biramous chaetiger). I, uncini, same segment, top view. J, uncini, same segment, side view. K, posterior end, ventral view. L, posterior end, lateral view. M, anus, posterior view. Abbreviations: b1, biramous segment 1; g1-2, groove between segment 1 and 2; u1-u3, uniramous segments 1 to 3. Registration numbers: A, AM W37195; B–D, F–J, W37191; E, AM W37192; K–M, AM W37193. Scale bars: A–C = 100 Mm; D, G, I, J = 2 Mm; E, H, K–M = 10 Mm; F = 4 Mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB8363FAB12D45BFF08FD0DA746FC07" blockId="12.[164,779,687,1015]" pageId="12" pageNumber="248">
|
||
<emphasis id="B973EA2DAB12D45BFF08FD0DA71FFD35" box="[164,354,687,709]" italics="true" pageId="12" pageNumber="248">Ecological notes:</emphasis>
|
||
This species has been found mostly in estuaries and sheltered bays in muddy and sandy sediments between 1 and
|
||
<quantity id="4CFF9BDAAB12D45BFE4DFD4FA464FCF2" box="[481,537,749,771]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="248" unit="m" value="60.0">60 m</quantity>
|
||
in depth, but some specimens were also found in deeper exposed environments along the temperate and tropical coast of eastern
|
||
<collectingCountry id="F31076AFAB12D45BFEA8FCEAA70DFCAE" box="[260,368,840,862]" name="Australia" pageId="12" pageNumber="248">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="133C2ABAAB12D45BFE29FCEAA7ABFCAF" box="[389,470,840,863]" captionStart="Figure 14" captionStartId="29.[144,223,1680,1699]" captionTargetBox="[149,1424,912,1650]" captionTargetId="figure-224@29.[144,1424,908,1650]" captionTargetPageId="29" captionText="Figure 14. Distribution of Australian oweniids except for members of genus Owenia." figureDoi="http://doi.org/10.5281/zenodo.5408827" httpUri="https://zenodo.org/record/5408827/files/figure.png" pageId="12" pageNumber="248">Fig. 14</figureCitation>
|
||
).
|
||
<taxonomicName id="4C074DBCAB12D45BFE43FCEAA4A3FCAD" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[495,734,840,861]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFE43FCEAA4A3FCAD" box="[495,734,840,861]" italics="true" pageId="12" pageNumber="248">Galathowenia quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFD46FCEBA6ACFC8C" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
cohabits with
|
||
<taxonomicName id="4C074DBCAB12D45BFE21FCC5A78EFC8D" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[397,499,871,893]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="annae" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFE21FCC5A78EFC8D" box="[397,499,871,893]" italics="true" pageId="12" pageNumber="248">G. annae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFDADFCC5A420FC8C" box="[513,605,871,892]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in some
|
||
<collectingRegion id="49C3F8DDAB12D45BFD7BFCC5A74EFC6C" country="Australia" name="New South Wales" pageId="12" pageNumber="248">New South Wales</collectingRegion>
|
||
estuaries. The
|
||
<typeStatus id="54BC889DAB12D45BFE44FC24A434FC6C" box="[488,585,902,924]" pageId="12" pageNumber="248" type="holotype">holotype</typeStatus>
|
||
and other specimens were found with one parasitic copepod living inside the tube and attached to the worm body by its anterior end.
|
||
</paragraph>
|
||
<paragraph id="8BB8363FAB12D45BFF08FBBDA73BFB82" blockId="12.[164,779,1053,1138]" pageId="12" pageNumber="248">
|
||
<emphasis id="B973EA2DAB12D45BFF08FBBDA75FFBC4" box="[164,290,1055,1076]" italics="true" pageId="12" pageNumber="248">Etymology:</emphasis>
|
||
This species is dedicated to Ángeles Iglesias-Díaz (nickname Quelis), who was supportive of the project.
|
||
</paragraph>
|
||
<paragraph id="8BB8363FAB12D45BFF08FB3BA4D5F8E2" blockId="12.[164,779,1177,1903]" pageId="12" pageNumber="248">
|
||
<emphasis id="B973EA2DAB12D45BFF08FB3BA464FB5E" box="[164,537,1177,1198]" italics="true" pageId="12" pageNumber="248">
|
||
Remarks:
|
||
<taxonomicName id="4C074DBCAB12D45BFE8FFB3BA464FB5E" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[291,537,1177,1198]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">Galathowenia quelis</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFD80FB3BA4F3FB5E" box="[556,654,1177,1198]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
could be another example of a species belonging to what we have named the
|
||
<taxonomicName id="4C074DBCAB12D45BFECFFB75A7AAFB1B" baseAuthorityName="Zachs" baseAuthorityYear="1923" box="[355,471,1238,1260]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="oculata">
|
||
<emphasis id="B973EA2DAB12D45BFECFFB75A7AAFB1B" box="[355,471,1238,1260]" italics="true" pageId="12" pageNumber="248">G. oculata</emphasis>
|
||
</taxonomicName>
|
||
complex, as it shares with this species the general combination of features mentioned above. However,
|
||
<taxonomicName id="4C074DBCAB12D45BFE14FAB6A461FAD8" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[440,540,1299,1321]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFE14FAB6A461FAD8" box="[440,540,1299,1321]" italics="true" pageId="12" pageNumber="248">G. quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFD89FAB6A401FAD9" box="[549,636,1300,1321]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from previous descriptions of
|
||
<taxonomicName id="4C074DBCAB12D45BFE6AFA90A444FAB7" baseAuthorityName="Zachs" baseAuthorityYear="1923" box="[454,569,1330,1351]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="oculata">
|
||
<emphasis id="B973EA2DAB12D45BFE6AFA90A444FAB7" box="[454,569,1330,1351]" italics="true" pageId="12" pageNumber="248">G. oculata</emphasis>
|
||
</taxonomicName>
|
||
in some features (see
|
||
<tableCitation id="C6850384AB12D45BFF72FAF3A74EFA97" box="[222,307,1361,1383]" captionStart="Table 2" captionStartId="9.[136,200,525,546]" captionText="Table 2. Comparison of three Australian species of Galathowenia and Galathowenia oculata from different geographical areas" pageId="12" pageNumber="248">Table 2</tableCitation>
|
||
). The RLUS (= 1:2:1), differs from the European, west
|
||
<collectingCountry id="F31076AFAB12D45BFEC2FACDA794FA75" box="[366,489,1391,1413]" name="Greenland" pageId="12" pageNumber="248">Greenland</collectingCountry>
|
||
, and Californian specimens (1:1:1,
|
||
<bibRefCitation id="EF964BCEAB12D45BFE97FA2CA43AFA54" author="Nilsen R & Holthe T" box="[315,583,1422,1444]" pageId="12" pageNumber="248" pagination="17 - 32" refId="ref23290" refString="Nilsen R, Holthe T. 1985. Arctic and Scandinavian Oweniidae (Polychaeta) with comments on the phylogeny of the family. Sarsia 70: 17 - 32." type="journal article" year="1985">Nilsen & Holthe, 1985</bibRefCitation>
|
||
; as per
|
||
<bibRefCitation id="EF964BCEAB12D45BFD06FA2CA758FA32" author="Blake JA & Dean D" pageId="12" pageNumber="248" pagination="31 - 39" refId="ref21217" refString="Blake JA, Dean D. 1973. Polychaetous annelids collected by the R / V Hero from Baffin Island, Davis Strait, and west Greenland in 1968. Bulletin of the Southern California Academy of Sciences 72: 31 - 39." type="journal article" year="1973">Blake & Dean, 1973</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF964BCEAB12D45BFEF3FA0EA797FA32" author="Blake JA" box="[351,490,1452,1474]" pageId="12" pageNumber="248" pagination="97 - 127" refId="ref21141" refString="Blake JA. 2000. Chapter 5. Family Oweniidae Rioja, 1917. In: Blake JA, Hilbig B, Scott PV, eds. Taxonomic atlas of the benthic fauna of the Santa Maria Basin and the Western Santa Barbara Channel. Vol. 7. The Annelida. Part 4: Polychaeta: Flabelligeridae to Sternaspidae. Santa Barbara, CA: Santa Barbara Museum Natural History, 97 - 127." type="book chapter" year="2000">Blake, 2000</bibRefCitation>
|
||
: fig. 5.1a) and those from
|
||
<collectingCountry id="F31076AFAB12D45BFF08FA69A697FA10" box="[164,234,1483,1504]" name="Japan" pageId="12" pageNumber="248">Japan</collectingCountry>
|
||
(1:1:0.5,
|
||
<bibRefCitation id="EF964BCEAB12D45BFEFAFA69A409FA11" author="Imajima M & Morita Y" box="[342,628,1483,1505]" pageId="12" pageNumber="248" pagination="85 - 102" refId="ref22660" refString="Imajima M, Morita Y. 1987. Oweniidae (Annelida, Polychaeta) from Japan. Bulletin of the National Science Museum, Tokyo, Series A 13: 85 - 102." type="journal article" year="1987">Imajima & Morita, 1987</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C074DBCAB12D45BFD25FA69A474F9F0" authority="(Gibbs, 1972)" baseAuthorityName="Gibbs" baseAuthorityYear="1972" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="haplosoma">
|
||
<emphasis id="B973EA2DAB12D45BFD25FA69A72BFA0F" italics="true" pageId="12" pageNumber="248">Galathowenia haplosoma</emphasis>
|
||
(
|
||
<bibRefCitation id="EF964BCEAB12D45BFEC3FA48A47CF9F0" author="Gibbs PE" box="[367,513,1514,1536]" pageId="12" pageNumber="248" pagination="199 - 220" refId="ref21995" refString="Gibbs PE. 1972. Polychaete annelids from the Cook Islands. Journal of Zoology, London 168: 199 - 220." type="journal article" year="1972">Gibbs, 1972</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
was described from the
|
||
<collectingCountry id="F31076AFAB12D45BFF78F9AAA711F9EE" box="[212,364,1544,1566]" name="Cook Islands" pageId="12" pageNumber="248">Cook Islands</collectingCountry>
|
||
as a
|
||
<taxonomicName id="4C074DBCAB12D45BFE1AF9AAA4E9F9EE" box="[438,660,1544,1566]" class="Polychaeta" family="Oweniidae" genus="Myriochele" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="B973EA2DAB12D45BFE1AF9AAA44CF9ED" box="[438,561,1544,1565]" italics="true" pageId="12" pageNumber="248">Myriochele</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
Owing to the shape of the head and the lacking references to the acicular chaetae, we consider that it should be moved to
|
||
<taxonomicName id="4C074DBCAB12D45BFEBAF9C6A7C8F989" authorityName="Kirkegaard" authorityYear="1959" box="[278,437,1636,1657]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="genus">
|
||
<emphasis id="B973EA2DAB12D45BFEBAF9C6A7C8F989" box="[278,437,1636,1657]" italics="true" pageId="12" pageNumber="248">Galathowenia</emphasis>
|
||
</taxonomicName>
|
||
. Members of this species are small (4.0 to 7.5 mm length) and share with
|
||
<taxonomicName id="4C074DBCAB12D45BFD0BF921A576F968" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[679,779,1667,1688]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFD0BF921A576F968" box="[679,779,1667,1688]" italics="true" pageId="12" pageNumber="248">G. quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFF08F900A686F947" box="[164,251,1698,1719]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
the presence of eyes and a pygidium divided by a small cleft (
|
||
<bibRefCitation id="EF964BCEAB12D45BFEC7F962A785F925" author="Gibbs PE" box="[363,504,1728,1750]" pageId="12" pageNumber="248" pagination="199 - 220" refId="ref21995" refString="Gibbs PE. 1972. Polychaete annelids from the Cook Islands. Journal of Zoology, London 168: 199 - 220." type="journal article" year="1972">Gibbs, 1972</bibRefCitation>
|
||
: fig. 8a–c), but differ in the relative length of uniramous segments (1:1:0.5 in
|
||
<taxonomicName id="4C074DBCAB12D45BFF08F95FA73DF8E2" baseAuthorityName="Gibbs" baseAuthorityYear="1972" box="[164,320,1789,1810]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="haplosoma">
|
||
<emphasis id="B973EA2DAB12D45BFF08F95FA73DF8E2" box="[164,320,1789,1810]" italics="true" pageId="12" pageNumber="248">G. haplosoma</emphasis>
|
||
</taxonomicName>
|
||
vs. 1:2:
|
||
<quantity id="4CFF9BDAAB12D45BFE31F95FA7B7F8E3" box="[413,458,1789,1811]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="248" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C074DBCAB12D45BFE78F95FA445F8E2" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[468,568,1789,1810]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFE78F95FA445F8E2" box="[468,568,1789,1810]" italics="true" pageId="12" pageNumber="248">G. quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFDEEF95FA4E0F8E2" box="[578,669,1789,1810]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB8363FAB12D45BFF10F8BEA26CFE1F" blockId="12.[164,779,1177,1903]" lastBlockId="12.[826,1441,198,495]" pageId="12" pageNumber="248">
|
||
<taxonomicName id="4C074DBCAB12D45BFF10F8BEA7C5F8C1" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[188,440,1820,1841]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFF10F8BEA7C5F8C1" box="[188,440,1820,1841]" italics="true" pageId="12" pageNumber="248">Galathowenia quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFE7EF8BEA444F8C1" box="[466,569,1820,1841]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
also resembles
|
||
<taxonomicName id="4C074DBCAB12D45BFF08F898A777F8A0" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[164,266,1850,1872]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="annae" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFF08F898A777F8A0" box="[164,266,1850,1872]" italics="true" pageId="12" pageNumber="248">G. annae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFEBEF899A714F8A0" box="[274,361,1851,1872]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
described above, as both have anterolateral eyespots, a groove between the first and second segments, a similar number and relative length of anterior segments (RLUS), and the same
|
||
<typeStatus id="54BC889DAB12D45BFC96FEA1A516FEE8" box="[826,875,259,280]" pageId="12" pageNumber="248">type</typeStatus>
|
||
of sediment attached to the tube. Moreover, these two species are sympatric in
|
||
<collectingRegion id="49C3F8DDAB12D45BFB78FE80A3DDFEC8" box="[1236,1440,290,312]" country="Australia" name="New South Wales" pageId="12" pageNumber="248">New South Wales</collectingRegion>
|
||
(even being found in the same samples). However, they are easily distinguished by conspicuous pigmentation on the head (an attribute of
|
||
<taxonomicName id="4C074DBCAB12D45BFB78FEDCA347FE63" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[1236,1338,382,403]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="annae" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFB78FEDCA347FE63" box="[1236,1338,382,403]" italics="true" pageId="12" pageNumber="248">G. annae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFAEFFEDCA3E7FE63" box="[1347,1434,382,403]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
) and by the pygidium (as a low rim or bilobed in
|
||
<taxonomicName id="4C074DBCAB12D45BFC96FE19A5E3FE20" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[826,926,443,464]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="quelis" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFC96FE19A5E3FE20" box="[826,926,443,464]" italics="true" pageId="12" pageNumber="248">G. quelis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFC00FE19A275FE20" box="[940,1032,443,464]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
against three digitiform lobes in
|
||
<taxonomicName id="4C074DBCAB12D45BFC96FE78A5DDFE1F" authority="Capa & Parapar & Hutchings, 2012" authorityName="Capa & Parapar & Hutchings" authorityYear="2012" box="[826,928,474,495]" class="Polychaeta" family="Oweniidae" genus="Galathowenia" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="248" phylum="Annelida" rank="species" species="annae" status="sp. nov.">
|
||
<emphasis id="B973EA2DAB12D45BFC96FE78A5DDFE1F" box="[826,928,474,495]" italics="true" pageId="12" pageNumber="248">G. annae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2405756AB12D45BFC06FE78A278FE1F" box="[938,1029,474,495]" pageId="12" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |