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<document ID-DOI="10.37520/aemnp.2023.003" ID-ISSN="1804-6487" ID-Zenodo-Dep="7887332" ID-ZooBank="937DF12A-BC2F-4406-802F-63838312B0C1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683048435184" checkinUser="felipe" docAuthor="Jiang, Zhuo-Yin, Zhao, Shuang, Mai, Zu-Qi, Jia, Feng-Long &amp; Hendrich, Lars" docDate="2023" docId="03975B72FFD1761B9459CDCD521EF8EF" docLanguage="en" docName="ActaEntMusNatPra.63.1.75-102.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 63 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2023.003" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Cybister convexus Sharp 1882" docType="treatment" docVersion="2" lastPageNumber="93" masterDocId="FFAE230AFFC376099545CC10505AFFDA" masterDocTitle="Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae)" masterLastPageNumber="102" masterPageNumber="75" pageNumber="93" updateTime="1683048975263" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Review of the genus Cybister in China, with description of a new species from Guangdong (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jiang, Zhuo-Yin</mods:namePart>
<mods:affiliation>Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou 510275, China; e-mail: zhuoyin _ jiang @ 126. com; ORCID ID https: // orcid. org / 0000 - 0001 - 6825 - 4283</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Shuang</mods:namePart>
<mods:affiliation>Guangzhou Customs Technology Center, Tianhe District, Guangzhou, China; e-mail: topzs @ 163. com;</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mai, Zu-Qi</mods:namePart>
<mods:affiliation>School of Agriculture, Sun Yat-sen University, Guangzhou, 511436, China; e-mail: maizq</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jia, Feng-Long</mods:namePart>
<mods:affiliation>Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou 510275, China; &amp; corresponding author</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hendrich, Lars</mods:namePart>
<mods:affiliation>@ mail &amp; corresponding author</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-12</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>63</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>75</mods:start>
<mods:end>102</mods:end>
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<mods:url>http://dx.doi.org/10.37520/aemnp.2023.003</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.37520/aemnp.2023.003</mods:identifier>
<mods:identifier type="ISSN">1804-6487</mods:identifier>
<mods:identifier type="Zenodo-Dep">7887332</mods:identifier>
<mods:identifier type="ZooBank">937DF12A-BC2F-4406-802F-63838312B0C1</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03975B72FFD1761B9459CDCD521EF8EF" httpUri="http://treatment.plazi.org/id/03975B72FFD1761B9459CDCD521EF8EF" lastPageNumber="93" pageId="18" pageNumber="93">
<subSubSection box="[284,626,477,501]" pageId="18" pageNumber="93" type="nomenclature">
<paragraph blockId="18.[284,626,477,501]" box="[284,626,477,501]" pageId="18" pageNumber="93">
<heading box="[284,626,477,501]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="93" reason="3">
<taxonomicName ID-CoL="7SVBP" ID-ENA="398589" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[284,626,477,501]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="93" phylum="Arthropoda" rank="species" species="convexus">
<emphasis bold="true" box="[284,626,477,501]" pageId="18" pageNumber="93">
<emphasis bold="true" box="[284,483,477,501]" italics="true" pageId="18" pageNumber="93">Cybister convexus</emphasis>
<bibRefCitation author="SHARP D." box="[490,626,478,501]" pageId="18" pageNumber="93" pagination="179 - 1003" refId="ref18695" refString="SHARP D. 1882: On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series II 2: 179 - 1003 + pls 7 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[326,585,508,527]" pageId="18" pageNumber="93" type="description">
<paragraph blockId="18.[151,761,508,870]" box="[326,585,508,527]" pageId="18" pageNumber="93">
(
<figureCitation box="[332,438,508,527]" captionStart="Fig" captionStartId="17.[150,181,2032,2051]" captionTargetBox="[299,1288,607,2004]" captionTargetId="figure-217@17.[286,1300,597,2014]" captionTargetPageId="17" captionText="Fig. 13. Cybister convexus Sharp, 1882 (J, Kunming, Yunnan). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887361" httpUri="https://zenodo.org/record/7887361/files/figure.png" pageId="18" pageNumber="93">Figs 13AE</figureCitation>
;
<figureCitation box="[447,487,508,527]" captionStart="Fig" captionStartId="23.[151,182,2008,2027]" captionTargetBox="[150,1435,190,1960]" captionTargetId="figure-13@23.[150,1435,190,1960]" captionTargetPageId="23" captionText="Fig. 18. Habitats of Cybister in China. AC lake, Shanxi (C. chinensis); DE lowland marsh, Shenzhen, Guangdong (C. rugosus, C. tripunctatus lateralis and C. sugillatus); F temporary pond in a valley, Danxiashan Mt., Guangdong (C. tripunctatus lateralis and C. limbatus); G natural pond, Lankeshan Mt., Zhejiang (C. rugosus and C. brevis); H terraced fields pools, Lvchun County, Yunnan (C. convexus and C. sugillatus)." figureDoi="http://doi.org/10.5281/zenodo.7887375" httpUri="https://zenodo.org/record/7887375/files/figure.png" pageId="18" pageNumber="93">18H</figureCitation>
;
<figureCitation box="[495,532,508,527]" captionStart="Fig" captionStartId="24.[151,182,1484,1503]" captionTargetBox="[151,1430,190,1468]" captionTargetId="figure-233@24.[148,1431,190,1471]" captionTargetPageId="24" captionText="Fig. 19. Live specimens of Chinese Cybister. A C. chinensis Motschulsky, 1854 (mating couple); B C. lateralimarginalis torquatus (Fischer von Waldheim, 1829); C C. rugosus (W. S. Macleay, 1825); D C. tripunctatus lateralis (Fabricius, 1798); E C. convexus Sharp, 1882; F C. sugillatus Erichson, 1834." figureDoi="http://doi.org/10.5281/zenodo.7887377" httpUri="https://zenodo.org/record/7887377/files/figure.png" pageId="18" pageNumber="93">19E</figureCitation>
;
<figureCitation box="[541,579,508,527]" captionStart="Fig" captionStartId="25.[150,181,2049,2068]" captionTargetBox="[211,732,1171,2024]" captionTargetId="figure-17@25.[167,744,1163,2032]" captionTargetPageId="25" captionText="Fig. 22. Map of distribution of Cybister in China." figureDoi="http://doi.org/10.5281/zenodo.7887381" httpUri="https://zenodo.org/record/7887381/files/figure.png" pageId="18" pageNumber="93">22A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="93" type="reference_group">
<paragraph blockId="18.[151,761,508,870]" pageId="18" pageNumber="93">
<treatmentCitationGroup pageId="18" pageNumber="93">
<treatmentCitation author="SHARP D." box="[151,464,542,561]" page="718" pageId="18" pageNumber="93" year="1882">
<taxonomicName ID-CoL="7SVBP" authority="Sharp, 1882: 718" authorityName="Sharp" authorityPageNumber="718" authorityYear="1882" box="[151,464,542,561]" class="Insecta" family="Dytiscidae" genus="Cybister" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="93" phylum="Arthropoda" rank="species" species="convexus">
<emphasis box="[151,308,542,561]" italics="true" pageId="18" pageNumber="93">Cybister convexus</emphasis>
<bibRefCitation author="SHARP D." box="[315,464,542,561]" pageId="18" pageNumber="93" pagination="179 - 1003" refId="ref18695" refString="SHARP D. 1882: On aquatic carnivorous Coleoptera or Dytiscidae. Scientific Transactions of the Royal Dublin Society, Series II 2: 179 - 1003 + pls 7 - 18." type="journal article" year="1882">Sharp, 1882: 718</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(orig. descr.); RḖGIMBART (1899): 353 (
<collectingRegion box="[228,299,569,588]" country="China" name="Yunnan" pageId="18" pageNumber="93">Yunnan</collectingRegion>
);
<treatmentCitation author="FENG H. T." box="[316,464,569,588]" page="35" pageId="18" pageNumber="93" year="1932">
<bibRefCitation author="FENG H. T." box="[316,432,569,588]" pageId="18" pageNumber="93" pagination="17 - 37" refId="ref17339" refString="FENG H. T. 1932: Aquatic insects of China. Article II. Catalogue of Chinese Dytiscidae. Peking Natural History Bulletin 7 (1932 - 1933): 17 - 37." type="journal article" year="1932">FENG (1932)</bibRefCitation>
: 35
</treatmentCitation>
(
<collectingRegion box="[478,549,569,588]" country="China" name="Yunnan" pageId="18" pageNumber="93">Yunnan</collectingRegion>
);
<treatmentCitation author="VAZIRANI T. G." box="[566,759,569,588]" page="284" pageId="18" pageNumber="93" year="1969">
<bibRefCitation author="VAZIRANI T. G." box="[566,717,569,588]" pageId="18" pageNumber="93" pagination="221 - 341" refId="ref18728" refString="VAZIRANI T. G. 1969: Contribution to the study of aquatic beetles (Coleoptera). 2. A review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India. Oriental Insects 2 (1968): 221 - 341." type="journal article" year="1969">VAZIRANI (1969)</bibRefCitation>
: 284
</treatmentCitation>
(
<collectingRegion box="[187,257,596,615]" country="China" name="Yunnan" pageId="18" pageNumber="93">Yunnan</collectingRegion>
);
<treatmentCitation author="ZENG H." box="[272,414,596,615]" page="93" pageId="18" pageNumber="93" year="1989">
<bibRefCitation author="ZENG H." box="[272,385,596,615]" pageId="18" pageNumber="93" refId="ref19071" refString="ZENG H. 1989: Classification of Dytiscidae (Insecta: Coleoptera) - A study of the collections of Dytiscidae from major museums in China. Unpublished PhD Thesis, Department of Biology, University of Sun Yat-sen University, Guangzhou, 152 pp (in Chinese with English abstract)." type="book" year="1989">ZENG (1989)</bibRefCitation>
: 93
</treatmentCitation>
(
<collectingRegion box="[426,496,596,615]" country="China" name="Yunnan" pageId="18" pageNumber="93">Yunnan</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="93" type="materials_examined">
<paragraph blockId="18.[151,761,508,870]" pageId="18" pageNumber="93">
<emphasis bold="true" box="[151,486,637,656]" pageId="18" pageNumber="93">
Material examined.
<collectingCountry box="[324,400,637,656]" name="China" pageId="18" pageNumber="93">CHINA</collectingCountry>
:
<collectingRegion box="[407,479,637,656]" country="China" name="Yunnan" pageId="18" pageNumber="93">YUNNAN</collectingRegion>
:
</emphasis>
1J, Tia Feng on (Grassy hill 3ml. N Yunnanfu).
<date box="[269,372,664,683]" pageId="18" pageNumber="93" value="1934-08-25">25.viii.1934</date>
, C. Brownell leg. (
<collectionCode box="[528,586,664,683]" pageId="18" pageNumber="93">SYSU</collectionCode>
);
<specimenCount box="[600,631,658,683]" pageId="18" pageNumber="93" type="female">1 ♀</specimenCount>
, Honghe Hani- Yi Autonomous Prefecture, Lvchun County, Niukong Township Section, terrace,
<geoCoordinate box="[216,300,717,736]" degrees="22.9873" direction="north" orientation="latitude" pageId="18" pageNumber="93" precision="5" value="22.9873">22.9873N</geoCoordinate>
,
<geoCoordinate box="[306,399,717,736]" degrees="102.2675" direction="east" orientation="longitude" pageId="18" pageNumber="93" precision="5" value="102.2675">102.2675E</geoCoordinate>
,
<quantity box="[405,468,717,736]" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" pageId="18" pageNumber="93" unit="m" value="1336.0">1336 m</quantity>
,
<date box="[475,563,717,736]" pageId="18" pageNumber="93" value="2021-04-30">30.iv.2021</date>
,Z.Y.Jiang, Z.M.Yang, B. P.Huang &amp; Z. Q. Mai leg. (
<collectionCode box="[401,459,744,763]" pageId="18" pageNumber="93">SYSU</collectionCode>
);
<specimenCount box="[472,502,738,763]" pageId="18" pageNumber="93" type="female">1 ♀</specimenCount>
, Dehong Dai Aut. Pref. mount. Range
<quantity box="[210,264,771,790]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="18" pageNumber="93" unit="km" value="31.0">31 km</quantity>
E Luxi,
<quantity box="[339,404,771,790]" metricMagnitude="3" metricUnit="m" metricValue="2.28" pageId="18" pageNumber="93" unit="m" value="2280.0">2280 m</quantity>
,
<geoCoordinate box="[414,513,771,790]" degrees="24" direction="north" minutes="29" orientation="latitude" pageId="18" pageNumber="93" precision="15" seconds="31" value="24.491945">24°2931″N</geoCoordinate>
<geoCoordinate box="[519,618,771,790]" degrees="98" direction="east" minutes="52" orientation="longitude" pageId="18" pageNumber="93" precision="15" seconds="58" value="98.88278">98°5258″E</geoCoordinate>
,
<date box="[627,707,771,790]" pageId="18" pageNumber="93" value="2007-06-03">3.vi.2007</date>
, D.W. Wrase leg. (
<collectionCode box="[250,316,797,816]" pageId="18" pageNumber="93">LHCM</collectionCode>
); 1 J, Yunnan, Baoshan Pref. mount. Range,
<quantity box="[691,743,797,816]" metricMagnitude="4" metricUnit="m" metricValue="2.2" pageId="18" pageNumber="93" unit="km" value="22.0">22 km</quantity>
S Tengchong,
<quantity box="[255,315,824,843]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="18" pageNumber="93" unit="m" value="1750.0">1750m</quantity>
,
<geoCoordinate box="[325,424,824,843]" degrees="24" direction="north" minutes="49" orientation="latitude" pageId="18" pageNumber="93" precision="15" seconds="29" value="24.824722">24°4929″N</geoCoordinate>
<geoCoordinate box="[431,528,824,843]" degrees="98" direction="east" minutes="29" orientation="longitude" pageId="18" pageNumber="93" precision="15" seconds="27" value="98.49083">98°2927″E</geoCoordinate>
(in fishpond),
<date box="[654,757,824,843]" pageId="18" pageNumber="93" value="2009-08-30">30.viii.2009</date>
, D. W. Wrase leg. (
<collectionCode box="[303,369,851,870]" pageId="18" pageNumber="93">LHCM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="93" type="diagnosis">
<paragraph blockId="18.[151,761,894,1845]" pageId="18" pageNumber="93">
<emphasis bold="true" box="[151,263,894,917]" pageId="18" pageNumber="93">Diagnosis.</emphasis>
TL
<quantity box="[302,414,894,917]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="3.0" metricValueMin="2.4" pageId="18" pageNumber="93" unit="mm" value="27.0" valueMax="30.0" valueMin="24.0">2430 mm</quantity>
, elongate, oblong-oval (
<figureCitation box="[655,748,894,917]" captionStart="Fig" captionStartId="17.[150,181,2032,2051]" captionTargetBox="[299,1288,607,2004]" captionTargetId="figure-217@17.[286,1300,597,2014]" captionTargetPageId="17" captionText="Fig. 13. Cybister convexus Sharp, 1882 (J, Kunming, Yunnan). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887361" httpUri="https://zenodo.org/record/7887361/files/figure.png" pageId="18" pageNumber="93">Fig. 13A</figureCitation>
). Head, pronotum and elytra dark green to black, apex of elytra with reddish-brown spots. Ventral surface black; abdominal ventrites IIIIV with a yellow spot on each side (
<figureCitation box="[653,747,990,1013]" captionStart="Fig" captionStartId="17.[150,181,2032,2051]" captionTargetBox="[299,1288,607,2004]" captionTargetId="figure-217@17.[286,1300,597,2014]" captionTargetPageId="17" captionText="Fig. 13. Cybister convexus Sharp, 1882 (J, Kunming, Yunnan). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887361" httpUri="https://zenodo.org/record/7887361/files/figure.png" pageId="18" pageNumber="93">Fig. 13B</figureCitation>
). Head and pronotum smooth, elytra densely covered with conspicuous small granules. Prosternal process lanceolate, with distinct lateral bead, surface flat, apex pointed; lateral parts of metaventrite tongue-shaped, slender, not reaching lateral margins; metacoxal lines distinct, well impressed, nearly parallel medially, divergent anteriorly, along metacoxal lines with a row of punctures. Legs black; protarsomeres 13 of male broadly expanded into a palette, with four rows of adhesive setae on their ventral side; male with a single metatarsal claw; female with two metatarsal claws, posterior claw small and sinuate. Median lobe of aedeagus ʻCʼ- shaped in lateral view, slightly curved, broadest at middle, strongly narrowing in apical fifth, apex rounded (
<figureCitation box="[582,679,1406,1429]" captionStart="Fig" captionStartId="17.[150,181,2032,2051]" captionTargetBox="[299,1288,607,2004]" captionTargetId="figure-217@17.[286,1300,597,2014]" captionTargetPageId="17" captionText="Fig. 13. Cybister convexus Sharp, 1882 (J, Kunming, Yunnan). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887361" httpUri="https://zenodo.org/record/7887361/files/figure.png" pageId="18" pageNumber="93">Fig. 13C</figureCitation>
); broad medially in ventral view, apical fifth slender like a tubular spine, apex truncated (
<figureCitation box="[398,500,1470,1493]" captionStart="Fig" captionStartId="17.[150,181,2032,2051]" captionTargetBox="[299,1288,607,2004]" captionTargetId="figure-217@17.[286,1300,597,2014]" captionTargetPageId="17" captionText="Fig. 13. Cybister convexus Sharp, 1882 (J, Kunming, Yunnan). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887361" httpUri="https://zenodo.org/record/7887361/files/figure.png" pageId="18" pageNumber="93">Fig. 13D</figureCitation>
). Parameres moderately broad, strap-like apically (
<figureCitation box="[424,518,1502,1525]" captionStart="Fig" captionStartId="17.[150,181,2032,2051]" captionTargetBox="[299,1288,607,2004]" captionTargetId="figure-217@17.[286,1300,597,2014]" captionTargetPageId="17" captionText="Fig. 13. Cybister convexus Sharp, 1882 (J, Kunming, Yunnan). A habitus in dorsal view; B habitus in ventral view; C median lobe of aedeagus in lateral view; D median lobe of aedeagus in ventral view; E paramere. Scale bars: 5.0 mm (A, B), 1.0 mm (CE)." figureDoi="http://doi.org/10.5281/zenodo.7887361" httpUri="https://zenodo.org/record/7887361/files/figure.png" pageId="18" pageNumber="93">Fig. 13E</figureCitation>
).
</paragraph>
<paragraph blockId="18.[151,761,894,1845]" pageId="18" pageNumber="93">
<emphasis bold="true" box="[151,244,1534,1557]" pageId="18" pageNumber="93">Habitat.</emphasis>
The habitat information is not well known for this species. It can be found from hilly to high mountain regions (
<bibRefCitation author="ATTHAKOR W. &amp; HENDRICH L. &amp; SANGPRADUB N. &amp; BALKE M." box="[158,400,1598,1621]" pageId="18" pageNumber="93" pagination="91 - 98" refId="ref16901" refString="ATTHAKOR W., HENDRICH L., SANGPRADUB N. &amp; BALKE M. 2018: Diving beetles of the Sakaerat Biosphere Reserve, Nakhon Ratchasima Province, with four new records for Thailand (Coleoptera, Dytiscidae). Spixiana 41: 91 - 98." type="journal article" year="2018">ATTHAKOR et al. 2018</bibRefCitation>
). In
<collectingRegion box="[448,531,1598,1621]" country="China" name="Yunnan" pageId="18" pageNumber="93">Yunnan</collectingRegion>
, it was collected in a water-filled rut on a forest track from Dehong (
<bibRefCitation author="HENDRICH L." pageId="18" pageNumber="93" pagination="37 - 41" refId="ref17630" refString="HENDRICH L. 2008: Rediscovery of Acilius sinensis Peschet, 1915 (Coleoptera: Dytiscidae). Koleopterologische Rundschau 78: 37 - 41." type="journal article" year="2008">HENDRICH 2008</bibRefCitation>
) and in a fish pond. In Lvchun County it was collected at night from terraced fields (
<figureCitation box="[455,552,1694,1717]" captionStart="Fig" captionStartId="23.[151,182,2008,2027]" captionTargetBox="[150,1435,190,1960]" captionTargetId="figure-13@23.[150,1435,190,1960]" captionTargetPageId="23" captionText="Fig. 18. Habitats of Cybister in China. AC lake, Shanxi (C. chinensis); DE lowland marsh, Shenzhen, Guangdong (C. rugosus, C. tripunctatus lateralis and C. sugillatus); F temporary pond in a valley, Danxiashan Mt., Guangdong (C. tripunctatus lateralis and C. limbatus); G natural pond, Lankeshan Mt., Zhejiang (C. rugosus and C. brevis); H terraced fields pools, Lvchun County, Yunnan (C. convexus and C. sugillatus)." figureDoi="http://doi.org/10.5281/zenodo.7887375" httpUri="https://zenodo.org/record/7887375/files/figure.png" pageId="18" pageNumber="93">Fig. 18H</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="18" pageNumber="93" type="distribution">
<paragraph blockId="18.[151,761,894,1845]" pageId="18" pageNumber="93">
<emphasis bold="true" box="[151,293,1726,1749]" pageId="18" pageNumber="93">Distribution.</emphasis>
<collectingCountry box="[301,364,1726,1749]" name="China" pageId="18" pageNumber="93">China</collectingCountry>
(
<collectingRegion box="[378,461,1726,1749]" country="China" name="Yunnan" pageId="18" pageNumber="93">Yunnan</collectingRegion>
) (
<figureCitation box="[483,582,1726,1749]" captionStart="Fig" captionStartId="25.[150,181,2049,2068]" captionTargetBox="[211,732,1171,2024]" captionTargetId="figure-17@25.[167,744,1163,2032]" captionTargetPageId="25" captionText="Fig. 22. Map of distribution of Cybister in China." figureDoi="http://doi.org/10.5281/zenodo.7887381" httpUri="https://zenodo.org/record/7887381/files/figure.png" pageId="18" pageNumber="93">Fig. 22A</figureCitation>
),
<collectingCountry box="[600,656,1726,1749]" name="India" pageId="18" pageNumber="93">India</collectingCountry>
,
<collectingCountry box="[667,760,1726,1749]" name="Thailand" pageId="18" pageNumber="93">Thailand</collectingCountry>
(RḖGIMBART 1899;
<bibRefCitation author="ZIMMERMANN A." box="[349,552,1758,1781]" pageId="18" pageNumber="93" refId="ref19188" refString="ZIMMERMANN A. 1920: Pars 71. Dytiscidae, Haliplidae, Hygrobiidae, Amphizoidae. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Volumen IV. Junk, Berlin, 326 pp." type="book" year="1920">ZIMMERMANN 1920</bibRefCitation>
;
<bibRefCitation author="FENG H. T." box="[561,678,1758,1781]" pageId="18" pageNumber="93" pagination="17 - 37" refId="ref17339" refString="FENG H. T. 1932: Aquatic insects of China. Article II. Catalogue of Chinese Dytiscidae. Peking Natural History Bulletin 7 (1932 - 1933): 17 - 37." type="journal article" year="1932">FENG 1932</bibRefCitation>
,
<bibRefCitation author="FENG H. T." box="[687,756,1758,1781]" pageId="18" pageNumber="93" pagination="81 - 146" refId="ref17415" refString="FENG H. T. 1933 b: Classification of Chinese Dytiscidae. Peking Natural History Bulletin 8 (1933 - 1934): 81 - 146 + 2 pls." type="journal article" year="1933">1933b</bibRefCitation>
;
<bibRefCitation author="GSCHWENDTNER L." box="[151,384,1790,1813]" pageId="18" pageNumber="93" pagination="33 - 76" refId="ref17576" refString="GSCHWENDTNER L. 1938: Monographie der palaarktischen Dytisciden. IX. Dytiscinae. Koleopterologische Rundschau 24: 33 - 76." type="journal article" year="1938">GSCHWENDTNER 1938</bibRefCitation>
;
<bibRefCitation author="VAZIRANI T. G." box="[392,550,1790,1813]" pageId="18" pageNumber="93" pagination="221 - 341" refId="ref18728" refString="VAZIRANI T. G. 1969: Contribution to the study of aquatic beetles (Coleoptera). 2. A review of the subfamilies Noterinae, Laccophilinae, Dytiscinae and Hydroporinae (in part) from India. Oriental Insects 2 (1968): 221 - 341." type="journal article" year="1969">VAZIRANI 1969</bibRefCitation>
;
<bibRefCitation author="ZENG H." box="[559,677,1790,1813]" pageId="18" pageNumber="93" refId="ref19071" refString="ZENG H. 1989: Classification of Dytiscidae (Insecta: Coleoptera) - A study of the collections of Dytiscidae from major museums in China. Unpublished PhD Thesis, Department of Biology, University of Sun Yat-sen University, Guangzhou, 152 pp (in Chinese with English abstract)." type="book" year="1989">ZENG 1989</bibRefCitation>
;
<bibRefCitation author="GHOSH S. K. &amp; NILSSON A. N." pageId="18" pageNumber="93" pagination="1 - 77" refId="ref17515" refString="GHOSH S. K. &amp; NILSSON A. N. 2012: Catalogue of the diving beetles of India and adjacent countries (Coleoptera: Dytiscidae). Skorvnopparn Supplement 3: 1 - 77." type="journal article" year="2012">GHOSH &amp; NILSSON 2012</bibRefCitation>
;
<bibRefCitation author="ATTHAKOR W. &amp; HENDRICH L. &amp; SANGPRADUB N. &amp; BALKE M." box="[336,569,1822,1845]" pageId="18" pageNumber="93" pagination="91 - 98" refId="ref16901" refString="ATTHAKOR W., HENDRICH L., SANGPRADUB N. &amp; BALKE M. 2018: Diving beetles of the Sakaerat Biosphere Reserve, Nakhon Ratchasima Province, with four new records for Thailand (Coleoptera, Dytiscidae). Spixiana 41: 91 - 98." type="journal article" year="2018">ATTHAKOR et al. 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>