161 lines
20 KiB
XML
161 lines
20 KiB
XML
<document id="6701C080BEFC47220585E4EF03B121CC" ID-CLB-Dataset="26556" ID-DOI="10.11646/zootaxa.4614.1.4" ID-GBIF-Dataset="2fdc41ca-97b1-4e90-bdd7-921797dff2c9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3241154" ID-ZooBank="B0378DC5-5B3F-4E64-9ADA-B50408BCF374" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1559897220234" checkinUser="plazi" docAuthor="Zilberman, Bruno" docDate="2019" docId="039087B9FF72FFF9FF49736EFB19FE9C" docLanguage="en" docName="zootaxa.4614.1.4.pdf" docOrigin="Zootaxa 4614 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Termitozophilus Silvestri 1901" docType="treatment" docVersion="10" lastPageNumber="96" masterDocId="FFA9FFC1FF73FFFBFFDE7102FFCFFFF9" masterDocTitle="Synopsis of the genus Termitozophilus Silvestri (Coleoptera, Staphylinidae, Aleocharinae, Corotocini)" masterLastPageNumber="116" masterPageNumber="95" pageNumber="96" updateTime="1698733478518" updateUser="ExternalLinkService">
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<mods:title id="13AB6B735022A9417D6A5956539BE4DF">Synopsis of the genus Termitozophilus Silvestri (Coleoptera, Staphylinidae, Aleocharinae, Corotocini)</mods:title>
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<mods:namePart id="84C447CE81A267B99EC1F7FAE1A7AD31">Zilberman, Bruno</mods:namePart>
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<mods:title id="E52B7CC706F4C9C99B2D6DF511D29F59">Zootaxa</mods:title>
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<mods:part id="04B783612A7528E85359787E4982B186">
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<mods:date id="05191532E8A57997DC91B99F8528387C">2019</mods:date>
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<mods:number id="ADCD000E9BBF3D9AFF688FF1C03E8948">2019-06-07</mods:number>
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<mods:detail id="97E4B8B98436E667AA7B0C63E009A579" type="volume">
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<mods:number id="EEBD5861E5F7A68EB9E8E10425BB17B5">4614</mods:number>
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<treatment id="039087B9FF72FFF9FF49736EFB19FE9C" ID-DOI="http://doi.org/10.5281/zenodo.5945169" ID-GBIF-Taxon="157436061" ID-Zenodo-Dep="5945169" LSID="urn:lsid:plazi:treatment:039087B9FF72FFF9FF49736EFB19FE9C" httpUri="http://treatment.plazi.org/id/039087B9FF72FFF9FF49736EFB19FE9C" lastPageId="2" lastPageNumber="96" pageId="1" pageNumber="96">
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<subSubSection id="C3236524FF72FFFAFF49736EFDDEFD7F" box="[151,529,620,646]" pageId="1" pageNumber="96" type="nomenclature">
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<paragraph id="8B8636AFFF72FFFAFF49736EFDDEFD7F" blockId="1.[151,529,620,646]" box="[151,529,620,646]" pageId="1" pageNumber="96">
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<heading id="D0CE81C3FF72FFFAFF49736EFDDEFD7F" bold="true" box="[151,529,620,646]" fontSize="11" level="1" pageId="1" pageNumber="96" reason="1">
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<taxonomicName id="4C394D2CFF72FFFAFF49736EFDDEFD7F" ID-CoL="8XQ9M" authority="Silvestri, 1901" authorityName="Silvestri" authorityYear="1901" box="[151,529,620,646]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="96" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEABDFF72FFFAFF49736EFDDEFD7F" bold="true" box="[151,529,620,646]" pageId="1" pageNumber="96">
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<emphasis id="B94DEABDFF72FFFAFF49736EFE95FD7F" bold="true" box="[151,346,620,646]" italics="true" pageId="1" pageNumber="96">Termitozophilus</emphasis>
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<bibRefCitation id="EFA84B5EFF72FFFAFEBE736EFDDEFD7F" author="Silvestri, F." box="[352,529,620,646]" pageId="1" pageNumber="96" pagination="1 - 24" refId="ref6361" refString="Silvestri, F. (1901) Descrizione di nuovi Termitofili e relazioni di essi con gli ospiti: III. Bollettino dei Musei di Zoologia e Anatomia Comparata della Reale Universita di Torino, 16 (398), 1 - 24." type="journal article" year="1901">Silvestri, 1901</bibRefCitation>
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</emphasis>
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</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C3236524FF72FFFAFF4973B0FD86FC4B" pageId="1" pageNumber="96" type="reference_group">
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<paragraph id="8B8636AFFF72FFFAFF4973B0FB6FFD1E" blockId="1.[151,1436,690,804]" pageId="1" pageNumber="96">
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<treatmentCitationGroup id="AB291181FF72FFFAFF4973B0FB6FFD1E" pageId="1" pageNumber="96">
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<taxonomicName id="4C394D2CFF72FFFAFF4973B0FE30FD31" ID-CoL="8XQ9M" authority="Silvestri, 1901: 7" authorityName="Silvestri" authorityPageNumber="7" authorityYear="1901" box="[151,511,690,712]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="96" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEABDFF72FFFAFF4973B0FEF6FD31" box="[151,313,690,712]" italics="true" pageId="1" pageNumber="96">Termitozophilus</emphasis>
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<treatmentCitation id="0A9810BEFF72FFFAFE9B73B0FE30FD31" author="Silvestri, F." box="[325,511,690,712]" page="7" pageId="1" pageNumber="96" year="1901">
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<bibRefCitation id="EFA84B5EFF72FFFAFE9B73B0FE30FD31" author="Silvestri, F." box="[325,511,690,712]" pageId="1" pageNumber="96" pagination="1 - 24" refId="ref6361" refString="Silvestri, F. (1901) Descrizione di nuovi Termitofili e relazioni di essi con gli ospiti: III. Bollettino dei Musei di Zoologia e Anatomia Comparata della Reale Universita di Torino, 16 (398), 1 - 24." type="journal article" year="1901">Silvestri, 1901: 7</bibRefCitation>
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(description,
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||
<typeStatus id="5482880DFF72FFFAFD4673B1FD0BFD31" box="[664,708,691,712]" pageId="1" pageNumber="96">type</typeStatus>
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species:
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<taxonomicName id="4C394D2CFF72FFFAFCF373B0FB0DFD31" ID-CoL="55H26" authority="Silvestri, 1901" authorityName="Silvestri" authorityYear="1901" box="[813,1218,690,712]" class="Insecta" family="Staphylinidae" genus="Termitozophilus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="96" phylum="Arthropoda" rank="species" species="laetus">
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<emphasis id="B94DEABDFF72FFFAFCF373B0FBD7FD31" box="[813,1048,690,712]" italics="true" pageId="1" pageNumber="96">Termitozophilus laetus</emphasis>
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<bibRefCitation id="EFA84B5EFF72FFFAFBFB73B0FB0DFD31" author="Silvestri, F." box="[1061,1218,690,712]" pageId="1" pageNumber="96" pagination="1 - 24" refId="ref6361" refString="Silvestri, F. (1901) Descrizione di nuovi Termitofili e relazioni di essi con gli ospiti: III. Bollettino dei Musei di Zoologia e Anatomia Comparata della Reale Universita di Torino, 16 (398), 1 - 24." type="journal article" year="1901">Silvestri, 1901</bibRefCitation>
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</taxonomicName>
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);
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<treatmentCitation id="0A9810BEFF72FFFAFB0573B0FA53FD31" author="Seevers, C. H." box="[1243,1436,690,712]" page="65" pageId="1" pageNumber="96" year="1957">
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<bibRefCitation id="EFA84B5EFF72FFFAFB0573B0FA53FD31" author="Seevers, C. H." box="[1243,1436,690,712]" pageId="1" pageNumber="96" pagination="1 - 334" refId="ref6319" refString="Seevers, C. H. (1957) A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology, 40, 1 - 334. https: // doi. org / 10.5962 / bhl. title. 3797" type="journal article" year="1957">Seevers, 1957: 65</bibRefCitation>
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</treatmentCitation>
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(redescription);
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||
<treatmentCitation id="0A9810BEFF72FFFAFEB773D3FDFEFD1E" author="Campbell, J. M." box="[361,561,721,743]" page="92" pageId="1" pageNumber="96" year="1973">
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<bibRefCitation id="EFA84B5EFF72FFFAFEB773D3FDFEFD1E" author="Campbell, J. M." box="[361,561,721,743]" pageId="1" pageNumber="96" pagination="83 - 94" refId="ref5964" refString="Campbell, J. M. (1973) New species and records of Neotropical termitophilous Staphylinidae: I. Subtribe Timeparthenina. Papeis Avulsos de Zoologia, 27 (2), 83 - 94." type="journal article" year="1973">Campbell, 1973: 92</bibRefCitation>
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</treatmentCitation>
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(characters in key);
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<treatmentCitation id="0A9810BEFF72FFFAFCDC73D3FC32FD1E" author="Jacobson, H. R. & Kistner, D. H. & Pasteels, J. M." box="[770,1021,721,743]" page="18" pageId="1" pageNumber="96" year="1986">
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<bibRefCitation id="EFA84B5EFF72FFFAFCDC73D3FC32FD1E" author="Jacobson, H. R. & Kistner, D. H. & Pasteels, J. M." box="[770,1021,721,743]" pageId="1" pageNumber="96" pagination="1 - 245" refId="ref6071" refString="Jacobson, H. R., Kistner, D. H. & Pasteels, J. M. (1986) Generic revision, phylogenetic classification, and phylogeny of the termitophilous tribe Corotocini (Coleoptera: Staphylinidae). Sociobiology, 12 (1), 1 - 245." type="journal article" year="1986">
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Jacobson
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<emphasis id="B94DEABDFF72FFFAFCBA73D0FC57FD1E" box="[868,920,721,743]" italics="true" pageId="1" pageNumber="96">et al.</emphasis>
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, 1986: 18
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</bibRefCitation>
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||
</treatmentCitation>
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||
(redescription).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8B8636AFFF72FFFAFF4973F2FE29FCDD" blockId="1.[151,1436,690,804]" pageId="1" pageNumber="96">
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||
<treatmentCitationGroup id="AB291181FF72FFFAFF4973F2FE29FCDD" pageId="1" pageNumber="96">
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||
<taxonomicName id="4C394D2CFF72FFFAFF4973F2FDCDFCFF" ID-CoL="8XKKG" authority="Mann, 1923: 346" authorityName="Mann" authorityPageNumber="346" authorityYear="1923" box="[151,514,752,774]" class="Insecta" family="Staphylinidae" genus="Corymbogaster" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="96" phylum="Arthropoda" rank="genus">
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<emphasis id="B94DEABDFF72FFFAFF4973F2FEFAFCFF" box="[151,309,752,774]" italics="true" pageId="1" pageNumber="96">Corymbogaster</emphasis>
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<treatmentCitation id="0A9810BEFF72FFFAFE9D73F3FDCDFCFF" author="Mann, W. M." box="[323,514,752,774]" page="346" pageId="1" pageNumber="96" year="1923">
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||
<bibRefCitation id="EFA84B5EFF72FFFAFE9D73F3FDCDFCFF" author="Mann, W. M." box="[323,514,752,774]" pageId="1" pageNumber="96" pagination="323 - 366" refId="ref6173" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from northern South America. Zoologica, 3 (17), 323 - 366." type="journal article" year="1923">Mann, 1923: 346</bibRefCitation>
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||
</treatmentCitation>
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||
</taxonomicName>
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||
(description,
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||
<typeStatus id="5482880DFF72FFFAFD4173F3FD04FCFF" box="[671,715,753,774]" pageId="1" pageNumber="96">type</typeStatus>
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||
species:
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||
<taxonomicName id="4C394D2CFF72FFFAFCE773F2FB01FCFF" ID-CoL="YQF2" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" box="[825,1230,752,774]" class="Insecta" family="Staphylinidae" genus="Corymbogaster" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="96" phylum="Arthropoda" rank="species" species="miranda">
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<emphasis id="B94DEABDFF72FFFAFCE773F2FBF4FCFF" box="[825,1083,752,774]" italics="true" pageId="1" pageNumber="96">Corymbogaster miranda</emphasis>
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<bibRefCitation id="EFA84B5EFF72FFFAFB9473F3FB01FCFF" author="Mann, W. M." box="[1098,1230,752,774]" pageId="1" pageNumber="96" pagination="323 - 366" refId="ref6173" refString="Mann, W. M. (1923) New genera and species of termitophilous Coleoptera from northern South America. Zoologica, 3 (17), 323 - 366." type="journal article" year="1923">Mann, 1923</bibRefCitation>
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</taxonomicName>
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||
) [synonymized by
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<treatmentCitation id="0A9810BEFF72FFFAFF19720CFE69FCDD" author="Borgmeier, T." box="[199,422,782,804]" page="638" pageId="1" pageNumber="96" year="1950">
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<bibRefCitation id="EFA84B5EFF72FFFAFF19720CFE69FCDD" author="Borgmeier, T." box="[199,422,782,804]" pageId="1" pageNumber="96" pagination="625 - 676" refId="ref5883" refString="Borgmeier, T. (1950) Neue Gattungen und Arten termitophiler Staphyliniden aus Brazilien nesbt einem Katalog aller bisher aus der neotropischen Region beschriebenen Arten (Col. Staph.). Revista de Entomologia, 21 (3), 625 - 676." type="journal article" year="1950">Borgmeier, 1950: 638</bibRefCitation>
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</treatmentCitation>
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, 640].
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B8636AFFF72FFFAFF497253FD86FC4B" blockId="1.[151,1436,849,2026]" pageId="1" pageNumber="96">Body moderated physogastric, partially sclerotized; chitinous portion light to dark brown; narrow basal band of pronotum black and lateral margins of sclerites dark-brown. Sclerotized areas with spine-like bristles, moderately dense and with moderate to long sizes.</paragraph>
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</subSubSection>
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<subSubSection id="C3236524FF72FFF9FF1972BFFB19FE9C" lastPageId="2" lastPageNumber="97" pageId="1" pageNumber="96" type="description">
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||
<paragraph id="8B8636AFFF72FFFAFF1972BFFD32FA37" blockId="1.[151,1436,849,2026]" pageId="1" pageNumber="96">
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<emphasis id="B94DEABDFF72FFFAFF1972BFFEC2FC2F" bold="true" box="[199,269,957,982]" pageId="1" pageNumber="96">Head.</emphasis>
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Elongate to subquadrate (figs. 6, 7, 27, 28, 37, 38, 55–58); gula subtriangular, widest at base, fused to mentum; foramen magnum short, occupying ½ of head width, with posterior sclerotized extensions present; dorsal surface mostly smooth and with a V-shaped strong impression in front, between the eyes.
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<emphasis id="B94DEABDFF72FFFAFB1B7507FAF7FBE7" bold="true" box="[1221,1336,1029,1054]" pageId="1" pageNumber="96">Antennae</emphasis>
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with 11 antennomeres (figs. 8, 39, 58–60).
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<emphasis id="B94DEABDFF72FFFAFDC0752BFCEFFBBB" bold="true" box="[542,800,1065,1090]" pageId="1" pageNumber="96">Mouthparts. Labrum</emphasis>
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more than twice wider than long; five main long bristles on each side, distributed in two pairs in a diagonal row (D2 - D1/ M2 - M1) and P1 with variable position; epiphrarynx covered with pores and small setae (figs. 12, 43, 61, 62).
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<emphasis id="B94DEABDFF72FFFAFC1C7573FB88FB73" bold="true" box="[962,1095,1137,1162]" pageId="1" pageNumber="96">Mandibles.</emphasis>
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Symmetrical, internal margin with a mesal wide tooth (figs. 13, 44, 45).
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<emphasis id="B94DEABDFF72FFFAFDB07597FD10FB57" bold="true" box="[622,735,1173,1198]" pageId="1" pageNumber="96">Maxillae:</emphasis>
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stipes almost as wide as long, with a long bristle at apex, close to first palpomere and four short bristles close to galea; galea subquadrate to elongate, covered with bristles of varied lengths at margins; lacinia narrow, as long as galea; surface scarcely covered with moderate long bristles; maxillary palpi densely setose and with four palpomeres with variable level of sclerotization: palpomere I short and subquadrate, and II–IV elongate; only palpomeres II and III covered with long bristles (figs. 11, 42).
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<emphasis id="B94DEABDFF72FFFAFAE87427FA53FAC7" bold="true" box="[1334,1436,1317,1342]" pageId="1" pageNumber="96">Labium.</emphasis>
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Prementum with labial palpi setose, 3-articulated; palpomeres I and II subquadrate; last elongate and narrow; ligula narrow at base and with apex round or bifid (figs. 10, 41); mentum and submentum distinct or not; bristles preapical (
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<emphasis id="B94DEABDFF72FFFAFF327490FEC7FA50" box="[236,264,1426,1449]" italics="true" pageId="1" pageNumber="96">pa</emphasis>
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) and proximal (
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<emphasis id="B94DEABDFF72FFFAFE1D7490FE12FA50" box="[451,477,1426,1449]" italics="true" pageId="1" pageNumber="96">px</emphasis>
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) always present and apical (
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<emphasis id="B94DEABDFF72FFFAFCF97490FC8CFA50" box="[807,835,1426,1449]" italics="true" pageId="1" pageNumber="96">ap</emphasis>
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) with presence variable. Bristle pre-apical long, and proximal short or moderate in size (figs. 9, 40, 63, 64).
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</paragraph>
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<paragraph id="8B8636AFFF72FFFAFF1974DBFC12F933" blockId="1.[151,1436,849,2026]" pageId="1" pageNumber="96">
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<emphasis id="B94DEABDFF72FFFAFF1974DBFE79FA0B" bold="true" box="[199,438,1497,1522]" pageId="1" pageNumber="96">Thorax. Prothorax.</emphasis>
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Pronotum almost as wide as long, wider at anterior region, and slightly narrowed posteriorly; discal area shiny and convex, covered with long bristles (figs. 2, 4, 25A, 30, 34).
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<emphasis id="B94DEABDFF72FFFAFAD874FFFA53F9EF" bold="true" box="[1286,1436,1533,1558]" pageId="1" pageNumber="96">Mesothorax.</emphasis>
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Mesonotum slightly shorter than metanotum. Mesosternum without carina and fused to metasternum.
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<emphasis id="B94DEABDFF72FFFAFAD97723FA53F9C3" bold="true" box="[1287,1436,1569,1594]" pageId="1" pageNumber="96">Metathorax.</emphasis>
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Metanotum: endosternite without extended articulating process, procoxal insertions close to each other; bristles covering all posterior region (figs. 14, 15, 46, 47).
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<emphasis id="B94DEABDFF72FFFAFD11776BFCC1F97B" bold="true" box="[719,782,1641,1666]" pageId="1" pageNumber="96">Legs.</emphasis>
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Well-developed, elongate and slender (figs. 1, 3, 24A, 29, 32).
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<emphasis id="B94DEABDFF72FFFAFF14778FFE8EF95F" bold="true" box="[202,321,1677,1702]" pageId="1" pageNumber="96">Scutellum</emphasis>
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slightly larger than long.
|
||
<emphasis id="B94DEABDFF72FFFAFDBE778FFD62F95F" bold="true" box="[608,685,1677,1702]" pageId="1" pageNumber="96">Elytra</emphasis>
|
||
trapezoidal or subquadrate, covering meso- and metatorax; surface covered with long bristles (figs. 16, 25, 48).
|
||
<emphasis id="B94DEABDFF72FFFAFD5677B3FCDFF933" bold="true" box="[648,784,1713,1738]" pageId="1" pageNumber="96">Hind wings</emphasis>
|
||
present or absent.
|
||
</paragraph>
|
||
<paragraph id="8B8636AFFF72FFF9FF1977D7FB19FE9C" blockId="1.[151,1436,849,2026]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="97" pageId="1" pageNumber="96">
|
||
<emphasis id="B94DEABDFF72FFFAFF1977D7FEF3F917" bold="true" box="[199,316,1749,1774]" pageId="1" pageNumber="96">Abdomen</emphasis>
|
||
expanded laterally, tergites and sternite with secondary sclerotization, varying in degree and size; inner and outer paratergites fusioned to a single sclerite by secondary sclerotization; each inner and outer paratergite with two rows of three to six bristles (figs. 1–4, 24A, 25A–B, 26A–B, 29–31); tergite I represented by a sclerotized sclerite attached to metanotum, with medial region projected backwards (figs. 15, 47); segments II-X each with tergite and sternite complete, except in female which sternite IX is absent; tergite VIII from slightly subquadrate to about twice as wide as long, with two rows of six main long bristles, one in medial and other in apical region; short bristles cover whole surface (figs. 17, 49); sternite VIII about three times wider than long, covered by short bristles, with six long bristles distributed symmetrically on apical margin, three on each side of sternite; bristles in a transversal row at median region also present (figs. 18, 50); sternite IX in male in one piece of irregular shape, moderated sclerotized, with two bristles on apical region (fig. 23); in female, sternite IX is presented as a pair of hemiesternite laterally attatched to tergite IX (fig. 51); tergite IX bilobed, each lobe with five to six long bristles (figs. 51, 19); tergite X not bilobed, with three long bristles each side and two at medial region (figs. 19, 51).
|
||
<emphasis id="B94DEABDFF71FFF9FE957025FE08FEB9" bold="true" box="[331,455,295,320]" pageId="2" pageNumber="97">Aedeagus:</emphasis>
|
||
lateral lobes broad in general aspect; median lobe bulbous and slighter shorter than lateral lobes (figs. 21, 22, 53, 54).
|
||
<emphasis id="B94DEABDFF71FFF9FDC6704EFD7AFE9C" bold="true" box="[536,693,332,357]" pageId="2" pageNumber="97">Spermatheca</emphasis>
|
||
with capsule large and sclerotized (figs. 20, 52).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |