treatments-xml/data/E9/1F/87/E91F87EDFF95FFBDD9DA8B19FF10FDB2.xml

<document id="42CED3B4545AC9F5734C3227F44D6EB7" ID-CLB-Dataset="37427" ID-DOI="10.5281/zenodo.293461" ID-GBIF-Dataset="ff02939e-3601-42a7-8eda-24e911e4b392" ID-ISSN="1175-5326" ID-Zenodo-Dep="293461" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481813413499" checkinUser="plazi" docAuthor="J. Pablo Jayat, Pablo E. Ortiz, Jorge Salazar-Bravo, Ulyses F. J. Pardiñas &amp; Guillermo D’Elía" docDate="2010" docId="E91F87EDFF95FFBDD9DA8B19FF10FDB2" docLanguage="en" docName="zt02409p061.pdf.imf" docOrigin="Zootaxa 2409" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Akodon caenosus Thomas 1918" docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="1526FF95FF83FFA6D94D8E7FFFD3FF95" masterDocTitle="The Akodon boliviensis species group (Rodentia: Cricetidae: Sigmodontinae) in Argentina: species limits and distribution, with the description of a new entity" masterLastPageNumber="61" masterPageNumber="1" pageNumber="23" updateTime="1698434547521" updateUser="ExternalLinkService">
<mods:mods id="380F382A91F6743EF0FC48950ACF88FD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="30537F848AF785F69B916AF8E2602A39">
<mods:title id="EED6784558965DB61E1737F615FF62AC">The Akodon boliviensis species group (Rodentia: Cricetidae: Sigmodontinae) in Argentina: species limits and distribution, with the description of a new entity</mods:title>
</mods:titleInfo>
<mods:name id="2D13E385A5B1C7E9D1908C7D4E17ACB0" type="personal">
<mods:role id="9741030CCE4AE2BFDA17FD605D09F122">
<mods:roleTerm id="16E7F926E2B7BBFA636D083A439647EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="32348005017F6C723CBDD3F97E5E5C20">J. Pablo Jayat</mods:namePart>
</mods:name>
<mods:name id="AB17ADC2191E565F7BB03CBE3FBC1070" type="personal">
<mods:role id="3540513B494CF4D2FF19F20383B21DE9">
<mods:roleTerm id="0E7DF366C96B5BE9A1D1A1243DC8EC3B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8129AFCF703122A3AE9781BD3AEFCFFD">Pablo E. Ortiz</mods:namePart>
</mods:name>
<mods:name id="DCA2E660B6CDF068AE001C8DB3E130AE" type="personal">
<mods:role id="3BE73FB15801519DF7CDD0B4F68ED567">
<mods:roleTerm id="86EB9C6D79FAC7E789396E8FFF80BC80">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77334F96A7EB43ED820D3A8A56566F99">Jorge Salazar-Bravo</mods:namePart>
</mods:name>
<mods:name id="0CC572AEBEA557B65296CD9602BDBAC8" type="personal">
<mods:role id="CC923A1C55AF7A0914BE9435F97B0AE2">
<mods:roleTerm id="55CEEF431CA9A44BC2EC2038F3857C1B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57E89D271AEF6DDFFF721CAA3B26CF75">Ulyses F. J. Pardiñas</mods:namePart>
</mods:name>
<mods:name id="B7118BA0BDCA88F8E7A076809DB212BD" type="personal">
<mods:role id="B98D5C6B27A62E54FDC938055A5AC406">
<mods:roleTerm id="6C37FC99411D96728DD1A82A99029EE8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08E8A433A2C7181310137A1BAE440EF7">Guillermo D’Elía</mods:namePart>
</mods:name>
<mods:typeOfResource id="0DC6CD188D1EE531F6B43CD333F87826">text</mods:typeOfResource>
<mods:relatedItem id="E08B6155C9FAA8B601B2C588C7EC2320" type="host">
<mods:titleInfo id="4D6060938159789CAE41032314A1A0BD">
<mods:title id="ECD4E35AC183A941964B842FA247E522">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8106B22FB8C710149D20FBE1D0393032">
<mods:date id="A6159DE3654A2CF196B03E2D41B61657">2010</mods:date>
<mods:detail id="C6606595CDC382C6AED77E05FAB4A1DE" type="volume">
<mods:number id="1F393E33AC03215D3695143F63DD9251">2409</mods:number>
</mods:detail>
<mods:extent id="C707FE83D0AC035EA541B8B56F33349C" unit="page">
<mods:start id="11FA89FD6D826CEF8C05E29E20E62BEB">1</mods:start>
<mods:end id="66D85408E225E883F243F83CD251C880">61</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9187BC579D2FD8AF119E114F9F2936F4">journal article</mods:classification>
<mods:identifier id="E544947C21FCD72FB0CF52F58F5EE7AC" type="CLB-Dataset">37427</mods:identifier>
<mods:identifier id="0CC555F0499B425F6C8D56E53A300B68" type="DOI">10.5281/zenodo.293461</mods:identifier>
<mods:identifier id="3AAA6F2663B16BD7161DA327D177E738" type="GBIF-Dataset">ff02939e-3601-42a7-8eda-24e911e4b392</mods:identifier>
<mods:identifier id="8900956BF241D3F1EF55F2BBBE280F44" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BAB1389B59AABF66F434F2D6398A4D29" type="Zenodo-Dep">293461</mods:identifier>
</mods:mods>
<treatment id="E91F87EDFF95FFBDD9DA8B19FF10FDB2" ID-DOI="http://doi.org/10.5281/zenodo.6196151" ID-GBIF-Taxon="125137299" ID-Zenodo-Dep="6196151" LSID="urn:lsid:plazi:treatment:E91F87EDFF95FFBDD9DA8B19FF10FDB2" httpUri="http://treatment.plazi.org/id/E91F87EDFF95FFBDD9DA8B19FF10FDB2" lastPageId="27" lastPageNumber="25" pageId="22" pageNumber="23">
<subSubSection id="29AC6570FF95FFB0D9DA8B19FBACFA70" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="610936FBFF95FFB0D9DA8B19FDF3FA15" blockId="22.[151,544,1382,1408]" box="[151,544,1382,1408]" pageId="22" pageNumber="23">
<heading id="3A418197FF95FFB0D9DA8B19FDF3FA15" bold="true" box="[151,544,1382,1408]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="A6B64D78FF95FFB0D9DA8B19FDF3FA15" authority="Thomas, 1918" authorityName="Thomas" authorityYear="1918" box="[151,544,1382,1408]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF95FFB0D9DA8B19FDF3FA15" bold="true" box="[151,544,1382,1408]" pageId="22" pageNumber="23">
<emphasis id="53C2EAE9FF95FFB0D9DA8B19FEBFFA15" bold="true" box="[151,364,1382,1408]" italics="true" pageId="22" pageNumber="23">Akodon caenosus</emphasis>
<bibRefCitation id="05274B0AFF95FFB0D83E8B19FDF3FA15" author="Thomas, O." box="[371,544,1382,1408]" journalOrPublisher="E. Budin. Annals and Magazine of Natural History" pageId="22" pageNumber="23" pagination="186 - 193" part="9" refString="Thomas, O. (1918) On small mammals form Salta and Jujuy collected by Mr. E. Budin. Annals and Magazine of Natural History, (9) 1, 186 - 193." title="On small mammals form Salta and Jujuy collected by Mr" type="journal article" year="1918">Thomas, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="610936FBFF95FFB0D9DA8BD0FBACFA70" blockId="22.[151,1151,1454,1510]" pageId="22" pageNumber="23">
<taxonomicName id="A6B64D78FF95FFB0D9DA8BD0FD6AFA50" authority="Barquez, Diaz and Goytia, 1994" authorityName="Barquez, Diaz and Goytia" authorityYear="1994" box="[151,697,1454,1478]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="diminutus">
<emphasis id="53C2EAE9FF95FFB0D9DA8BD0FE8BFA53" box="[151,344,1455,1478]" italics="true" pageId="22" pageNumber="23">Akodon diminutus</emphasis>
Barquez, Díaz and Goytia, 1994
</taxonomicName>
. Nomen nudum (see 
<bibRefCitation id="05274B0AFF95FFB0DAD68BD1FBA3FA50" author="Galliari, C. A. &amp; Pardinas, U. F. J. &amp; Goin, F. J." box="[923,1136,1454,1478]" journalOrPublisher="Mastozoologia Neotropical" pageId="22" pageNumber="23" pagination="39 - 61" part="3" refString="Galliari, C. A., Pardinas, U. F. J. &amp; Goin, F. J. (1996) Lista comentada de los mamiferos argentinos. Mastozoologia Neotropical, 3, 39 - 61." title="Lista comentada de los mamiferos argentinos" type="journal article" year="1996">
Galliari 
<emphasis id="53C2EAE9FF95FFB0DABE8BD0FBF1FA53" box="[1011,1058,1455,1478]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
., 1996
</bibRefCitation>
). 
<taxonomicName id="A6B64D78FF95FFB0D9DA8BB0FCCCFA70" authority="Diaz, Barquez, Braun and Mares, 1999" authorityName="Diaz, Barquez, Braun and Mares" authorityYear="1999" box="[151,799,1486,1510]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF95FFB0D9DA8BB0FEA7FA73" box="[151,372,1487,1510]" italics="true" pageId="22" pageNumber="23">Akodon aliquantulus</emphasis>
<bibRefCitation id="05274B0AFF95FFB0D8368BB1FCCCFA70" author="Diaz, M. M. &amp; Barquez, R. M. &amp; Braun, J. K. &amp; Mares, M. A." box="[379,799,1486,1509]" journalOrPublisher="Journal of Mammalogy" pageId="22" pageNumber="23" pagination="786 - 798" part="80" refString="Diaz, M. M., Barquez, R. M., Braun, J. K. &amp; Mares, M. A. (1999) A new species of Akodon (Muridae: Sigmodontinae) from Northwestern Argentina. Journal of Mammalogy, 80, 786 - 798." title="A new species of Akodon (Muridae: Sigmodontinae) from Northwestern Argentina" type="journal article" year="1999">Díaz, Barquez, Braun and Mares, 1999</bibRefCitation>
</taxonomicName>
. Journal of Mammalogy, 80:788.
</paragraph>
</subSubSection>
<subSubSection id="29AC6570FF95FFBDD9DA886BFF10FDB2" lastPageId="27" lastPageNumber="28" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="610936FBFF95FFB0D9DA886BFDA1F9BB" blockId="22.[151,1436,1556,2030]" box="[151,626,1556,1582]" pageId="22" pageNumber="23">
<typeStatus id="BE0D8859FF95FFB0D9DA886BFED7F9BB" box="[151,260,1556,1582]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
: 
<collectionCode id="07A7AE3EFF95FFB0D85E886BFEB8F9BB" box="[275,363,1556,1582]" pageId="22" pageNumber="23">BMNH</collectionCode>
18.1.1.38, adult male.
</paragraph>
<paragraph id="610936FBFF95FFB0D98B8846FCFAF9C6" blockId="22.[151,1436,1556,2030]" box="[198,809,1593,1619]" pageId="22" pageNumber="23">
<typeStatus id="BE0D8859FF95FFB0D98B8846FED3F9C6" box="[198,256,1593,1619]" pageId="22" pageNumber="23">Type</typeStatus>
locality: León, 
<collectingRegion id="A372F819FF95FFB0D8F78846FE29F9C6" box="[442,506,1593,1619]" country="Argentina" name="Jujuy" pageId="22" pageNumber="23">Jujuy</collectingRegion>
, 
<quantity id="A64E9B1EFF95FFB0DB4B8846FD8CF9C6" box="[518,607,1593,1619]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="23" unit="m" value="1500.0">1500 m</quantity>
(
<bibRefCitation id="05274B0AFF95FFB0DB238846FCCFF9C6" author="Thomas, O." box="[622,796,1593,1619]" journalOrPublisher="E. Budin. Annals and Magazine of Natural History" pageId="22" pageNumber="23" pagination="186 - 193" part="9" refString="Thomas, O. (1918) On small mammals form Salta and Jujuy collected by Mr. E. Budin. Annals and Magazine of Natural History, (9) 1, 186 - 193." title="On small mammals form Salta and Jujuy collected by Mr" type="journal article" year="1918">Thomas, 1918</bibRefCitation>
).
</paragraph>
<paragraph id="610936FBFF95FFB0D98B8821FEC7F89B" blockId="22.[151,1436,1556,2030]" pageId="22" pageNumber="23">
Description: Detailed morphological description in 
<bibRefCitation id="05274B0AFF95FFB0DA688821FBDAF9ED" author="Myers, P. &amp; Patton, J. L. &amp; Smith, M. F." box="[805,1033,1630,1656]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="22" pageNumber="23" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. &amp; Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
Myers 
<emphasis id="53C2EAE9FF95FFB0DA358820FC7FF9ED" box="[888,940,1631,1656]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (1990)
</bibRefCitation>
(as 
<taxonomicName id="A6B64D78FF95FFB0DD718820FAD6F9ED" box="[1084,1285,1630,1656]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="puer" subSpecies="caenosus">
<emphasis id="53C2EAE9FF95FFB0DD718820FB9DF9ED" box="[1084,1102,1631,1656]" italics="true" pageId="22" pageNumber="23">A</emphasis>
. 
<emphasis id="53C2EAE9FF95FFB0DD118820FAD6F9ED" box="[1116,1285,1631,1656]" italics="true" pageId="22" pageNumber="23">puer caenosus</emphasis>
</taxonomicName>
). Additional morphological and morphometric data for some populations of northwestern 
<collectingCountry id="19A1766BFF95FFB0DD1E88FBFB1BF90B" box="[1107,1224,1668,1694]" name="Argentina" pageId="22" pageNumber="23">Argentina</collectingCountry>
can be found in 
<bibRefCitation id="05274B0AFF95FFB0D9DA88D6FC97F956" author="Thomas, O." box="[151,836,1705,1731]" journalOrPublisher="E. Budin. Annals and Magazine of Natural History" pageId="22" pageNumber="23" pagination="186 - 193" part="9" refString="Thomas, O. (1918) On small mammals form Salta and Jujuy collected by Mr. E. Budin. Annals and Magazine of Natural History, (9) 1, 186 - 193." title="On small mammals form Salta and Jujuy collected by Mr" type="journal article" year="1918">
Thomas (1918; original description of 
<taxonomicName id="A6B64D78FF95FFB0DB2688D6FCEAF957" box="[619,825,1705,1731]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="subSpecies" species="puer" subSpecies="caenosus">
<emphasis id="53C2EAE9FF95FFB0DB2688D6FDAEF957" box="[619,637,1705,1730]" italics="true" pageId="22" pageNumber="23">A</emphasis>
. 
<emphasis id="53C2EAE9FF95FFB0DBC388D6FCEAF957" box="[654,825,1705,1730]" italics="true" pageId="22" pageNumber="23">puer caenosus</emphasis>
</taxonomicName>
)
</bibRefCitation>
, 
<bibRefCitation id="05274B0AFF95FFB0DA1988D6FB85F956" author="Barquez, R. M. &amp; Williams, D. F. &amp; Mares, M. A. &amp; Genoways, H. H." box="[852,1110,1705,1731]" journalOrPublisher="Annals of Carnegie Museum" pageId="22" pageNumber="23" pagination="379 - 403" part="49" refString="Barquez, R. M., Williams, D. F., Mares, M. A. &amp; Genoways, H. H. (1980) Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina. Annals of Carnegie Museum, 49, 379 - 403." title="Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina" type="journal article" year="1980">
Barquez 
<emphasis id="53C2EAE9FF95FFB0DA8C88D6FC24F957" box="[961,1015,1705,1730]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
. (1980)
</bibRefCitation>
and 
<bibRefCitation id="05274B0AFF95FFB0DDD888D6FAFBF956" author="Diaz, M. M." bookContentInfo="640 pp." box="[1173,1320,1705,1731]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="22" pageNumber="23" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">Díaz (1999)</bibRefCitation>
. Here we summarize the characteristics of populations along the latitudinal and altitudinal gradients of northwestern 
<collectingCountry id="19A1766BFF95FFB0D9DA888BFEC3F89B" box="[151,272,1780,1806]" name="Argentina" pageId="22" pageNumber="23">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="610936FBFF95FFB1D98B8966FC8EFF42" blockId="22.[151,1436,1556,2030]" lastBlockId="23.[151,1437,152,2007]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<taxonomicName id="A6B64D78FF95FFB0D98B8966FE41F8A7" box="[198,402,1817,1842]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF95FFB0D98B8966FE41F8A7" box="[198,402,1817,1842]" italics="true" pageId="22" pageNumber="23">Akodon caenosus</emphasis>
</taxonomicName>
is the smallest species of the 
<taxonomicName id="A6B64D78FF95FFB0DBA38966FCB8F8A7" box="[750,875,1817,1842]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF95FFB0DBA38966FCB8F8A7" box="[750,875,1817,1842]" italics="true" pageId="22" pageNumber="23">boliviensis</emphasis>
</taxonomicName>
group in northwestern 
<collectingCountry id="19A1766BFF95FFB0DD338966FB25F8A6" box="[1150,1270,1817,1843]" name="Argentina" pageId="22" pageNumber="23">Argentina</collectingCountry>
. Dorsal coloration uniform and highly variable, ochraceous brown with yellowish, rufous, or olivaceous casts. Ears of the same color than the dorsum and with a tuft of hairs in front. Laterals clearer than dorsum and more richly colored in some specimens. Venter paler, whitish gray, buffy gray, yellowish or even ruddy, clearly contrasting with dorsum. Chin with few white hairs that do not form a conspicuous patch. Inguinal region with a more intense tinge than the rest of the venter in some specimens. Fore and hind feet covered with whitish or buffy hairs. Tail strongly bicolored, dorsally blackish-brown and ventrally whitish or buffy, densely or sparsely covered depending on the analyzed populations and individuals.
</paragraph>
<paragraph id="610936FBFF94FFB1D98B8E9DFD2BFD77" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
The skull is the smallest among the 
<taxonomicName id="A6B64D78FF94FFB1DB3D8E9DFD1AFF6E" box="[624,713,226,251]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF94FFB1DB3D8E9DFD1AFF6E" box="[624,713,226,251]" italics="true" pageId="23" pageNumber="24">Akodon</emphasis>
</taxonomicName>
species of northwestern 
<collectingCountry id="19A1766BFF94FFB1DABD8E9DFBBAFF69" box="[1008,1129,226,252]" name="Argentina" pageId="23" pageNumber="24">Argentina</collectingCountry>
. Rostrum short, with very narrow and shallow zygomatic notches and frontal sinuses not well developed. Interorbital region comparatively broad, with hourglass shaped and with margins rounded or slightly squared. Zygomatic arches not flared, braincase small and inflated, with temporal and lambdoid crests not well developed. Zygomatic plate narrow with anterior border straight or slightly concave and generally slopes gently backward from bottom to top. In most of the studied specimens the posterior ascending process of alisphenoid does not reach the squamoso-alisphenoid groove or only touch their lower margin. Hamular process of squamosal delicate, but in a few individuals it is strongly built. Incisive foramina generally extended to the protocone of M1, but in some individuals its reach the hypoflexus or even further back. Mesopterygoid fossa very narrow, its anterior margin rounded or slightly squared and with lateral margins straight and slightly divergent backward. Parapterygoid fossa generally broader than mesopterygoid, with convex border and slightly divergent backward. Mandibular ramus very delicate. Masseteric crest extends slightly behind the anterior margin of m1. Capsular projection generally poorly developed and situated at the same level or behind the coronoid process. Angular process is less extended backward than the condyle.
</paragraph>
<paragraph id="610936FBFF94FFB1D98B8C92FD9BFB99" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">Upper incisors orthodont or slightly opistodont. The procingulum and anteromedian flexus of M1 are well developed. Anteroloph and mesoloph are always present and in some individuals an enteroloph can be observed. The M2 has a conspicuous mesoloph and the M3 shows great variability. In old individuals it is completely oval shaped but in young ones anteroflexus, metaflexus and hipoflexus are present. The m1 has the procingulum and anteromedian flexid well developed and presents protostylid and ectostylid. In some individuals a remnant of metastylid and mesostylid can be observed. The m2 retains only the labial stylid although some specimens have a very small ectostylid. The m3 presents deep labial and lingual flexids. Some individuals show a small protostylid.</paragraph>
<paragraph id="610936FBFF94FFB1D98B8A67FCAFFBC2" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Karyotype: 2n = 34, FN = 40, based on six specimens from León, 
<collectingRegion id="A372F819FF94FFB1DD408A67FB9DFBA7" box="[1037,1102,1048,1074]" country="Argentina" name="Jujuy" pageId="23" pageNumber="24">Jujuy</collectingRegion>
, and two from El Cadillal, 
<collectingRegion id="A372F819FF94FFB1D9DA8A42FED6FBC2" box="[151,261,1085,1111]" country="Argentina" name="Tucuman" pageId="23" pageNumber="24">Tucumán</collectingRegion>
(
<bibRefCitation id="05274B0AFF94FFB1D8598A42FDD5FBC2" author="Barquez, R. M. &amp; Williams, D. F. &amp; Mares, M. A. &amp; Genoways, H. H." box="[276,518,1085,1111]" journalOrPublisher="Annals of Carnegie Museum" pageId="23" pageNumber="24" pagination="379 - 403" part="49" refString="Barquez, R. M., Williams, D. F., Mares, M. A. &amp; Genoways, H. H. (1980) Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina. Annals of Carnegie Museum, 49, 379 - 403." title="Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina" type="journal article" year="1980">
Barquez 
<emphasis id="53C2EAE9FF94FFB1D8338A42FE61FBC3" box="[382,434,1085,1110]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 1980
</bibRefCitation>
; 
<bibRefCitation id="05274B0AFF94FFB1DB5F8A42FD3DFBC2" author="Vitullo, A. D. &amp; Merani, M. S. &amp; Reig, O. A. &amp; Kajon, A. E. &amp; Scaglia, O. &amp; Espinosa, M. B. &amp; Perez-Zapata, A." box="[530,750,1085,1111]" journalOrPublisher="Journal of Mammalogy" pageId="23" pageNumber="24" pagination="69 - 80" part="67" refString="Vitullo, A. D., Merani, M. S., Reig, O. A., Kajon, A. E., Scaglia, O., Espinosa, M. B. &amp; Perez-Zapata, A. (1986) Cytogenetics of South American Akodont Rodents (Cricetidae): new kariotypes and chromosomal banding patterns of argentinian and uruguayan forms. Journal of Mammalogy, 67, 69 - 80." title="Cytogenetics of South American Akodont Rodents (Cricetidae): new kariotypes and chromosomal banding patterns of argentinian and uruguayan forms" type="journal article" year="1986">
Vitullo 
<emphasis id="53C2EAE9FF94FFB1DB278A42FD4DFBC3" box="[618,670,1085,1110]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
., 1986
</bibRefCitation>
as 
<taxonomicName id="A6B64D78FF94FFB1DA588A42FCBFFBC3" box="[789,876,1085,1111]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF94FFB1DA588A42FCF4FBC3" box="[789,807,1085,1110]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DA7B8A42FCBFFBC3" box="[822,876,1085,1110]" italics="true" pageId="23" pageNumber="24">puer</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="610936FBFF94FFB1D98B8A1DFD71F982" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Variation: We have observed substantial morphological variation for this species. Within the same population, we find pale and dark individuals, some of them with striking rufous tones, particularly lactating females. Among the cranial characters, the development of zygomatic notches, zygomatic plate, and the thickness of hamular process show some variability. We also observed morphometric variations for individuals of the same age and population, with remarkable differences among some of the studied localities. In addition, consistent variation in the coloration pattern with elevation and some geographic variation for some morphometric variables are also apparent. The specimens from high altitude populations are conspicuously paler than those from lower localitites, showing more buffy tinges in the coloration of the fur and more developed eyerings. These populations come from open environments characterized as the ecotone between humid grasslands and semiarid areas (e.g., the Prepuna or impoverished grasslands of high-Andean region). Specimens from central 
<collectingRegion id="A372F819FF94FFB1D87D8BA7FEB8FA67" box="[304,363,1496,1522]" country="Argentina" name="Salta" pageId="23" pageNumber="24">Salta</collectingRegion>
and central 
<collectingRegion id="A372F819FF94FFB1D8B38BA7FDBFFA67" box="[510,620,1496,1522]" country="Argentina" name="Tucuman" pageId="23" pageNumber="24">Tucumán</collectingRegion>
provinces are on average smaller than individuals from Bárcena and Reyes, southern 
<collectingRegion id="A372F819FF94FFB1D8148B82FE49F982" box="[345,410,1533,1559]" country="Argentina" name="Jujuy" pageId="23" pageNumber="24">Jujuy</collectingRegion>
, near the 
<typeStatus id="BE0D8859FF94FFB1DB478B82FDEFF982" box="[522,572,1533,1559]" pageId="23" pageNumber="24">type</typeStatus>
locality.
</paragraph>
<paragraph id="610936FBFF94FFB1D98B885DFDFEF912" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Comparisons: Externally this species can be confused with 
<taxonomicName id="A6B64D78FF94FFB1DA31885CFBCAF9A9" box="[892,1049,1570,1596]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF94FFB1DA31885CFC5DF9A9" box="[892,910,1571,1596]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DAD1885CFBCAF9A9" box="[924,1049,1571,1596]" italics="true" pageId="23" pageNumber="24">boliviensis</emphasis>
</taxonomicName>
and young individuals of 
<taxonomicName id="A6B64D78FF94FFB1DC06885CFF21F9F4" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="spegazzinii">
<emphasis id="53C2EAE9FF94FFB1DC06885CFA8EF9A9" box="[1355,1373,1571,1596]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DC26885CFF21F9F4" italics="true" pageId="23" pageNumber="24">spegazzinii</emphasis>
</taxonomicName>
. The differences between 
<taxonomicName id="A6B64D78FF94FFB1DB758837FD14F9F4" box="[568,711,1608,1634]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF94FFB1DB758837FD99F9F4" box="[568,586,1608,1633]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DB118837FD14F9F4" box="[604,711,1608,1633]" italics="true" pageId="23" pageNumber="24">caenosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF94FFB1DA448837FC78F9F4" box="[777,939,1608,1634]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF94FFB1DA448837FCC8F9F4" box="[777,795,1608,1633]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DA638837FC78F9F4" box="[814,939,1608,1633]" italics="true" pageId="23" pageNumber="24">boliviensis</emphasis>
</taxonomicName>
were described in detail by 
<bibRefCitation id="05274B0AFF94FFB1DC438837FF37F912" author="Myers, P. &amp; Patton, J. L. &amp; Smith, M. F." journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="23" pageNumber="24" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. &amp; Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
Myers 
<emphasis id="53C2EAE9FF94FFB1DC298837FA4FF9F4" box="[1380,1436,1608,1633]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
(1990)
</bibRefCitation>
and are summarized above.
</paragraph>
<paragraph id="610936FBFF94FFB1D98B88EDFEC4F842" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
In areas of sympatry 
<taxonomicName id="A6B64D78FF94FFB1D8F688ECFD94F939" box="[443,583,1682,1708]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF94FFB1D8F688ECFE1EF939" box="[443,461,1683,1708]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1D89688ECFD94F939" box="[475,583,1683,1708]" italics="true" pageId="23" pageNumber="24">caenosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF94FFB1DB3288ECFCF2F939" box="[639,801,1682,1708]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="spegazzinii">
<emphasis id="53C2EAE9FF94FFB1DB3288ECFD42F939" box="[639,657,1683,1708]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DBD288ECFCF2F939" box="[671,801,1683,1708]" italics="true" pageId="23" pageNumber="24">spegazzinii</emphasis>
</taxonomicName>
follow similar variation patterns in coloration through altitudinal gradients. However, many morphometric values and several cranial characters are useful in specimen determination. In 15 of the 20 analyzed morphometric variables, 
<taxonomicName id="A6B64D78FF94FFB1DA8B88A2FB82F963" box="[966,1105,1757,1783]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF94FFB1DA8B88A2FC0BF963" box="[966,984,1757,1782]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1DAAB88A2FB82F963" box="[998,1105,1757,1782]" italics="true" pageId="23" pageNumber="24">caenosus</emphasis>
</taxonomicName>
is significantly smaller than 
<taxonomicName id="A6B64D78FF94FFB1D9DA897CFEE9F889" box="[151,314,1794,1820]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="spegazzinii">
<emphasis id="53C2EAE9FF94FFB1D9DA897CFF7AF889" box="[151,169,1795,1820]" italics="true" pageId="23" pageNumber="24">A</emphasis>
. 
<emphasis id="53C2EAE9FF94FFB1D9F5897CFEE9F889" box="[184,314,1795,1820]" italics="true" pageId="23" pageNumber="24">spegazzinii</emphasis>
</taxonomicName>
(
<tableCitation id="2C340340FF94FFB1D804897DFE7DF889" box="[329,430,1794,1820]" captionStart="TABLE 1" captionStartId="10.[119,196,733,753]" captionTargetBox="[127,1864,857,1460]" captionTargetPageId="10" captionText="TABLE 1. Comparative measurements of adult specimens (age classes 3 and 4) of 5 species of the Akodon boliviensis group in Argentina: A. caenosus, A. boliviensis, A. spegazzinii, A. sylvanus, and Akodon new species. X: mean; SD: standard deviation; r: range; n: sample size." httpUri="http://table.plazi.org/id/35C96673FF89FFACD93A8CA2FC05FC84" pageId="23" pageNumber="24" tableUuid="35C96673FF89FFACD93A8CA2FC05FC84">Tables 1</tableCitation>
and 
<tableCitation id="2C340340FF94FFB1D8AA897DFE2AF889" box="[487,505,1794,1820]" captionStart="TABLE 2" captionStartId="11.[151,242,157,181]" captionTargetBox="[165,1405,254,1084]" captionText="TABLE 2. Significative average values among adult specimens (age classes 3 and 4) of 5 species of the Akodon boliviensis group in Argentina according the N HDS test. Abbreviations as provided in text." httpUri="http://table.plazi.org/id/35C96673FF88FFADD9DA8EE2FBBFFF4B" pageId="23" pageNumber="24" tableUuid="35C96673FF88FFADD9DA8EE2FBBFFF4B">2</tableCitation>
). This is also verified by the minimum overlap in the PCA analysis (
<figureCitation id="F98D2A7EFF94FFB1DC6C897DFAB2F889" box="[1313,1377,1794,1820]" captionStart="FIGURE 2" captionStartId="9.[151,255,893,917]" captionTargetBox="[255,1327,159,878]" captionTargetId="figure@9.[249,1333,155,884]" captionTargetPageId="9" captionText="FIGURE 2. Individual specimen scores based on log-transformed values of 11 cranial measurements, projected onto the first and second principal components extracted from analysis of adult specimens (age class 3 and 4, n = 199) of 5 species of the genus Akodon: A. boliviensis, A. caenosus, Akodon n. sp., A. spegazzinii and A. sylvanus. Results of principal components analysis in table 3." httpUri="https://zenodo.org/record/293855/files/figure.png" pageId="23" pageNumber="24">Fig 2</figureCitation>
) and the lack of missclasifications in the DA. In addition, 
<taxonomicName id="A6B64D78FF94FFB1DA5D8957FC0EF8D4" box="[784,989,1832,1857]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF94FFB1DA5D8957FC0EF8D4" box="[784,989,1832,1857]" italics="true" pageId="23" pageNumber="24">Akodon caenosus</emphasis>
</taxonomicName>
shows narrower and shallower zygomatic notches, comparatively broader interorbital constriction, temporal and mastoid ridges less developed, narrower mesopterygoid fossa and less hypsodont molars. The average percentage of genetic divergence between these species is the greatest (7.7%) observed for the 
<taxonomicName id="A6B64D78FF94FFB1DAC789E7FBD4F824" box="[906,1031,1944,1969]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF94FFB1DAC789E7FBD4F824" box="[906,1031,1944,1969]" italics="true" pageId="23" pageNumber="24">boliviensis</emphasis>
</taxonomicName>
group in northwestern 
<collectingCountry id="19A1766BFF94FFB1DC6A89E7FA48F827" box="[1319,1435,1944,1970]" name="Argentina" pageId="23" pageNumber="24">Argentina</collectingCountry>
(
<tableCitation id="2C340340FF94FFB1D9D389C2FED9F842" box="[158,266,1981,2007]" captionStart="TABLE 12" captionStartId="22.[151,239,151,175]" captionTargetBox="[165,1421,351,713]" captionText="TABLE 12. Average percentage of observerd sequence divergence within and between seven species of the Akodon boliviensis species group. Standard error estimates are shown between parentheses and were obtained by a bootstrap procedure (500 replicates) in MEGA 4. Positions containing missing data were excluded only in pairwise sequence comparisons. Intraspecific values for A. lutescens and A. subfuscus were not calculated because a single sequence was available for each species." httpUri="http://table.plazi.org/id/35C96673FF95FFB0D9DA8EE8FE63FEAC" pageId="23" pageNumber="24" tableUuid="35C96673FF95FFB0D9DA8EE8FE63FEAC">Table 12</tableCitation>
).
</paragraph>
<paragraph id="610936FBFF9BFFBED98B8EE7FCBCFE07" blockId="24.[151,1437,152,2007]" pageId="24" pageNumber="25">
<taxonomicName id="A6B64D78FF9BFFBED98B8EE7FE40FF24" box="[198,403,152,177]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED98B8EE7FE40FF24" box="[198,403,152,177]" italics="true" pageId="24" pageNumber="25">Akodon caenosus</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF9BFFBED8838EE7FD87FF24" box="[462,596,152,178]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="53C2EAE9FF9BFFBED8838EE7FE33FF24" box="[462,480,152,177]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED8A28EE7FD87FF24" box="[495,596,152,177]" italics="true" pageId="24" pageNumber="25">sylvanus</emphasis>
</taxonomicName>
represent two extremes in the morphometric variation observed within the 
<taxonomicName id="A6B64D78FF9BFFBED98E8EC2FE93FF43" box="[195,320,189,214]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF9BFFBED98E8EC2FE93FF43" box="[195,320,189,214]" italics="true" pageId="24" pageNumber="25">boliviensis</emphasis>
</taxonomicName>
group in northwestern 
<collectingCountry id="19A1766BFF9BFFBEDB1B8EC2FD18FF42" box="[598,715,189,215]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
(
<tableCitation id="2C340340FF9BFFBEDB978EC2FC93FF42" box="[730,832,189,215]" captionStart="TABLE 1" captionStartId="10.[119,196,733,753]" captionTargetBox="[127,1864,857,1460]" captionTargetPageId="10" captionText="TABLE 1. Comparative measurements of adult specimens (age classes 3 and 4) of 5 species of the Akodon boliviensis group in Argentina: A. caenosus, A. boliviensis, A. spegazzinii, A. sylvanus, and Akodon new species. X: mean; SD: standard deviation; r: range; n: sample size." httpUri="http://table.plazi.org/id/35C96673FF89FFACD93A8CA2FC05FC84" pageId="24" pageNumber="25" tableUuid="35C96673FF89FFACD93A8CA2FC05FC84">Tables 1</tableCitation>
and 
<tableCitation id="2C340340FF9BFFBEDA378EC2FC5FFF42" box="[890,908,189,215]" captionStart="TABLE 2" captionStartId="11.[151,242,157,181]" captionTargetBox="[165,1405,254,1084]" captionText="TABLE 2. Significative average values among adult specimens (age classes 3 and 4) of 5 species of the Akodon boliviensis group in Argentina according the N HDS test. Abbreviations as provided in text." httpUri="http://table.plazi.org/id/35C96673FF88FFADD9DA8EE2FBBFFF4B" pageId="24" pageNumber="25" tableUuid="35C96673FF88FFADD9DA8EE2FBBFFF4B">2</tableCitation>
; figure 2). 
<taxonomicName id="A6B64D78FF9BFFBEDD5D8EC2FB0EFF43" box="[1040,1245,189,214]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDD5D8EC2FB0EFF43" box="[1040,1245,189,214]" italics="true" pageId="24" pageNumber="25">Akodon caenosus</emphasis>
</taxonomicName>
is, furthermore, paler than 
<taxonomicName id="A6B64D78FF9BFFBED85E8E9DFE4AFF6E" box="[275,409,226,252]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="53C2EAE9FF9BFFBED85E8E9DFEF6FF6E" box="[275,293,226,251]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED8798E9DFE4AFF6E" box="[308,409,226,251]" italics="true" pageId="24" pageNumber="25">sylvanus</emphasis>
</taxonomicName>
, showing a more evident contrast between dorsum and venter, and has more developed eyerings. The skull of 
<taxonomicName id="A6B64D78FF9BFFBED8ED8F77FDFEFEB4" box="[416,557,264,290]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED8ED8F77FE61FEB4" box="[416,434,264,289]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED88F8F77FDFEFEB4" box="[450,557,264,289]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
has a shorter and narrower rostrum, less swollen frontal sinuses, narrower braincase and interorbital region and a narrower mesopterygoid and parapterygoid fossa. The upper incisors in 
<taxonomicName id="A6B64D78FF9BFFBED9DA8F2DFEF6FEFE" box="[151,293,338,364]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED9DA8F2DFF7AFEFE" box="[151,169,338,363]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED9F78F2DFEF6FEFE" box="[186,293,338,363]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
are less orthodont and the molars less hypsodont than 
<taxonomicName id="A6B64D78FF9BFFBEDA8E8F2DFB9FFEFE" box="[963,1100,338,364]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="53C2EAE9FF9BFFBEDA8E8F2DFC06FEFE" box="[963,981,338,363]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDAAA8F2DFB9FFEFE" box="[999,1100,338,363]" italics="true" pageId="24" pageNumber="25">sylvanus</emphasis>
</taxonomicName>
. Additionally, these species show a high average percentage of genetic divergence (6.7%).
</paragraph>
<paragraph id="610936FBFF9BFFBED98B8FE2FA88FE22" blockId="24.[151,1437,152,2007]" box="[198,1371,413,439]" pageId="24" pageNumber="25">
The differences between 
<taxonomicName id="A6B64D78FF9BFFBED8A08FE2FDABFE23" box="[493,632,413,439]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED8A08FE2FE2CFE23" box="[493,511,413,438]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDB408FE2FDABFE23" box="[525,632,413,438]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
and the new species are listed under the treatment of the latter.
</paragraph>
<paragraph id="610936FBFF9BFFBED98B8FBDFE95FDB2" blockId="24.[151,1437,152,2007]" pageId="24" pageNumber="25">
Distribution: 
<taxonomicName id="A6B64D78FF9BFFBED8288FBDFDE0FE4E" box="[357,563,450,475]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED8288FBDFDE0FE4E" box="[357,563,450,475]" italics="true" pageId="24" pageNumber="25">Akodon caenosus</emphasis>
</taxonomicName>
is the more broadly distributed species of the 
<taxonomicName id="A6B64D78FF9BFFBEDD2D8FBDFB0EFE4E" box="[1120,1245,450,475]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF9BFFBEDD2D8FBDFB0EFE4E" box="[1120,1245,450,475]" italics="true" pageId="24" pageNumber="25">boliviensis</emphasis>
</taxonomicName>
group in northwestern 
<collectingCountry id="19A1766BFF9BFFBED9B78F97FEA1FD97" box="[250,370,488,514]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
, with records from northernmost 
<collectingRegion id="A372F819FF9BFFBEDBB28F97FCE9FD97" box="[767,826,488,514]" country="Argentina" name="Salta" pageId="24" pageNumber="25">Salta</collectingRegion>
to southern 
<collectingRegion id="A372F819FF9BFFBEDA808F97FB9DFD97" box="[973,1102,488,514]" country="Argentina" name="Catamarca" pageId="24" pageNumber="25">Catamarca</collectingRegion>
, from 
<quantity id="A64E9B1EFF9BFFBEDDD18F97FB34FD97" box="[1180,1255,488,514]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="25" unit="m" value="400.0">400 m</quantity>
to 
<quantity id="A64E9B1EFF9BFFBEDC438F97FABBFD97" box="[1294,1384,488,514]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="24" pageNumber="25" unit="m" value="3100.0">3100 m</quantity>
elevation (
<figureCitation id="F98D2A7EFF9BFFBED9BD8C72FEEBFDB2" box="[240,312,525,551]" captionStart="FIGURE 7" captionStartId="25.[151,255,1971,1995]" captionTargetBox="[219,1379,136,1965]" captionTargetId="figure@25.[219,1379,136,1969]" captionTargetPageId="25" captionText="FIGURE 7. Recorded localities of A. caenosus in Northwestern Argentina. Localities and geographic position in Appedix II." httpUri="https://zenodo.org/record/293858/files/figure.png" pageId="24" pageNumber="25">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="610936FBFF9BFFBED98B8C4DFBF1FD02" blockId="24.[151,1437,152,2007]" pageId="24" pageNumber="25">Habitat: Most of the records come from Yungas environments, from the lower altitudinal belt to high altitude grasslands. Notwithstanding, we have recorded the species in Chacoan environments near the ecotone with Yungas and in the lower altitudinal limit of the High Andean grasslands.</paragraph>
<paragraph id="610936FBFF9BFFBED98B8CDDFEA4FB37" blockId="24.[151,1437,152,2007]" pageId="24" pageNumber="25">
Natural history: Although we have recorded specimens in reproductive condition throughout the year, most of the individuals were active between November and January. We observed the highest proportion of molting animals in fall and winter. In their broad distributional range 
<taxonomicName id="A6B64D78FF9BFFBEDAA98C92FBA1FC93" box="[996,1138,749,775]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDAA98C92FC25FC93" box="[996,1014,749,774]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDD4A8C92FBA1FC93" box="[1031,1138,749,774]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
can be found coexisting with many different species depending on the altitudinal or latitudinal sector in northwestern 
<collectingCountry id="19A1766BFF9BFFBEDD9A8D6DFA83FCB9" box="[1239,1360,786,812]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
. Thus, in the north 
<taxonomicName id="A6B64D78FF9BFFBED8628D47FE6DFCC4" box="[303,446,824,850]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED8628D47FE92FCC4" box="[303,321,824,849]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED81E8D47FE6DFCC4" box="[339,446,824,849]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
was captured alongside 
<taxonomicName id="A6B64D78FF9BFFBEDBA68D47FC60FCC4" box="[747,947,824,849]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="sylvanus">
<emphasis id="53C2EAE9FF9BFFBEDBA68D47FC60FCC4" box="[747,947,824,849]" italics="true" pageId="24" pageNumber="25">Akodon sylvanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDA888D47FB8BFCC4" box="[965,1112,824,850]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="simulator">
<emphasis id="53C2EAE9FF9BFFBEDA888D47FC04FCC4" box="[965,983,824,849]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDAA48D47FB8BFCC4" box="[1001,1112,824,849]" italics="true" pageId="24" pageNumber="25">simulator</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDD278D47FA47FCC4" box="[1130,1428,824,849]" class="Mammalia" family="Muridae" genus="Oxymycterus" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="paramensis">
<emphasis id="53C2EAE9FF9BFFBEDD278D47FA47FCC4" box="[1130,1428,824,849]" italics="true" pageId="24" pageNumber="25">Oxymycterus paramensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED9DA8D22FEE9FCE3" box="[151,314,861,886]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBED9DA8D22FEE9FCE3" box="[151,314,861,886]" italics="true" pageId="24" pageNumber="25">Oligoryzomys</emphasis>
</taxonomicName>
cf. 
<taxonomicName id="A6B64D78FF9BFFBED8288D22FDD3FCE3" box="[357,512,861,887]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="flavescens">
<emphasis id="53C2EAE9FF9BFFBED8288D22FEA9FCE3" box="[357,378,861,886]" italics="true" pageId="24" pageNumber="25">O</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED8C58D22FDD3FCE3" box="[392,512,861,886]" italics="true" pageId="24" pageNumber="25">flavescens</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF9BFFBEDB748D22FD0FFCE3" box="[569,732,861,886]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBEDB748D22FD0FFCE3" box="[569,732,861,886]" italics="true" pageId="24" pageNumber="25">Oligoryzomys</emphasis>
</taxonomicName>
sp. in high altitudinal grasslands at 
<quantity id="A64E9B1EFF9BFFBEDD308D22FB0BFCE2" box="[1149,1240,861,887]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="24" pageNumber="25" unit="m" value="1400.0">1400 m</quantity>
. At the same latitude in upper belts above 
<quantity id="A64E9B1EFF9BFFBED89C8DFDFDFDFC09" box="[465,558,898,924]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="m" value="2000.0">2000 m</quantity>
, 
<taxonomicName id="A6B64D78FF9BFFBEDB778DFDFD14FC0E" box="[570,711,898,924]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDB778DFDFD9FFC0E" box="[570,588,898,923]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDB118DFDFD14FC0E" box="[604,711,898,923]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
is sympatric with 
<taxonomicName id="A6B64D78FF9BFFBEDAEE8DFDFB92FC0E" box="[931,1089,898,924]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF9BFFBEDAEE8DFDFC66FC0E" box="[931,949,898,923]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDA898DFDFB92FC0E" box="[964,1089,898,923]" italics="true" pageId="24" pageNumber="25">boliviensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDD1C8DFDFAF3FC0E" box="[1105,1312,898,923]" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="lactens">
<emphasis id="53C2EAE9FF9BFFBEDD1C8DFDFAF3FC0E" box="[1105,1312,898,923]" italics="true" pageId="24" pageNumber="25">Necromys lactens</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDC7D8DFDFF13FC54" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="amoenus">
<emphasis id="53C2EAE9FF9BFFBEDC7D8DFDFA90FC0E" box="[1328,1347,898,923]" italics="true" pageId="24" pageNumber="25">N</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDC1E8DFDFF13FC54" italics="true" pageId="24" pageNumber="25">amoenus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED9838DD7FDEFFC57" authority="Thomas" authorityName="Thomas" box="[206,572,936,962]" class="Mammalia" family="Muridae" genus="Calomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="musculinus">
<emphasis id="53C2EAE9FF9BFFBED9838DD7FE13FC54" box="[206,448,936,961]" italics="true" pageId="24" pageNumber="25">Calomys musculinus</emphasis>
(Thomas)
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDB0A8DD7FC56FC57" authority="Pearson" authorityName="Pearson" box="[583,901,936,962]" class="Mammalia" family="Muridae" genus="Phyllotis" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caprinus">
<emphasis id="53C2EAE9FF9BFFBEDB0A8DD7FCCEFC54" box="[583,797,936,961]" italics="true" pageId="24" pageNumber="25">Phyllotis caprinus</emphasis>
Pearson
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDADD8DD7FC20FC54" box="[912,1011,936,962]" class="Mammalia" family="Muridae" genus="Phyllotis" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="osilae">
<emphasis id="53C2EAE9FF9BFFBEDADD8DD7FC4CFC54" box="[912,927,936,961]" italics="true" pageId="24" pageNumber="25">P</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDAE08DD7FC20FC54" box="[941,1011,936,961]" italics="true" pageId="24" pageNumber="25">osilae</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF9BFFBEDD618DD7FB0DFC54" box="[1068,1246,936,962]" class="Mammalia" family="Muridae" genus="Phyllotis" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="xanthopygus">
<emphasis id="53C2EAE9FF9BFFBEDD618DD7FBE8FC54" box="[1068,1083,936,961]" italics="true" pageId="24" pageNumber="25">P</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDD078DD7FB0DFC54" box="[1098,1246,936,961]" italics="true" pageId="24" pageNumber="25">xanthopygus</emphasis>
</taxonomicName>
. In the south, in Yungas areas of 
<collectingRegion id="A372F819FF9BFFBED8138DB2FDECFC72" box="[350,575,973,999]" country="Argentina" name="Tucuman" pageId="24" pageNumber="25">Tucumán province</collectingRegion>
, the species was caught together with 
<taxonomicName id="A6B64D78FF9BFFBEDD408DB2FB21FC73" box="[1037,1266,973,998]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spegazzinii">
<emphasis id="53C2EAE9FF9BFFBEDD408DB2FB21FC73" box="[1037,1266,973,998]" italics="true" pageId="24" pageNumber="25">Akodon spegazzinii</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDC4E8DB2FA47FC73" box="[1283,1428,973,999]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="simulator">
<emphasis id="53C2EAE9FF9BFFBEDC4E8DB2FAC6FC73" box="[1283,1301,973,998]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDC688DB2FA47FC73" box="[1317,1428,973,998]" italics="true" pageId="24" pageNumber="25">simulator</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED9DA8D8DFEB5FB9E" box="[151,358,1010,1035]" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="lactens">
<emphasis id="53C2EAE9FF9BFFBED9DA8D8DFEB5FB9E" box="[151,358,1010,1035]" italics="true" pageId="24" pageNumber="25">Necromys lactens</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED83B8D8DFD80FB99" authority="Lund" authorityName="Lund" box="[374,595,1010,1036]" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="lasiurus">
<emphasis id="53C2EAE9FF9BFFBED83B8D8DFE24FB9E" box="[374,503,1010,1035]" italics="true" pageId="24" pageNumber="25">N. lasiurus</emphasis>
(Lund)
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDB2C8D8DFC5AFB9E" box="[609,905,1010,1035]" class="Mammalia" family="Muridae" genus="Oxymycterus" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="paramensis">
<emphasis id="53C2EAE9FF9BFFBEDB2C8D8DFC5AFB9E" box="[609,905,1010,1035]" italics="true" pageId="24" pageNumber="25">Oxymycterus paramensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDAD48D8DFADFFB99" authority="Jayat et al." authorityName="Jayat et al." box="[921,1292,1010,1036]" class="Mammalia" family="Muridae" genus="Oxymycterus" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="wayku">
<emphasis id="53C2EAE9FF9BFFBEDAD48D8DFB56FB9E" box="[921,1157,1010,1035]" italics="true" pageId="24" pageNumber="25">Oxymycterus wayku</emphasis>
Jayat 
<emphasis id="53C2EAE9FF9BFFBEDD9F8D8DFAD5FB9E" box="[1234,1286,1010,1035]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
.
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDC508D8DFF16FBA4" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBEDC508D8DFF16FBA4" italics="true" pageId="24" pageNumber="25">Oligoryzomys</emphasis>
</taxonomicName>
cf. 
<taxonomicName id="A6B64D78FF9BFFBED9BF8A67FE5EFBA4" box="[242,397,1048,1074]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="flavescens">
<emphasis id="53C2EAE9FF9BFFBED9BF8A67FED4FBA4" box="[242,263,1048,1073]" italics="true" pageId="24" pageNumber="25">O</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED8588A67FE5EFBA4" box="[277,397,1048,1073]" italics="true" pageId="24" pageNumber="25">flavescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED8D08A67FD93FBA4" box="[413,576,1048,1073]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBED8D08A67FD93FBA4" box="[413,576,1048,1073]" italics="true" pageId="24" pageNumber="25">Oligoryzomys</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="A6B64D78FF9BFFBEDB3A8A67FCE3FBA4" box="[631,816,1048,1073]" class="Mammalia" family="Muridae" genus="Phyllotis" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="anitae">
<emphasis id="53C2EAE9FF9BFFBEDB3A8A67FCE3FBA4" box="[631,816,1048,1073]" italics="true" pageId="24" pageNumber="25">Phyllotis anitae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDA728A67FC70FBA4" box="[831,931,1048,1074]" class="Mammalia" family="Muridae" genus="Phyllotis" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="osilae">
<emphasis id="53C2EAE9FF9BFFBEDA728A67FC9DFBA4" box="[831,846,1048,1073]" italics="true" pageId="24" pageNumber="25">P</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDA108A67FC70FBA4" box="[861,931,1048,1073]" italics="true" pageId="24" pageNumber="25">osilae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDAFF8A67FBA0FBA4" box="[946,1139,1048,1073]" class="Mammalia" family="Muridae" genus="Andinomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="edax">
<emphasis id="53C2EAE9FF9BFFBEDAFF8A67FBA0FBA4" box="[946,1139,1048,1073]" italics="true" pageId="24" pageNumber="25">Andinomys edax</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF9BFFBEDDE18A67FAA3FBA4" box="[1196,1392,1048,1073]" class="Mammalia" family="Cricetidae" genus="Abrothrix" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="illutea">
<emphasis id="53C2EAE9FF9BFFBEDDE18A67FAA3FBA4" box="[1196,1392,1048,1073]" italics="true" pageId="24" pageNumber="25">Abrothrix illutea</emphasis>
</taxonomicName>
. At this same latitude in Chacoan environments in transition with Yungas 
<taxonomicName id="A6B64D78FF9BFFBEDA958A42FBB7FBC3" box="[984,1124,1085,1111]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDA958A42FC39FBC3" box="[984,1002,1085,1110]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDAB48A42FBB7FBC3" box="[1017,1124,1085,1110]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
is sympatric with 
<taxonomicName id="A6B64D78FF9BFFBEDC0F8A42FECAFBE9" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spegazzinii">
<emphasis id="53C2EAE9FF9BFFBEDC0F8A42FECAFBE9" italics="true" pageId="24" pageNumber="25">Akodon spegazzinii</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED8648A1CFE68FBE9" box="[297,443,1122,1148]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="simulator">
<emphasis id="53C2EAE9FF9BFFBED8648A1CFEE8FBE9" box="[297,315,1123,1148]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED8018A1CFE68FBE9" box="[332,443,1123,1148]" italics="true" pageId="24" pageNumber="25">simulator</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBED8868A1CFDECFBE9" box="[459,575,1123,1148]" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBED8868A1CFDECFBE9" box="[459,575,1123,1148]" italics="true" pageId="24" pageNumber="25">Necromys</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="A6B64D78FF9BFFBEDB378A1CFCCEFBE9" box="[634,797,1123,1148]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBEDB378A1CFCCEFBE9" box="[634,797,1123,1148]" italics="true" pageId="24" pageNumber="25">Oligoryzomys</emphasis>
</taxonomicName>
cf. 
<taxonomicName id="A6B64D78FF9BFFBEDA018A1CFC38FBE9" box="[844,1003,1122,1148]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="flavescens">
<emphasis id="53C2EAE9FF9BFFBEDA018A1CFCB2FBE9" box="[844,865,1123,1148]" italics="true" pageId="24" pageNumber="25">O</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDA3F8A1CFC38FBE9" box="[882,1003,1123,1148]" italics="true" pageId="24" pageNumber="25">flavescens</emphasis>
</taxonomicName>
, 
<taxonomicName id="A6B64D78FF9BFFBEDAB78A1CFB4EFBE9" box="[1018,1181,1123,1148]" class="Mammalia" family="Muridae" genus="Oligoryzomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBEDAB78A1CFB4EFBE9" box="[1018,1181,1123,1148]" italics="true" pageId="24" pageNumber="25">Oligoryzomys</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="A6B64D78FF9BFFBEDD958A1CFAECFBE9" box="[1240,1343,1123,1148]" class="Mammalia" family="Muridae" genus="Calomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="53C2EAE9FF9BFFBEDD958A1CFAECFBE9" box="[1240,1343,1123,1148]" italics="true" pageId="24" pageNumber="25">Calomys</emphasis>
</taxonomicName>
sp. and 
<taxonomicName id="A6B64D78FF9BFFBED9DA8AF7FEA3FB34" box="[151,368,1160,1185]" class="Mammalia" family="Muridae" genus="Graomys" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="centralis">
<emphasis id="53C2EAE9FF9BFFBED9DA8AF7FEA3FB34" box="[151,368,1160,1185]" italics="true" pageId="24" pageNumber="25">Graomys centralis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="610936FBFF9BFFBED98B8AD2FCF1F912" blockId="24.[151,1437,152,2007]" pageId="24" pageNumber="25">
Comments: Although originally described as a subspecies of 
<taxonomicName id="A6B64D78FF9BFFBEDAF08AD2FAF1FB52" authority="Thomas 1918" authorityName="Thomas" authorityYear="1918" box="[957,1314,1197,1223]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF9BFFBEDAF08AD2FB89FB53" box="[957,1114,1197,1222]" italics="true" pageId="24" pageNumber="25">Akodon puer</emphasis>
(
<bibRefCitation id="05274B0AFF9BFFBEDD238AD2FACAFB52" author="Thomas, O." box="[1134,1305,1197,1223]" journalOrPublisher="E. Budin. Annals and Magazine of Natural History" pageId="24" pageNumber="25" pagination="186 - 193" part="9" refString="Thomas, O. (1918) On small mammals form Salta and Jujuy collected by Mr. E. Budin. Annals and Magazine of Natural History, (9) 1, 186 - 193." title="On small mammals form Salta and Jujuy collected by Mr" type="journal article" year="1918">Thomas 1918</bibRefCitation>
)
</taxonomicName>
, the form 
<taxonomicName id="A6B64D78FF9BFFBED9DA8AACFED0FB79" box="[151,259,1235,1260]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED9DA8AACFED0FB79" box="[151,259,1235,1260]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
was soon considered as a valid species by 
<bibRefCitation id="05274B0AFF9BFFBEDA408AADFC14FB79" author="Thomas, O." box="[781,967,1234,1260]" journalOrPublisher="Annals and Magazine of Natural History" pageId="24" pageNumber="25" pagination="195 - 196" part="9" refString="Thomas, O. (1920 a) A further collection of mammals from Jujuy. Annals and Magazine of Natural History, (9) 5, 195 - 196." title="A further collection of mammals from Jujuy" type="journal article" year="1920">Thomas (1920)</bibRefCitation>
; this latter suggestion was followed by 
<bibRefCitation id="05274B0AFF9BFFBED9DA8A87FE9EFA87" author="Cabrera, A." box="[151,333,1272,1298]" journalOrPublisher="Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot;, Zoologia" pageId="24" pageNumber="25" pagination="309 - 732" part="4" refString="Cabrera, A. (1961) Catalogo de los mamiferos de America del Sur. Parte II. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot;, Zoologia, 4 (2), 309 - 732." title="Catalogo de los mamiferos de America del Sur. Parte II" type="journal article" year="1961">Cabrera (1961)</bibRefCitation>
, 
<bibRefCitation id="05274B0AFF9BFFBED8168A87FD88FA87" author="Barquez, R. M. &amp; Williams, D. F. &amp; Mares, M. A. &amp; Genoways, H. H." box="[347,603,1272,1298]" journalOrPublisher="Annals of Carnegie Museum" pageId="24" pageNumber="25" pagination="379 - 403" part="49" refString="Barquez, R. M., Williams, D. F., Mares, M. A. &amp; Genoways, H. H. (1980) Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina. Annals of Carnegie Museum, 49, 379 - 403." title="Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina" type="journal article" year="1980">
Barquez 
<emphasis id="53C2EAE9FF9BFFBED8888A87FE29FA84" box="[453,506,1272,1297]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1980)
</bibRefCitation>
, and 
<bibRefCitation id="05274B0AFF9BFFBEDBD68A87FC50FA87" author="Mares, M. A &amp; Ojeda, R. A. &amp; Kosco, M. P." box="[667,899,1272,1298]" journalOrPublisher="Annals of Carnegie Museum" pageId="24" pageNumber="25" pagination="151 - 206" part="50" refString="Mares, M. A, Ojeda, R. A. &amp; Kosco, M. P. (1981) Observation on the distribution and ecology of the mammals of Salta Province, Argentina. Annals of Carnegie Museum, 50, 151 - 206." title="Observation on the distribution and ecology of the mammals of Salta Province, Argentina" type="journal article" year="1981">
Mares 
<emphasis id="53C2EAE9FF9BFFBEDBA08A87FCF1FA84" box="[749,802,1272,1297]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1981)
</bibRefCitation>
. 
<bibRefCitation id="05274B0AFF9BFFBEDAC28A87FBAFFA87" author="Vitullo, A. D. &amp; Merani, M. S. &amp; Reig, O. A. &amp; Kajon, A. E. &amp; Scaglia, O. &amp; Espinosa, M. B. &amp; Perez-Zapata, A." box="[911,1148,1272,1298]" journalOrPublisher="Journal of Mammalogy" pageId="24" pageNumber="25" pagination="69 - 80" part="67" refString="Vitullo, A. D., Merani, M. S., Reig, O. A., Kajon, A. E., Scaglia, O., Espinosa, M. B. &amp; Perez-Zapata, A. (1986) Cytogenetics of South American Akodont Rodents (Cricetidae): new kariotypes and chromosomal banding patterns of argentinian and uruguayan forms. Journal of Mammalogy, 67, 69 - 80." title="Cytogenetics of South American Akodont Rodents (Cricetidae): new kariotypes and chromosomal banding patterns of argentinian and uruguayan forms" type="journal article" year="1986">
Vitullo 
<emphasis id="53C2EAE9FF9BFFBEDAA78A87FBCCFA84" box="[1002,1055,1272,1297]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1986)
</bibRefCitation>
and 
<bibRefCitation id="05274B0AFF9BFFBEDDF58A87FF37FAA2" author="Apfelbaum, L. I. &amp; Reig O. A." journalOrPublisher="Biological Journal of the Linnean Society" pageId="24" pageNumber="25" pagination="257 - 280" part="38" refString="Apfelbaum, L. I. &amp; Reig O. A. (1989) Allozyme genetic distances and evolutionary relationships in species of akodontine rodents (Cricetidae: Sigmodontinae). Biological Journal of the Linnean Society, 38, 257 - 280." title="Allozyme genetic distances and evolutionary relationships in species of akodontine rodents (Cricetidae: Sigmodontinae)" type="journal article" year="1989">Apfelbaum &amp; Reig (1989)</bibRefCitation>
considered specimens from León, 
<collectingRegion id="A372F819FF9BFFBEDBC88B62FCE7FAA2" box="[645,820,1309,1335]" country="Argentina" name="Jujuy" pageId="24" pageNumber="25">Jujuy Province</collectingRegion>
(
<typeStatus id="BE0D8859FF9BFFBEDA088B62FCAAFAA2" box="[837,889,1309,1335]" pageId="24" pageNumber="25">type</typeStatus>
locality of 
<taxonomicName id="A6B64D78FF9BFFBEDD4E8B62FBBDFAA3" box="[1027,1134,1309,1334]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDD4E8B62FBBDFAA3" box="[1027,1134,1309,1334]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
) as 
<taxonomicName id="A6B64D78FF9BFFBEDDEC8B62FB2AFAA3" box="[1185,1273,1309,1335]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF9BFFBEDDEC8B62FB60FAA3" box="[1185,1203,1309,1334]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDD8E8B62FB2AFAA3" box="[1219,1273,1309,1334]" italics="true" pageId="24" pageNumber="25">puer</emphasis>
</taxonomicName>
and 
<bibRefCitation id="05274B0AFF9BFFBEDC798B62FEDEFAC9" author="Myers, P. &amp; Patton, J. L. &amp; Smith, M. F." journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="24" pageNumber="25" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. &amp; Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
Myers 
<emphasis id="53C2EAE9FF9BFFBEDCCA8B62FF7DFAC9" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1990)
</bibRefCitation>
followed this position (but see 
<bibRefCitation id="05274B0AFF9BFFBEDBDD8B3DFCBBFAC9" author="Hershkovitz, P." box="[656,872,1346,1372]" journalOrPublisher="Fieldiana, Zoology, new series" pageId="24" pageNumber="25" pagination="1 - 35" part="57" refString="Hershkovitz, P. (1990) Mice of the Akodon boliviensis size class (Sigmodontinae, Cricetidae), with the description of two species from Brazil. Fieldiana, Zoology, new series, 57, 1 - 35." title="Mice of the Akodon boliviensis size class (Sigmodontinae, Cricetidae), with the description of two species from Brazil" type="journal article" year="1990">Hershkovitz 1990</bibRefCitation>
), considering 
<taxonomicName id="A6B64D78FF9BFFBEDD5F8B3CFBAEFAC9" box="[1042,1149,1347,1372]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDD5F8B3CFBAEFAC9" box="[1042,1149,1347,1372]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
as the subspecies of 
<taxonomicName id="A6B64D78FF9BFFBEDCCE8B3CFF1EFA14" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF9BFFBEDCCE8B3CFA46FAC9" box="[1411,1429,1347,1372]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED9DA8B17FF1EFA14" box="[151,205,1384,1409]" italics="true" pageId="24" pageNumber="25">puer</emphasis>
</taxonomicName>
inhabiting northwestern 
<collectingCountry id="19A1766BFF9BFFBED8B88B17FDB9FA17" box="[501,618,1384,1410]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
(see also 
<bibRefCitation id="05274B0AFF9BFFBEDBAF8B17FC68FA17" author="Barros, M. A. &amp; Liascovich, R. C. &amp; Gonzalez, L. &amp; Lizarralde, M. S. &amp; Reig, O. A." box="[738,955,1384,1410]" journalOrPublisher="Zeitschrift fur Saugetierkunde" pageId="24" pageNumber="25" pagination="115 - 127" part="55" refString="Barros, M. A., Liascovich, R. C., Gonzalez, L., Lizarralde, M. S. &amp; Reig, O. A. (1990) Banding pattern comparison between Akodon iniscatus, and Akodon puer (Rodentia, Cricetidae). Zeitschrift fur Saugetierkunde, 55, 115 - 127." title="Banding pattern comparison between Akodon iniscatus, and Akodon puer (Rodentia, Cricetidae)" type="journal article" year="1990">
Barros 
<emphasis id="53C2EAE9FF9BFFBEDA758B17FCBFFA14" box="[824,876,1384,1409]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. 1990
</bibRefCitation>
). 
<bibRefCitation id="05274B0AFF9BFFBEDA9D8B17FB4AFA17" author="Anderson, S." box="[976,1177,1384,1410]" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="24" pageNumber="25" pagination="1 - 652" part="231" refString="Anderson, S. (1997) Mammals of Bolivia, taxonomy and distribution. Bulletin of the American Museum of Natural History, 231, 1 - 652." title="Mammals of Bolivia, taxonomy and distribution" type="journal article" year="1997">Anderson (1997)</bibRefCitation>
also recognized three subspecies for 
<taxonomicName id="A6B64D78FF9BFFBED8048BF2FE71FA33" box="[329,418,1421,1446]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF9BFFBED8048BF2FE71FA33" box="[329,418,1421,1446]" italics="true" pageId="24" pageNumber="25">A. puer</emphasis>
</taxonomicName>
but noted the priority of the name 
<taxonomicName id="A6B64D78FF9BFFBEDA068BF2FC65FA33" box="[843,950,1421,1446]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="lutescens">
<emphasis id="53C2EAE9FF9BFFBEDA068BF2FC65FA33" box="[843,950,1421,1446]" italics="true" pageId="24" pageNumber="25">lutescens</emphasis>
</taxonomicName>
over 
<taxonomicName id="A6B64D78FF9BFFBEDAB08BF2FBE2FA33" box="[1021,1073,1421,1446]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF9BFFBEDAB08BF2FBE2FA33" box="[1021,1073,1421,1446]" italics="true" pageId="24" pageNumber="25">puer</emphasis>
</taxonomicName>
. Since then, the nomenclature used for this species has been constantly changing. 
<bibRefCitation id="05274B0AFF9BFFBEDA408BCDFBF4FA59" author="Capllonch, P. &amp; Autino, A. &amp; Diaz, M. M. &amp; Barquez, R. M. &amp; Goytia, M." box="[781,1063,1458,1484]" journalOrPublisher="Mastozoologia Neotropical" pageId="24" pageNumber="25" pagination="49 - 71" part="4" refString="Capllonch, P., Autino, A., Diaz, M. M., Barquez, R. M. &amp; Goytia, M. (1997) Los mamiferos del Parque Biologico Sierra de San Javier, Tucuman, Argentina: observaciones sobre su sistematica y distribucion. Mastozoologia Neotropical, 4, 49 - 71." title="Los mamiferos del Parque Biologico Sierra de San Javier, Tucuman, Argentina: observaciones sobre su sistematica y distribucion" type="journal article" year="1997">
Capllonch 
<emphasis id="53C2EAE9FF9BFFBEDADD8BCCFC14FA59" box="[912,967,1459,1484]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1997)
</bibRefCitation>
and 
<bibRefCitation id="05274B0AFF9BFFBEDD2B8BCDFA82FA59" author="Diaz, M. M. &amp; Braun, J. K. &amp; Mares, M. A. &amp; Barquez, R. M." box="[1126,1361,1458,1484]" journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="24" pageNumber="25" pagination="1 - 10" part="2" refString="Diaz, M. M., Braun, J. K., Mares, M. A. &amp; Barquez, R. M. (1997) Key of Mammals of Salta Province, Argentina. Occasional Papers of the Oklahoma Museum of Natural History, 2, 1 - 10." title="Key of Mammals of Salta Province, Argentina" type="journal article" year="1997">
Mares 
<emphasis id="53C2EAE9FF9BFFBEDDF68BCCFB21FA59" box="[1211,1266,1459,1484]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1997)
</bibRefCitation>
listed 
<taxonomicName id="A6B64D78FF9BFFBED9DA8BA7FED1FA64" box="[151,258,1496,1521]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED9DA8BA7FED1FA64" box="[151,258,1496,1521]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
as valid species whereas 
<bibRefCitation id="05274B0AFF9BFFBEDB7D8BA7FD2AFA67" author="Diaz, M. M." bookContentInfo="640 pp." box="[560,761,1496,1522]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="24" pageNumber="25" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">
Díaz 
<emphasis id="53C2EAE9FF9BFFBEDB228BA7FD71FA64" box="[623,674,1496,1521]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1999
</bibRefCitation>
, 
<bibRefCitation id="05274B0AFF9BFFBEDA488BA7FC92FA67" author="Diaz, M. M. &amp; Braun, J. K. &amp; Mares, M. A. &amp; Barquez, R. M." box="[773,833,1496,1522]" journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="24" pageNumber="25" pagination="1 - 52" part="10" refString="Diaz, M. M., Braun, J. K., Mares, M. A. &amp; Barquez, R. M. (2000) An update of the taxonomy, systematics, and distribution of the mammals of Salta province, Argentina. Occasional Papers of the Oklahoma Museum of Natural History, 10, 1 - 52." title="An update of the taxonomy, systematics, and distribution of the mammals of Salta province, Argentina" type="journal article" year="2000">2000</bibRefCitation>
) recognized 
<taxonomicName id="A6B64D78FF9BFFBEDA948BA7FB97FA64" box="[985,1092,1496,1521]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDA948BA7FB97FA64" box="[985,1092,1496,1521]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
as subspecies of 
<taxonomicName id="A6B64D78FF9BFFBEDC5C8BA7FA94FA64" box="[1297,1351,1496,1521]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="puer">
<emphasis id="53C2EAE9FF9BFFBEDC5C8BA7FA94FA64" box="[1297,1351,1496,1521]" italics="true" pageId="24" pageNumber="25">puer</emphasis>
</taxonomicName>
for the populations of the northwestern 
<collectingCountry id="19A1766BFF9BFFBEDB5C8B82FD5AF982" box="[529,649,1533,1559]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
. Notwithstanding, 
<bibRefCitation id="05274B0AFF9BFFBEDA2B8B82FC21F982" author="Diaz, M. M." bookContentInfo="640 pp." box="[870,1010,1533,1559]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="24" pageNumber="25" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">Díaz (1999)</bibRefCitation>
and 
<bibRefCitation id="05274B0AFF9BFFBEDD668B82FAEDF982" author="Diaz, M. M. &amp; Barquez, R. M." box="[1067,1342,1533,1559]" editor="Kelt, D. A." journalOrPublisher="University of California Publications in Zoology 134" pageId="24" pageNumber="25" pagination="417 - 578" refString="Diaz, M. M. &amp; Barquez, R. M. (2007) The Wild Mammals of Jujuy Province, Argentina: Systematics and Distribution. In: Kelt, D. A., Lessa, E. P., Salazar-Bravo, J. &amp; Patton, J. L. (Eds.), The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson. University of California Publications in Zoology 134, pp. 417 - 578." title="The Wild Mammals of Jujuy Province, Argentina: Systematics and Distribution" type="book chapter" volumeTitle="The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson" year="2007">Díaz &amp; Barquez (2007)</bibRefCitation>
considered again the nominal form 
<taxonomicName id="A6B64D78FF9BFFBED8A5885CFD80F9A9" box="[488,595,1571,1596]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED8A5885CFD80F9A9" box="[488,595,1571,1596]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
as representing a valid biological species including 
<taxonomicName id="A6B64D78FF9BFFBEDDF9885CFAAEF9A9" box="[1204,1405,1570,1596]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="subSpecies" species="lutescens" subSpecies="puer">
<emphasis id="53C2EAE9FF9BFFBEDDF9885CFB15F9A9" box="[1204,1222,1571,1596]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDD99885CFAAEF9A9" box="[1236,1405,1571,1596]" italics="true" pageId="24" pageNumber="25">lutescens puer</emphasis>
</taxonomicName>
as an additional sigmodontine species in northwestern 
<collectingCountry id="19A1766BFF9BFFBEDBB18837FCA6F9F7" box="[764,885,1608,1634]" name="Argentina" pageId="24" pageNumber="25">Argentina</collectingCountry>
. 
<bibRefCitation id="05274B0AFF9BFFBEDACD8837FB6AF9F7" author="Musser, G. M. &amp; Carleton, M. D." box="[896,1209,1608,1634]" editor="Wilson, D. E." journalOrPublisher="Baltimore: John Hopkins University Press" pageId="24" pageNumber="25" pagination="894 - 1531" refString="Musser, G. M. &amp; Carleton, M. D. (2005) Superfamily Muroidea .. In: Wilson, D. E. &amp; Reeder, D. M. (Eds.) Mammal species of the world: A taxonomic and geographic reference. Third ed. Baltimore: John Hopkins University Press, pp. 894 - 1531." title="Superfamily Muroidea .." type="book chapter" volumeTitle="Mammal species of the world: A taxonomic and geographic reference. Third ed" year="2005">Musser &amp; Carleton (2005)</bibRefCitation>
and 
<bibRefCitation id="05274B0AFF9BFFBEDDB98837FF36F912" author="Jayat, J. P. &amp; Ortiz, P. E. &amp; Teta, P. &amp; Pardinas, U. F. J. &amp; D'Elia, G." journalOrPublisher="Mastozoologia Neotropical" pageId="24" pageNumber="25" pagination="51 - 67" part="13" refString="Jayat, J. P., Ortiz, P. E., Teta, P., Pardinas, U. F. J. &amp; D'Elia, G. (2006) Nuevas localidades argentinas para algunos roedores sigmodontinos (Rodentia: Cricetidae). Mastozoologia Neotropical, 13, 51 - 67." title="Nuevas localidades argentinas para algunos roedores sigmodontinos (Rodentia: Cricetidae)" type="journal article" year="2006">
Pardiñas 
<emphasis id="53C2EAE9FF9BFFBEDC2D8837FA46F9F4" box="[1376,1429,1608,1633]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (2006)
</bibRefCitation>
followed the conclusions of 
<bibRefCitation id="05274B0AFF9BFFBEDB758812FCCDF912" author="Myers, P. &amp; Patton, J. L. &amp; Smith, M. F." box="[568,798,1645,1671]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="24" pageNumber="25" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. &amp; Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
Myers 
<emphasis id="53C2EAE9FF9BFFBEDBC78812FD6EF913" box="[650,701,1645,1670]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1990)
</bibRefCitation>
.
</paragraph>
<paragraph id="610936FBFF9BFFBDD98B88EDFF10FDB2" blockId="24.[151,1437,152,2007]" lastBlockId="27.[151,1437,152,551]" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
<bibRefCitation id="05274B0AFF9BFFBED98B88EDFE73F939" author="Diaz, M. M." bookContentInfo="640 pp." box="[198,416,1682,1708]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="24" pageNumber="25" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">
Díaz 
<emphasis id="53C2EAE9FF9BFFBED84588ECFEECF939" box="[264,319,1683,1708]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1999)
</bibRefCitation>
described 
<taxonomicName id="A6B64D78FF9BFFBEDB6B88ECFCCEF939" box="[550,797,1683,1708]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF9BFFBEDB6B88ECFCCEF939" box="[550,797,1683,1708]" italics="true" pageId="24" pageNumber="25">Akodon aliquantulus</emphasis>
</taxonomicName>
, from the ecotone between upper Yungas forest and highland grassland in 
<collectingRegion id="A372F819FF9BFFBED8D488C7FDA6F947" box="[409,629,1720,1746]" country="Argentina" name="Tucuman" pageId="24" pageNumber="25">Tucumán province</collectingRegion>
(Las Agüitas, Cumbres del Taficillo, 
<quantity id="A64E9B1EFF9BFFBEDD6288C7FB5FF947" box="[1071,1164,1720,1746]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="25" unit="m" value="1700.0">1700 m</quantity>
, 
<geoCoordinate id="0482503CFF9BFFBEDDDA88C7FAD5F947" box="[1175,1286,1720,1746]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="925" value="-26.7">26º 42’ S</geoCoordinate>
, 
<geoCoordinate id="0482503CFF9BFFBEDC5E88C7FA5DF947" box="[1299,1422,1720,1746]" direction="west" orientation="longitude" pageId="24" pageNumber="25" precision="925" value="-65.36667">65º 22’ W</geoCoordinate>
), and related it to the 
<taxonomicName id="A6B64D78FF9BFFBED8CC88A2FDCCF963" box="[385,543,1757,1783]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="boliviensis">
<emphasis id="53C2EAE9FF9BFFBED8CC88A2FE40F963" box="[385,403,1757,1782]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED8EF88A2FDCCF963" box="[418,543,1757,1782]" italics="true" pageId="24" pageNumber="25">boliviensis</emphasis>
</taxonomicName>
group on the basis of its morphology. The unimpressive morphological distinction from 
<taxonomicName id="A6B64D78FF9BFFBED80E897CFE07F889" box="[323,468,1794,1820]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="lutescens">
<emphasis id="53C2EAE9FF9BFFBED80E897CFE86F889" box="[323,341,1795,1820]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBED824897CFE07F889" box="[361,468,1795,1820]" italics="true" pageId="24" pageNumber="25">lutescens</emphasis>
</taxonomicName>
, based only on multivariate ordinations and univariate overlap, induced some authors to recommend a revision of the status of this form (
<bibRefCitation id="05274B0AFF9BFFBEDA078957FBA2F8D7" author="Musser, G. M. &amp; Carleton, M. D." box="[842,1137,1832,1858]" editor="Wilson, D. E." journalOrPublisher="Baltimore: John Hopkins University Press" pageId="24" pageNumber="25" pagination="894 - 1531" refString="Musser, G. M. &amp; Carleton, M. D. (2005) Superfamily Muroidea .. In: Wilson, D. E. &amp; Reeder, D. M. (Eds.) Mammal species of the world: A taxonomic and geographic reference. Third ed. Baltimore: John Hopkins University Press, pp. 894 - 1531." title="Superfamily Muroidea .." type="book chapter" volumeTitle="Mammal species of the world: A taxonomic and geographic reference. Third ed" year="2005">Musser &amp; Carleton 2005</bibRefCitation>
; 
<bibRefCitation id="05274B0AFF9BFFBEDD308957FA87F8D7" author="Jayat, J. P. &amp; Ortiz, P. E. &amp; Miotti, M. D." box="[1149,1364,1832,1858]" journalOrPublisher="Acta Zoologica Mexicana" pageId="24" pageNumber="25" pagination="137 - 177" part="24" refString="Jayat, J. P., Ortiz, P. E. &amp; Miotti, M. D. (2008 a) Distribucion de sigmodontinos (Rodentia: Cricetidae) en pastizales de neblina del noroeste argentino. Acta Zoologica Mexicana, 24, 137 - 177." title="Distribucion de sigmodontinos (Rodentia: Cricetidae) en pastizales de neblina del noroeste argentino" type="journal article" year="2008" yearSuffix="a">
Jayat 
<emphasis id="53C2EAE9FF9BFFBEDDF28957FB29F8D4" box="[1215,1274,1832,1857]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
, 2008a
</bibRefCitation>
). This species was diagnosed and differentiated from 
<taxonomicName id="A6B64D78FF9BFFBEDB878932FC8BF8F3" box="[714,856,1869,1895]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDB878932FD0FF8F3" box="[714,732,1869,1894]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDBA08932FC8BF8F3" box="[749,856,1869,1894]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
(as 
<taxonomicName id="A6B64D78FF9BFFBEDAC08932FB88F8F3" box="[909,1115,1869,1895]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="subSpecies" species="puer" subSpecies="caenosus">
<emphasis id="53C2EAE9FF9BFFBEDAC08932FC4CF8F3" box="[909,927,1869,1894]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDAFD8932FB88F8F3" box="[944,1115,1869,1894]" italics="true" pageId="24" pageNumber="25">puer caenosus</emphasis>
</taxonomicName>
) and 
<taxonomicName id="A6B64D78FF9BFFBEDDEE8932FA94F8F3" box="[1187,1351,1869,1895]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="spegazzinii">
<emphasis id="53C2EAE9FF9BFFBEDDEE8932FB66F8F3" box="[1187,1205,1869,1894]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDD888932FA94F8F3" box="[1221,1351,1869,1894]" italics="true" pageId="24" pageNumber="25">spegazzinii</emphasis>
</taxonomicName>
on the account of being smaller in a number of measurements and the “weakly developed eye ring” of two individuals originally preserved in fluid and then prepared as skin plus skull. We could not distinguish the 
<typeStatus id="BE0D8859FF9BFFBEDC5189E7FA9DF827" box="[1308,1358,1944,1970]" pageId="24" pageNumber="25">type</typeStatus>
specimens of 
<taxonomicName id="A6B64D78FF9BFFBED9B689C2FE7DF843" box="[251,430,1981,2006]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF9BFFBED9B689C2FE7DF843" box="[251,430,1981,2006]" italics="true" pageId="24" pageNumber="25">A. aliquantulus</emphasis>
</taxonomicName>
from 
<taxonomicName id="A6B64D78FF9BFFBED8B989C2FDACF843" box="[500,639,1981,2007]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF9BFFBED8B989C2FDD5F843" box="[500,518,1981,2006]" italics="true" pageId="24" pageNumber="25">A</emphasis>
. 
<emphasis id="53C2EAE9FF9BFFBEDB5989C2FDACF843" box="[532,639,1981,2006]" italics="true" pageId="24" pageNumber="25">caenosus</emphasis>
</taxonomicName>
following the diagnosis provided by 
<bibRefCitation id="05274B0AFF9BFFBEDD7E89C2FAD7F842" author="Diaz, M. M." bookContentInfo="640 pp." box="[1075,1284,1981,2007]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="24" pageNumber="25" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">
Díaz 
<emphasis id="53C2EAE9FF9BFFBEDD3C89C2FB77F843" box="[1137,1188,1981,2006]" italics="true" pageId="24" pageNumber="25">et al</emphasis>
. (1999)
</bibRefCitation>
. Most of the characters used for the diagnosis of 
<taxonomicName id="A6B64D78FF99FFBCDB0A8EE7FD2FFF24" box="[583,764,152,178]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF99FFBCDB0A8EE7FD8AFF24" box="[583,601,152,177]" italics="true" pageId="26" pageNumber="27">A</emphasis>
. 
<emphasis id="53C2EAE9FF99FFBCDB248EE7FD2FFF24" box="[617,764,152,177]" italics="true" pageId="26" pageNumber="27">aliquantulus</emphasis>
</taxonomicName>
(centered, as said, on a few external and cranial dimensions) are subtle, highly variable in many sigmodontine species, and what is most important are age-dependent. Although 
<bibRefCitation id="05274B0AFF99FFBCD8138E9DFD97FF69" author="Diaz, M. M." bookContentInfo="640 pp." box="[350,580,226,252]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="26" pageNumber="27" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">
Díaz 
<emphasis id="53C2EAE9FF99FFBCD8E98E9DFE33FF6E" box="[420,480,226,251]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. (1999)
</bibRefCitation>
considered the two specimens as old individuals (age-class 5) our observations indicate that they must be assigned to the age class 4 of 
<bibRefCitation id="05274B0AFF99FFBCDA8C8F77FBB2FEB7" author="Myers, P." box="[961,1121,264,290]" editor="Redford, K. H." journalOrPublisher="Gainesville, Florida: Sandhill Crane Press, Inc" pageId="26" pageNumber="27" pagination="5 - 54" refString="Myers, P. (1989) A preliminary revision of the varius group of Akodon (A. dayi, dolores, molinae, neocenus, simulator, toba and varius). In: Redford, K. H. &amp; Eisenberg, J. F. (Eds.), Advances in Neotropical Mammalogy. Gainesville, Florida: Sandhill Crane Press, Inc., pp. 5 - 54." title="A preliminary revision of the varius group of Akodon (A. dayi, dolores, molinae, neocenus, simulator, toba and varius)" type="book chapter" volumeTitle="Advances in Neotropical Mammalogy" year="1989">Myers (1989)</bibRefCitation>
(
<figureCitation id="F98D2A7EFF99FFBCDD3C8F77FB04FEB7" box="[1137,1239,264,290]" captionStart="FIGURE 8" captionStartId="26.[151,255,1930,1954]" captionTargetBox="[307,1263,338,1890]" captionTargetId="figure@26.[307,1264,330,1890]" captionTargetPageId="26" captionText="FIGURE 8. Comparison between the toothwear classes of Myers (1989) (A) and Akodon aliquantulus left upper molar rows (left; Holotype, CML 6122 and rigth; paratype, CML 6121) (B). Scale bar = 1 mm." httpUri="https://zenodo.org/record/293469/files/figure.png" pageId="26" pageNumber="27">Figure 8</figureCitation>
). The morphometric distinction between 
<taxonomicName id="A6B64D78FF98FFBDD8998EE7FD54FF24" box="[468,647,152,178]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF98FFBDD8998EE7FE35FF24" box="[468,486,152,177]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDD8B98EE7FD54FF24" box="[500,647,152,177]" italics="true" pageId="27" pageNumber="28">aliquantulus</emphasis>
</taxonomicName>
and 
<taxonomicName id="A6B64D78FF98FFBDDBF38EE7FC9AFF24" box="[702,841,152,178]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF98FFBDDBF38EE7FD03FF24" box="[702,720,152,177]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDDB938EE7FC9AFF24" box="[734,841,152,177]" italics="true" pageId="27" pageNumber="28">caenosus</emphasis>
</taxonomicName>
was based on a PCA analysis with a relatively low explained variance (63.3%, 7.9% and 5.1% for components 1, 2 and 3 respectively) and the statistic significance of the observed differences was not provided by the authors. We think that the erroneous assignment of age for the two specimens by 
<bibRefCitation id="05274B0AFF98FFBDD8BF8F77FD6DFEB7" author="Diaz, M. M." bookContentInfo="640 pp." box="[498,702,264,290]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="27" pageNumber="28" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">
Díaz 
<emphasis id="53C2EAE9FF98FFBDDB7D8F77FDB0FEB4" box="[560,611,264,289]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. (1999)
</bibRefCitation>
misinform their conclusions. The analysis of table 1 of 
<bibRefCitation id="05274B0AFF98FFBDDC058F77FED9FED2" author="Diaz, M. M." bookContentInfo="640 pp." journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="27" pageNumber="28" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">
Díaz 
<emphasis id="53C2EAE9FF98FFBDDCCA8F77FF7DFED3" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. (1999)
</bibRefCitation>
clearly indicates the overlap between 
<taxonomicName id="A6B64D78FF98FFBDDB878F52FCADFED3" box="[714,894,301,327]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF98FFBDDB878F52FD0FFED3" box="[714,732,301,326]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDDBA68F52FCADFED3" box="[747,894,301,326]" italics="true" pageId="27" pageNumber="28">aliquantulus</emphasis>
</taxonomicName>
specimens and 
<taxonomicName id="A6B64D78FF98FFBDDD7A8F52FB10FED3" box="[1079,1219,301,327]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF98FFBDDD7A8F52FB9AFED3" box="[1079,1097,301,326]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDDD158F52FB10FED3" box="[1112,1219,301,326]" italics="true" pageId="27" pageNumber="28">caenosus</emphasis>
</taxonomicName>
exemplars of ageclass 4 (see also 
<tableCitation id="2C340340FF98FFBDD8288F2DFE6DFEF9" box="[357,446,338,364]" captionStart="TABLE 9" captionStartId="15.[151,239,932,956]" captionTargetBox="[160,1410,1247,2025]" captionTargetPageId="15" captionText="TABLE 9. Comparative measurements of adult specimens (age classes 3 and 4) of A. caenosus and holotype and paratype of A. aliquantulus (age class 4). The values for specimens of around the type locality of A. lutescens caenosus from Jujuy province are distinguished from those of remaining localities of Northwestern Argentina. Measurements for A. aliquantulus were extracted from Díaz et al. (1999), except for those indicated with an asterisk; these were taken by us. X: mean; SD: standard deviation; r: range; n: sample size." httpUri="http://table.plazi.org/id/35C96673FF8CFFA9D9DA8DDBFD33FBD3" pageId="27" pageNumber="28" tableUuid="35C96673FF8CFFA9D9DA8DDBFD33FBD3">Table 9</tableCitation>
of this work). Remarkably, even some of the diagnostic skull measurements (e.g. braincase breadth, maxillary toothrow length) are included in the observed range for age-class 5 individuals of 
<taxonomicName id="A6B64D78FF98FFBDD9DA8FE2FEF1FE23" box="[151,290,413,439]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF98FFBDD9DA8FE2FF7AFE23" box="[151,169,413,438]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDD9FA8FE2FEF1FE23" box="[183,290,413,438]" italics="true" pageId="27" pageNumber="28">caenosus</emphasis>
</taxonomicName>
. Moreover, all the characters mentioned in the description of 
<taxonomicName id="A6B64D78FF98FFBDDAB88FE2FB7BFE23" box="[1013,1192,413,439]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF98FFBDDAB88FE2FBD4FE23" box="[1013,1031,413,438]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDDD588FE2FB7BFE23" box="[1045,1192,413,438]" italics="true" pageId="27" pageNumber="28">aliquantulus</emphasis>
</taxonomicName>
are within the levels of morphological variation observed in 
<taxonomicName id="A6B64D78FF98FFBDDB248FBDFD26FE4E" box="[617,757,450,476]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF98FFBDDB248FBDFDA8FE4E" box="[617,635,450,475]" italics="true" pageId="27" pageNumber="28">A</emphasis>
. 
<emphasis id="53C2EAE9FF98FFBDDBC78FBDFD26FE4E" box="[650,757,450,475]" italics="true" pageId="27" pageNumber="28">caenosus</emphasis>
</taxonomicName>
of Northwestern 
<collectingCountry id="19A1766BFF98FFBDDA888FBDFBEDFE49" box="[965,1086,450,476]" name="Argentina" pageId="27" pageNumber="28">Argentina</collectingCountry>
. Therefore, we find no defensible argument to consider 
<taxonomicName id="A6B64D78FF98FFBDD8988F97FD5AFD94" box="[469,649,488,513]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="aliquantulus">
<emphasis id="53C2EAE9FF98FFBDD8988F97FD5AFD94" box="[469,649,488,513]" italics="true" pageId="27" pageNumber="28">A. aliquantulus</emphasis>
</taxonomicName>
as a distinct species and consider it a junior synonym of 
<taxonomicName id="A6B64D78FF98FFBDDC668F97FF6FFDB3" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="caenosus">
<emphasis id="53C2EAE9FF98FFBDDC668F97FF6FFDB3" italics="true" pageId="27" pageNumber="28">A. caenosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>