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<document ID-DOI="http://doi.org/10.5281/zenodo.5646354" ID-GBIF-Dataset="5566188f-1bd9-49b8-8862-d5a49752355f" ID-GBIF-Taxon="189991411" ID-ISSN="2147-7612" ID-Zenodo-Dep="5646354" LSID="urn:lsid:zoobank.org:act:E7B4F251-9690-4022-A96F-C97A97849A97" approvalRequired="3" approvalRequired_for_document="1" approvalRequired_for_taxonomicNames="1" approvalRequired_for_treatments="1" checkinTime="1636056240689" checkinUser="felipe" docAuthor="Löbl, Ivan &amp; Smetana, Aleš" docDate="2021" docId="D0225B07FFA4FF9AD6DB819E0B9209E3" docLanguage="en" docName="JInsectBiodiv.023.2.023-042.pdf" docOrigin="J. Insect Biodiversity 23 (2)" docSource="https://doi.org/10.12976/jib/2021.23.2.1" docTitle="Baeocera segregata Lobl 2021, sp. nov." docType="treatment" docUuid="E7B4F251-9690-4022-A96F-C97A97849A97" docUuidSource="ZooBank" docVersion="9" lastPageNumber="34" masterDocId="2C1B237FFFAFFF91D653807D08350F27" masterDocTitle="On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity" masterLastPageNumber="42" masterPageNumber="23" pageId="11" pageNumber="34" updateTime="1636080418144" updateUser="felipe">
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<mods:titleInfo>
<mods:title>On the Baeocera Erichson (Coleoptera: Staphylinidae: Scaphidiinae) of Sabah, Malaysia, and a tale on mystified biodiversity</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Löbl, Ivan</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-8747-3702</mods:nameIdentifier>
<mods:affiliation>Muséum dHistoire Naturelle, Case postale 6434, CH- 1211 Genève, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">ivan.lobl@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smetana, Aleš</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7804-4706</mods:nameIdentifier>
<mods:affiliation>Canadian National Collection of Insects, Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, ON KIA 0 C 6, Canada</mods:affiliation>
<mods:nameIdentifier type="email">ales.smetana7@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>J. Insect Biodiversity</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-16</mods:number>
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<mods:detail type="volume">
<mods:number>23</mods:number>
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<mods:number>2</mods:number>
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<mods:start>23</mods:start>
<mods:end>42</mods:end>
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<mods:url>https://doi.org/10.12976/jib/2021.23.2.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.12976/jib/2021.23.2.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">5566188f-1bd9-49b8-8862-d5a49752355f</mods:identifier>
<mods:identifier type="ISSN">2147-7612</mods:identifier>
<mods:identifier type="Zenodo-Dep">5646470</mods:identifier>
<mods:identifier type="ZooBank">70420B00-3668-4A16-9970-395070A02D64</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5646354" ID-GBIF-Taxon="189991411" ID-Zenodo-Dep="5646354" LSID="urn:lsid:zoobank.org:act:E7B4F251-9690-4022-A96F-C97A97849A97" httpUri="http://treatment.plazi.org/id/D0225B07FFA4FF9AD6DB819E0B9209E3" lastPageNumber="34" pageId="11" pageNumber="34">
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<paragraph blockId="11.[136,942,482,581]" box="[136,529,482,509]" pageId="11" pageNumber="34">
<heading bold="true" box="[136,529,482,509]" fontSize="11" level="1" pageId="11" pageNumber="34" reason="1">
<taxonomicName authority="Lobl" authorityName="Lobl" authorityYear="2021" box="[136,425,482,509]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="segregata" status="sp. nov.">
<emphasis bold="true" box="[136,425,482,509]" pageId="11" pageNumber="34">
<emphasis bold="true" box="[136,362,483,509]" italics="true" pageId="11" pageNumber="34">Baeocera segregata</emphasis>
Löbl
</emphasis>
</taxonomicName>
,
<emphasis bold="true" box="[439,529,482,508]" pageId="11" pageNumber="34">
<taxonomicNameLabel box="[439,529,482,508]" pageId="11" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="34" type="description">
<paragraph blockId="11.[136,942,482,581]" box="[136,942,518,545]" pageId="11" pageNumber="34">
<uri box="[136,942,518,545]" pageId="11" pageNumber="34">
urn:lsid:zoobank.org:act:
<uuid box="[423,942,518,545]" pageId="11" pageNumber="34">E7B4F251-9690-4022-A96F-C97A97849A97</uuid>
</uri>
</paragraph>
<paragraph blockId="11.[136,942,482,581]" box="[136,280,554,581]" pageId="11" pageNumber="34">
(
<figureCitation box="[144,272,554,581]" captionStart="Figures 815" captionStartId="10.[136,213,1945,1967]" captionTargetBox="[199,1394,206,1899]" captionTargetId="figure-31@10.[171,1416,187,1923]" captionText="Figures 815. Baeocera spp., aedeagi; (89) B. secreta sp. nov.. in dorsal and lateral views, scale = 0.1 mm; (10) ditto, paramere in ventral view, scale = 0.05 mm; (11) ditto, internal sac in dorsal view, scale = 0.05 mm. (12, 13) B. segregata sp. nov., in dorsal and lateral views (left paramere missing), scale = 0.1 mm; (14) ditto, paramere in ventral view, scale = 0.05 mm; (15) ditto, internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5646474" httpUri="https://zenodo.org/record/5646474/files/figure.png" pageId="11" pageNumber="34">Figs 1215</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[136,1348,626,653]" pageId="11" pageNumber="34" type="materials_examined">
<paragraph blockId="11.[136,1452,626,1733]" box="[136,1348,626,653]" pageId="11" pageNumber="34">
<materialsCitation ID-GBIF-Occurrence="3398245415" box="[136,1344,626,653]" collectingDate="1987-05-11" collectorName="Burckhardt &amp; Lobl" country="Malaysia" elevation="500" location="Poring Hot Springs" pageId="11" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis bold="true" box="[136,370,626,652]" pageId="11" pageNumber="34">Material examined.</emphasis>
<typeStatus box="[376,480,626,652]" pageId="11" pageNumber="34">Holotype</typeStatus>
<specimenCount box="[488,511,627,653]" pageId="11" pageNumber="34" type="male"></specimenCount>
,
<collectingRegion box="[523,596,626,652]" country="Malaysia" name="Sabah" pageId="11" pageNumber="34">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:D0225B07FFA4FF9AD6DB819E0B9209E3:5D54BCCAFFA4FF9AD40D820F0B030DAB" box="[606,822,626,653]" country="Malaysia" name="Poring Hot Springs" pageId="11" pageNumber="34" stateProvince="Sabah">Poring Hot Springs</location>
<quantity box="[829,900,626,653]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="34" unit="m" value="500.0">
<elevation box="[829,900,626,653]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="34" unit="m" value="500.0">500 m</elevation>
</quantity>
<date box="[907,1020,626,652]" pageId="11" pageNumber="34" value="1987-05-11">
<collectingDate box="[907,1020,626,652]" pageId="11" pageNumber="34" value="1987-05-11">11.V.1987</collectingDate>
</date>
<collectorName box="[1027,1153,626,652]" pageId="11" pageNumber="34">Burckhardt</collectorName>
-
<collectorName box="[1177,1230,626,652]" pageId="11" pageNumber="34">Löbl</collectorName>
(
<collectionCode box="[1244,1336,626,652]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="11" pageNumber="34" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="34" type="description">
<paragraph blockId="11.[136,1452,626,1733]" pageId="11" pageNumber="34">
<emphasis bold="true" box="[136,282,662,688]" pageId="11" pageNumber="34">Description.</emphasis>
Length
<quantity box="[384,490,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="34" unit="mm" value="1.15">1.15 mm</quantity>
, width
<quantity box="[582,689,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="11" pageNumber="34" unit="mm" value="0.76">0.76 mm</quantity>
. Body light reddish-brown, mesoventrite, metaventrite and elytra slightly darker than pronotum and abdomen, femora and tibiae as pronotum, tarsi and antennae yellowish. Head with interocular distance slightly smaller than dorsoventral eye diameter. Lateral contours of pronotum and elytra nearly continuously arcuate.Pronotum not microsculptured, with punctation very fine, hardly visible at 40 times magnification, lateral margin carinae not visible in dorsal view. Tip of scutellum exposed, triangular. Elytra not microsculptured, with lateral margins rounded, lateral margin carinae concealed in dorsal view, sutural striae parallel with suture, curved at bases and extended along basal margins to form basal striae joined with lateral striae; adsutural areas flat, finely punctate in basal halves, impunctate in apical halves. Discal punctation coarse anterior elytral mid-length, punctures well delimited, mostly somewhat smaller than puncture intervals. Punctation hardly visible (
<date box="[1189,1241,950,977]" pageId="11" pageNumber="34" value="1980-10">80 x</date>
magnification) on apical halves of elytra. Hind wings fully developed. Hypomera appearing impunctate. Mesoventrite without mesal ridge, intercoxal process punctate. Mesepimera nearly four times as long as wide and about three times as long as intervals to coxae. Centre of metaventrite flat, with scattered, fairly coarse punctures, to part impunctate. Lateral areas of metaventrite densely and coarsely punctate, punctures round, much larger than puncture intervals and larger than punctures on centre of metaventrite. Submesocoxal areas
<quantity box="[849,951,1130,1157]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="34" unit="mm" value="0.01">0.01 mm</quantity>
long, densely margined by round punctures as large as those on centre of metaventrite. Metanepisterna concealed. Tibiae straight. Ventrite I lacking striae, with punctures margining base not elongate, smaller than most punctures on sides of metaventrite, not interrupted at middle. Remaining abdominal punctation very fine and sparse, hardly visible at
<date box="[943,992,1238,1265]" pageId="11" pageNumber="34" value="1980-10">80 x</date>
magnification.
</paragraph>
<paragraph blockId="11.[136,1452,626,1733]" pageId="11" pageNumber="34">
<emphasis box="[136,321,1274,1300]" italics="true" pageId="11" pageNumber="34">Male characters</emphasis>
. Protarsomeres I to III hardly widened. Aedeagus (
<figureCitation box="[893,1021,1274,1301]" captionStart="Figures 815" captionStartId="10.[136,213,1945,1967]" captionTargetBox="[199,1394,206,1899]" captionTargetId="figure-31@10.[171,1416,187,1923]" captionText="Figures 815. Baeocera spp., aedeagi; (89) B. secreta sp. nov.. in dorsal and lateral views, scale = 0.1 mm; (10) ditto, paramere in ventral view, scale = 0.05 mm; (11) ditto, internal sac in dorsal view, scale = 0.05 mm. (12, 13) B. segregata sp. nov., in dorsal and lateral views (left paramere missing), scale = 0.1 mm; (14) ditto, paramere in ventral view, scale = 0.05 mm; (15) ditto, internal sac in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5646474" httpUri="https://zenodo.org/record/5646474/files/figure.png" pageId="11" pageNumber="34">Figs 1215</figureCitation>
)
<quantity box="[1036,1136,1274,1301]" metricMagnitude="-4" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="34" unit="mm" value="0.34">0.34 mm</quantity>
long. Median lobe in dorsal view abruptly narrowed apically, with tip acute and bent; apical process strongly inflexed, about as long as third of basal bulb, ventral contour of apical process in lateral view arcuate. Articular process not prominent. Parameres fairly wide anterior mid-length, narrowed at mid-length, straight in dorsal view, weakly bent in lateral view. Internal sac with flagellar-guide sclerite elongate, not joined with flagellum and overlapped by strongly sclerotized proximal lobelike process. Accessory transverse sclerite strongly sclerotized, acute. Membranes lacking denticulate or squamous vesicles, ejaculatory duck not vesicular anterior entering flagellum.
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="34" type="diagnosis">
<paragraph blockId="11.[136,1452,626,1733]" pageId="11" pageNumber="34">
<emphasis bold="true" box="[136,399,1526,1552]" pageId="11" pageNumber="34">Differential diagnosis.</emphasis>
This species resembles
<taxonomicName authority="Lobl, 1990" authorityName="Lobl" authorityYear="1990" box="[669,960,1526,1552]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="suthepensis">
<emphasis box="[669,829,1526,1552]" italics="true" pageId="11" pageNumber="34">B. suthepensis</emphasis>
<bibRefCitation author="Lobl I." box="[837,960,1526,1552]" journalOrPublisher="Revue suisse de Zoologie" pageId="11" pageNumber="34" pagination="505 - 621" part="97" refId="ref15659" refString="Lobl I. 1990. Review of the Scaphidiidae (Coleoptera) of Thailand. Revue suisse de Zoologie 97: 505 - 621. https: // doi. org / 10.5962 / bhl. part. 79752" title="Review of the Scaphidiidae (Coleoptera) of Thailand" type="journal article" year="1990">Löbl, 1990</bibRefCitation>
</taxonomicName>
by the shape of the parameres, though their narrowed apical section is notably longer. It may be distinguished from
<emphasis box="[956,1114,1562,1589]" italics="true" pageId="11" pageNumber="34">
S.
<taxonomicName authorityName="Lobl" authorityYear="1990" box="[985,1114,1562,1588]" class="Insecta" family="Staphylinidae" genus="Baeocera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="34" phylum="Arthropoda" rank="species" species="suthepensis">suthepensis</taxonomicName>
</emphasis>
by the smaller body-size, the lighter colour, the membranes of the internal sac lacking scale-like structures, the flagellar-guide sclerite widened and bent apically, and the distinctive shape of the proximal end of the sclerotized complex.
</paragraph>
</subSubSection>
<subSubSection box="[136,545,1670,1697]" pageId="11" pageNumber="34" type="distribution">
<paragraph blockId="11.[136,1452,626,1733]" box="[136,545,1670,1697]" pageId="11" pageNumber="34">
<emphasis bold="true" box="[136,290,1670,1696]" pageId="11" pageNumber="34">Distribution.</emphasis>
East Malaysia: Sabah.
</paragraph>
</subSubSection>
<subSubSection box="[136,935,1706,1733]" pageId="11" pageNumber="34" type="etymology">
<paragraph blockId="11.[136,1452,626,1733]" box="[136,935,1706,1733]" pageId="11" pageNumber="34">
<emphasis bold="true" box="[136,271,1706,1732]" pageId="11" pageNumber="34">Etymology.</emphasis>
The species epithet is a Latin adjective meaning separated.
</paragraph>
</subSubSection>
</treatment>
</document>