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<document ID-DOI="10.1206/0003-0090.457.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="6971356" approvalRequired="59" approvalRequired_for_taxonomicNames="59" checkinTime="1659882917898" checkinUser="felipe" docAuthor="Beck, Robin M. D., Voss, Robert S. &amp; Jansa, Sharon A." docDate="2022" docId="03EFDD5DF7756966D95FFA171BBBFA9A" docLanguage="en" docName="BulAmeMusNatHist.2022.457.1-350.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (457)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.7:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="7" docTitle="Hypsiprymnodon bartholomaii Ramsay 1876" docType="treatment" docVersion="8" lastPageNumber="335" masterDocId="FFD6A525F63B6829DA0BFFAE1A62FFC7" masterDocTitle="Craniodental Morphology And Phylogeny Of Marsupials" masterLastPageNumber="353" masterPageNumber="1" pageNumber="334" updateTime="1661891901480" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Craniodental Morphology And Phylogeny Of Marsupials</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beck, Robin M. D.</mods:namePart>
<mods:affiliation>School of Science, Engineering and Environment University of Salford, U. K. &amp; School of Biological, Earth &amp; Environmental Sciences University of New South Wales, Australia &amp; Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Voss, Robert S.</mods:namePart>
<mods:affiliation>Division of Vertebrate Zoology (Mammalogy) American Museum of Natural History</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jansa, Sharon A.</mods:namePart>
<mods:affiliation>Bell Museum and Department of Ecology, Evolution, and Behavior University of Minnesota</mods:affiliation>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-28</mods:number>
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<mods:detail type="volume">
<mods:number>2022</mods:number>
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<mods:detail type="issue">
<mods:number>457</mods:number>
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<mods:url>https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-457/issue-1/0003-0090.457.1.1/Craniodental-Morphology-and-Phylogeny-of-Marsupials/10.1206/0003-0090.457.1.1.full</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1206/0003-0090.457.1.1</mods:identifier>
<mods:identifier type="ISSN">0003-0090</mods:identifier>
<mods:identifier type="Zenodo-Dep">6971356</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6974511" ID-Zenodo-Dep="6974511" LSID="urn:lsid:plazi:treatment:03EFDD5DF7756966D95FFA171BBBFA9A" httpUri="http://treatment.plazi.org/id/03EFDD5DF7756966D95FFA171BBBFA9A" lastPageId="335" lastPageNumber="335" pageId="334" pageNumber="334">
<subSubSection box="[852,1047,1465,1489]" pageId="334" pageNumber="334" type="multiple">
<paragraph blockId="334.[852,1047,1465,1489]" box="[852,1047,1465,1489]" pageId="334" pageNumber="334">
<heading box="[852,1047,1465,1489]" centered="true" fontSize="9" level="2" pageId="334" pageNumber="334" reason="2">
<taxonomicName box="[852,1047,1465,1489]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="334" pageNumber="334" phylum="Chordata" rank="species" species="undetermined">
<emphasis bold="true" box="[852,1047,1465,1489]" italics="true" pageId="334" pageNumber="334">Hypsiprymnodon</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="334" pageNumber="334" type="nomenclature">
<paragraph blockId="334.[684,1212,1515,1738]" pageId="334" pageNumber="334">
SPECIES SCORED:
<taxonomicName authorityName="Ramsay" authorityYear="1876" baseAuthorityName="Flannery and Archer" baseAuthorityYear="1987" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="334" pageNumber="334" phylum="Chordata" rank="species" species="bartholomaii">
<emphasis box="[916,1101,1515,1539]" italics="true" pageId="334" pageNumber="334">Hypsiprymnodon</emphasis>
<emphasis italics="true" pageId="334" pageNumber="334">bartholomaii.</emphasis>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="334" pageNumber="334" type="materials_examined">
<paragraph blockId="334.[684,1212,1515,1738]" pageId="334" pageNumber="334">
GEOLOGICAL PROVENANCE OF SCORED
<collectionCode box="[684,744,1615,1639]" pageId="334" pageNumber="334">SPECIMENS</collectionCode>
: Riversleigh Faunal Zone C, Riversleigh World Heritage Area,
<collectingRegion box="[926,1058,1647,1671]" country="Australia" name="Queensland" pageId="334" pageNumber="334">Queensland</collectingRegion>
,
<collectingCountry box="[1070,1172,1647,1671]" name="Australia" pageId="334" pageNumber="334">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="335" lastPageNumber="335" pageId="334" pageNumber="334" type="description">
<paragraph blockId="334.[684,1212,1515,1738]" lastBlockId="335.[108,637,226,1374]" lastPageId="335" lastPageNumber="335" pageId="334" pageNumber="334">AGES OF SCORED SPECIMENS: Riversleigh Faunal Zone C is interpreted to be middle Miocene based on biostratigraphy (see above). In the absence of radiometric dates, we have assumed the entire span of the middle Miocene (Langhian to Serravallian; Cohen et al., 2013 [updated]) for this terminal.</paragraph>
<paragraph blockId="335.[108,637,226,1374]" box="[140,595,391,415]" pageId="335" pageNumber="335">
ASSIGNED AGE
<collectionCode box="[311,388,391,415]" pageId="335" pageNumber="335">RANGE</collectionCode>
: 15.9711.63 Mya.
</paragraph>
</subSubSection>
<subSubSection pageId="335" pageNumber="335" type="discussion">
<paragraph blockId="335.[108,637,226,1374]" pageId="335" pageNumber="335">
REMARKS:
<taxonomicName authorityName="Ramsay" authorityYear="1876" box="[271,463,424,448]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis box="[271,463,424,448]" italics="true" pageId="335" pageNumber="335">Hypsiprymnodon</emphasis>
</taxonomicName>
<taxonomicName baseAuthorityName="Flannery and Archer" baseAuthorityYear="1987" box="[492,636,424,448]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="bartholomaii">
<emphasis box="[492,636,424,448]" italics="true" pageId="335" pageNumber="335">bartholomaii</emphasis>
</taxonomicName>
was described by Flannery and Archer (1987a) based on a partial cranium and two isolated molars, all from the Gag Site, which is part of Riversleigh Faunal Zone C. If this taxon is indeed referable to
<taxonomicName authorityName="Ramsay" authorityYear="1876" box="[247,434,589,613]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis box="[247,434,589,613]" italics="true" pageId="335" pageNumber="335">Hypsiprymnodon</emphasis>
</taxonomicName>
, then it indicates that the genus originated prior to the middle Miocene. However, Flannery and Archer (1987a) noted a number of striking craniodental differences between
<emphasis box="[277,304,722,745]" italics="true" pageId="335" pageNumber="335">H.</emphasis>
<taxonomicName baseAuthorityName="Flannery and Archer" baseAuthorityYear="1987" box="[323,462,722,746]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="bartholomaii">
<emphasis box="[323,462,722,746]" italics="true" pageId="335" pageNumber="335">bartholomaii</emphasis>
</taxonomicName>
and the Recent species
<taxonomicName authorityName="Ramsay" authorityYear="1876" box="[196,346,755,779]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="species" species="moschatus">
<emphasis box="[196,346,755,779]" italics="true" pageId="335" pageNumber="335">H. moschatus</emphasis>
</taxonomicName>
, notably parietal-alisphenoid versus frontal-squamosal contact and presence versus absence of a distinct postglenoid process. Three further fossil
<taxonomicName pageId="335" pageNumber="335">
<emphasis box="[447,636,854,878]" italics="true" pageId="335" pageNumber="335">Hypsiprymnodon</emphasis>
species
</taxonomicName>
have recently been described from Riversleigh Faunal zones B and C sites (Bates et al., 2014), but we did not examine these for scoring purposes. Some phylogenetic analyses have found
<taxonomicName authorityName="Ramsay" authorityYear="1876" box="[179,363,1019,1043]" class="Mammalia" family="Hypsiprymnodontidae" genus="Hypsiprymnodon" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis box="[179,363,1019,1043]" italics="true" pageId="335" pageNumber="335">Hypsiprymnodon</emphasis>
</taxonomicName>
to be polyphyletic (Black et al., 2014c; den Boer and Kear, 2018: fig. S11), others have found it to be paraphyletic (den Boer and Kear, 2018: figs. S910), and still others have failed to unambiguously support its monophyly (Bates et al., 2014; Butler et al., 2016, 2018; den Boer and Kear, 2018: supplemental data), but that of Travouillon et al. (2016) placed Recent and fossil
<taxonomicName box="[215,481,1284,1308]" pageId="335" pageNumber="335">
<emphasis box="[215,398,1284,1308]" italics="true" pageId="335" pageNumber="335">Hypsiprymnodon</emphasis>
species
</taxonomicName>
in a clade that also included the propleopines, †
<taxonomicName baseAuthorityName="Archer and Flannery" baseAuthorityYear="1985" box="[511,629,1317,1341]" class="Mammalia" family="Macropodidae" genus="Ekaltadeta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis box="[511,629,1317,1341]" italics="true" pageId="335" pageNumber="335">Ekaltadeta</emphasis>
</taxonomicName>
, †
<taxonomicName authorityName="Ride" authorityYear="1993" box="[121,275,1350,1374]" class="Mammalia" family="Macropodidae" genus="Jackmahoneya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis box="[121,275,1350,1374]" italics="true" pageId="335" pageNumber="335">Jackmahoneya</emphasis>
</taxonomicName>
, and †
<taxonomicName authorityName="Longman" authorityYear="1924" box="[351,468,1350,1374]" class="Mammalia" family="Macropodidae" genus="Propleopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="335" pageNumber="335" phylum="Chordata" rank="genus">
<emphasis box="[351,468,1350,1374]" italics="true" pageId="335" pageNumber="335">Propleopus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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