277 lines
45 KiB
XML
277 lines
45 KiB
XML
<document id="01E6E218BCA428E65B283C4D54852E4B" ID-DOI="10.1080/00222930500218664" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629351132246" checkinUser="felipe" docAuthor="Schmelz, R. M. & Römbke, J." docDate="2005" docId="03E3594E584202616DBC0285A8C2FB1C" docLanguage="en" docName="JNatHist.39.32.2967-2986.pdf" docOrigin="Journal of Natural History 39 (32)" docSource="http://dx.doi.org/10.1080/00222930500218664" docStyle="DocumentStyle:5A9EB2ED5589A523F0236C66DE2A6BA8.4:JNatHist.2005-2007.journal_article.1cover" docStyleId="5A9EB2ED5589A523F0236C66DE2A6BA8" docStyleName="JNatHist.2005-2007.journal_article.1cover" docStyleVersion="4" docTitle="Hemienchytraeus tanjae Schmelz & Römbke 2005, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2979" masterDocId="FFDA21365848026C6C0B0632AA02FF9E" masterDocTitle="Three new species of Hemienchytraeus (Enchytraeidae, Oligochaeta) from Amazonian forest soil" masterLastPageNumber="2986" masterPageNumber="2967" pageNumber="2976" updateTime="1699069100524" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
|
||
<mods:mods id="8153DFCEA8087059A7D28FF76C4159A5" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="0B4AFFA32FCC8A10749DB00DD632B6EA">
|
||
<mods:title id="A8D84A7E17B7B8D9C8AC584A4DC3C65E">Three new species of Hemienchytraeus (Enchytraeidae, Oligochaeta) from Amazonian forest soil</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="4534F702A75405479B4EAD5F2E032020" type="personal">
|
||
<mods:role id="D2987F52141516FA53F2965834A988D0">
|
||
<mods:roleTerm id="0CAF757EB0F1ACFAB2AB906D2E57DE3E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0171F08F8244340FCCC13546FE853601">Schmelz, R. M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="1870EA891618420D0BE9CE288ADA143D" type="personal">
|
||
<mods:role id="87DAEDC6AA2FCD868A1A78A18EEF74F8">
|
||
<mods:roleTerm id="05C4FE8DBA64D154B71F939F42A77EBA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0B85C3CF0B3DABE9F05866070AA01962">Römbke, J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3026905625D4F25C9652323F1F835073">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F63813EA03589FA3371C343A360D5C2C" type="host">
|
||
<mods:titleInfo id="E8952A2657DE806F978459F9B72E6947">
|
||
<mods:title id="08D6730585457C8FC0AEE09604A1BB27">Journal of Natural History</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="61921F6AC051E6E5BAA462459BC190C1">
|
||
<mods:date id="059CA2205B574D4247CF4BE7A58D48AF">2005</mods:date>
|
||
<mods:detail id="E4DFC4177B0F59B4C8812310A8259D18" type="pubDate">
|
||
<mods:number id="A696F0DEB36F372268FF7FA99B23745C">2005-09-27</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="27E96EA22B9B9342DFA91E89959FAFC7" type="volume">
|
||
<mods:number id="55439B2FF154C8384C3AFCA9A9CB95A9">39</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="D3E435AB1576F07E3C3EFCD64769F096" type="issue">
|
||
<mods:number id="58723329045EA7CE74283407AB8CC1AF">32</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="96A69A394E59043FDFC19AAA79687036" unit="page">
|
||
<mods:start id="6D4C6800BD73E2CBF7BF8BA29527B4D8">2967</mods:start>
|
||
<mods:end id="512B8FAB273AFAE2FDCE2BC0BEF31B57">2986</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="009BBB1FE25B4CD7AB5A8EE639C6D99C">
|
||
<mods:url id="38D392462929673252CF3239FEA5E746">http://dx.doi.org/10.1080/00222930500218664</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="7D3CF46AFB69AF44FE442CF79A43B542">journal article</mods:classification>
|
||
<mods:identifier id="0051BD634303518BC99F4BE2D855A10F" type="DOI">10.1080/00222930500218664</mods:identifier>
|
||
<mods:identifier id="64CE66A11871BEB291BA2550CCFB43A2" type="ISSN">1464-5262</mods:identifier>
|
||
<mods:identifier id="5D50124704F6A24B15231659A2100822" type="Zenodo-Dep">5220042</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E3594E584202616DBC0285A8C2FB1C" LSID="urn:lsid:plazi:treatment:03E3594E584202616DBC0285A8C2FB1C" httpUri="http://treatment.plazi.org/id/03E3594E584202616DBC0285A8C2FB1C" lastPageId="13" lastPageNumber="2979" pageId="10" pageNumber="2976">
|
||
<subSubSection id="C350BBD3584202666DBC0285A958FB51" box="[439,858,1207,1231]" pageId="10" pageNumber="2976" type="nomenclature">
|
||
<paragraph id="8BF5E858584202666DBC0285A958FB51" blockId="10.[439,858,1207,1264]" box="[439,858,1207,1231]" pageId="10" pageNumber="2976">
|
||
<heading id="D0BD5F34584202666DBC0285A958FB51" box="[439,858,1207,1231]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="2976" reason="2">
|
||
<taxonomicName id="4C4A93DB584202666DBC0285A8F7FB51" ID-CoL="852QW" authority="Schmelz & Römbke, 2005" authorityName="Schmelz & Römbke" authorityYear="2005" box="[439,757,1207,1231]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="10" pageNumber="2976" phylum="Annelida" rank="species" species="tanjae" status="sp. nov.">
|
||
<emphasis id="B93E344A584202666DBC0285A8F7FB51" bold="true" box="[439,757,1207,1231]" italics="true" pageId="10" pageNumber="2976">Hemienchytraeus tanjae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20D8931584202666EF50285A958FB51" box="[766,858,1207,1231]" pageId="10" pageNumber="2976" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C350BBD3584202616E4A02EAA8C2FB1C" lastPageId="13" lastPageNumber="2979" pageId="10" pageNumber="2976" type="description">
|
||
<paragraph id="8BF5E858584202666E4A02EAA8B6FB6E" blockId="10.[439,858,1207,1264]" box="[577,692,1240,1264]" pageId="10" pageNumber="2976">
|
||
(
|
||
<figureCitation id="1371F4DD584202666E4202EAA8A8FB6E" box="[585,682,1240,1264]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="10" pageNumber="2976">Figure 3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584202666C76031BAB17FADE" blockId="10.[125,277,1320,1344]" box="[125,277,1320,1344]" pageId="10" pageNumber="2976">
|
||
<emphasis id="B93E344A584202666C76031BAB17FADE" box="[125,277,1320,1344]" italics="true" pageId="10" pageNumber="2976">Type material</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584202666C76036BA99EFA4D" blockId="10.[125,1172,1369,1491]" pageId="10" pageNumber="2976">
|
||
<materialsCitation id="3B22E205584202666C76036BAABEFA0C" collectingDate="2000-12" collectionCode="INPA" pageId="10" pageNumber="2976" specimenCode="INPA 125" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54F156FA584202666C76036BAAEFFAEF" box="[125,237,1369,1393]" pageId="10" pageNumber="2976" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<specimenCode id="DBEC4023584202666CF2036BAB76FAEF" box="[249,372,1369,1393]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="10" pageNumber="2976">INPA 125</specimenCode>
|
||
, stained and whole-mounted specimen, fully mature, coll.
|
||
<date id="FFF4CE98584202666817036BAABEFA0C" pageId="10" pageNumber="2976" value="2000-12">
|
||
<collectingDate id="EFB03770584202666817036BAABEFA0C" pageId="10" pageNumber="2976" value="2000-12">December 2000</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B22E205584202666CC30348A880FA0C" box="[200,642,1402,1426]" collectionCode="INPA" pageId="10" pageNumber="2976" specimenCode="INPA 126" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54F156FA584202666CC30348AB3EFA0C" box="[200,316,1402,1426]" pageId="10" pageNumber="2976" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCode id="DBEC4023584202666D430348ABC1FA0C" box="[328,451,1402,1426]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="10" pageNumber="2976">INPA 126</specimenCode>
|
||
,
|
||
<specimenCount id="9D4C23D1584202666DDB0348A880FA0C" box="[464,642,1402,1426]" pageId="10" pageNumber="2976" type="generic">four specimens</specimenCount>
|
||
</materialsCitation>
|
||
, coll.
|
||
<date id="FFF4CE98584202666ECC0348A980FA0C" box="[711,898,1402,1426]" pageId="10" pageNumber="2976" value="1997-12">December 1997</date>
|
||
(
|
||
<specimenCount id="9D4C23D1584202666F990348AE4BFA0C" box="[914,1097,1402,1426]" pageId="10" pageNumber="2976" type="generic">four specimens</specimenCount>
|
||
);
|
||
<materialsCitation id="3B22E2055842026668560348ABB8FA2C" collectionCode="ZIM" pageId="10" pageNumber="2976" specimenCode="OL 14284" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="ED5B709D5842026668560348AE96FA0C" box="[1117,1172,1402,1426]" country="Slovenia" httpUri="http://grbio.org/cool/3263-rkr7" name="ZIM Culture Collection of Industrial Microorganisms" pageId="10" pageNumber="2976">ZIM</collectionCode>
|
||
<specimenCode id="DBEC4023584202666C7603A8AAF9FA2C" box="[125,251,1434,1458]" collectionCode="ZIM" country="Slovenia" httpUri="http://grbio.org/cool/3263-rkr7" name="ZIM Culture Collection of Industrial Microorganisms" pageId="10" pageNumber="2976">OL 14284</specimenCode>
|
||
,
|
||
<specimenCount id="9D4C23D1584202666D0303A8ABB8FA2C" box="[264,442,1434,1458]" pageId="10" pageNumber="2976" type="generic">four specimens</specimenCount>
|
||
</materialsCitation>
|
||
, coll.
|
||
<date id="FFF4CE98584202666E0903A8A8BCFA2C" box="[514,702,1434,1458]" pageId="10" pageNumber="2976" value="1997-12">December 1997</date>
|
||
(
|
||
<specimenCount id="9D4C23D1584202666EC503A8A977FA2C" box="[718,885,1434,1458]" pageId="10" pageNumber="2976" type="generic">one specimen</specimenCount>
|
||
),
|
||
<date id="FFF4CE98584202666F8103A8AE45FA2C" box="[906,1095,1434,1458]" pageId="10" pageNumber="2976" value="2000-12">December 2000</date>
|
||
(
|
||
<specimenCount id="9D4C23D158420266685C03A8AAF9FA4D" pageId="10" pageNumber="2976" type="generic">three specimens</specimenCount>
|
||
). All
|
||
<typeStatus id="54F156FA584202666D310389ABAAFA4D" box="[314,424,1467,1491]" pageId="10" pageNumber="2976" type="paratype">paratypes</typeStatus>
|
||
stained and whole mounted, fully mature.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584202666C760039AAF7F9BD" blockId="10.[125,245,1547,1571]" box="[125,245,1547,1571]" pageId="10" pageNumber="2976">
|
||
<emphasis id="B93E344A584202666C760039AAF7F9BD" box="[125,245,1547,1571]" italics="true" pageId="10" pageNumber="2976">Description</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584202606C76000EA833FE1D" blockId="10.[125,1172,1596,1685]" lastBlockId="12.[125,1173,137,1685]" lastPageId="12" lastPageNumber="2978" pageId="10" pageNumber="2976">
|
||
Length ca
|
||
<quantity id="4CB245BD584202666D0B000EAB43F9CA" box="[256,321,1596,1620]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="2976" unit="mm" value="5.0">5 mm</quantity>
|
||
(viv) or
|
||
<quantity id="4CB245BD584202666DBA000EA80DF9CA" box="[433,527,1596,1620]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="10" pageNumber="2976" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
(fix), diameter
|
||
<quantity id="4CB245BD584202666EDB000EA947F9CA" box="[720,837,1596,1620]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="10" pageNumber="2976" unit="mm" value="0.125">0.125 mm</quantity>
|
||
(viv) or ca
|
||
<quantity id="4CB245BD584202666FD1000EAE41F9CA" box="[986,1091,1596,1620]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="2976" unit="mm" value="0.12">0.12 mm</quantity>
|
||
, up to
|
||
<quantity id="4CB245BD584202666C76006FAAE6F9EB" box="[125,228,1629,1653]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="2976" unit="mm" value="0.15">0.15 mm</quantity>
|
||
in XII/XIII, down to
|
||
<quantity id="4CB245BD584202666DEB006FA83AF9EB" box="[480,568,1629,1653]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="2976" unit="mm" value="0.1">0.1 mm</quantity>
|
||
in terminal segments (fix). Segment number 31–34. Segments following clitellum 2–3× as long as wide when stretched, segments shortening towards posterior end. Two chaetae per bundle throughout, present laterally in XII also in mature specimens. Chaetae without nodulus, pointed distally, blunt proximally, only faintly sigmoid, distal curve rather slight, often inconspicuous (
|
||
<figureCitation id="1371F4DD584402606F6306FBA9E3FF7F" box="[872,993,201,225]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3C</figureCitation>
|
||
). Chaetae only slightly enlarged in posterior segments; ventrals
|
||
<quantity id="4CB245BD584402606EB306D8A902FE9C" box="[696,768,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="2978" unit="mm" value="28.0">28 mm</quantity>
|
||
long and
|
||
<quantity id="4CB245BD584402606F7706D8A9CEFE9C" box="[892,972,232,258]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="2978" unit="mm" value="2.5">2.5 mm</quantity>
|
||
thick in anterior segments,
|
||
<quantity id="4CB245BD584402606CF00738AB41FEBC" box="[251,323,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="12" pageNumber="2978" unit="mm" value="33.0">33 mm</quantity>
|
||
long and
|
||
<quantity id="4CB245BD584402606DCF0738ABFFFEBC" box="[452,509,264,290]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="2978" unit="mm" value="3.0">3 mm</quantity>
|
||
thick in terminal segments. Head pore (
|
||
<figureCitation id="1371F4DD584402606FE70738AE60FEBC" box="[1004,1122,266,290]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3A</figureCitation>
|
||
) on prostomium in form of a small transverse slit. Epidermal gland cells in three to four rows in preclitellar segments, more cells on dorsal half than on ventral half; in posterior segments one row per segment at chaetal level.
|
||
</paragraph>
|
||
<caption id="DF35B8D0584302676C8103EDA82CF9E9" ID-DOI="http://doi.org/10.5281/zenodo.5220048" ID-Zenodo-Dep="5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="11" pageNumber="2977" startId="11.[138,198,1503,1522]" targetBox="[142,1181,135,1462]" targetPageId="11">
|
||
<paragraph id="8BF5E858584302676C8103EDA82CF9E9" blockId="11.[138,1186,1503,1655]" pageId="11" pageNumber="2977">
|
||
Figure 3.
|
||
<taxonomicName id="4C4A93DB584302676CE103EDABBBFA6C" authority="Schmelz & Römbke, 2005" authorityName="Schmelz & Römbke" authorityYear="2005" box="[234,441,1503,1522]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="11" pageNumber="2977" phylum="Annelida" rank="species" species="tanjae" status="sp. nov.">
|
||
<emphasis id="B93E344A584302676CE103EDABBBFA6C" box="[234,441,1503,1522]" italics="true" pageId="11" pageNumber="2977">Hemienchytraeus tanjae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20D8931584302676DB403EDA80BFA6C" box="[447,521,1503,1522]" pageId="11" pageNumber="2977" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF5E858584402606C9307BEAABAFD19" blockId="12.[125,1173,137,1685]" pageId="12" pageNumber="2978">
|
||
Clitellum (
|
||
<figureCitation id="1371F4DD584402606D1107BEABB8FE3A" box="[282,442,396,420]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3D, E</figureCitation>
|
||
) girdle-shaped, i.e. fully developed ventrally, and short, i.e. extending over little more than one segment length, ending at some distance before chaetae of XIII; cellular arrangement reticulate or in indefinite rows on all sides except ventrally; hyalocytes about twice as wide (diameter ca
|
||
<quantity id="4CB245BD584402606EAD07DFA8F3FD9B" box="[678,753,491,517]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="2978" unit="mm" value="20.0">20 mm</quantity>
|
||
, viv) as granulocytes (diameter ca
|
||
<quantity id="4CB245BD584402606C76043CAACAFDB8" box="[125,200,524,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="2978" unit="mm" value="12.0">12 mm</quantity>
|
||
, viv); difference more marked in fixed specimens. Hyalocytes largest in specimens with fully developed egg (
|
||
<figureCitation id="1371F4DD584402606DA3041CA826FDD8" box="[424,548,558,582]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3D</figureCitation>
|
||
). Ventrally, in a field as wide as the distance between bursal slits, only granulocytes, arranged in dense transverse rows, clitellum less elevated here.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584402606C9304A2ABC3FB50" blockId="12.[125,1173,137,1685]" pageId="12" pageNumber="2978">
|
||
<materialsCitation id="3B22E205584402606C9304A2ABBEFB50" collectionCode="V, IV, VI" pageId="12" pageNumber="2978" specimenCount="1">
|
||
Body wall usually
|
||
<quantity id="4CB245BD584402606D7204A2ABCDFD36" box="[377,463,654,680]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="12" pageNumber="2978" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
thick, up to
|
||
<quantity id="4CB245BD584402606E7D04A2A8BCFD36" box="[630,702,654,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="2978" unit="mm" value="10.0">10 mm</quantity>
|
||
ventrally in preclitellar segments (fix), cuticle thin (,
|
||
<quantity id="4CB245BD584402606D2D0482AB66FD56" box="[294,356,686,712]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="2978" unit="mm" value="1.0">1 mm</quantity>
|
||
), preclitellar septa not thickened. Brain (
|
||
<figureCitation id="1371F4DD584402606F320482A9D2FD56" box="[825,976,688,712]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3A, B</figureCitation>
|
||
) ca twice as long as wide (e.g.
|
||
<quantity id="4CB245BD584402606D1504E3AB64FD77" box="[286,358,719,745]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="2978" unit="mm" value="65.0">65 mm</quantity>
|
||
×
|
||
<quantity id="4CB245BD584402606D8B04E3ABC9FD77" box="[384,459,719,745]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="2978" unit="mm" value="35.0">35 mm</quantity>
|
||
, fix), deeply incised anteriorly, slightly indented posteriorly, sides parallel or slightly converging anteriad. Post-pharyngeal bulbs conspicuous (
|
||
<figureCitation id="1371F4DD584402606C8E0523AAFFFCB7" box="[133,253,785,809]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3B</figureCitation>
|
||
). Ventral nerve cord perikarya concentrated in segmental ganglia from
|
||
<collectionCode id="ED5B709D58440260685A0523AE66FCB7" box="[1105,1124,785,809]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="2978" type="Museum">V</collectionCode>
|
||
on, i.e. no perikarya in the region of the septa. Oesophageal appendage (
|
||
<figureCitation id="1371F4DD584402606FEE0500AE84FCD4" box="[997,1158,818,842]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3A, B</figureCitation>
|
||
): unpaired root with moderately large proximal chamber; root and primary branches of about equal length and equal canal width. Two secondary branches on each side, short, with narrow canal lumen. Diameter of appendage decreasing from unpaired root over primary to secondary branches. Pharyngeal glands (
|
||
<figureCitation id="1371F4DD584402606E5F0581A8EEFC55" box="[596,748,947,971]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3A, B</figureCitation>
|
||
) all united dorsally in
|
||
<collectionCode id="ED5B709D584402606FFF0581AE16FC55" box="[1012,1044,947,971]" pageId="12" pageNumber="2978">IV</collectionCode>
|
||
,
|
||
<collectionCode id="ED5B709D58440260682A0581AE36FC55" box="[1057,1076,947,971]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="2978" type="Museum">V</collectionCode>
|
||
and
|
||
<collectionCode id="ED5B709D5844026068640581AE8DFC55" box="[1135,1167,947,971]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="2978">VI</collectionCode>
|
||
, connection widest in
|
||
<collectionCode id="ED5B709D584402606D7A05E6AB93FC72" box="[369,401,980,1004]" pageId="12" pageNumber="2978">IV</collectionCode>
|
||
, narrow in
|
||
<collectionCode id="ED5B709D584402606E1905E6A830FC72" box="[530,562,980,1004]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="2978">VI</collectionCode>
|
||
; primary ventral lobes in
|
||
<collectionCode id="ED5B709D584402606F5F05E6A965FC72" box="[852,871,980,1004]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="2978" type="Museum">V</collectionCode>
|
||
and
|
||
<collectionCode id="ED5B709D584402606F9405E6A9BDFC72" box="[927,959,980,1004]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="12" pageNumber="2978">VI</collectionCode>
|
||
, secondary ventral lobes in
|
||
<collectionCode id="ED5B709D584402606CED05C6AAFEFB92" box="[230,252,1012,1036]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="2978" type="Museum">V</collectionCode>
|
||
, small, attached to septum 4/5, apparently absent in
|
||
<specimenCount id="9D4C23D1584402606F8905C6AE2AFB92" box="[898,1064,1012,1036]" pageId="12" pageNumber="2978" type="generic">one specimen</specimenCount>
|
||
. Efferent fascicles (
|
||
<figureCitation id="1371F4DD584402606CE40226AB61FBB2" box="[239,355,1044,1068]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3B</figureCitation>
|
||
) not conspicuously enlarged. Chloragocytes as a thin layer from
|
||
<collectionCode id="ED5B709D5844026068720226AE8DFBB2" box="[1145,1167,1044,1068]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="2978" type="Museum">V</collectionCode>
|
||
, filled with light brown vesicles (same colour as gut contents). Dorsal blood vessel from XIV. Inflated ventral gut epithelium inconspicuous, from 1/2 XIX–XXI, only slightly higher than epithelium in adjacent anterior and posterior segments, higher than dorsal epithelium in same segments, recognizable mainly by granular texture and less intensive staining in fixed specimens
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584402606C9302E5A93EF9CA" blockId="12.[125,1173,137,1685]" pageId="12" pageNumber="2978">
|
||
Nephridia (
|
||
<figureCitation id="1371F4DD584402606D1702E5ABB0FB71" box="[284,434,1239,1263]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3A, F</figureCitation>
|
||
) in preclitellar segments: four pairs, from 5/6 to 8/9; anteseptale globular, with minute and numerous brownish granules at periphery; funnel orientated obliquely ventrad, with small and narrow anterior projection; postseptale elongate, ca twice (1.7–2.5×) as long and about as high as anteseptale; length of nephridia
|
||
<quantity id="4CB245BD584402606FD4030AAE50FACE" box="[991,1106,1335,1361]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="8.5" metricValueMin="5.0" pageId="12" pageNumber="2978" unit="mm" value="67.5" valueMax="85.0" valueMin="50.0">50–85 mm</quantity>
|
||
(fix), mostly subterminal rise of efferent duct (also medial and terminal); terminal vesicle small, inconspicuous. Postclitellar segments: first pair at 14/15, nephridia reduced in number in following segments, often unpaired, many segments without nephridia; nephridia in posterior body half larger than in preclitellar segments, longer (up to
|
||
<quantity id="4CB245BD584402606FCA0388AE19FA4C" box="[961,1051,1464,1490]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="12" pageNumber="2978" unit="mm" value="140.0">140 mm</quantity>
|
||
, viv) than body diameter, shape as in preclitellar segments, with mostly terminal rise of efferent duct. Coelomocytes small, length ca
|
||
<quantity id="4CB245BD584402606DE003C9A836F98D" box="[491,564,1529,1555]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="2978" unit="mm" value="15.0">15 mm</quantity>
|
||
(viv, fix:
|
||
<quantity id="4CB245BD584402606EAD03C9A91FF98D" box="[678,797,1529,1555]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="12" pageNumber="2978" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
|
||
), longer than wide; colour pale, margin wavy, texture with blurred vesicles (viv); cells not numerous, often aggregations postero-dorsally in V, VI and VII and in terminal segments.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584402616C93006FA97CFEDE" blockId="12.[125,1173,137,1685]" lastBlockId="13.[138,1185,135,705]" lastPageId="13" lastPageNumber="2979" pageId="12" pageNumber="2978">
|
||
Seminal vesicle absent. Few sperm on top of sperm funnel collar, spermatozoa longer than funnel (ca
|
||
<quantity id="4CB245BD584402606D3C004FAB80F90B" box="[311,386,1659,1685]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="12" pageNumber="2978" unit="mm" value="70.0">70 mm</quantity>
|
||
, fix), heads short. Sperm funnel (
|
||
<figureCitation id="1371F4DD584402606F01004FA97CF90B" box="[778,894,1661,1685]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="12" pageNumber="2978">Figure 3E</figureCitation>
|
||
) small, ca twice as long as wide, less than half as long as body diameter (e.g. 53×
|
||
<quantity id="4CB245BD584502616F5B06BBA999FF3F" box="[848,923,135,161]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="2979" unit="mm" value="22.0">22 mm</quantity>
|
||
, viv, 50×
|
||
<quantity id="4CB245BD58450261681C06BBAE60FF3F" box="[1047,1122,135,161]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="2979" unit="mm" value="19.0">19 mm</quantity>
|
||
, fix), funnel body as wide as or narrower than collar, tapering distad. Vas deferens in XII, usually coiled in small, irregular, consecutive loops ventro-laterally; diameter
|
||
<quantity id="4CB245BD584502616FD106FAAE11FF7E" box="[986,1043,198,224]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="2979" unit="mm" value="3.0">3 mm</quantity>
|
||
(fix). Male copulatory organ (
|
||
<figureCitation id="1371F4DD584502616D5606DAABD6FE9E" box="[349,468,232,256]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="13" pageNumber="2979">Figure 3E</figureCitation>
|
||
): glandular body globular, diameter
|
||
<quantity id="4CB245BD584502616F7306DAA9E8FE9E" box="[888,1002,230,256]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" metricValueMax="2.8" metricValueMin="2.4" pageId="13" pageNumber="2979" unit="mm" value="26.0" valueMax="28.0" valueMin="24.0">24–28 mm</quantity>
|
||
(fix), bursal slit longitudinal (viv), staple-shaped or broadly U-shaped in fixed specimens, shorter than diameter of glandular body. Accessory copulatory glands absent.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584502616CAE077BA843FD5F" blockId="13.[138,1185,135,705]" pageId="13" pageNumber="2979">
|
||
Spermatheca (
|
||
<figureCitation id="1371F4DD584502616D41077BABE2FEFF" box="[330,480,329,353]" captionStart="Figure 3" captionStartId="11.[138,198,1503,1522]" captionTargetBox="[142,1181,135,1462]" captionTargetId="figure-1@11.[138,1185,130,1466]" captionTargetPageId="11" captionText="Figure 3. Hemienchytraeus tanjae sp. nov. (A) Anterior body region, lateral view; (B) anterior body region, dorsal view; lateral chaetae and spermatheca to the left not shown; (C) chaetae in preclitellar (left) and in terminal (right) segment; (D) clitellum, dorsal view; (E) clitellum and male reproductive system, ventral view. Note size difference of hyalocytes between (D) and (E): (E) is drawn from slightly less mature specimen (egg not fully developed); (F) nephridia in preclitellar (left) and terminal (right) segments. All figures from stained whole mounts. Scale bars: 100 mm (A, B, D, E); 25 mm (C); 50 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.5220048" httpUri="https://zenodo.org/record/5220048/files/figure.png" pageId="13" pageNumber="2979">Figure 3A, B</figureCitation>
|
||
) inconspicuous, often difficult to see (viv, fix), extending into VI or VII, consisting of ectal duct and ampulla, the latter subdivided into ectal dilatation, connecting tube and ental reservoir. Ectal dilatation not seen in all specimens, here ectal duct continuously merging into connecting tube. Ectal duct ca one segment length long, canal and wall epithelia of about the same diameter; ectal pore
|
||
<quantity id="4CB245BD584502616F7407FBA9D6FE7F" box="[895,980,455,481]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="13" pageNumber="2979" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
|
||
wide (fix), up to
|
||
<quantity id="4CB245BD584502616C8107DBAAC1FD9F" box="[138,195,487,513]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="13" pageNumber="2979" unit="mm" value="4.0">4 mm</quantity>
|
||
in living specimens, canal lined with cuticle distally, cuticle thinning out proximad. Ectal dilatation of ampulla
|
||
<quantity id="4CB245BD584502616DC9043BA837FDBF" box="[450,565,519,545]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="13" pageNumber="2979" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
|
||
wide (fix), connecting tube with inconspicuous canal, ental reservoir thin-walled,
|
||
<quantity id="4CB245BD584502616DCF041BA835FDDF" box="[452,567,551,577]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="13" pageNumber="2979" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
|
||
wide (fix). Sperm present in ectal dilatation and ental reservoir; in ectal dilatation, sperm arranged side-by-side in a wisp with nuclei orientated distad and flagella extending straight into proximal tube; in ental reservoir, spermatozoa arranged in a dense and irregular coil. One mature egg at a time, extending over two to three segments when fully developed.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584502616C8104C8AAEBFC8C" blockId="13.[138,233,762,786]" box="[138,233,762,786]" pageId="13" pageNumber="2979">
|
||
<emphasis id="B93E344A584502616C8104C8AAEBFC8C" box="[138,233,762,786]" italics="true" pageId="13" pageNumber="2979">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584502616C810518A8C2FB1C" blockId="13.[138,1185,810,1154]" pageId="13" pageNumber="2979">
|
||
The secondary pharyngeal gland lobes are quite small, and apparently absent (i.e. not seen) in one of the
|
||
<typeStatus id="54F156FA584502616D350578ABA3FCFC" box="[318,417,842,866]" pageId="13" pageNumber="2979" type="paratype">paratype</typeStatus>
|
||
specimens. Other characters (especially details of oesophageal appendages, nephridia, and clitellum) are without exception. Two further specimens found at the type locality, one submature,
|
||
<specimenCount id="9D4C23D1584502616E2F05B8A8B0FC3C" box="[548,690,906,930]" pageId="13" pageNumber="2979" type="juvenile">one juvenile</specimenCount>
|
||
, agree with the diagnosis of
|
||
<taxonomicName id="4C4A93DB584502616FF905B9AE5BFC3C" authorityName="Schmelz & Römbke" authorityYear="2005" box="[1010,1113,906,930]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="13" pageNumber="2979" phylum="Annelida" rank="species" species="tanjae">
|
||
<emphasis id="B93E344A584502616FF905B9AE5BFC3C" box="[1010,1113,906,930]" italics="true" pageId="13" pageNumber="2979">H. tanjae</emphasis>
|
||
</taxonomicName>
|
||
in the pattern of pharyngeal glands (one pair of secondary lobes in V) and in the distribution of preclitellar nephridia (four pairs, from 5/6 to 8/9). However, the secondary pharyngeal gland lobes are large and conspicuous, terminal chaetae are almost twice as large as preclitellar chaetae and the submature specimen is larger (body diameter
|
||
<quantity id="4CB245BD584502616FF80238AE69FBBC" box="[1011,1131,1034,1058]" metricMagnitude="-4" metricUnit="m" metricValue="2.25" pageId="13" pageNumber="2979" unit="mm" value="0.225">0.225 mm</quantity>
|
||
, fix) than mature specimens of
|
||
<taxonomicName id="4C4A93DB584502616DC50219A838FBDC" authorityName="Schmelz & Römbke" authorityYear="2005" box="[462,570,1066,1090]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="13" pageNumber="2979" phylum="Annelida" rank="species" species="tanjae">
|
||
<emphasis id="B93E344A584502616DC50219A838FBDC" box="[462,570,1066,1090]" italics="true" pageId="13" pageNumber="2979">H. tanjae</emphasis>
|
||
</taxonomicName>
|
||
. In view of the clearcut differences it would seem unjustified to include these specimens into
|
||
<taxonomicName id="4C4A93DB584502616E760279A8E6FBFC" authorityName="Schmelz & Römbke" authorityYear="2005" box="[637,740,1098,1122]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="13" pageNumber="2979" phylum="Annelida" rank="species" species="tanjae">
|
||
<emphasis id="B93E344A584502616E760279A8E6FBFC" box="[637,740,1098,1122]" italics="true" pageId="13" pageNumber="2979">H. tanjae</emphasis>
|
||
</taxonomicName>
|
||
. Their identity must remain uncertain until more and mature material has been found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |