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<document id="52D90722240D315309635608B80D84B6" ID-DOI="10.11646/zootaxa.3925.4.1" ID-GBIF-Dataset="4ec28816-af9e-450f-a2b6-aa07fe6478d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="244795" ID-ZooBank="69DADAD6-84DA-4CBE-B653-6527293BA560" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461004229777" checkinUser="plazi" docAuthor="Silva, Fernando A. B., Louzada, Júlio &amp; Vaz-De-Mello, Fernando" docDate="2015" docId="03DE87D58524FF965991F9CE56B6FB11" docLanguage="en" docName="zt03925p504.pdf" docOrigin="Zootaxa 3925 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Deltochilum (Aganhyboma) schefflerorum Silva, Louzada &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="10" lastPageNumber="474" masterDocId="FFE7FFAD8532FF815906FFF9554FFFD4" masterDocTitle="A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="504" masterPageNumber="451" pageNumber="473" updateTime="1698565278120" updateUser="plazi">
<mods:mods id="F45B68042B748B4C446997A10068714F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2E585B0112EA9647ED5A6609A450FC1C">A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="569C0EEE5E10AFE1181848A90FE05A34">Silva, Fernando A. B.</mods:namePart>
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<mods:namePart id="9F8017F3CBC01A8BAA9234A176543628">Louzada, Júlio</mods:namePart>
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<mods:roleTerm id="319E2E84B2447FB589E1620E71D6D7E1">Author</mods:roleTerm>
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<mods:namePart id="E3FAD325DC4522834E58887C1A7847D2">Vaz-De-Mello, Fernando</mods:namePart>
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<mods:title id="CC9FC9730EA03A59CA42FC5871E94B9B">Zootaxa</mods:title>
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<mods:part id="EDEAEBAA010221999A60C338F035C828">
<mods:date id="3ACEC0F831C0F0FD640E57F2322FF639">2015</mods:date>
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<mods:number id="948AC34EC8825C61EBF70DA652ADC911">3925</mods:number>
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<treatment id="03DE87D58524FF965991F9CE56B6FB11" ID-DOI="http://doi.org/10.5281/zenodo.6115722" ID-GBIF-Taxon="119598952" ID-Zenodo-Dep="6115722" LSID="urn:lsid:plazi:treatment:03DE87D58524FF965991F9CE56B6FB11" httpUri="http://treatment.plazi.org/id/03DE87D58524FF965991F9CE56B6FB11" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
<subSubSection id="C36D65488524FF975991F9CE5493F9A7" pageId="22" pageNumber="473" type="nomenclature">
<paragraph id="8BC836C38524FF975991F9CE567EF986" blockId="22.[151,817,1591,1651]" box="[151,817,1591,1618]" pageId="22" pageNumber="473">
<heading id="D08081AF8524FF975991F9CE567EF986" bold="true" box="[151,817,1591,1618]" fontSize="11" level="1" pageId="22" pageNumber="473" reason="1">
<emphasis id="B903EAD18524FF975991F9CE567EF986" bold="true" box="[151,817,1591,1618]" pageId="22" pageNumber="473">
<taxonomicName id="4C774D408524FF975991F9CE57D4F985" ID-CoL="6CJ5B" box="[151,667,1591,1618]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="schefflerorum" status="sp. nov." subGenus="Aganhyboma">
<emphasis id="B903EAD18524FF975991F9CE5462F985" bold="true" box="[151,301,1591,1617]" italics="true" pageId="22" pageNumber="473">Deltochilum</emphasis>
(
<emphasis id="B903EAD18524FF97583BF9CE5491F985" bold="true" box="[317,478,1591,1617]" italics="true" pageId="22" pageNumber="473">Aganhyboma</emphasis>
)
<emphasis id="B903EAD18524FF9758F6F9CE57D4F985" bold="true" box="[496,667,1591,1617]" italics="true" pageId="22" pageNumber="473">schefflerorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23057AA8524FF975BA4F9C1567EF986" box="[674,817,1592,1618]" pageId="22" pageNumber="473" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC836C38524FF975991F9A35493F9A7" blockId="22.[151,817,1591,1651]" box="[151,476,1626,1651]" pageId="22" pageNumber="473">
(
<figureCitation id="134C2A468524FF975999F9A355BAF9A7" box="[159,245,1626,1651]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="22" pageNumber="473">Figs 12</figureCitation>
,
<figureCitation id="134C2A468524FF975807F9A2546FF9A7" box="[257,288,1627,1651]" captionStart="FIGURES 28 – 32" captionStartId="41.[151,264,1863,1886]" captionTargetBox="[163,1427,199,1832]" captionTargetId="figure@41.[151,1435,193,1843]" captionTargetPageId="41" captionText="FIGURES 28 – 32. Genital segment in Deltochilum (Aganhyboma). 28. D. finestriatum new species. 29. D. schefflerorum new species 30. D. valgum. 31. D. icaroides. 32. D. icariforme. Figures 33 – 36. Lamella copulatrix sclerites in D. (Aganhyboma). 33. D. cupreicolle. 34. D. viridescens new status and D. viridicatum new species 35. D. trisignatum. 36. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244800/files/figure.png" pageId="22" pageNumber="473">29</figureCitation>
, 54, 81, 91, 108)
</paragraph>
</subSubSection>
<subSubSection id="C36D65488524FF975991F95B545CF90B" pageId="22" pageNumber="473" type="etymology">
<paragraph id="8BC836C38524FF975991F95B545CF90B" blockId="22.[151,1437,1698,2011]" pageId="22" pageNumber="473">
<emphasis id="B903EAD18524FF975991F95B5451F96F" bold="true" box="[151,286,1698,1723]" pageId="22" pageNumber="473">Etymology.</emphasis>
Named in honour of Pamela Scheffler and Timothy Scheffler who collected a part of the
<typeStatus id="54CC88618524FF975C6AF95A50D3F96F" box="[1388,1436,1699,1723]" pageId="22" pageNumber="473">type</typeStatus>
specimens.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488524FF9659C1F9135107FD50" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473" type="description">
<paragraph id="8BC836C38524FF9759C1F91350D2F80F" blockId="22.[151,1437,1698,2011]" pageId="22" pageNumber="473">
<emphasis id="B903EAD18524FF9759C1F91354E7F8D7" bold="true" box="[199,424,1770,1795]" pageId="22" pageNumber="473">Description. Body.</emphasis>
Head, pronotum, elytra and ventral surface black or dark brown, with a silky sheen. Length 12 to 13.5 mm.
<emphasis id="B903EAD18524FF9758A8F8F754BAF8F3" bold="true" box="[430,501,1806,1831]" pageId="22" pageNumber="473">Head.</emphasis>
Punctures extremely dense and touching; around clypeus shallower, smaller and more dispersed than those on the dorsal interocular surface. Lateral margin of gena, around eyes, without row of setae or with almost inconspicuous setae; margin with central portion clearly convex. Dorsal interocular distance approximately 2.5 times eye width.
<emphasis id="B903EAD18524FF975B2EF88357C7F847" bold="true" box="[552,648,1914,1939]" pageId="22" pageNumber="473">Thorax.</emphasis>
Surface of pronotum with dense foveiform punctures, nearly touching on the sides, anterior angles and posterior portion. Discal punctures only somewhat smaller and more dispersed than those of lateral portion. Density, size and proximity of anteromedial punctures similar to those on disk.
</paragraph>
<paragraph id="8BC836C38525FF965991FF6E5107FD50" blockId="23.[151,1437,151,1221]" pageId="23" pageNumber="474">
Posteromedial punctures usually separated by less than one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles with length somewhat smaller than that of margin between anterior and lateral angles. Hypomera with foveiform punctures dense at anterior, median and posterior portions (
<figureCitation id="134C2A468525FF965885FEFD54AEFEC8" box="[387,481,260,285]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="23" pageNumber="474">Fig. 81</figureCitation>
); anteromedial punctures nearly touching. Diagonal band of hypomera from posteromedial portion to middle of lateral margin with obvious punctures (
<figureCitation id="134C2A468525FF965D12FEDE5122FE94" box="[1044,1133,295,320]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="23" pageNumber="474">Fig. 81</figureCitation>
).
<emphasis id="B903EAD18525FF965D80FEDE5195FE94" bold="true" box="[1158,1242,295,320]" pageId="23" pageNumber="474">Elytra.</emphasis>
Lateral margins slightly curved outward (
<figureCitation id="134C2A468525FF9658BAFEB5576DFEB0" box="[444,546,332,357]" captionStart="FIGURES 103 – 110" captionStartId="49.[151,264,1951,1974]" captionTargetBox="[197,1421,212,1926]" captionTargetId="figure@49.[165,1421,193,1931]" captionTargetPageId="49" captionText="FIGURES 103 – 110. Species of D. (Aganhyboma). 103. D. viridescens new status. 104. D. viridicatum new species. 105. D. viridicatum new species. 106. D. acropyge new status. 107. D. feeri new species. 108. D. schefflerorum new species. 109. D. streblopodum new species. 110. D. longiceps new status. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244808/files/figure.png" pageId="23" pageNumber="474">Fig. 108</figureCitation>
), and surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated. Interstriae flattened, with foveiform punctures surrounded by approximately four well-defined shiny microtubercles. Microtubercles with sheen more evident than surrounding surface. Foveiform punctures separated on elytral disc by one diameter or less. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length (as in Fig. 80). Apical carina or tubercle of second elytral interstria absent. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria less than the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina incomplete, effaced medially (as in Fig. 80).
</paragraph>
</subSubSection>
<subSubSection id="C36D65488525FF9659C1FD7650D2FC1C" pageId="23" pageNumber="474" type="materials_examined">
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<emphasis id="B903EAD18525FF9659C1FD7657DEFD7C" bold="true" box="[199,657,655,680]" pageId="23" pageNumber="474">
Material studied.
<typeStatus id="54CC88618525FF9658A2FD76575AFD7C" box="[420,533,655,680]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
:
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</emphasis>
:
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GROSSO: Cotriguaçu, Faz. S. Nicolau, (
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,
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),
<date id="FFC910038525FF965817FD4D54C5FD18" box="[273,394,692,716]" pageId="23" pageNumber="474" value="2009-10-12">12.X.2009</date>
, F. Vaz-de-Mello (
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).
</paragraph>
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<typeStatus id="54CC88618525FF9659C1FD2E5470FD24" box="[199,319,727,752]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
[9]:
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</emphasis>
:
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GROSSO: Alta Floresta, (
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,
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),
<date id="FFC910038525FF965D9BFD2151B3FD24" box="[1181,1276,728,752]" pageId="23" pageNumber="474" value="2008-06">VI.2008</date>
, E. Berenguer (
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<collectionCode id="ED66AE068525FF9659D4FD05546AFCC0" box="[210,293,764,788]" pageId="23" pageNumber="474">CEMT</collectionCode>
); Cotriguaçu, Faz. São Nicolau, (
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,
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),
<date id="FFC910038525FF965ACBFD055177FCC0" box="[973,1080,764,788]" pageId="23" pageNumber="474" value="2009-10-09">9.X.2009</date>
, F. Vaz-de-Mello (
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<collectionCode id="ED66AE068525FF965C3BFD0550DFFCC0" box="[1341,1424,764,788]" pageId="23" pageNumber="474">CEMT</collectionCode>
); same, (
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,
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),
<date id="FFC910038525FF965B22FCD957FFFCEC" box="[548,688,800,824]" pageId="23" pageNumber="474" value="2010-12-08">08.XII.2010</date>
, F. Vaz-de-Mello (
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1 ♀
<collectionCode id="ED66AE068525FF965D02FCD95118FCEC" box="[1028,1111,800,824]" pageId="23" pageNumber="474">CEMT</collectionCode>
); Cotriguaçu,
<date id="FFC910038525FF965DFBFCD95001FCEC" box="[1277,1358,800,824]" pageId="23" pageNumber="474" value="2011-05">V.2011</date>
, R. E. Vicente (
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);
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, Redenção, (
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,
<geoCoordinate id="EE4350048525FF965BD3FCBD5674FC89" box="[725,827,836,861]" direction="west" orientation="longitude" pageId="23" pageNumber="474" precision="925" value="-51.966667">51°58'W</geoCoordinate>
),
<date id="FFC910038525FF965A49FCBD56E1FC88" box="[847,942,836,860]" pageId="23" pageNumber="474" value="1998-11">XI.1998</date>
, P &amp; T. Scheffler (
<specimenCount id="9D71FD4A8525FF965D86FCBD51E5FC88" box="[1152,1194,836,860]" pageId="23" pageNumber="474" type="generic">2 ♂</specimenCount>
2 ♀
<collectionCode id="ED66AE068525FF965DE4FCBD507AFC88" box="[1250,1333,836,860]" pageId="23" pageNumber="474">CEMT</collectionCode>
).
</paragraph>
<paragraph id="8BC836C38525FF9659C1FC9E55F1FC70" blockId="23.[151,1437,151,1221]" pageId="23" pageNumber="474">
<emphasis id="B903EAD18525FF9659C1FC9E54E2FC54" bold="true" box="[199,429,871,896]" pageId="23" pageNumber="474">Sampling methods.</emphasis>
flight interception trap [
<specimenCount id="9D71FD4A8525FF965BC4FC915619FC54" box="[706,854,871,896]" pageId="23" pageNumber="474" type="generic">4 specimens</specimenCount>
]; at light-trap [1]; collected manually on millipedes [2].
</paragraph>
<paragraph id="8BC836C38525FF9659C1FC5650D2FC1C" blockId="23.[151,1437,151,1221]" box="[199,1437,943,968]" pageId="23" pageNumber="474">
<emphasis id="B903EAD18525FF9659C1FC565465FC1C" bold="true" box="[199,298,943,968]" pageId="23" pageNumber="474">Habitat.</emphasis>
<collectingCountry id="F36076538525FF965836FC56543BFC1C" box="[304,372,943,968]" name="Brazil" pageId="23" pageNumber="474">Brazil</collectingCountry>
(Mato Grosso and Pará) (
<figureCitation id="134C2A468525FF965B91FC5657A6FC1C" box="[663,745,943,968]" captionStart="FIGURES 89 – 92" captionStartId="46.[151,264,1748,1771]" captionTargetBox="[150,1406,193,1727]" captionTargetId="figure@46.[150,1437,193,1728]" captionTargetPageId="46" captionText="FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex." httpUri="https://zenodo.org/record/244805/files/figure.png" pageId="23" pageNumber="474">Fig. 91</figureCitation>
), from available data this species inhabits Amazon rainforest.
</paragraph>
</subSubSection>
<subSubSection id="C36D65488525FF9659C1FC2D56B6FB11" pageId="23" pageNumber="474" type="diagnosis">
<paragraph id="8BC836C38525FF9659C1FC2D56B6FB11" blockId="23.[151,1437,151,1221]" pageId="23" pageNumber="474">
<emphasis id="B903EAD18525FF9659C1FC2D54ADFC39" bold="true" box="[199,482,980,1005]" pageId="23" pageNumber="474">Diagnosis and remarks</emphasis>
(within the
<taxonomicName id="4C774D408525FF965B73FC2C5793FC38" box="[629,732,981,1004]" class="Insecta" family="Scarabaeidae" genus="Deltochilum" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="acropyge">
<emphasis id="B903EAD18525FF965B73FC2C5793FC38" box="[629,732,981,1004]" italics="true" pageId="23" pageNumber="474">acropyge</emphasis>
</taxonomicName>
complex): foveiform punctures of dorsal interocular surface touching; anteromedial punctures of pronotum separated by less than one diameter; diagonal band of hypomera with obvious punctures (
<figureCitation id="134C2A468525FF9658BFFBE5575EFBE0" box="[441,529,1052,1077]" captionStart="FIGURES 81 – 83" captionStartId="45.[151,264,1841,1864]" captionTargetBox="[156,1435,197,1820]" captionTargetId="figure@45.[151,1435,193,1821]" captionTargetPageId="45" captionText="FIGURES 81 – 83. Illustrations of hypomera in D. (Aganhyboma). 81. D. schefflerorum new species. 82. D. arturoi new species. 83. D. longiceps new status. Figure 84. Right elytron of D. longiceps new status (arrows indicate the irregular spacing among the points of the elytral striae). Figure 85. Pronotum in D. icaroides. Figure 86. Apex of third interestria in D. subrubrum new species. Figures 87 – 88. Detail of the apical tubercle of third interstria in D. (Aganhyboma). 87. D. icaroides. 88. D. valgum." httpUri="https://zenodo.org/record/244804/files/figure.png" pageId="23" pageNumber="474">Fig. 81</figureCitation>
); apical carina or tubercle of second elytral interstria absent; pseudepipleural carina incomplete, effaced medially (as in Fig. 80). The following aedeagus characters are also diagnostic: parameres symmetric and elongate, in lateral view forming an angle of approximately 45° with phallobase; apex acuminate (
<figureCitation id="134C2A468525FF96581CFB7E5421FB74" box="[282,366,1159,1184]" captionStart="FIGURES 12 – 19" captionStartId="39.[151,264,1845,1868]" captionTargetBox="[172,1432,201,1788]" captionTargetId="figure@39.[151,1435,193,1793]" captionTargetPageId="39" captionText="FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244798/files/figure.png" pageId="23" pageNumber="474">Fig. 12</figureCitation>
). Genital segment with superior margin weakly sclerotized (
<figureCitation id="134C2A468525FF965D1EFB7E5122FB74" box="[1048,1133,1159,1184]" captionStart="FIGURES 28 – 32" captionStartId="41.[151,264,1863,1886]" captionTargetBox="[163,1427,199,1832]" captionTargetId="figure@41.[151,1435,193,1843]" captionTargetPageId="41" captionText="FIGURES 28 – 32. Genital segment in Deltochilum (Aganhyboma). 28. D. finestriatum new species. 29. D. schefflerorum new species 30. D. valgum. 31. D. icaroides. 32. D. icariforme. Figures 33 – 36. Lamella copulatrix sclerites in D. (Aganhyboma). 33. D. cupreicolle. 34. D. viridescens new status and D. viridicatum new species 35. D. trisignatum. 36. D. finestriatum new species. Figures scale = 1 mm." httpUri="https://zenodo.org/record/244800/files/figure.png" pageId="23" pageNumber="474">Fig. 29</figureCitation>
). Lamella copulatrix (LC): subrectangular sclerites. Fronto-lateral peripheral sclerite (FLP) as in Fig. 54.
</paragraph>
</subSubSection>
</treatment>
</document>
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