154 lines
17 KiB
XML
154 lines
17 KiB
XML
<document id="23AF745AE6866B702705A2A0E11C2769" ID="10.11646/zootaxa.4258.1.1SLASH10897" ID-CLB-Dataset="33167" ID-DOI="10.11646/zootaxa.4258.1.1" ID-GBIF-Dataset="fdbe32e3-f5b6-4537-b870-6d295865da73" ID-ISSN="1175-5326" ID-Zenodo-Dep="569114" ID-ZooBank="B1F45C56-A43A-4A8C-9190-2B861A37D33C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493194184337" checkinUser="plazi" docAuthor="Judson, Mark L. I." docDate="2017" docId="03D65C0B65683357FF333E8C66D1FCAD" docLanguage="en" docName="zootaxa.4258.1.1SLASH10897.pdf" docOrigin="Zootaxa 4258 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybella" docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="FFEF24736573334CFFA43E146234FFEF" masterDocTitle="A new subfamily of Feaellidae (Arachnida, Chelonethi, Feaelloidea) from Southeast Asia" masterLastPageNumber="33" masterPageNumber="1" pageNumber="28" updateTime="1698439057093" updateUser="ExternalLinkService">
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<mods:titleInfo id="756A1F2B8F723DFF735AE2FEB82F1E17">
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<mods:title id="A69A5E3EA47C5C4BC7C65E0EBD8D4AAE">A new subfamily of Feaellidae (Arachnida, Chelonethi, Feaelloidea) from Southeast Asia</mods:title>
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<mods:name id="E98188B64366B67589685FBE34CEB51E" type="personal">
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<mods:roleTerm id="9CEFEDC580AA873217B05A4A1BFD5C94">Author</mods:roleTerm>
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<mods:namePart id="345709395CD3C4E7A507CF162C348C46">Judson, Mark L. I.</mods:namePart>
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<mods:typeOfResource id="F76E7B4F84274F15CB3D672F03CA3ADE">text</mods:typeOfResource>
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<mods:titleInfo id="6806B941124C62635F561760D1FF25A0">
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<mods:title id="E3FDFAA495AEAFF0A1AF30BD51D669EE">Zootaxa</mods:title>
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<mods:part id="9B988101738DE88E4377B46B84B3E8B3">
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<mods:date id="4D66E5B481501F676CCBE5CB5AF481AE">2017</mods:date>
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<mods:detail id="B673D4A2D5882BC6F08DB49F85C2883F" type="volume">
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<mods:number id="EF02AF3C0B4DAD1B0A0626D991C25526">4258</mods:number>
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<mods:number id="FD3A46CA9B29BBB0EA74E3DF954C6408">1</mods:number>
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<mods:identifier id="9E1B84B96CFC4C26121EF73908EF603B" type="DOI">10.11646/zootaxa.4258.1.1</mods:identifier>
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<treatment id="03D65C0B65683357FF333E8C66D1FCAD" ID-DOI="http://doi.org/10.5281/zenodo.6023809" ID-GBIF-Taxon="128699668" ID-Zenodo-Dep="6023809" LSID="urn:lsid:plazi:treatment:03D65C0B65683357FF333E8C66D1FCAD" httpUri="http://treatment.plazi.org/id/03D65C0B65683357FF333E8C66D1FCAD" lastPageNumber="28" pageId="27" pageNumber="28">
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<subSubSection id="C365BE9665683357FF333E8C63CCFF5D" box="[151,504,151,178]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BC0ED1D65683357FF333E8C63CCFF5D" blockId="27.[151,504,151,178]" box="[151,504,151,178]" pageId="27" pageNumber="28">
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<heading id="D0885A7165683357FF333E8C63CCFF5D" bold="true" box="[151,504,151,178]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
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<emphasis id="B90B310F65683357FF333E8C63CCFF5D" bold="true" box="[151,504,151,178]" pageId="27" pageNumber="28">
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Ontogeny of
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<taxonomicName id="4C7F969E65683357FE9C3E8363A7FF5E" ID-CoL="3XSK" box="[312,403,151,177]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90B310F65683357FE9C3E8363A7FF5E" bold="true" box="[312,403,151,177]" italics="true" pageId="27" pageNumber="28">Cybella</emphasis>
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</taxonomicName>
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species.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C365BE9665683357FF333ECA66D1FCAD" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BC0ED1D65683357FF333ECA605BFDD4" blockId="27.[151,1436,222,834]" pageId="27" pageNumber="28">
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Although only the tritonymph of
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<taxonomicName id="4C7F969E65683357FDAF3ECB609EFF18" box="[523,682,222,247]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
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<emphasis id="B90B310F65683357FDAF3ECB609EFF18" box="[523,682,222,247]" italics="true" pageId="27" pageNumber="28">C. deharvengi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FD143ECA60DBFF18" box="[688,751,222,247]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FD143ECA60DBFF18" bold="true" box="[688,751,222,247]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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and the protonymph of
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<taxonomicName id="4C7F969E65683357FC583ECB664DFF18" box="[1020,1145,222,247]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
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<emphasis id="B90B310F65683357FC583ECB664DFF18" box="[1020,1145,222,247]" italics="true" pageId="27" pageNumber="28">C. bedosae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FB243ECA668BFF18" box="[1152,1215,222,247]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FB243ECA668BFF18" bold="true" box="[1152,1215,222,247]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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are known, the two species are so similar that these provide a general idea of the ontogeny of
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<taxonomicName id="4C7F969E65683357FBA33F16666AFEF4" box="[1031,1118,258,283]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90B310F65683357FBA33F16666AFEF4" box="[1031,1118,258,283]" italics="true" pageId="27" pageNumber="28">Cybella</emphasis>
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</taxonomicName>
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. An unusual feature of the growth of
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<taxonomicName id="4C7F969E65683357FEB43F336385FED0" box="[272,433,294,319]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
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<emphasis id="B90B310F65683357FEB43F336385FED0" box="[272,433,294,319]" italics="true" pageId="27" pageNumber="28">C. deharvengi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FE1F3F3263C8FED0" box="[443,508,294,319]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FE1F3F3263C8FED0" bold="true" box="[443,508,294,319]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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(and, presumably,
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<taxonomicName id="4C7F969E65683357FD7D3F33616CFED0" box="[729,856,294,319]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
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<emphasis id="B90B310F65683357FD7D3F33616CFED0" box="[729,856,294,319]" italics="true" pageId="27" pageNumber="28">C. bedosae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FCC63F326197FED0" box="[866,931,294,319]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FCC63F326197FED0" bold="true" box="[866,931,294,319]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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) is the small difference in size between the tritonymph and adult. This might be related to the small size of
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<taxonomicName id="4C7F969E65683357FCF93F5E6180FE8C" box="[861,948,330,355]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="21" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90B310F65683357FCF93F5E6180FE8C" box="[861,948,330,355]" italics="true" pageId="27" pageNumber="28">Cybella</emphasis>
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</taxonomicName>
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species compared to most other
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<taxonomicName id="4C7F969E65683357FA863F5E67ACFE8C" box="[1314,1432,330,355]" class="Arachnida" family="Feaellidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Feaellidae</taxonomicName>
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, although in
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<taxonomicName id="4C7F969E65683357FE873F7A63FAFE69" box="[291,462,366,391]" class="Arachnida" family="Feaellidae" genus="Feaella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="perreti">
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<emphasis id="B90B310F65683357FE873F7A63FAFE69" box="[291,462,366,391]" italics="true" pageId="27" pageNumber="28">Feaella perreti</emphasis>
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</taxonomicName>
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—which is even smaller than
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<taxonomicName id="4C7F969E65683357FC8F3F7B61F9FE68" box="[811,973,366,391]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
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<emphasis id="B90B310F65683357FC8F3F7B61F9FE68" box="[811,973,366,391]" italics="true" pageId="27" pageNumber="28">C. deharvengi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FC733F7A662DFE68" box="[983,1049,366,391]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FC733F7A662DFE68" bold="true" box="[983,1049,366,391]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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—adults are distinctly larger than tritonymphs (
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<bibRefCitation id="EFEE90EC65683357FE893F8663EBFE44" author="Mahnert" box="[301,479,402,427]" pageId="27" pageNumber="28" refString="Mahnert, V. (1982 a) Die Pseudoskorpione (Arachnida) Kenyas II. Feaellidae; Cheiridiidae. Revue Suisse de Zoologie, 89, 115 - 134." type="journal article" year="1982" yearSuffix="a">Mahnert 1982a</bibRefCitation>
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). Because males are generally smaller than females in
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<taxonomicName id="4C7F969E65683357FBE83F8666F6FE44" box="[1100,1218,402,427]" class="Arachnida" family="Feaellidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Feaellidae</taxonomicName>
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, it is likely that the difference in size between the tritonymphs and females will be more marked when the latter become known in Cybellinae. If the non-type tritonymph of
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<taxonomicName id="4C7F969E65683357FD383FCF6176FE1C" box="[668,834,474,499]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
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<emphasis id="B90B310F65683357FD383FCF6176FE1C" box="[668,834,474,499]" italics="true" pageId="27" pageNumber="28">C. deharvengi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FCF53FCE61ACFE1C" box="[849,920,474,499]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FCF53FCE61ACFE1C" bold="true" box="[849,920,474,499]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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recorded here is correctly identified, it is conceivable that there might be a dimorphism in the size of tritonymphs, with the paratype tritonymph being a female and the non-type specimen a male.
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</paragraph>
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<paragraph id="8BC0ED1D65683357FF633C526619FD3C" blockId="27.[151,1436,222,834]" pageId="27" pageNumber="28">
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The presence of only one proximal sensillum (
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<emphasis id="B90B310F65683357FD6A3C5360E8FDB1" box="[718,732,583,606]" italics="true" pageId="27" pageNumber="28">p</emphasis>
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1) in the protonymph of
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<taxonomicName id="4C7F969E65683357FC433C536650FDB0" box="[999,1124,582,607]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
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<emphasis id="B90B310F65683357FC433C536650FDB0" box="[999,1124,582,607]" italics="true" pageId="27" pageNumber="28">C. bedosae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2388C7465683357FBCE3C52669CFDB0" box="[1130,1192,582,607]" pageId="27" pageNumber="28" rank="species">
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<emphasis id="B90B310F65683357FBCE3C52669CFDB0" bold="true" box="[1130,1192,582,607]" pageId="27" pageNumber="28">n. sp.</emphasis>
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</taxonomicNameLabel>
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follows a pattern that seems to be typical of all pseudoscorpions. Sensillum (
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<emphasis id="B90B310F65683357FC8F3C7A610DFD6A" box="[811,825,622,645]" italics="true" pageId="27" pageNumber="28">p</emphasis>
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2) usually appears in the deutonymph, regardless of whether it is coupled with
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<emphasis id="B90B310F65683357FE723C8163D0FD43" box="[470,484,661,684]" italics="true" pageId="27" pageNumber="28">p</emphasis>
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1 or separate (
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<bibRefCitation id="EFEE90EC65683357FD2C3C806111FD42" author="Gardini" box="[648,805,660,685]" pageId="27" pageNumber="28" refString="Gardini, G. (1991) Tre nuove specie di Roncus L. Koch, 1873 della Liguria occidentale (Pseudoscorpionida Neobisiidae). Memorie della Societ entomologica Italiana, 70, 313 - 334." type="journal article" year="1991">Gardini 1991</bibRefCitation>
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,
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<bibRefCitation id="EFEE90EC65683357FC913C816144FD42" author="Gardini" box="[821,880,661,685]" pageId="27" pageNumber="28" refString="Gardini, G. (1993) Roncus caprai n. sp. della Liguria orientale e note su R. gestroi Beier, 1930 (Pseudoscorpionida, Neobisiidae). Memorie della Societ entomologica Italiana, 71, 409 - 425." type="journal article" year="1993">1993</bibRefCitation>
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; pers. obs.). Although not explained by
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<bibRefCitation id="EFEE90EC65683357FAEA3C8062C4FD3C" author="Judson" pageId="27" pageNumber="28" refString="Judson, M. L. I. (2007 a) A new and endangered pseudoscorpion of the genus Lagynochthonius (Arachnida, Chelonethi, Chthoniidae) from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini. Zootaxa, 1627, 53 - 68." type="journal article" year="2007" yearSuffix="a">Judson (2007a)</bibRefCitation>
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, the notations
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<emphasis id="B90B310F65683357FE373CA86395FD3C" box="[403,417,700,723]" italics="true" pageId="27" pageNumber="28">p</emphasis>
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1 and
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<emphasis id="B90B310F65683357FE7B3CA863D9FD3C" box="[479,493,700,723]" italics="true" pageId="27" pageNumber="28">p</emphasis>
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2 were selected to reflect this ontogenetic sequence.
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</paragraph>
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<paragraph id="8BC0ED1D65683357FF633CF566D1FCAD" blockId="27.[151,1436,222,834]" pageId="27" pageNumber="28">
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The trichobothria are evidently added in the same sequence and positions as those of Feaellinae, even if deutonymphs were not available. Thus Cybellinae do not provide any additional information about the remarkable rearrangements in position that characterize
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<taxonomicName id="4C7F969E65683357FD2C3D3D60CFFCAD" box="[648,763,809,834]" class="Arachnida" family="Feaellidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="family">Feaellidae</taxonomicName>
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as a whole (
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<bibRefCitation id="EFEE90EC65683357FC223D3D660CFCAD" author="Mahnert" box="[902,1080,809,834]" pageId="27" pageNumber="28" refString="Mahnert, V. (1982 b) Sigles trichobothriaux chez les pseudoscorpions (Arachnida). Atti della Societ Toscana di Scienze naturali, Memorie (B), 88 (Supplement), 185 - 192. [1981]" type="journal article" year="1982" yearSuffix="b">Mahnert 1982b</bibRefCitation>
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;
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<bibRefCitation id="EFEE90EC65683357FBE73D3E66ECFCAD" author="Harvey" box="[1091,1240,810,834]" pageId="27" pageNumber="28" refString="Harvey, M. S. (1992) The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy, 6, 1373 - 1435." type="journal article" year="1992">Harvey 1992</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |