203 lines
28 KiB
XML
203 lines
28 KiB
XML
<document id="D84F5E6BAA238E1B0F88C2901E3FC31D" ID-CLB-Dataset="26407" ID-DOI="10.11646/zootaxa.4623.2.8" ID-GBIF-Dataset="db2b3bd4-2e27-4918-8fff-226b303c65b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3255499" ID-ZooBank="EEDC9982-ED9C-40BD-8584-CDADA572DDE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561449672174" checkinUser="plazi" docAuthor="Bocek, Matej & Adamkova, Kristyna" docDate="2019" docId="03AF335EC3606350FF06FA29FA44FD6C" docLanguage="en" docName="zootaxa.4623.2.8.pdf" docOrigin="Zootaxa 4623 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eniclases moluccanus Kleine 1930" docType="treatment" docVersion="9" lastPageNumber="346" masterDocId="FF964B26C3646355FF91FF95FFD3FFEB" masterDocTitle="New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae)" masterLastPageNumber="350" masterPageNumber="341" pageNumber="345" updateTime="1698736235721" updateUser="ExternalLinkService">
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<mods:title id="B932966FA2BBBB92B0FF5D702D5D6B73">New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae)</mods:title>
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<mods:namePart id="AA0DAC93E0AD28FF3E7E18694213A895">Bocek, Matej</mods:namePart>
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<mods:namePart id="AD00A7AC3ED77203CCBA1EBC4E970F17">Adamkova, Kristyna</mods:namePart>
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<mods:title id="84EFDE7D9BE877EC7C5BE7D029EE9739">Zootaxa</mods:title>
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<mods:part id="067EC09B9BCD965906CDF490486227C7">
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<mods:date id="6C40576545C2952CF002F6325EF824C3">2019</mods:date>
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<mods:number id="055F010040A57A496FB49B458CFA3444">2019-06-25</mods:number>
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<mods:number id="30B65132A6B6696A3C2D0D1720485ED9">4623</mods:number>
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<treatment id="03AF335EC3606350FF06FA29FA44FD6C" ID-GBIF-Taxon="158380874" LSID="urn:lsid:plazi:treatment:03AF335EC3606350FF06FA29FA44FD6C" httpUri="http://treatment.plazi.org/id/03AF335EC3606350FF06FA29FA44FD6C" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345">
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<subSubSection id="C31CD1C3C3606351FF06FA29FD57FA3D" box="[151,644,1468,1495]" pageId="4" pageNumber="345" type="nomenclature">
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<paragraph id="8BB98248C3606351FF06FA29FD57FA3D" blockId="4.[151,644,1468,1531]" box="[151,644,1468,1495]" pageId="4" pageNumber="345">
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<heading id="D0F13524C3606351FF06FA29FD57FA3D" bold="true" box="[151,644,1468,1495]" fontSize="11" level="1" pageId="4" pageNumber="345" reason="1">
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<treatmentCitationGroup id="AB16A566C3606351FF06FA29FD57FA3D" box="[151,644,1468,1495]" pageId="4" pageNumber="345">
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<taxonomicName id="4C06F9CBC3606351FF06FA29FD57FA3D" authority="Kleine, 1930: 328" authorityName="Kleine" authorityPageNumber="328" authorityYear="1930" box="[151,644,1468,1495]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="moluccanus">
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<emphasis id="B9725E5AC3606351FF06FA29FD57FA3D" bold="true" box="[151,644,1468,1495]" pageId="4" pageNumber="345">
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<emphasis id="B9725E5AC3606351FF06FA29FE76FA3C" bold="true" box="[151,421,1468,1495]" italics="true" pageId="4" pageNumber="345">Eniclases moluccanus</emphasis>
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<treatmentCitation id="0AA7A459C3606351FE3DFA29FD57FA3D" author="Kleine, R." box="[428,644,1468,1494]" page="328" pageId="4" pageNumber="345" year="1930">
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<bibRefCitation id="EF97FFB9C3606351FE3DFA29FD57FA3D" author="Kleine, R." box="[428,644,1468,1494]" pageId="4" pageNumber="345" pagination="325 - 340" refId="ref5974" refString="Kleine, R. (1930) Bestimmungstabelle der Trichalusverwandschaft. Treubia, 9, 325 - 340." type="journal article" year="1930">Kleine, 1930: 328</bibRefCitation>
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</treatmentCitation>
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</emphasis>
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</taxonomicName>
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</treatmentCitationGroup>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3606351FF06FA75FE9FFA11" box="[151,332,1504,1531]" pageId="4" pageNumber="345" type="description">
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<paragraph id="8BB98248C3606351FF06FA75FE9FFA11" blockId="4.[151,644,1468,1531]" box="[151,332,1504,1531]" pageId="4" pageNumber="345">
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<figureCitation id="133D9ECDC3606351FF06FA75FE9FFA11" box="[151,332,1504,1531]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="4" pageNumber="345">Figs 5, 8, 12, 16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3606351FF06F9BDFD1FF98C" pageId="4" pageNumber="345" type="materials_examined">
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<paragraph id="8BB98248C3606351FF06F9BDFD1FF98C" blockId="4.[151,1437,1576,2035]" pageId="4" pageNumber="345">
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<emphasis id="B9725E5AC3606351FF06F9BDFE52F9A8" bold="true" box="[151,385,1576,1603]" pageId="4" pageNumber="345">Material examined.</emphasis>
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Male,
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<specimenCount id="9D0049C1C3606351FE5EF9BCFDE8F9A9" box="[463,571,1576,1603]" pageId="4" pageNumber="345" type="female">4 females</specimenCount>
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(A02451,
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<accessionNumber id="94551FABC3606351FD23F9BCFCDCF9A8" box="[690,783,1577,1603]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/A02437" pageId="4" pageNumber="345" type="EnaNcbi">A02437</accessionNumber>
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, A02452–54,
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<figureCitation id="133D9ECDC3606351FC3AF9BDFC3DF9A8" box="[939,1006,1576,1603]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="4" pageNumber="345">Fig. 5</figureCitation>
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),
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<collectingCountry id="F311C2D8C3606351FB95F9BDFBA7F9A9" box="[1028,1140,1576,1602]" name="Indonesia" pageId="4" pageNumber="345">Indonesia</collectingCountry>
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, Halmahera, road Weda to Tobaru,
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<quantity id="4CFE2FADC3606351FF65F9D8FEEEF98C" box="[244,317,1613,1639]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="4" pageNumber="345" unit="m" value="190.0">190 m</quantity>
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,
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<geoCoordinate id="EE32E48FC3606351FED8F9D9FE12F98C" box="[329,449,1612,1639]" degrees="0" direction="north" minutes="19.58" orientation="latitude" pageId="4" pageNumber="345" precision="9" value="0.32633334">0°19.58’N</geoCoordinate>
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,
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<geoCoordinate id="EE32E48FC3606351FE5CF9D9FD88F98D" box="[461,603,1612,1638]" degrees="127" direction="east" minutes="49.68" orientation="longitude" pageId="4" pageNumber="345" precision="9" value="127.828">127°49.68’E</geoCoordinate>
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(
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<collectionCode id="ED171A8DC3606351FDFBF9D8FD13F98C" box="[618,704,1613,1639]" pageId="4" pageNumber="345">LMBC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3606351FF56F9E5FDAEF888" pageId="4" pageNumber="345" type="diagnosis">
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<paragraph id="8BB98248C3606351FF56F9E5FDAEF888" blockId="4.[151,1437,1576,2035]" pageId="4" pageNumber="345">
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<emphasis id="B9725E5AC3606351FF56F9E5FE91F960" bold="true" box="[199,322,1648,1675]" pageId="4" pageNumber="345">Diagnosis.</emphasis>
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<taxonomicName id="4C06F9CBC3606351FED8F9E5FD92F961" authorityName="Kleine" authorityYear="1930" box="[329,577,1648,1674]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="moluccanus">
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<emphasis id="B9725E5AC3606351FED8F9E5FD92F961" box="[329,577,1648,1674]" italics="true" pageId="4" pageNumber="345">Eniclases moluccanus</emphasis>
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</taxonomicName>
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shares similar color pattern with
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<taxonomicName id="4C06F9CBC3606351FC28F9E4FAF9F961" authority="Bocek & Bocak, 2016" authorityName="Bocek & Bocak" authorityYear="2016" box="[953,1322,1648,1675]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="bicolor">
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<emphasis id="B9725E5AC3606351FC28F9E4FBF9F961" box="[953,1066,1648,1674]" italics="true" pageId="4" pageNumber="345">E. bicolor</emphasis>
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<bibRefCitation id="EF97FFB9C3606351FBA0F9E5FAF9F961" author="Bocek, M. & Bocak, L." box="[1073,1322,1648,1675]" pageId="4" pageNumber="345" pagination="15 - 35" refId="ref5333" refString="Bocek, M. & Bocak, L. (2016) Species limits in polymorphic mimetic Eniclases net-winged beetles from New Guinean mountains (Coleoptera, Lycidae). ZooKeys, 593, 15 - 35. https: // doi. org / 10.3897 / zookeys. 593.7728" type="journal article" year="2016">Bocek & Bocak, 2016</bibRefCitation>
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</taxonomicName>
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and some specimens of
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<taxonomicName id="4C06F9CBC3606351FEA0F900FD6EF945" authority="Bocek & Bocak, 2016" authorityName="Bocek & Bocak" authorityYear="2016" box="[305,701,1684,1711]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="variabilis">
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<emphasis id="B9725E5AC3606351FEA0F900FE68F945" box="[305,443,1684,1710]" italics="true" pageId="4" pageNumber="345">E. variabilis</emphasis>
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<bibRefCitation id="EF97FFB9C3606351FE53F901FD6EF945" author="Bocek, M. & Bocak, L." box="[450,701,1684,1711]" pageId="4" pageNumber="345" pagination="15 - 35" refId="ref5333" refString="Bocek, M. & Bocak, L. (2016) Species limits in polymorphic mimetic Eniclases net-winged beetles from New Guinean mountains (Coleoptera, Lycidae). ZooKeys, 593, 15 - 35. https: // doi. org / 10.3897 / zookeys. 593.7728" type="journal article" year="2016">Bocek & Bocak, 2016</bibRefCitation>
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</taxonomicName>
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, however, these species are unrelated and distributed only in New
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<collectingCountry id="F311C2D8C3606351FF06F92DFF3BF939" box="[151,232,1720,1746]" name="Guinea-Bissau" pageId="4" pageNumber="345">Guinea</collectingCountry>
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(
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<figureCitation id="133D9ECDC3606351FF66F92DFEE9F938" box="[247,314,1720,1747]" captionStart="FIGURE 1" captionStartId="2.[151,250,1336,1361]" captionTargetBox="[156,1429,352,1316]" captionTargetId="figure@2.[151,1436,345,1317]" captionTargetPageId="2" captionText="FIGURE 1. Molecular phylogeny of Eniclases and Schizotrichalus based on the maximum likelihood analysis of cox1 mtDNA fragment. Branch values indicate ultrafast bootstrap support (UFboot)." httpUri="https://zenodo.org/record/3255501/files/figure.png" pageId="4" pageNumber="345">Fig. 1</figureCitation>
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,
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<bibRefCitation id="EF97FFB9C3606351FED9F92DFDECF938" author="Bocek, M. & Bocak, L." box="[328,575,1720,1747]" pageId="4" pageNumber="345" pagination="15 - 35" refId="ref5333" refString="Bocek, M. & Bocak, L. (2016) Species limits in polymorphic mimetic Eniclases net-winged beetles from New Guinean mountains (Coleoptera, Lycidae). ZooKeys, 593, 15 - 35. https: // doi. org / 10.3897 / zookeys. 593.7728" type="journal article" year="2016">Bocek & Bocak 2016</bibRefCitation>
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).
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<taxonomicName id="4C06F9CBC3606351FDC0F92CFD0FF939" authorityName="Bocek & Bocak" authorityYear="2016" box="[593,732,1720,1746]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="variabilis">
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<emphasis id="B9725E5AC3606351FDC0F92CFD0FF939" box="[593,732,1720,1746]" italics="true" pageId="4" pageNumber="345">E. variabilis</emphasis>
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</taxonomicName>
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has smaller eyes (Ediam/Edist 0.83–0.95). Both New Guinean species have completely dark colored abdomen unlike
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<taxonomicName id="4C06F9CBC3606351FC9FF948FC66F91D" authorityName="Kleine" authorityYear="1930" box="[782,949,1756,1782]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="moluccanus">
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<emphasis id="B9725E5AC3606351FC9FF948FC66F91D" box="[782,949,1756,1782]" italics="true" pageId="4" pageNumber="345">E. moluccanus</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="4C06F9CBC3606351FC54F948FBBEF91D" authorityName="Kleine" authorityYear="1930" box="[965,1133,1756,1782]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="moluccanus">
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<emphasis id="B9725E5AC3606351FC54F948FBBEF91D" box="[965,1133,1756,1782]" italics="true" pageId="4" pageNumber="345">E. moluccanus</emphasis>
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</taxonomicName>
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occurs sympatrically with
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<taxonomicName id="4C06F9CBC3606351FF06F895FE3DF8F1" authorityName="Bocek & Adamkova" authorityYear="2019" box="[151,494,1792,1818]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="halmaherensis">
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<emphasis id="B9725E5AC3606351FF06F895FE3DF8F1" box="[151,494,1792,1818]" italics="true" pageId="4" pageNumber="345">Schizotrichalus halmaherensis</emphasis>
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</taxonomicName>
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and both species belong to the same mimicry ring (
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<figureCitation id="133D9ECDC3606351FBABF895FB6FF8F0" box="[1082,1212,1792,1819]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="4" pageNumber="345">Figs 2–3, 5</figureCitation>
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||
). These species can be easily recognized by the different shape of male antennae, arrangement of pronotal carinae and by unique shape of male genitalia (
|
||
<figureCitation id="133D9ECDC3606351FEF3F8DDFDA3F888" box="[354,624,1864,1891]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="4" pageNumber="345">Figs 6–8, 10, 12–14, 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3606350FF56F8F9FDD2FF10" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345" type="description">
|
||
<paragraph id="8BB98248C3606350FF56F8F9FDD2FF10" blockId="4.[151,1437,1576,2035]" lastBlockId="5.[151,1437,153,647]" lastPageId="5" lastPageNumber="346" pageId="4" pageNumber="345">
|
||
<emphasis id="B9725E5AC3606351FF56F8F9FEA6F86C" bold="true" box="[199,373,1900,1927]" pageId="4" pageNumber="345">Redescription.</emphasis>
|
||
Male. Body length
|
||
<quantity id="4CFE2FADC3606351FDC9F8F8FD31F86C" box="[600,738,1901,1927]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="8.9" metricValueMin="7.8" pageId="4" pageNumber="345" unit="mm" value="8.35" valueMax="8.9" valueMin="7.8">7.8–8.9 mm</quantity>
|
||
, head and antennae black, almost three fifths of elytra yellow to orange, apical two fifths black, apical part of abdomen yellow. Legs uniformly black, only basal part of femora orange. Head small with prominent eyes, eye diameter 1.05 times interocular distance, antennomeres 3–10 slightly serrate, flat, slender, similar in shape, compressed. Pronotum transverse, its length 0.66 times width at base, basal margin with slightly projected posterior angles. Elytra with robust secondary costae and regular dense cells. Male genitalia with well-sclerotized phallus, dorsal keel conspicuous, phallobase with extensive membranous sac, apical part of phallus slender (
|
||
<figureCitation id="133D9ECDC3616350FE31FF74FE20FF10" box="[416,499,225,251]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="5" pageNumber="346">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3616350FF56FE91FD04FEF4" box="[199,727,260,287]" pageId="5" pageNumber="346" type="distribution">
|
||
<paragraph id="8BB98248C3616350FF56FE91FD04FEF4" blockId="5.[151,1437,153,647]" box="[199,727,260,287]" pageId="5" pageNumber="346">
|
||
<emphasis id="B9725E5AC3616350FF56FE91FDD1FEF4" bold="true" box="[199,514,260,287]" pageId="5" pageNumber="346">Distribution and diversity.</emphasis>
|
||
Halmahera Island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3616350FF56FEBCFDD6FE8C" pageId="5" pageNumber="346" type="description">
|
||
<paragraph id="8BB98248C3616350FF56FEBCFDD6FE8C" blockId="5.[151,1437,153,647]" pageId="5" pageNumber="346">
|
||
<emphasis id="B9725E5AC3616350FF56FEBCFEADFEA8" bold="true" box="[199,382,297,323]" pageId="5" pageNumber="346">Measurements.</emphasis>
|
||
BL
|
||
<quantity id="4CFE2FADC3616350FE3EFEBCFDEBFEA8" box="[431,568,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="8.35" metricValueMax="8.9" metricValueMin="7.8" pageId="5" pageNumber="346" unit="mm" value="8.35" valueMax="8.9" valueMin="7.8">7.8–8.9 mm</quantity>
|
||
, EL
|
||
<quantity id="4CFE2FADC3616350FDFCFEBCFD25FEA8" box="[621,758,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="6.95" metricValueMax="7.2" metricValueMin="6.7" pageId="5" pageNumber="346" unit="mm" value="6.95" valueMax="7.2" valueMin="6.7">6.7–7.2 mm</quantity>
|
||
, WH
|
||
<quantity id="4CFE2FADC3616350FCA9FEBDFC0EFEA8" box="[824,989,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.295" metricValueMax="2.34" metricValueMin="2.25" pageId="5" pageNumber="346" unit="mm" value="2.295" valueMax="2.34" valueMin="2.25">2.25–2.34 mm</quantity>
|
||
, PL
|
||
<quantity id="4CFE2FADC3616350FB81FEBCFB66FEA8" box="[1040,1205,297,323]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" metricValueMax="1.37" metricValueMin="1.31" pageId="5" pageNumber="346" unit="mm" value="1.34" valueMax="1.37" valueMin="1.31">1.31–1.37 mm</quantity>
|
||
, PW
|
||
<quantity id="4CFE2FADC3616350FB63FEBDFA4BFEA8" box="[1266,1432,296,323]" metricMagnitude="-3" metricUnit="m" metricValue="2.01" metricValueMax="2.05" metricValueMin="1.97" pageId="5" pageNumber="346" unit="mm" value="2.01" valueMax="2.05" valueMin="1.97">1.97–2.05 mm</quantity>
|
||
, Edist
|
||
<quantity id="4CFE2FADC3616350FF49FED9FEEDFE8C" box="[216,318,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="346" unit="mm" value="0.55">0.55 mm</quantity>
|
||
, Ediam
|
||
<quantity id="4CFE2FADC3616350FE08FED9FDD3FE8C" box="[409,512,332,359]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="5" pageNumber="346" unit="mm" value="0.58">0.58 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3616350FF56FEE4FA44FD6C" pageId="5" pageNumber="346" type="discussion">
|
||
<paragraph id="8BB98248C3616350FF56FEE4FA44FD6C" blockId="5.[151,1437,153,647]" pageId="5" pageNumber="346">
|
||
<emphasis id="B9725E5AC3616350FF56FEE4FEE3FE60" bold="true" box="[199,304,369,395]" pageId="5" pageNumber="346">Remark.</emphasis>
|
||
Only a female was cited in the original description by
|
||
<bibRefCitation id="EF97FFB9C3616350FC31FEE4FBEEFE60" author="Kleine, R." box="[928,1085,369,395]" pageId="5" pageNumber="346" pagination="325 - 340" refId="ref5974" refString="Kleine, R. (1930) Bestimmungstabelle der Trichalusverwandschaft. Treubia, 9, 325 - 340." type="journal article" year="1930">Kleine (1930)</bibRefCitation>
|
||
and we did not localize the female in Kleine’s collection (
|
||
<bibRefCitation id="EF97FFB9C3616350FE45FE00FDB7FE44" author="Kleine, R." box="[468,612,405,431]" pageId="5" pageNumber="346" pagination="325 - 340" refId="ref5974" refString="Kleine, R. (1930) Bestimmungstabelle der Trichalusverwandschaft. Treubia, 9, 325 - 340." type="journal article" year="1930">Kleine 1930</bibRefCitation>
|
||
). Therefore, our identification is based on similarity in color patterns and occurrence in the same region. The identification of species based on female semaphoront is problematic in netwinged beetles and often impossible in
|
||
<taxonomicName id="4C06F9CBC3616350FDDDFE48FC10FE13" authority="(Bocek & Bocak 2016)" authorityName="Bocek & Bocak" authorityYear="2016" baseAuthorityName="Bocek & Bocak" baseAuthorityYear="2016" box="[588,963,477,504]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3616350FDDDFE48FD64FE1C" box="[588,695,477,503]" italics="true" pageId="5" pageNumber="346">Eniclases</emphasis>
|
||
(
|
||
<bibRefCitation id="EF97FFB9C3616350FD54FE48FC68FE13" author="Bocek, M. & Bocak, L." box="[709,955,477,504]" pageId="5" pageNumber="346" pagination="15 - 35" refId="ref5333" refString="Bocek, M. & Bocak, L. (2016) Species limits in polymorphic mimetic Eniclases net-winged beetles from New Guinean mountains (Coleoptera, Lycidae). ZooKeys, 593, 15 - 35. https: // doi. org / 10.3897 / zookeys. 593.7728" type="journal article" year="2016">Bocek & Bocak 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. The male genitalia of
|
||
<taxonomicName id="4C06F9CBC3616350FB2CFE4BFAB1FE1C" authorityName="Kleine" authorityYear="1930" box="[1213,1378,477,503]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="moluccanus">
|
||
<emphasis id="B9725E5AC3616350FB2CFE4BFAB1FE1C" box="[1213,1378,477,503]" italics="true" pageId="5" pageNumber="346">E. moluccanus</emphasis>
|
||
</taxonomicName>
|
||
have a unique shape of phallic apex. Unlike all known
|
||
<taxonomicName id="4C06F9CBC3616350FD2DFD94FCF4FDF0" baseAuthorityName="Bocek & Bocak" baseAuthorityYear="2016" box="[700,807,513,539]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3616350FD2DFD94FCF4FDF0" box="[700,807,513,539]" italics="true" pageId="5" pageNumber="346">Eniclases</emphasis>
|
||
</taxonomicName>
|
||
, this species does not have the characteristic cup-shaped phallic apex. The generic placement is supported by the presence of characteristic V-shaped pronotal carinae and by the present molecular analysis. We can hypothesize that
|
||
<taxonomicName id="4C06F9CBC3616350FCA7FDDFFC08FD88" authorityName="Kleine" authorityYear="1930" box="[822,987,585,611]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="moluccanus">
|
||
<emphasis id="B9725E5AC3616350FCA7FDDFFC08FD88" box="[822,987,585,611]" italics="true" pageId="5" pageNumber="346">E. moluccanus</emphasis>
|
||
</taxonomicName>
|
||
is involved in mimicry rings with other net-winged beetles in Halmahera as their body coloration and shape are very similar and they occur sympatrically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |