416 lines
34 KiB
XML
416 lines
34 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.872.35361" ID-GBIF-Dataset="ed1a1322-b118-4434-b9ff-503a0fb64792" ID-PMC="PMC6711934" ID-Pensoft-Pub="1313-2970-872-13" ID-Pensoft-UUID="A9BE568D9BF65A179464AB8C7CB8BEB2" ID-PubMed="31496884" ID-ZooBank="DCDEFCB1E33D476F90D7178BC08C311A" ModsDocID="1313-2970-872-13" checkinTime="1566364211746" checkinUser="pensoft" docAuthor="Astafurova, Yulia V., Proshchalykin, Maxim Yu. & Schwarz, Maximilian" docDate="2019" docId="F3062642680F57EA9FB353FA0E5078C1" docLanguage="en" docName="ZooKeys 872: 13-40" docOrigin="ZooKeys 872" docSource="http://dx.doi.org/10.3897/zookeys.872.35361" docTitle="Sphecodes atlanticus Warncke 1992" docType="treatment" docVersion="4" id="A9BE568D9BF65A179464AB8C7CB8BEB2" lastPageNumber="16" masterDocId="A9BE568D9BF65A179464AB8C7CB8BEB2" masterDocTitle="The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010" masterLastPageNumber="40" masterPageNumber="13" pageNumber="16" updateTime="1668167741671" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The distribution of the genus Sphecodes Latreille (Hymenoptera, Halictidae) of the Arabian Peninsula and surrounding countries with description of hitherto unknown female of S. atlanticus Warncke, 1992 and male of S. dathei Schwarz, 2010</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Astafurova, Yulia V.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Proshchalykin, Maxim Yu.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schwarz, Maximilian</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>872</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>13</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.872.35361</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.872.35361</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-872-13</mods:identifier>
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<mods:identifier type="ZooBank">DCDEFCB1E33D476F90D7178BC08C311A</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159217122" LSID="urn:lsid:plazi:treatment:F3062642680F57EA9FB353FA0E5078C1" httpUri="http://treatment.plazi.org/id/F3062642680F57EA9FB353FA0E5078C1" lastPageNumber="16" pageId="3" pageNumber="16">
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<subSubSection pageId="3" pageNumber="16" type="nomenclature">
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<paragraph pageId="3" pageNumber="16">
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<taxonomicName LSID="f3062642-680f-57ea-9fb3-53fa0e5078c1" authority="Warncke, 1992" authorityName="Warncke" authorityYear="1992" class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes atlanticus" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="atlanticus">Sphecodes atlanticus Warncke, 1992</taxonomicName>
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<figureCitation captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Head, females, frontal view. 2 Sphecodes majalis Perez 3 S. scabricollis Wesmael 4 S. barbatus Bluethgen 5 S. atlanticus Warncke 6 S. rubripes Spinola 7 S. albilabris (Fabricius). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16">Figures 5</figureCitation>
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,
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">9</figureCitation>
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,
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">10</figureCitation>
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,
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<figureCitation captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16 - 19. T 1 (16, 17), metasoma (18, 19), females, dorsal view. 16 Sphecodes majalis Perez 17 S. barbatus Bluethgen 18 S. scabricollis Wesmael 19 S. atlanticus Warncke. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures16-19" httpUri="https://binary.pensoft.net/fig/327084" pageId="3" pageNumber="16">19</figureCitation>
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,
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<figureCitation captionStart="Figures 22–25" captionStartId="F7" captionText="Figures 22 - 25. Genitalia, males, dorsal view. 22 Sphecodes atlanticus Warncke 23 S. scabri collis Wesmael 24 S. dathei Schwarz 25 S. crassus Thomson. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.872.35361.figures22-25" httpUri="https://binary.pensoft.net/fig/327086" pageId="3" pageNumber="16">22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="16" type="reference_group">
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<paragraph pageId="3" pageNumber="16">
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<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes atlanticus" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="atlanticus">Sphecodes atlanticus</taxonomicName>
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Warncke, 1992: 25, Abb. 17, ♂ (holotype: ♂, Algeria: Hoggar-Geb., Guelta; OLBL/PCMS), examined.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="16" type="diagnosis">
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<paragraph pageId="3" pageNumber="16">Diagnosis.</paragraph>
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<paragraph pageId="3" pageNumber="16">
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This species is similar to the Trans-Palaearctic
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<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes scabricollis" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="scabricollis">
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<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes scabricollis</emphasis>
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</taxonomicName>
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Wesmael, 1835 owing to the flat genal area, the developed preoccipital lateral carina, the densely punctate head and mesoscutum, the size and shape of male antennal tyloids, and in the similar gonostylar shape. However,
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<taxonomicName lsidName="S. atlanticus" pageId="3" pageNumber="16" rank="species" species="atlanticus">
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<emphasis italics="true" pageId="3" pageNumber="16">S. atlanticus</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName lsidName="S. scabricollis" pageId="3" pageNumber="16" rank="species" species="scabricollis">
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<emphasis italics="true" pageId="3" pageNumber="16">S. scabricollis</emphasis>
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</taxonomicName>
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by a number of characters outlined in Table 2. In addition to presence of preoccipital lateral carina,
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<taxonomicName lsidName="S. atlanticus" pageId="3" pageNumber="16" rank="species" species="atlanticus">
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<emphasis italics="true" pageId="3" pageNumber="16">S. atlanticus</emphasis>
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</taxonomicName>
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clearly differs from the
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<taxonomicName lsidName="gibbus" pageId="3" pageNumber="16" rank="species" species="gibbus">
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<emphasis italics="true" pageId="3" pageNumber="16">gibbus</emphasis>
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</taxonomicName>
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species group (
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<taxonomicName lsidName="S. anatolicus" pageId="3" pageNumber="16" rank="species" species="anatolicus">
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<emphasis italics="true" pageId="3" pageNumber="16">S. anatolicus</emphasis>
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</taxonomicName>
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Warncke, 1922,
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<taxonomicName lsidName="S. gibbus" pageId="3" pageNumber="16" rank="species" species="gibbus">
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<emphasis italics="true" pageId="3" pageNumber="16">S. gibbus</emphasis>
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</taxonomicName>
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(Linnaeus, 1758),
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<taxonomicName lsidName="S. nippon" pageId="3" pageNumber="16" rank="species" species="nippon">
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<emphasis italics="true" pageId="3" pageNumber="16">S. nippon</emphasis>
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</taxonomicName>
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Meyer, 1922,
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<taxonomicName lsidName="S. rufiventris" pageId="3" pageNumber="16" rank="species" species="rufiventris">
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<emphasis italics="true" pageId="3" pageNumber="16">S. rufiventris</emphasis>
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</taxonomicName>
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(Panzer, 1798),
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<taxonomicName lsidName="S. schenckii" pageId="3" pageNumber="16" rank="species" species="schenckii">
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<emphasis italics="true" pageId="3" pageNumber="16">S. schenckii</emphasis>
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</taxonomicName>
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Hagens, 1882,
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<taxonomicName lsidName="S. tadschicus" pageId="3" pageNumber="16" rank="species" species="tadschicus">
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<emphasis italics="true" pageId="3" pageNumber="16">S. tadschicus</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
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in Popov, 1935; see
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<bibRefCitation DOI="https://doi.org/10.25221/fee.369.1" author="Astafurova, YuV" journalOrPublisher="Far Eastern Entomologist" pageId="22" pageNumber="35" pagination="1 - 47" refId="B13" refString="Astafurova, YuV, Proshchalykin, MYu, Engel, MS, 2018a. The cuckoo bee genus Sphecodes Latreille, 1804 in Kazakhstan (Hymenoptera: Halictidae). . Far Eastern Entomologist 369: 1 - 47" title="The cuckoo bee genus Sphecodes Latreille, 1804 in Kazakhstan (Hymenoptera: Halictidae)." url="https://doi.org/10.25221/fee.369.1" volume="369" year="2018 a">Astafurova et al. 2018a</bibRefCitation>
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) by a short distance from top of head to upper margin of lateral ocellus (2 lateral ocellar diameters as seen in dorsal view, versus those with a long vertex where this distance is at most 2.5-3.0 diameters).
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</paragraph>
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<caption ID-Table-UUID="19C1A8A5C2F2DF9539693FAC7C1738C7" httpUri="http://table.plazi.org/id/19C1A8A5C2F2DF9539693FAC7C1738C7" pageId="3" pageNumber="16">
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<paragraph pageId="3" pageNumber="16">
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Table 2. Differences between
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<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes atlanticus" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="atlanticus">
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<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes atlanticus</emphasis>
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</taxonomicName>
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Warncke, 1992 and
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<taxonomicName lsidName="S. scabricollis" pageId="3" pageNumber="16" rank="species" species="scabricollis">
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<emphasis italics="true" pageId="3" pageNumber="16">S. scabricollis</emphasis>
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</taxonomicName>
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Wesmael, 1835.
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</paragraph>
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</caption>
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<paragraph pageId="3" pageNumber="16">
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<table pageId="3" pageNumber="16">
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<tr pageId="3" pageNumber="16">
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<th colspan="1" pageId="3" pageNumber="16" rowspan="1">Characters</th>
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<th colspan="1" pageId="3" pageNumber="16" rowspan="1">
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<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes atlanticus" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="atlanticus">
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<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes atlanticus</emphasis>
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</taxonomicName>
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</th>
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<th colspan="1" pageId="3" pageNumber="16" rowspan="1">
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<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes scabricollis" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="scabricollis">
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<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes scabricollis</emphasis>
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</taxonomicName>
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</th>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="3" pageId="3" pageNumber="16" rowspan="1">
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<emphasis bold="true" pageId="3" pageNumber="16">Both sexes</emphasis>
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</td>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">Distance from top of head to upper margin of lateral ocellus as seen in frontal and dorsal views</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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about one lateral ocellar diameters (
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<figureCitation captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Head, females, frontal view. 2 Sphecodes majalis Perez 3 S. scabricollis Wesmael 4 S. barbatus Bluethgen 5 S. atlanticus Warncke 6 S. rubripes Spinola 7 S. albilabris (Fabricius). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16">Fig. 5</figureCitation>
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)
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</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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about two lateral ocellar diameters (
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<figureCitation captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Head, females, frontal view. 2 Sphecodes majalis Perez 3 S. scabricollis Wesmael 4 S. barbatus Bluethgen 5 S. atlanticus Warncke 6 S. rubripes Spinola 7 S. albilabris (Fabricius). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16">Fig. 3</figureCitation>
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)
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</td>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">Propodeal triangle/metaposnotum</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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equal (in female) or longer (in male) than mesoscutellum (
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">Figs 9</figureCitation>
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,
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">10</figureCitation>
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)
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</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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distinctly shorter than mesoscutellum (
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">Figs 8</figureCitation>
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)
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</td>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">Metasomal terga</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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with coarser and denser punctures (
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<figureCitation captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16 - 19. T 1 (16, 17), metasoma (18, 19), females, dorsal view. 16 Sphecodes majalis Perez 17 S. barbatus Bluethgen 18 S. scabricollis Wesmael 19 S. atlanticus Warncke. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures16-19" httpUri="https://binary.pensoft.net/fig/327084" pageId="3" pageNumber="16">Fig. 19</figureCitation>
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)
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</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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with fine and sparser punctures, especially on T1 (
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<figureCitation captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16 - 19. T 1 (16, 17), metasoma (18, 19), females, dorsal view. 16 Sphecodes majalis Perez 17 S. barbatus Bluethgen 18 S. scabricollis Wesmael 19 S. atlanticus Warncke. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures16-19" httpUri="https://binary.pensoft.net/fig/327084" pageId="3" pageNumber="16">Fig. 18</figureCitation>
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)
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</td>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="3" pageId="3" pageNumber="16" rowspan="1">
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<emphasis bold="true" pageId="3" pageNumber="16">Male</emphasis>
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</td>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">Mesoscutum</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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punctures separated by at most 1.5-2.0 puncture diameters; polished between punctures (
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">Fig. 9</figureCitation>
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)
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</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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areolate (
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<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">Fig. 8</figureCitation>
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)
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</td>
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</tr>
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<tr pageId="3" pageNumber="16">
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">Genitalia</td>
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<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
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gonocoxite dorsally with weak impression; gonostylar process longer (
|
||
<figureCitation captionStart="Figures 22–25" captionStartId="F7" captionText="Figures 22 - 25. Genitalia, males, dorsal view. 22 Sphecodes atlanticus Warncke 23 S. scabri collis Wesmael 24 S. dathei Schwarz 25 S. crassus Thomson. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.872.35361.figures22-25" httpUri="https://binary.pensoft.net/fig/327086" pageId="3" pageNumber="16">Fig. 22</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
|
||
gonocoxite dorsally without impression; gonostylar process shorter (
|
||
<figureCitation captionStart="Figures 22–25" captionStartId="F7" captionText="Figures 22 - 25. Genitalia, males, dorsal view. 22 Sphecodes atlanticus Warncke 23 S. scabri collis Wesmael 24 S. dathei Schwarz 25 S. crassus Thomson. Scale bars: 0.25 mm." figureDoi="10.3897/zookeys.872.35361.figures22-25" httpUri="https://binary.pensoft.net/fig/327086" pageId="3" pageNumber="16">Fig. 23</figureCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="16">
|
||
<td colspan="3" pageId="3" pageNumber="16" rowspan="1">
|
||
<emphasis bold="true" pageId="3" pageNumber="16">Female</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="3" pageNumber="16">
|
||
<td colspan="1" pageId="3" pageNumber="16" rowspan="1">Paraocular areas</td>
|
||
<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
|
||
with dense pubescence obscuring integument (
|
||
<figureCitation captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Head, females, frontal view. 2 Sphecodes majalis Perez 3 S. scabricollis Wesmael 4 S. barbatus Bluethgen 5 S. atlanticus Warncke 6 S. rubripes Spinola 7 S. albilabris (Fabricius). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16">Fig. 5</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="3" pageNumber="16" rowspan="1">
|
||
with sparse pubescence not obscuring integument (
|
||
<figureCitation captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Head, females, frontal view. 2 Sphecodes majalis Perez 3 S. scabricollis Wesmael 4 S. barbatus Bluethgen 5 S. atlanticus Warncke 6 S. rubripes Spinola 7 S. albilabris (Fabricius). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16">Fig. 3</figureCitation>
|
||
)
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="16" type="description">
|
||
<paragraph pageId="3" pageNumber="16">Description of hitherto unknown female.</paragraph>
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Total body length 6.5-8.5 mm. Head (
|
||
<figureCitation captionStart="Figures 2–7" captionStartId="F2" captionText="Figures 2 - 7. Head, females, frontal view. 2 Sphecodes majalis Perez 3 S. scabricollis Wesmael 4 S. barbatus Bluethgen 5 S. atlanticus Warncke 6 S. rubripes Spinola 7 S. albilabris (Fabricius). Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16">Fig. 5</figureCitation>
|
||
) black (except reddish mouthparts); transverse, 1.3 times as wide as long; vertex elevated, distance from top of head to upper margin of lateral ocellus ca. one lateral ocellar diameter as seen in frontal view and ca. two lateral ocellar diameters as seen in dorsal view; F1 and F2 transverse, 0.7-0.8 times as long as wide; F3 as long as wide; face with fine contiguous punctures (10-20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
), clypeus with shiny interspaces between punctures separated by 0.1-0.5 of a puncture diameter; mandible with an inner tooth; paraocular areas and upper part of gena with dense adpressed, snow-white, plumose pubescence obscuring the integument.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.872.35361.figures2-7" httpUri="https://binary.pensoft.net/fig/327081" pageId="3" pageNumber="16" start="Figures 2–7" startId="F2">
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Figures 2-7. Head, females, frontal view.
|
||
<emphasis bold="true" pageId="3" pageNumber="16">2</emphasis>
|
||
<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes majalis" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="majalis">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes majalis</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
|
||
<emphasis bold="true" pageId="3" pageNumber="16">3</emphasis>
|
||
<taxonomicName lsidName="S. scabricollis" pageId="3" pageNumber="16" rank="species" species="scabricollis">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. scabricollis</emphasis>
|
||
</taxonomicName>
|
||
Wesmael
|
||
<emphasis bold="true" pageId="3" pageNumber="16">4</emphasis>
|
||
<taxonomicName lsidName="S. barbatus" pageId="3" pageNumber="16" rank="species" species="barbatus">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. barbatus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
|
||
<emphasis bold="true" pageId="3" pageNumber="16">5</emphasis>
|
||
<taxonomicName lsidName="S. atlanticus" pageId="3" pageNumber="16" rank="species" species="atlanticus">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. atlanticus</emphasis>
|
||
</taxonomicName>
|
||
Warncke
|
||
<emphasis bold="true" pageId="3" pageNumber="16">6</emphasis>
|
||
<taxonomicName lsidName="S. rubripes" pageId="3" pageNumber="16" rank="species" species="rubripes">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. rubripes</emphasis>
|
||
</taxonomicName>
|
||
Spinola
|
||
<emphasis bold="true" pageId="3" pageNumber="16">7</emphasis>
|
||
<taxonomicName lsidName="S. albilabris" pageId="3" pageNumber="16" rank="species" species="albilabris">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. albilabris</emphasis>
|
||
</taxonomicName>
|
||
(Fabricius). Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Mesosoma black; mesoscutum with coarse punctures (25-50
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
) separated by at most a puncture diameter (
|
||
<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">Fig. 10</figureCitation>
|
||
); mesoscutellum with irregular punctures separated by 0.1-4 puncture diameters; mesepisternum densely reticulate-rugose; propodeal triangle coarsely reticulate-rugose with large shiny, smooth interspaces between wrinkles (
|
||
<figureCitation captionStart="Figures 8–11" captionStartId="F3" captionText="Figures 8 - 11. Mesosoma (8 - 10), dorsal view; lectotype labels (11). 8 Sphecodes scabricollis Wesmael, male 9, 10 S. atlanticus Warncke (9 - male, 10 - female) 11 S. intermedius Bluethgen, label of lectotype. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16">Fig. 10</figureCitation>
|
||
); lateral parts of propodeum finely and densely strigate or strigate-rugose with granulate interspaces between wrinkles; vertical part of propodeum smooth with coarse and dense punctures; legs reddish or dark brown. Hind wing costal margin with 9-10 hamuli.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.872.35361.figures8-11" httpUri="https://binary.pensoft.net/fig/327082" pageId="3" pageNumber="16" start="Figures 8–11" startId="F3">
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Figures 8-11. Mesosoma (
|
||
<emphasis bold="true" pageId="3" pageNumber="16">8-10</emphasis>
|
||
), dorsal view; lectotype labels (11).
|
||
<emphasis bold="true" pageId="3" pageNumber="16">8</emphasis>
|
||
<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes scabricollis" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="scabricollis">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes scabricollis</emphasis>
|
||
</taxonomicName>
|
||
Wesmael, male
|
||
<emphasis bold="true" pageId="3" pageNumber="16">9, 10</emphasis>
|
||
<taxonomicName lsidName="S. atlanticus" pageId="3" pageNumber="16" rank="species" species="atlanticus">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. atlanticus</emphasis>
|
||
</taxonomicName>
|
||
Warncke (
|
||
<emphasis bold="true" pageId="3" pageNumber="16">9</emphasis>
|
||
- male,
|
||
<emphasis bold="true" pageId="3" pageNumber="16">10</emphasis>
|
||
- female)
|
||
<emphasis bold="true" pageId="3" pageNumber="16">11</emphasis>
|
||
<taxonomicName lsidName="S. intermedius" pageId="3" pageNumber="16" rank="species" species="intermedius">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. intermedius</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
|
||
, label of lectotype. Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Metasoma (
|
||
<figureCitation captionStart="Figures 16–19" captionStartId="F5" captionText="Figures 16 - 19. T 1 (16, 17), metasoma (18, 19), females, dorsal view. 16 Sphecodes majalis Perez 17 S. barbatus Bluethgen 18 S. scabricollis Wesmael 19 S. atlanticus Warncke. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.872.35361.figures16-19" httpUri="https://binary.pensoft.net/fig/327084" pageId="3" pageNumber="16">Fig. 19</figureCitation>
|
||
) with colouration varying from red on T1-T4 to entirely dark-brown; tergal discs with coarse and dense punctures (20-30
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
/ 0.5-2, sparser on anterior third of T1), marginal zone impunctate except on T1 with dense punctures (10-20
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
/ 0.5-2); sterna finely tessellate with coarse setae pores; pygidial plate dull, as wide as metabasitarsus.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.872.35361.figures12-15" httpUri="https://binary.pensoft.net/fig/327083" pageId="3" pageNumber="16" start="Figures 12–15" startId="F4">
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Figures 12-15.
|
||
<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes dathei" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="dathei">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes dathei</emphasis>
|
||
</taxonomicName>
|
||
Schwarz, male.
|
||
<emphasis bold="true" pageId="3" pageNumber="16">12</emphasis>
|
||
Head, frontal view
|
||
<emphasis bold="true" pageId="3" pageNumber="16">13</emphasis>
|
||
antenna, frontal view
|
||
<emphasis bold="true" pageId="3" pageNumber="16">14</emphasis>
|
||
mesosoma, dorsal view
|
||
<emphasis bold="true" pageId="3" pageNumber="16">15</emphasis>
|
||
T1, dorsal view. Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.872.35361.figures16-19" httpUri="https://binary.pensoft.net/fig/327084" pageId="3" pageNumber="16" start="Figures 16–19" startId="F5">
|
||
<paragraph pageId="3" pageNumber="16">
|
||
Figures 16-19. T1 (
|
||
<emphasis bold="true" pageId="3" pageNumber="16">16, 17</emphasis>
|
||
), metasoma (
|
||
<emphasis bold="true" pageId="3" pageNumber="16">18, 19</emphasis>
|
||
), females, dorsal view.
|
||
<emphasis bold="true" pageId="3" pageNumber="16">16</emphasis>
|
||
<taxonomicName class="Insecta" family="Halictidae" genus="Sphecodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphecodes majalis" order="Hymenoptera" pageId="3" pageNumber="16" phylum="Arthropoda" rank="species" species="majalis">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">Sphecodes majalis</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Pérez">Perez</normalizedToken>
|
||
<emphasis bold="true" pageId="3" pageNumber="16">17</emphasis>
|
||
<taxonomicName lsidName="S. barbatus" pageId="3" pageNumber="16" rank="species" species="barbatus">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. barbatus</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Blüthgen">Bluethgen</normalizedToken>
|
||
<emphasis bold="true" pageId="3" pageNumber="16">18</emphasis>
|
||
<taxonomicName lsidName="S. scabricollis" pageId="3" pageNumber="16" rank="species" species="scabricollis">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. scabricollis</emphasis>
|
||
</taxonomicName>
|
||
Wesmael
|
||
<emphasis bold="true" pageId="3" pageNumber="16">19</emphasis>
|
||
<taxonomicName lsidName="S. atlanticus" pageId="3" pageNumber="16" rank="species" species="atlanticus">
|
||
<emphasis italics="true" pageId="3" pageNumber="16">S. atlanticus</emphasis>
|
||
</taxonomicName>
|
||
Warncke. Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="16" type="materials_examined">
|
||
<paragraph pageId="3" pageNumber="16">Material examined.</paragraph>
|
||
<paragraph pageId="3" pageNumber="16">
|
||
SAUDI ARABIA: 6 ♂♂, Wadi Majarish (below Taif), 12.II.1983, K. Guichard (NHMUK 013380451, 013380453, 013380460, 013380462, 013380466, 013380459); 4 ♂♂, Fayfa, 200 m, 29.I. 1983, K. Guichard (NHMUK 013380452, 013380461, 013380463, 013380464); 1 ♂, Lodar, 800 m, 16.V.1967, K. Guichard (NHMUK 0133804446); 1 ♂, Abu Arish, 26.III.1980, K. Guichard (NHMUK 013380465); 2 ♀♀, 1 ♂, Abu Arish, Jizzan Hot Springs, 25.III.1980, K. Guichard (NHMUK 013380458, 013380441, 013380454), 1 ♂, idem, 28.I.1983 (NHMUK 013380450); 1 ♀, Wadi Maraba, 25.I.1983, 1000 m, K. Guichard (NHMUK 013380457); 1 ♀, Jeddah, Locust Research Station, 17.I.1972, A. Basha (NHMUK 013380442); YEMEN: 1 ♂, Usaifira, 1 mile N
|
||
<normalizedToken originalValue="Ta’izz">Ta'izz</normalizedToken>
|
||
, 4.500 ft, 21.XII.1937, H. Scott, E. Britton (NHMUK 013380468), 1 ♂, Wadi Maytam, 12 km SE Ibb, 1600 m,
|
||
<geoCoordinate degrees="13" direction="north" minutes="53" orientation="latitude" precision="925" value="13.883333">13°53'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44" direction="east" minutes="18" orientation="longitude" precision="925" value="44.3">44°18'E</geoCoordinate>
|
||
, 27.X.2005, J. Halada (OLBL/PCMS); 2 ♀♀, 3 ♂♂, Hawf NE Albhaydah, 200-730 m,
|
||
<geoCoordinate degrees="16" direction="north" minutes="53" orientation="latitude" precision="925" value="16.883333">16°53'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="53" direction="east" minutes="05" orientation="longitude" precision="925" value="53.083332">53°05'E</geoCoordinate>
|
||
, 14.X.2005, J. Halada (OLBL/PCMS); 2 ♀♀, 4 ♂♂, 20 km S Taizz, 1200 m,
|
||
<geoCoordinate degrees="13" direction="north" minutes="30" orientation="latitude" precision="925" value="13.5">13°30'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="43" direction="east" minutes="57" orientation="longitude" precision="925" value="43.95">43°57'E</geoCoordinate>
|
||
, 24.X.2005, J. Halada (OLBL/PCMS); 2 ♂♂, Jabal Bura, NEE Al Hudaydah, 200-800 m,
|
||
<geoCoordinate degrees="14" direction="north" minutes="52" orientation="latitude" precision="925" value="14.866667">14°52'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="43" direction="east" minutes="24" orientation="longitude" precision="925" value="43.4">43°24'E</geoCoordinate>
|
||
, 30.X-1.XI.2005, J. Halada (OLBL/PCMS); 1 ♂, Wadi Aniz, SSW Sana, 1520 m,
|
||
<geoCoordinate degrees="14" direction="north" minutes="60" orientation="latitude" precision="925" value="15.0">14°60'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="44" direction="east" minutes="09" orientation="longitude" precision="925" value="44.15">44°09'E</geoCoordinate>
|
||
, 7.X.2005, J. Halada (OLBL/PCMS).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="16" type="distribution">
|
||
<paragraph pageId="3" pageNumber="16">Distribution.</paragraph>
|
||
<paragraph pageId="3" pageNumber="16">*Saudi Arabia, *Yemen; Algeria, the Canary Islands.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |