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<document id="21B58808F092C12CCE2DB313435BAF76" ID-CLB-Dataset="29929" ID-DOI="10.11646/zootaxa.4433.1.13" ID-GBIF-Dataset="6537c69f-8419-4a67-94fd-e0433b3fd47a" ID-ISSN="1175-5326" ID-Zenodo-Dep="1287085" ID-ZooBank="5D1E2ACB-C252-411E-B6AB-F447D1971164" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1528732616621" checkinUser="plazi" docAuthor="Trivedi, Jigneshkumar N., Campos, Ernesto &amp; Vachhrajani, Kauresh D." docDate="2018" docId="96247A4783207C034C9EFE59FD52F877" docLanguage="en" docName="zootaxa.4433.1.13.pdf" docOrigin="Zootaxa 4433 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arcotheres shahi Trivedi &amp; Campos &amp; Vachhrajani 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="199" masterDocId="6A1D023F83217C074C09FFAEFFDEFF85" masterDocTitle="A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India" masterLastPageNumber="200" masterPageNumber="195" pageNumber="196" updateTime="1698548246911" updateUser="ExternalLinkService">
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<mods:title id="AC7EA53D15515728BEA7D3ADCCB28B90">A new pea crab species of the genus Arcotheres Manning, 1993 (Crustacea: Decapoda: Brachyura: Pinnotheridae) from India</mods:title>
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<mods:namePart id="01509CC6B037A7ABEA8AF138C3AFE0D8">Trivedi, Jigneshkumar N.</mods:namePart>
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<mods:namePart id="C53C738331E6E7F25D3A59F83029237E">Campos, Ernesto</mods:namePart>
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<mods:namePart id="F1E384AB830353FC5B34751D4C406788">Vachhrajani, Kauresh D.</mods:namePart>
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<mods:date id="58F956906EDE2D32D8D6DB73434C0984">2018</mods:date>
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<treatment id="96247A4783207C034C9EFE59FD52F877" ID-DOI="http://doi.org/10.5281/zenodo.5988831" ID-GBIF-Taxon="145939528" ID-Zenodo-Dep="5988831" LSID="urn:lsid:plazi:treatment:96247A4783207C034C9EFE59FD52F877" httpUri="http://treatment.plazi.org/id/96247A4783207C034C9EFE59FD52F877" lastPageId="4" lastPageNumber="199" pageId="1" pageNumber="196">
<subSubSection id="569798DA83207C064C9EFE59FECFFDB6" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="1E32CB5183207C064C9EFE59FE70FD97" blockId="1.[151,430,503,563]" box="[151,430,503,530]" pageId="1" pageNumber="196">
<heading id="457A7C3D83207C064C9EFE59FE70FD97" bold="true" box="[151,430,503,530]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
<emphasis id="2CF9174383207C064C9EFE59FE70FD97" bold="true" box="[151,430,503,530]" pageId="1" pageNumber="196">
<taxonomicName id="D98DB0D283207C064C9EFE59FEBBFD94" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[151,357,503,529]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383207C064C9EFE59FEBBFD94" bold="true" box="[151,357,503,529]" italics="true" pageId="1" pageNumber="196">Arcotheres shahi</emphasis>
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<taxonomicNameLabel id="37CAAA3883207C064D65FE56FE70FD97" box="[364,430,504,530]" pageId="1" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="1E32CB5183207C064C9EFDB4FECFFDB6" blockId="1.[151,430,503,563]" box="[151,273,538,563]" pageId="1" pageNumber="196">
(
<figureCitation id="86B6D7D483207C064C96FDB4FED7FDB6" box="[159,265,538,563]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1295,1317]" captionStartId-1="3.[151,250,1553,1575]" captionStartId-2="4.[151,250,1269,1291]" captionTargetBox-0="[348,1277,402,1273]" captionTargetBox-1="[338,1264,642,1506]" captionTargetBox-2="[288,1255,362,1232]" captionTargetId-0="figure@2.[304,1282,383,1290]" captionTargetId-1="figure@3.[305,1282,625,1532]" captionTargetId-2="figure@4.[269,1317,341,1248]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." captionText-1="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." captionText-2="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri-0="https://zenodo.org/record/1287087/files/figure.png" httpUri-1="https://zenodo.org/record/1287089/files/figure.png" httpUri-2="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Figs. 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="569798DA83207C064C9EFDCCFE30FD62" pageId="1" pageNumber="196" type="materials_examined">
<paragraph id="1E32CB5183207C064C9EFDCCFE30FD62" blockId="1.[151,1437,610,2003]" pageId="1" pageNumber="196">
<materialsCitation id="AEE5C10C83207C064C9EFDCCFEFDFD47" ID-GBIF-Occurrence="1848743982" collectingDate="2016-02-02" collectorName="J. Trivedi" country="India" latitude="9.282222" location="Material" longLatPrecision="21" longitude="79.20861" municipality="Pamban Fishing Harbor" pageId="1" pageNumber="196" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<emphasis id="2CF9174383207C064C9EFDCCFE5EFDFE" bold="true" box="[151,384,610,635]" pageId="1" pageNumber="196">
<location id="1B529D8A83207C064C9EFDCCFF21FDFE" LSID="urn:lsid:plazi:treatment:96247A4783207C034C9EFE59FD52F877:1B529D8A83207C064C9EFDCCFF21FDFE" box="[151,255,610,635]" country="India" latitude="9.282222" longLatPrecision="21" longitude="79.20861" municipality="Pamban Fishing Harbor" name="Material" pageId="1" pageNumber="196" stateProvince="Tamil Nadu">Material</location>
examined.
</emphasis>
<typeStatus id="C13675F383207C064D8EFDCCFE2CFDFE" box="[391,498,610,635]" pageId="1" pageNumber="196" type="holotype">Holotype</typeStatus>
, female (CL
<quantity id="D97566B483207C064E8CFDCCFD35FDFF" box="[645,747,610,635]" metricMagnitude="-3" metricUnit="m" metricValue="6.13" pageId="1" pageNumber="196" unit="mm" value="6.13">6.13 mm</quantity>
, CW
<quantity id="D97566B483207C064F22FDCCFC4FFDFE" box="[811,913,610,635]" metricMagnitude="-3" metricUnit="m" metricValue="6.77" pageId="1" pageNumber="196" unit="mm" value="6.77">6.77 mm</quantity>
) (ZL-AR-CR-97), in
<taxonomicName id="D98DB0D283207C06488AFDCDFE9BFD1A" authority="(Preston, 1916)" authorityName="Preston" authorityYear="1916" baseAuthorityName="Preston" baseAuthorityYear="1916" class="Bivalvia" family="Ostreidae" genus="Crassostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pterioida" pageId="1" pageNumber="196" phylum="Mollusca" rank="species" species="madrasensis">
<emphasis id="2CF9174383207C06488AFDCDFA42FDFE" box="[1155,1436,611,635]" italics="true" pageId="1" pageNumber="196">Crassostrea madrasensis</emphasis>
(Preston, 1916)
</taxonomicName>
, from
<collectingMunicipality id="FE56512B83207C064D84FD28FD47FD1A" box="[397,665,646,671]" pageId="1" pageNumber="196">Pamban Fishing Harbor</collectingMunicipality>
,
<collectingRegion id="DC4905B383207C064EADFD28FCF5FD1A" box="[676,811,646,671]" country="India" name="Tamil Nadu" pageId="1" pageNumber="196">Tamil Nadu</collectingRegion>
,
<collectingCountry id="669A8BC183207C064F3CFD28FCAFFD1A" box="[821,881,646,671]" name="India" pageId="1" pageNumber="196">India</collectingCountry>
,
<geoCoordinate id="7BB9AD9683207C064F75FD28FC2AFD1A" box="[892,1012,646,671]" direction="north" orientation="latitude" pageId="1" pageNumber="196" precision="15" value="9.282222">9°16'56&quot;N</geoCoordinate>
,
<geoCoordinate id="7BB9AD9683207C064FF6FD29FB5DFD1A" box="[1023,1155,647,671]" direction="east" orientation="longitude" pageId="1" pageNumber="196" precision="15" value="79.20861">79°12'31&quot;E</geoCoordinate>
,
<collectingDate id="7A77147983207C064887FD29FA96FD1A" box="[1166,1352,646,671]" pageId="1" pageNumber="196" value="2016-02-02">
<date id="6A33ED9183207C064887FD29FA96FD1A" box="[1166,1352,646,671]" pageId="1" pageNumber="196" value="2016-02-02">2 February 2016</date>
</collectingDate>
, coll.
<collectorName id="B378AE8783207C064983FD29FF37FD46" pageId="1" pageNumber="196">J. Trivedi</collectorName>
&amp; D
</materialsCitation>
. Patel. Paratypes: 1 female (CL
<quantity id="D97566B483207C064E83FD05FD2EFD47" box="[650,752,683,707]" metricMagnitude="-3" metricUnit="m" metricValue="3.43" pageId="1" pageNumber="196" unit="mm" value="3.43">3.43 mm</quantity>
, CW
<quantity id="D97566B483207C064F27FD05FC49FD46" box="[814,919,683,707]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="196" unit="mm" value="3.3">3.30 mm</quantity>
),
<materialsCitation id="AEE5C10C83207C064FA0FD05FE34FD62" ID-GBIF-Occurrence="1848744096" collectingDate="2016-02-02" collectorName="J. Trivedi" country="India" latitude="9.282222" location="Material" longLatPrecision="21" longitude="79.20861" municipality="Pamban Fishing Harbor" pageId="1" pageNumber="196" specimenCount="1" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<specimenCount id="088B00D883207C064FA0FD05FC2AFD46" box="[937,1012,682,707]" pageId="1" pageNumber="196" type="male">1 male</specimenCount>
(CL
<quantity id="D97566B483207C064826FD04FB50FD46" box="[1071,1166,682,707]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="196" unit="mm" value="3.5">3.50mm</quantity>
, CW
<quantity id="D97566B483207C0648C4FD05FAF0FD46" box="[1229,1326,683,707]" metricMagnitude="-3" metricUnit="m" metricValue="4.14" pageId="1" pageNumber="196" unit="mm" value="4.14">4.14mm</quantity>
), ZL-AR- CR-98, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="569798DA83207C054CCEFD5CFA89FEE0" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196" type="description">
<paragraph id="1E32CB5183207C064CCEFD5CFF1AFB3E" blockId="1.[151,1437,610,2003]" pageId="1" pageNumber="196">
<emphasis id="2CF9174383207C064CCEFD5CFE87FC8E" bold="true" box="[199,345,754,779]" pageId="1" pageNumber="196">Description.</emphasis>
Female: carapace sub-trapezoidal (
<figureCitation id="86B6D7D483207C064F0FFD5CFCB8FC8E" box="[774,870,754,779]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1A</figureCitation>
), slightly wider than long, regions moderately defined, anterolateral margin cristate, front slightly projecting, relatively convex, posterior margin slightly concave. Eyes small, pigmented, not visible in dorsal view; orbits oval. Antennules folded obliquely, antennular cavities slightly larger than ocular cavities (
<figureCitation id="86B6D7D483207C064E86FCF0FD36FCF2" box="[655,744,862,887]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1B</figureCitation>
); antennae 6-segmented, basal segment with large, quadrate urinal article (
<figureCitation id="86B6D7D483207C064D3EFC2CFE50FC1E" box="[311,398,898,923]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1B</figureCitation>
), fused with epistome, distal segment setose, scarcely reaching beyond inner edge of orbit. Epistome, endostome separated, margin of first thoracic sternite emarginated. MXP3 obliquely placed in buccal cavity, propodus rhomboidal about twice as long as high, longer than carpus, dorsal margin with fringe of short setae, digitiform dactylus distally setose, inserted medially in notch on ventral margin of propodus, distally not overreaching tip of propodus but overreaching anteromesial angle of merus; ischiomerus completely fused (
<figureCitation id="86B6D7D483207C064CA9FB98FF33FBCA" box="[160,237,1078,1103]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig.1F</figureCitation>
), 1.80 times as long as wide, lateral margin convex, with short, scattered setae; mesial margin angular at the widest point forming anteromesial angle, with long, slender setae. Exopod slightly longer than half length of ischiomerus, flagellum with three articles before aesthetascs, small distal segment inconspicuously articulated (
<figureCitation id="86B6D7D483207C064966FBD0FF69FB3E" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1F</figureCitation>
).
</paragraph>
<paragraph id="1E32CB5183207C064CCEFB68FB8FFA33" blockId="1.[151,1437,610,2003]" pageId="1" pageNumber="196">
Chelipeds (P1) stouter than ambulatory legs (P25), merus slightly longer than carpus (
<figureCitation id="86B6D7D483207C06489EFB68FB2FFB5A" box="[1175,1265,1222,1247]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1A</figureCitation>
); chela slightly globose (
<figureCitation id="86B6D7D483207C064D08FB44FE52FA86" box="[257,396,1258,1283]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Figs. 1D, E</figureCitation>
), ventral margin gently convex, dactylus shorter than dorsal margin of palm, longer than greatest height of palm, small scattered patches of granules present on outer surface, distal tip pointed, strongly curved (
<figureCitation id="86B6D7D483207C064CFFFA9CFE96FACE" box="[246,328,1330,1355]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig.1D</figureCitation>
), cutting edge with one large proximal tooth, few small teeth distally, with long setae on inner surface. Pollex with two large, proximal teeth separated by denticulate notch, patch of short setae present just below cutting edge visible on outer surface, row of long setae running parallel to cutting edge visible on inner surface (
<figureCitation id="86B6D7D483207C064CFEFA30FE92FA32" box="[247,332,1438,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1E</figureCitation>
), outer surface of palm slightly convex, inner surface highly convex.
</paragraph>
<paragraph id="1E32CB5183207C064CCEFA6CFCB3F89A" blockId="1.[151,1437,610,2003]" pageId="1" pageNumber="196">
Ambulatory legs (P25) asymmetrical in length (
<figureCitation id="86B6D7D483207C064EF7FA6CFC56FA5E" box="[766,904,1474,1499]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Figs. 3AD</figureCitation>
), those on left side longer than corresponding legs on right side (
<figureCitation id="86B6D7D483207C064D6CFA48FE63FA7A" box="[357,445,1510,1535]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1A</figureCitation>
,
<figureCitation id="86B6D7D483207C064DC1FA48FD90FA7A" box="[456,590,1510,1535]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Figs. 3AD</figureCitation>
; AC); P2, P3 (
<figureCitation id="86B6D7D483207C064F04FA48FC4CFA7A" box="[781,914,1510,1535]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Figs. 3AB</figureCitation>
; AB) similar, with merus, carpus, propodus, dactylus glabrous; propodus longer than carpus, slightly curved on both margins; dactylus conical, shorter than carpus, propodus, hooked at tip. Fourth pereopods markedly asymmetrical (
<figureCitation id="86B6D7D483207C064803F980FBAEF9C2" box="[1034,1136,1582,1607]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Figs. 3C</figureCitation>
, C), longer P4 (
<figureCitation id="86B6D7D483207C064933F980FA4DF9C2" box="[1338,1427,1582,1607]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Fig. 3C</figureCitation>
) slender than P2, P3; carpus outer margin curved; dactylus sickle-like, ending in small curved tip, slightly smaller than propodus, longer than dactyli of P2, P3, P5, ventral margin with long fine setae, few short setae also present on dorsal distal margin of curved tip. P5 more slender than P2P4, all articles similar in shape to those of P2, P3; except longer sickle-like dactylus, dactylus much longer than those of P2and P3 but shorter than that of P4, few long setae present on distal half of ventral (inner) margin. Relative length of ambulatory legs P4&gt;P3&gt;P2&gt;P5 and dactyli of longer pereopods P4&gt;P5&gt;P3&gt;P2 (
<figureCitation id="86B6D7D483207C064E8FF8A8FCD3F89A" box="[646,781,1798,1823]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="1" pageNumber="196">Figs. 3AD</figureCitation>
; AC).
</paragraph>
<paragraph id="1E32CB5183207C064CCEF884FC31F8E2" blockId="1.[151,1437,610,2003]" pageId="1" pageNumber="196">
Pleon wide (
<figureCitation id="86B6D7D483207C064D5FF884FE70F8C6" box="[342,430,1834,1859]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="1" pageNumber="196">Fig. 1C</figureCitation>
), with six free somites and telson, margin with tomentum, pleonal somites increasing in length distally, widening from somite 1 to 4, then decreasing towards telson.
</paragraph>
<paragraph id="1E32CB5183207C054CCEF8DCFA89FEE0" blockId="1.[151,1437,610,2003]" lastBlockId="2.[151,1436,151,357]" lastPageId="2" lastPageNumber="197" pageId="1" pageNumber="196">
Male: carapace subovate (
<figureCitation id="86B6D7D483207C064E0DF8DCFDB8F80E" box="[516,614,1906,1931]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="1" pageNumber="196">Fig. 2A</figureCitation>
), slightly longer than wide, dorsal surface smooth, front projecting anteriorly, emarginated, eyes visible in dorsal view. MXP3 as in female. Chela stouter than in female, dorsal and ventral margins strongly convex (
<figureCitation id="86B6D7D483207C064E13F814FDABF856" box="[538,629,1978,2003]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="1" pageNumber="196">Fig. 2B</figureCitation>
), inner margin of fingers with long setae, outer surface without setae, cutting edges of both fingers same as female chela. Ambulatory legs symmetrical, length of ambulatory legs P4&gt;P3&gt;P2&gt;P5 (
<figureCitation id="86B6D7D483237C054D44FF12FE0AFF51" box="[333,468,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="2" pageNumber="197">Figs. 3EH</figureCitation>
). P2P4 with a fringe of long swimming setae on outer face of carpus and propodus increasing in size distally. P5 with propodus ellipsoidal, dactylus very curved, ventral margin of both with long setae. Relative length of dactyli of ambulatory legs P5&gt;P4&gt;P3&gt;P2. Pleon with six free somites and telson (
<figureCitation id="86B6D7D483237C054966FEAAFF65FEC5" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="2" pageNumber="197">Fig. 2C</figureCitation>
), widest at somites 2, 3, decreasing towards evenly rounded telson. G1 long (
<figureCitation id="86B6D7D483237C054836FE89FB44FEC5" box="[1087,1178,295,320]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="2" pageNumber="197">Fig. 2D</figureCitation>
), sharply curved, with short angular tip, ventral surface with long setae; G2 (
<figureCitation id="86B6D7D483237C054EFAFEE2FC99FEE1" box="[755,839,332,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="2" pageNumber="197">Fig. 2E</figureCitation>
) stout with pointed angular tip, exopod absent.
</paragraph>
</subSubSection>
<caption id="4AF29BD983237C054C9EFAA1FAF3FAC1" httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="2" pageNumber="197" startId="2.[151,250,1295,1317]" targetBox="[348,1277,402,1273]" targetPageId="2">
<paragraph id="1E32CB5183237C054C9EFAA1FAF3FAC1" blockId="2.[151,1436,1295,1348]" pageId="2" pageNumber="197">
<emphasis id="2CF9174383237C054C9EFAA1FECDFAA1" bold="true" box="[151,275,1295,1317]" pageId="2" pageNumber="197">FIGURE 1.</emphasis>
<taxonomicName id="D98DB0D283237C054D13FAA1FE1BFAA0" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[282,453,1295,1317]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383237C054D13FAA1FE1BFAA0" box="[282,453,1295,1317]" italics="true" pageId="2" pageNumber="197">Arcotheres shahi</emphasis>
</taxonomicName>
<emphasis id="2CF9174383237C054DC5FABEFDD5FAA0" bold="true" box="[460,523,1296,1317]" pageId="2" pageNumber="197">
<taxonomicNameLabel id="37CAAA3883237C054DC5FABEFDDBFAA0" box="[460,517,1296,1317]" pageId="2" pageNumber="197" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view.
</paragraph>
</caption>
<subSubSection id="569798DA83237C054CCEFAC5FF01FA2D" pageId="2" pageNumber="197" type="biology_ecology">
<paragraph id="1E32CB5183237C054CCEFAC5FF01FA2D" blockId="2.[151,1436,1387,2024]" pageId="2" pageNumber="197">
<emphasis id="2CF9174383237C054CCEFAC5FED8FA01" bold="true" box="[199,262,1387,1412]" pageId="2" pageNumber="197">Host.</emphasis>
The specimens of
<taxonomicName id="D98DB0D283237C054DEEFAC2FD99FA01" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[487,583,1388,1412]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383237C054DEEFAC2FD99FA01" box="[487,583,1388,1412]" italics="true" pageId="2" pageNumber="197">A. shahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37CAAA3883237C054E58FAC5FD4DFA01" box="[593,659,1387,1412]" pageId="2" pageNumber="197" rank="species">
<emphasis id="2CF9174383237C054E58FAC5FD4DFA01" bold="true" box="[593,659,1387,1412]" pageId="2" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
were collected from the oyster,
<taxonomicName id="D98DB0D283237C054819FAC2FF05FA2D" authority="(Preston, 1916)" authorityName="Preston" authorityYear="1916" baseAuthorityName="Preston" baseAuthorityYear="1916" class="Bivalvia" family="Ostreidae" genus="Crassostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pterioida" pageId="2" pageNumber="197" phylum="Mollusca" rank="species" species="madrasensis">
<emphasis id="2CF9174383237C054819FAC2FAF3FA01" box="[1040,1325,1388,1412]" italics="true" pageId="2" pageNumber="197">Crassostrea madrasensis</emphasis>
(Preston, 1916)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="569798DA83237C054CCEFA1DFA44FA49" box="[199,1434,1459,1484]" pageId="2" pageNumber="197" type="distribution">
<paragraph id="1E32CB5183237C054CCEFA1DFA44FA49" blockId="2.[151,1436,1387,2024]" box="[199,1434,1459,1484]" pageId="2" pageNumber="197">
<emphasis id="2CF9174383237C054CCEFA1DFEBEFA49" bold="true" box="[199,352,1459,1484]" pageId="2" pageNumber="197">Distribution.</emphasis>
So far known only from the coastal waters of Pamban, Gulf of Mannar,
<collectingRegion id="DC4905B383237C054885FA1DFACFFA49" box="[1164,1297,1459,1484]" country="India" name="Tamil Nadu" pageId="2" pageNumber="197">Tamil Nadu</collectingRegion>
state,
<collectingCountry id="669A8BC183237C054950FA1DFA48FA49" box="[1369,1430,1459,1484]" name="India" pageId="2" pageNumber="197">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="569798DA83237C054CCEFA79FDEEF9BD" pageId="2" pageNumber="197" type="etymology">
<paragraph id="1E32CB5183237C054CCEFA79FDEEF9BD" blockId="2.[151,1436,1387,2024]" pageId="2" pageNumber="197">
<emphasis id="2CF9174383237C054CCEFA79FE90FA75" bold="true" box="[199,334,1495,1520]" pageId="2" pageNumber="197">Etymology.</emphasis>
The new species is named in honor of Professor Raman Vadilal Shah, former Head of the Department of Zoology, The Maharaja Sayajirao University of Baroda, in Vadodara,
<collectingRegion id="DC4905B383237C05484EFA55FB40F991" box="[1095,1182,1531,1556]" country="India" name="Gujarat" pageId="2" pageNumber="197">Gujarat</collectingRegion>
,
<collectingCountry id="669A8BC183237C0548A0FA55FB3BF991" box="[1193,1253,1531,1556]" name="India" pageId="2" pageNumber="197">India</collectingCountry>
, for his valuable contributions in the field of zoology.
</paragraph>
</subSubSection>
<subSubSection id="569798DA83237C054CCEF9EDFD7BF86D" pageId="2" pageNumber="197" type="discussion">
<paragraph id="1E32CB5183237C054CCEF9EDFD7BF86D" blockId="2.[151,1436,1387,2024]" pageId="2" pageNumber="197">
<emphasis id="2CF9174383237C054CCEF9EDFEE5F9D9" bold="true" box="[199,315,1603,1628]" pageId="2" pageNumber="197">Remarks.</emphasis>
On the basis of the relative lengths of the dactyli of the longer P4 and P5, it is possible to recognize two subgroups within the genus
<taxonomicName id="D98DB0D283237C054E07F9C6FD59F905" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[526,647,1640,1664]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="genus">
<emphasis id="2CF9174383237C054E07F9C6FD59F905" box="[526,647,1640,1664]" italics="true" pageId="2" pageNumber="197">Arcotheres</emphasis>
</taxonomicName>
. The larger subgroup, contains 23 species: viz.
<taxonomicName id="D98DB0D283237C0548B8F9C6FF06F921" authority="(Rathbun, 1909)" authorityName="Rathbun" authorityYear="1909" baseAuthorityName="Rathbun" baseAuthorityYear="1909" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="2CF9174383237C0548B8F9C6FAFDF905" box="[1201,1315,1640,1664]" italics="true" pageId="2" pageNumber="197">A. alcocki</emphasis>
(Rathbun, 1909)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054CE1F922FDEDF921" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[232,563,1675,1700]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="acrophilus">
<emphasis id="2CF9174383237C054CE1F922FE5FF921" box="[232,385,1676,1700]" italics="true" pageId="2" pageNumber="197">A. acrophilus</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054D98F922FDF4F921" box="[401,554,1675,1700]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054E48F922FC86F921" authority="(Sakai, 1939)" authorityName="Sakai" authorityYear="1939" baseAuthorityName="Sakai" baseAuthorityYear="1939" box="[577,856,1675,1700]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="atrinae">
<emphasis id="2CF9174383237C054E48F922FD6DF926" box="[577,691,1676,1699]" italics="true" pageId="2" pageNumber="197">A. atrinae</emphasis>
(Sakai, 1939)
</taxonomicName>
;
<taxonomicName id="D98DB0D283237C054F6FF922FB6BF921" authority="(Nobili, 1905)" authorityName="Nobili" authorityYear="1905" baseAuthorityName="Nobili" baseAuthorityYear="1905" box="[870,1205,1675,1700]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="borradailei">
<emphasis id="2CF9174383237C054F6FF922FBD6F921" box="[870,1032,1676,1700]" italics="true" pageId="2" pageNumber="197">A. borradailei</emphasis>
(Nobili, 1905)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C0548CAF922FEBBF94D" authority="Antony &amp; Kuttyamma, 1971" authorityName="Antony &amp; Kuttyamma" authorityYear="1971" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="casta">
<emphasis id="2CF9174383237C0548CAF922FAC1F926" box="[1219,1311,1676,1699]" italics="true" pageId="2" pageNumber="197">A. casta</emphasis>
Antony &amp; Kuttyamma, 1971
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054D7BF91EFD6EF94D" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[370,688,1711,1736]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="coarcatus">
<emphasis id="2CF9174383237C054D7BF91EFDDEF94D" box="[370,512,1712,1736]" italics="true" pageId="2" pageNumber="197">A. coarcatus</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054E19F91EFD76F94D" box="[528,680,1711,1736]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054EB7F91EFC09F94D" authority="(Shen, 1932)" authorityName="Shen" authorityYear="1932" baseAuthorityName="Shen" baseAuthorityYear="1932" box="[702,983,1711,1736]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="cyclinus">
<emphasis id="2CF9174383237C054EB7F91EFCE4F94D" box="[702,826,1712,1736]" italics="true" pageId="2" pageNumber="197">A. cyclinus</emphasis>
(Shen, 1932)
</taxonomicName>
;
<taxonomicName id="D98DB0D283237C054FECF91EFAD2F94D" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[997,1292,1711,1736]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="exiguus">
<emphasis id="2CF9174383237C054FECF91EFB85F942" box="[997,1115,1712,1735]" italics="true" pageId="2" pageNumber="197">A. exiguus</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054862F91EFADAF94D" box="[1131,1284,1711,1736]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054910F91EFEE0F969" authority="Campos, 2001" authorityName="Campos" authorityYear="2001" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="guinotae">
<emphasis id="2CF9174383237C054910F91EFA42F942" box="[1305,1436,1712,1735]" italics="true" pageId="2" pageNumber="197">A. guinotae</emphasis>
Campos, 2001
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054D45F97AFDA0F969" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[332,638,1747,1772]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="2CF9174383237C054D45F97AFE15F969" box="[332,459,1748,1772]" italics="true" pageId="2" pageNumber="197">A. latifrons</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054DD5F97AFDA8F969" box="[476,630,1747,1772]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054E84F97AFC46F969" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[653,920,1747,1772]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="2CF9174383237C054E84F97AFD3AF969" box="[653,740,1748,1772]" italics="true" pageId="2" pageNumber="197">A. latus</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054EFFF97AFC4EF969" box="[758,912,1747,1772]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054FAEF97AFADFF969" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[935,1281,1747,1772]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="modiolicola">
<emphasis id="2CF9174383237C054FAEF97AFB91F969" box="[935,1103,1748,1772]" italics="true" pageId="2" pageNumber="197">A. modiolicola</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054856F97AFB27F969" box="[1119,1273,1747,1772]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054919F97AFE9CF895" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="nudifrons">
<emphasis id="2CF9174383237C054919F97AFA45F969" box="[1296,1435,1748,1772]" italics="true" pageId="2" pageNumber="197">A. nudifrons</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054C97F956FEE7F895" box="[158,313,1783,1808]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054D5BF956FD4AF895" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[338,660,1783,1808]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="palaensis">
<emphasis id="2CF9174383237C054D5BF956FE01F895" box="[338,479,1784,1808]" italics="true" pageId="2" pageNumber="197">A. palaensis</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054DF8F956FD52F895" box="[497,652,1783,1808]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054EADF956FB55F895" authority="(Hornell &amp; Southwell, 1909)" authorityName="Hornell &amp; Southwell" authorityYear="1909" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[676,1163,1783,1808]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383237C054EADF956FCF0F895" box="[676,814,1784,1808]" italics="true" pageId="2" pageNumber="197">A. placunae</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054F49F959FB5CF895" box="[832,1154,1783,1808]" pageId="2" pageNumber="197" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell, 1909</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054892F956FF04F8B1" authority="(Alcock, 1900)" authorityName="Alcock" authorityYear="1900" baseAuthorityName="Alcock" baseAuthorityYear="1900" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="purpureus">
<emphasis id="2CF9174383237C054892F956FAEEF895" box="[1179,1328,1784,1808]" italics="true" pageId="2" pageNumber="197">A. purpureus</emphasis>
(Alcock, 1900)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054CE2F8B2FDE9F8B1" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[235,567,1819,1844]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="rhombifer">
<emphasis id="2CF9174383237C054CE2F8B2FEA0F8B1" box="[235,382,1820,1844]" italics="true" pageId="2" pageNumber="197">A. rhombifer</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054D9AF8B2FDF0F8B1" box="[403,558,1819,1844]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054E41F8B2FC66F8B1" authority="(Southwell, 1911)" authorityName="Southwell" authorityYear="1911" baseAuthorityName="Southwell" baseAuthorityYear="1911" box="[584,952,1819,1844]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="ridgewayi">
<emphasis id="2CF9174383237C054E41F8B2FD02F8B1" box="[584,732,1820,1844]" italics="true" pageId="2" pageNumber="197">A. ridgewayi</emphasis>
(Southwell, 1911)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054FC0F8B2FAC0F8B1" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[969,1310,1819,1844]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="rotundatus">
<emphasis id="2CF9174383237C054FC0F8B2FBBBF8B1" box="[969,1125,1820,1844]" italics="true" pageId="2" pageNumber="197">A. rotundatus</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054870F8B2FACBF8B1" box="[1145,1301,1819,1844]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054927F8B2FE98F8DD" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="2CF9174383237C054927F8B2FA42F8B1" box="[1326,1436,1820,1844]" italics="true" pageId="2" pageNumber="197">A. similis</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054C97F8EEFEE0F8DD" box="[158,318,1855,1880]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054D52F8EEFD58F8DD" authority="(Shen, 1932)" authorityName="Shen" authorityYear="1932" baseAuthorityName="Shen" baseAuthorityYear="1932" box="[347,646,1855,1880]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="2CF9174383237C054D52F8EEFE05F8D2" box="[347,475,1856,1879]" italics="true" pageId="2" pageNumber="197">A. sinensis</emphasis>
(Shen, 1932)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054E93F8EEFBC2F8DD" authority="(Gordon, 1936)" authorityName="Gordon" authorityYear="1936" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[666,1052,1855,1880]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="spinidactylus">
<emphasis id="2CF9174383237C054E93F8EEFC88F8DD" box="[666,854,1856,1880]" italics="true" pageId="2" pageNumber="197">A. spinidactylus</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054F64F891FBCAF8DD" box="[877,1044,1855,1880]" pageId="2" pageNumber="197" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon, 1936</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D98DB0D283237C05486AF8EEFA42F8DD" authority="(Gordon, 1936)" authorityName="Gordon" authorityYear="1936" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[1123,1436,1855,1880]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="tivelae">
<emphasis id="2CF9174383237C05486AF8EEFB08F8DD" box="[1123,1238,1856,1880]" italics="true" pageId="2" pageNumber="197">A. tivelae</emphasis>
(
<bibRefCitation id="7A1CB6A083237C0548E4F891FA4AF8DD" box="[1261,1428,1855,1880]" pageId="2" pageNumber="197" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon, 1936</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="7A1CB6A083237C054C96F8CDFEF7F8F9" box="[159,297,1891,1916]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger1895</bibRefCitation>
;
<bibRefCitation id="7A1CB6A083237C054D33F8CDFDA7F8F9" box="[314,633,1891,1916]" pageId="2" pageNumber="197" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
;
<bibRefCitation id="7A1CB6A083237C054E82F8CDFCF7F8F9" box="[651,809,1891,1916]" pageId="2" pageNumber="197" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
;
<bibRefCitation id="7A1CB6A083237C054F32F8CDFBF1F8F9" box="[827,1071,1891,1916]" pageId="2" pageNumber="197" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
; Ng
<emphasis id="2CF9174383237C054879F8CAFB71F8F9" box="[1136,1199,1892,1916]" italics="true" pageId="2" pageNumber="197">et al.</emphasis>
2017). The smaller subgroup contains only three species —
<taxonomicName id="D98DB0D283237C054E6FF826FC7DF825" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[614,931,1927,1952]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383237C054E6FF826FD2DF825" box="[614,755,1928,1952]" italics="true" pageId="2" pageNumber="197">A. pernicola</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054F0AF826FC45F825" box="[771,923,1927,1952]" pageId="2" pageNumber="197" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="D98DB0D283237C054FB8F826FACBF825" authority="(Gordon, 1936)" authorityName="Gordon" authorityYear="1936" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[945,1301,1927,1952]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383237C054FB8F826FB83F825" box="[945,1117,1928,1952]" italics="true" pageId="2" pageNumber="197">A. winckworthi</emphasis>
(
<bibRefCitation id="7A1CB6A083237C054864F829FAD2F825" box="[1133,1292,1927,1952]" pageId="2" pageNumber="197" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon, 1936</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="D98DB0D283237C054947F826FDE2F841" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="2CF9174383237C054947F826FA42F81A" box="[1358,1436,1928,1951]" italics="true" pageId="2" pageNumber="197">A. rayi</emphasis>
<bibRefCitation id="7A1CB6A083237C054C9EF805FE5FF841" box="[151,385,1963,1988]" pageId="2" pageNumber="197" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng, 2007</bibRefCitation>
(Campos 2001)
</taxonomicName>
.
<taxonomicName id="D98DB0D283237C054E41F802FCD8F841" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[584,774,1964,1988]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="197" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383237C054E41F802FCD8F841" box="[584,774,1964,1988]" italics="true" pageId="2" pageNumber="197">Arcotheres shahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37CAAA3883237C054F07F805FC90F841" box="[782,846,1963,1988]" pageId="2" pageNumber="197" rank="species">
<emphasis id="2CF9174383237C054F07F805FC90F841" bold="true" box="[782,846,1963,1988]" pageId="2" pageNumber="197">n. sp.</emphasis>
</taxonomicNameLabel>
belongs to the smaller subgroup on the basis of the relative lengths of the dactyli of the P4 and P5.
</paragraph>
</subSubSection>
<subSubSection id="569798DA83227C034CCEFF36FD52F877" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198" type="reference_group">
<paragraph id="1E32CB5183227C044CCEFF36FDCCFDB9" blockId="3.[151,1436,151,573]" pageId="3" pageNumber="198">
<taxonomicName id="D98DB0D283227C044CCEFF36FE5AFF35" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[199,388,152,176]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383227C044CCEFF36FE5AFF35" box="[199,388,152,176]" italics="true" pageId="3" pageNumber="198">Arcotheres shahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37CAAA3883227C044D82FF39FE14FF35" box="[395,458,151,176]" pageId="3" pageNumber="198" rank="species">
<emphasis id="2CF9174383227C044D82FF39FE14FF35" bold="true" box="[395,458,151,176]" pageId="3" pageNumber="198">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="D98DB0D283227C044E69FF36FC46FF35" ID-CoL="5W29W" authority="(Burger, 1895)" authorityName="Burger" authorityYear="1895" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[608,920,151,176]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C044E69FF36FD35FF35" box="[608,747,152,176]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
(
<bibRefCitation id="7A1CB6A083227C044EF3FF36FC51FF35" box="[762,911,151,176]" pageId="3" pageNumber="198" refId="ref4007" refString="Burger, O. (1895) Ein Beitragzur Kenntniss der Pinnotherinen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 8, 361 - 390." type="journal article">Bürger, 1895</bibRefCitation>
)
</taxonomicName>
(from Ubay, northeastern
<collectingRegion id="DC4905B383227C0448CBFF39FAD7FF35" box="[1218,1289,151,176]" country="Philippines" name="Bohol" pageId="3" pageNumber="198">Bohol</collectingRegion>
,
<collectingCountry id="669A8BC183227C04491CFF39FA4BFF35" box="[1301,1429,151,176]" name="Philippines" pageId="3" pageNumber="198">Philippines</collectingCountry>
) in the following characters: in the female, the carapace is subtrapezoidal (
<figureCitation id="86B6D7D483227C044FD9FF12FBF8FF51" box="[976,1062,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="3" pageNumber="198">Fig. 1A</figureCitation>
) (vs. subquadrate in
<taxonomicName id="D98DB0D283227C044918FF12FA45FF51" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1297,1435,188,212]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C044918FF12FA45FF51" box="[1297,1435,188,212]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044CC5FF71FE6DFF7D" box="[204,435,223,248]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
; fig. 10F), the front is slightly produced (
<figureCitation id="86B6D7D483227C044F84FF71FC3BFF7D" box="[909,997,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="3" pageNumber="198">Fig. 1A</figureCitation>
) (vs. not produced in
<taxonomicName id="D98DB0D283227C0448EAFF4EFAB1FF7D" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1251,1391,224,248]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C0448EAFF4EFAB1FF7D" box="[1251,1391,224,248]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044C9EFEAAFEA1FE99" box="[151,383,260,285]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
; fig. 10F), the flagellum of the MXP3 exopod has three articles before aesthetascs (
<figureCitation id="86B6D7D483227C044936FEAAFA4DFE99" box="[1343,1427,260,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="3" pageNumber="198">Fig. 1F</figureCitation>
) (vs. only two articles in
<taxonomicName id="D98DB0D283227C044DA0FE86FDEBFEC5" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[425,565,296,320]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C044DA0FE86FDEBFEC5" box="[425,565,296,320]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044E61FE89FC90FEC5" box="[616,846,295,320]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10G), the dactylus of the P1 chela is more than half the length of the palm (
<figureCitation id="86B6D7D483227C044DDBFEE2FD8BFEE1" box="[466,597,332,357]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="3" pageNumber="198">Figs. 1D, E</figureCitation>
) (vs. dactylus is half the length of the palm in
<taxonomicName id="D98DB0D283227C04486CFEE2FB2EFEE1" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1125,1264,332,356]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C04486CFEE2FB2EFEE1" box="[1125,1264,332,356]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C04492DFEE2FF20FE0D" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10A), the pollex has two large proximal teeth and is covered with long setae (
<figureCitation id="86B6D7D483227C0448AFFEC1FAF7FE0D" box="[1190,1321,367,392]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="3" pageNumber="198">Figs. 1D, E</figureCitation>
) (vs. with single large proximal tooth, pollex glabrous in
<taxonomicName id="D98DB0D283227C044EA1FE3AFCECFE29" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[680,818,404,428]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C044EA1FE3AFCECFE29" box="[680,818,404,428]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044F6FFE3AFB92FE29" box="[870,1100,404,429]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10A), the dactylus of the longer P4 is 1.26 times longer than the dactyli of the longer P5 (
<figureCitation id="86B6D7D483227C044F7BFE19FC26FE55" box="[882,1016,439,464]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="3" pageNumber="198">Figs. 3C, D</figureCitation>
) (vs. dactylus of the longer P4 about twice as long as dactyli of longer P
<quantity id="D97566B483227C044E28FE72FD95FE70" box="[545,587,476,501]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="198" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="D98DB0D283227C044E58FE72FD05FE71" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[593,731,476,500]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C044E58FE72FD05FE71" box="[593,731,476,500]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044F07FE72FC2CFE71" box="[782,1010,476,501]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10E, F), and the dactyli of the P2 and P3 are conical and terminally hooked (
<figureCitation id="86B6D7D483227C044E74FE51FCDFFD9D" box="[637,769,511,536]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="3" pageNumber="198">Figs. 3A, B</figureCitation>
) (vs. conical but not hooked terminally in
<taxonomicName id="D98DB0D283227C0448EDFDAEFAAEFD9D" ID-CoL="5W29W" baseAuthorityName="Burger" baseAuthorityYear="1895" box="[1252,1392,512,536]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="pernicola">
<emphasis id="2CF9174383227C0448EDFDAEFAAEFD9D" box="[1252,1392,512,536]" italics="true" pageId="3" pageNumber="198">A. pernicola</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044C9EFD8AFEA4FDB9" box="[151,378,548,573]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
: fig. 10B, C).
</paragraph>
<caption id="4AF29BD983227C044C9EF9BFFBE2F9C3" httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="3" pageNumber="198" startId="3.[151,250,1553,1575]" targetBox="[338,1264,642,1506]" targetPageId="3">
<paragraph id="1E32CB5183227C044C9EF9BFFBE2F9C3" blockId="3.[151,1436,1553,1606]" pageId="3" pageNumber="198">
<emphasis id="2CF9174383227C044C9EF9BFFECCF9A3" bold="true" box="[151,274,1553,1575]" pageId="3" pageNumber="198">FIGURE 2.</emphasis>
<taxonomicName id="D98DB0D283227C044D10F9BFFE1DF9A2" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[281,451,1553,1575]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383227C044D10F9BFFE1DF9A2" box="[281,451,1553,1575]" italics="true" pageId="3" pageNumber="198">Arcotheres shahi</emphasis>
</taxonomicName>
<emphasis id="2CF9174383227C044DC3F9BCFDD7F9A2" bold="true" box="[458,521,1554,1575]" pageId="3" pageNumber="198">
<taxonomicNameLabel id="37CAAA3883227C044DC3F9BCFDDDF9A2" box="[458,515,1554,1575]" pageId="3" pageNumber="198" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view.
</paragraph>
</caption>
<paragraph id="1E32CB5183227C044CCEF9C3FC72F823" blockId="3.[151,1436,1645,2030]" pageId="3" pageNumber="198">
<taxonomicName id="D98DB0D283227C044CCEF9C3FE59F900" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[199,391,1645,1669]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383227C044CCEF9C3FE59F900" box="[199,391,1645,1669]" italics="true" pageId="3" pageNumber="198">Arcotheres shahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37CAAA3883227C044D98F9C3FE0DF903" box="[401,467,1645,1670]" pageId="3" pageNumber="198" rank="species">
<emphasis id="2CF9174383227C044D98F9C3FE0DF903" bold="true" box="[401,467,1645,1670]" pageId="3" pageNumber="198">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="D98DB0D283227C044E79F9C0FC07F903" ID-CoL="67PY7" authority="(Gordon, 1936)" authorityName="Gordon" authorityYear="1936" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[624,985,1645,1670]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383227C044E79F9C0FCC3F900" box="[624,797,1645,1669]" italics="true" pageId="3" pageNumber="198">A. winckworthi</emphasis>
(
<bibRefCitation id="7A1CB6A083227C044F26F9C3FC0EF903" box="[815,976,1645,1670]" pageId="3" pageNumber="198" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon, 1936</bibRefCitation>
)
</taxonomicName>
(from
<collectingRegion id="DC4905B383227C044824F9C0FB5CF903" box="[1069,1154,1646,1670]" country="Malaysia" name="Pulau Pinang" pageId="3" pageNumber="198">Penang</collectingRegion>
,
<collectingCountry id="669A8BC183227C044899F9C3FB26F903" box="[1168,1272,1645,1670]" name="Malaysia" pageId="3" pageNumber="198">Malaysia</collectingCountry>
) (known only from females) in the following characters: the posterolateral margin of carapace is gently convex (vs. posterolateral margin concave in
<taxonomicName id="D98DB0D283227C044D67F918FDC6F948" ID-CoL="67PY7" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[366,536,1717,1741]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383227C044D67F918FDC6F948" box="[366,536,1717,1741]" italics="true" pageId="3" pageNumber="198">A. winckworthi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044E45F91BFD38F94B" box="[588,742,1717,1742]" pageId="3" pageNumber="198" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 7a), the chelae are globose, with the palms height about 1.7 times its length (
<figureCitation id="86B6D7D483227C044D70F977FE27F977" box="[377,505,1753,1778]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="3" pageNumber="198">Figs. 1D, E</figureCitation>
) (vs. chelae slender, palm height being about 2.4 times its length in
<taxonomicName id="D98DB0D283227C0448E2F974FA4AF977" ID-CoL="67PY7" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[1259,1428,1754,1778]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383227C0448E2F974FA4AF977" box="[1259,1428,1754,1778]" italics="true" pageId="3" pageNumber="198">A. winckworthi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044CB5F953FE89F893" box="[188,343,1789,1814]" pageId="3" pageNumber="198" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 7c), the relative lengths of ambulatory legs are ranked as P4&gt;P3&gt;P2&gt;P5 (
<figureCitation id="86B6D7D483227C0448D2F953FABCF893" box="[1243,1378,1789,1814]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="3" pageNumber="198">Figs. 3AD</figureCitation>
) (vs. P4&gt;P5&gt;P3&gt;P
<quantity id="D97566B483227C044D26F88FFE82F8BF" box="[303,348,1825,1850]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="198" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="D98DB0D283227C044D6FF88CFDCCF8BF" ID-CoL="67PY7" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[358,530,1826,1850]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383227C044D6FF88CFDCCF8BF" box="[358,530,1826,1850]" italics="true" pageId="3" pageNumber="198">A. winckworthi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044E42F88FFD38F8BF" box="[587,742,1825,1850]" pageId="3" pageNumber="198" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 7a), the propodi of the P3P5 are without setae on the ventral margin (
<figureCitation id="86B6D7D483227C044D45F8EBFE1BF8DB" box="[332,453,1861,1886]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="3" pageNumber="198">Fig. 3BD</figureCitation>
) (vs. propodi of P3P5 with setae in
<taxonomicName id="D98DB0D283227C044F67F8E8FBC7F8D8" ID-CoL="67PY7" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[878,1049,1861,1885]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383227C044F67F8E8FBC7F8D8" box="[878,1049,1861,1885]" italics="true" pageId="3" pageNumber="198">A. winckworthi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044844F8EBFB38F8DB" box="[1101,1254,1861,1886]" pageId="3" pageNumber="198" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 7a, d, e, f), and the dactyli of the P4 and P5 are sickle-like and the tip is not hooked (
<figureCitation id="86B6D7D483227C044FC2F8C7FBC8F807" box="[971,1046,1897,1922]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="3" pageNumber="198">Figs. 3</figureCitation>
C-D) (vs. dactyli of P4, P5 sickle - like, with hooked at tip in
<taxonomicName id="D98DB0D283227C044DB4F820FDB9F820" ID-CoL="67PY7" baseAuthorityName="Gordon" baseAuthorityYear="1936" box="[445,615,1933,1957]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="winckworthi">
<emphasis id="2CF9174383227C044DB4F820FDB9F820" box="[445,615,1933,1957]" italics="true" pageId="3" pageNumber="198">A. winckworthi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083227C044E93F823FCECF823" box="[666,818,1933,1958]" pageId="3" pageNumber="198" refId="ref4175" refString="Gordon, I. (1936) On a few Indo-Pacific species of Pinnotheres, with special reference to asymmetry of the walking legs. Journal of Linnaean Society London, Zoology, 40 (209), 163 - 180. https: // doi. org / 10.1111 / j. 1096 - 3642.1936. tb 01683. x" type="journal article">Gordon 1936</bibRefCitation>
: fig. 7e, f).
</paragraph>
<paragraph id="1E32CB5183227C034CCEF81CFE97FEC5" blockId="3.[151,1436,1645,2030]" lastBlockId="4.[151,1436,151,320]" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198">
<taxonomicName id="D98DB0D283227C044CCEF81CFE59F84F" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[199,391,1970,1994]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383227C044CCEF81CFE59F84F" box="[199,391,1970,1994]" italics="true" pageId="3" pageNumber="198">Arcotheres shahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37CAAA3883227C044D9AF81FFE08F84F" box="[403,470,1969,1994]" pageId="3" pageNumber="198" rank="species">
<emphasis id="2CF9174383227C044D9AF81FFE08F84F" bold="true" box="[403,470,1969,1994]" pageId="3" pageNumber="198">n. sp.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="D98DB0D283227C044E71F81CFC16F84F" ID-CoL="67PY8" authority="Ahyong &amp; Ng, 2007" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[632,968,1969,1994]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="2CF9174383227C044E71F81CFD17F84C" box="[632,713,1970,1993]" italics="true" pageId="3" pageNumber="198">A. rayi</emphasis>
<bibRefCitation id="7A1CB6A083227C044EDDF81FFC16F84F" box="[724,968,1969,1994]" pageId="3" pageNumber="198" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng, 2007</bibRefCitation>
</taxonomicName>
(from Mariveles or
<collectingRegion id="DC4905B383227C0448B4F81FFADAF84F" box="[1213,1284,1969,1994]" country="Philippines" name="Bohol" pageId="3" pageNumber="198">Bohol</collectingRegion>
,
<collectingCountry id="669A8BC183227C04491DF81FFA4AF84F" box="[1300,1428,1969,1994]" name="Philippines" pageId="3" pageNumber="198">Philippines</collectingCountry>
) (known only from females) in the following characters: the frontal margin of the carapace is gently convex (
<figureCitation id="86B6D7D483227C034966F87BFF67FF35" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" lastPageId="4" lastPageNumber="199" pageId="3" pageNumber="198">Fig. 1A</figureCitation>
) (vs. flattened in
<taxonomicName id="D98DB0D283257C034D8FFF36FE0AFF2A" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[390,468,152,175]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="2CF9174383257C034D8FFF36FE0AFF2A" box="[390,468,152,175]" italics="true" pageId="4" pageNumber="199">A. rayi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083257C034E00FF39FD2EFF35" box="[521,752,151,176]" pageId="4" pageNumber="199" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
; fig. 11A), the cheliped palm is relatively shorter, about 1.6 times as long as the fingers (
<figureCitation id="86B6D7D483257C034DEFFF12FDB3FF51" box="[486,621,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="4" pageNumber="199">Figs. 1D, E</figureCitation>
) (vs. palm elongated, about twice as long as the fingers in
<taxonomicName id="D98DB0D283257C034916FF12FAB0FF51" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1311,1390,188,212]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="2CF9174383257C034916FF12FAB0FF51" box="[1311,1390,188,212]" italics="true" pageId="4" pageNumber="199">A. rayi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083257C034C9EFF71FEA6FF7D" box="[151,376,223,248]" pageId="4" pageNumber="199" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
; fig. 11D), the telson is rhomboidal with the proximal and distal margins convex (
<figureCitation id="86B6D7D483257C034901FF71FA82FF7D" box="[1288,1372,223,248]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="4" pageNumber="199">Fig. 1C</figureCitation>
) (vs. telson crescent-shaped, the proximal margin is concave and the distal margin convex in
<taxonomicName id="D98DB0D283257C034870FEAAFB18FE99" ID-CoL="67PY8" authorityName="Ahyong &amp; Ng" authorityYear="2007" box="[1145,1222,260,284]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="rayi">
<emphasis id="2CF9174383257C034870FEAAFB18FE99" box="[1145,1222,260,284]" italics="true" pageId="4" pageNumber="199">A. rayi</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083257C0348F3FEAAFF0DFEC5" pageId="4" pageNumber="199" refId="ref3948" refString="Ahyong, S. T. &amp; Ng, P. K. L. (2007) The pinnotherid type material of Semper (1880), Nauck (1880) and Burger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 16 (Supplement), 191 - 226." type="journal article">Ahyong &amp; Ng 2007</bibRefCitation>
; fig. 11C).
</paragraph>
<caption id="4AF29BD983257C034C9EFB5BFE7DFACC" httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="4" pageNumber="199" startId="4.[151,250,1269,1291]" targetBox="[288,1255,362,1232]" targetPageId="4">
<paragraph id="1E32CB5183257C034C9EFB5BFE7DFACC" blockId="4.[151,1436,1269,1353]" pageId="4" pageNumber="199">
<emphasis id="2CF9174383257C034C9EFB5BFECDFA8E" bold="true" box="[151,275,1269,1291]" pageId="4" pageNumber="199">FIGURE 3.</emphasis>
<taxonomicName id="D98DB0D283257C034D12FB5BFE18FA8E" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[283,454,1269,1291]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383257C034D12FB5BFE18FA8E" box="[283,454,1269,1291]" italics="true" pageId="4" pageNumber="199">Arcotheres shahi</emphasis>
</taxonomicName>
<emphasis id="2CF9174383257C034DC7FB58FDD0FA8E" bold="true" box="[462,526,1270,1291]" pageId="4" pageNumber="199">
<taxonomicNameLabel id="37CAAA3883257C034DC7FB58FDD6FA8E" box="[462,520,1270,1291]" pageId="4" pageNumber="199" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view.
<taxonomicName id="D98DB0D283257C034E1AFABAFD63FAAF" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[531,701,1300,1322]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383257C034E1AFABAFD63FAAF" box="[531,701,1300,1322]" italics="true" pageId="4" pageNumber="199">Arcotheres shahi</emphasis>
</taxonomicName>
<emphasis id="2CF9174383257C034ECAFABBFCDCFAAF" bold="true" box="[707,770,1301,1322]" pageId="4" pageNumber="199">
<taxonomicNameLabel id="37CAAA3883257C034ECAFABBFD22FAAF" box="[707,764,1301,1322]" pageId="4" pageNumber="199" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view.
</paragraph>
</caption>
<paragraph id="1E32CB5183257C034CCEFADBFD52F877" blockId="4.[151,1437,1397,2034]" pageId="4" pageNumber="199">
<taxonomicName id="D98DB0D283257C034CCEFADBFE5DFA08" ID-CoL="84BWP" authority="Trivedi &amp; Campos &amp; Vachhrajani, 2018" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[199,387,1397,1421]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="shahi" status="sp. nov.">
<emphasis id="2CF9174383257C034CCEFADBFE5DFA08" box="[199,387,1397,1421]" italics="true" pageId="4" pageNumber="199">Arcotheres shahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="37CAAA3883257C034D83FAD8FE16FA0B" box="[394,456,1397,1422]" pageId="4" pageNumber="199" rank="species">
<emphasis id="2CF9174383257C034D83FAD8FE47FA08" bold="true" box="[394,409,1398,1421]" italics="true" pageId="4" pageNumber="199">n</emphasis>
. sp.
</taxonomicNameLabel>
is superficially similar to
<taxonomicName id="D98DB0D283257C034EE5FAD8FB64FA0B" ID-CoL="67PXW" authority="(Hornell &amp; Southwell, 1909)" authorityName="Hornell &amp; Southwell" authorityYear="1909" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[748,1210,1397,1422]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034EE5FAD8FCAFFA08" box="[748,881,1397,1421]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
(
<bibRefCitation id="7A1CB6A083257C034F76FADBFB6CFA0B" box="[895,1202,1397,1422]" pageId="4" pageNumber="199" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell, 1909</bibRefCitation>
)
</taxonomicName>
(from
<collectingRegion id="DC4905B383257C03490EFADBFA85FA0B" box="[1287,1371,1397,1422]" country="India" name="Gujarat" pageId="4" pageNumber="199">Gujarat</collectingRegion>
state,
<collectingCountry id="669A8BC183257C034C9EFA37FF0BFA37" box="[151,213,1433,1458]" name="India" pageId="4" pageNumber="199">India</collectingCountry>
), of the larger subgroup of
<taxonomicName id="D98DB0D283257C034E1AFA34FD52FA37" ID-CoL="7NQY7" authorityName="Trivedi &amp; Campos &amp; Vachhrajani" authorityYear="2018" box="[531,652,1434,1458]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="genus">
<emphasis id="2CF9174383257C034E1AFA34FD52FA37" box="[531,652,1434,1458]" italics="true" pageId="4" pageNumber="199">Arcotheres</emphasis>
</taxonomicName>
, in having a subtrapezoidal carapace and having the dactylus of the MXP3 inserted on the ventral margin of the propodus and not reaching its tip. It differs in the following characters: in the female, the frontal margin of the carapace is gently convex (
<figureCitation id="86B6D7D483257C034F7EFA4FFC12FA7F" box="[887,972,1505,1530]" captionStart="FIGURE 1" captionStartId="2.[151,250,1295,1317]" captionTargetBox="[348,1277,402,1273]" captionTargetId="figure@2.[304,1282,383,1290]" captionTargetPageId="2" captionText="FIGURE 1. Arcotheres shahi n. sp., holotype, female (CL 6.13 mm, CW 6.77 mm) (ZL-AR-CR-97). A, habitus, dorsal view; B, frontal view; C, pleon and telson; D, right chela, outer view; E, right chela, inner view; F, left MXP3, inner view." httpUri="https://zenodo.org/record/1287087/files/figure.png" pageId="4" pageNumber="199">Fig. 1A</figureCitation>
) (vs. flattened in
<taxonomicName id="D98DB0D283257C034899FA4CFACBFA7F" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[1168,1301,1506,1530]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034899FA4CFACBFA7F" box="[1168,1301,1506,1530]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; cf.
<bibRefCitation id="7A1CB6A083257C03494FFA4FFEB7F99B" pageId="4" pageNumber="199" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
: fig. 5; Trivedi
<emphasis id="2CF9174383257C034E1EF9A8FD8EF998" box="[535,592,1541,1565]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press: figs. 1A, 2A), the dactyli of the left P2 and P3 are glabrous (
<figureCitation id="86B6D7D483257C03496AF9ABFF03F9C7" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="4" pageNumber="199">Figs. 3A, B</figureCitation>
) (vs. ventral margin covered with short setae in
<taxonomicName id="D98DB0D283257C034F1DF984FC43F9C7" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[788,925,1578,1602]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034F1DF984FC43F9C7" box="[788,925,1578,1602]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; cf. Trivedi
<emphasis id="2CF9174383257C034839F984FBB5F9C7" box="[1072,1131,1578,1602]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press: figs. 4A, B), the propodi of the left P2P5 are glabrous (
<figureCitation id="86B6D7D483257C034E68F9E3FD37F9E3" box="[609,745,1613,1638]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="4" pageNumber="199">Figs. 3AD</figureCitation>
) (vs. left P2P5 propodi ventral margin with scattered short setae distally in
<taxonomicName id="D98DB0D283257C034D44F9DCFE0CF90F" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[333,466,1650,1674]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034D44F9DCFE0CF90F" box="[333,466,1650,1674]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; cf. Trivedi
<emphasis id="2CF9174383257C034E57F9DCFD49F90F" box="[606,663,1650,1674]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press: figs. 4AD), the dactylus of the left P5 has short setae only on the ventral margin distally (vs. entire dactylus covered with short setae in
<taxonomicName id="D98DB0D283257C034FF8F938FBA8F928" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[1009,1142,1685,1709]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034FF8F938FBA8F928" box="[1009,1142,1685,1709]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; cf. Trivedi
<emphasis id="2CF9174383257C034909F938FAE9F928" box="[1280,1335,1685,1709]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press: fig. 4D); in the male, the carapace is subovate and longer than broad (
<figureCitation id="86B6D7D483257C034F92F917FC2DF957" box="[923,1011,1721,1746]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="4" pageNumber="199">Fig. 2A</figureCitation>
) (vs. subhexagonal, broader than long in
<taxonomicName id="D98DB0D283257C034CBFF970FEE3F970" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[182,317,1757,1781]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034CBFF970FEE3F970" box="[182,317,1757,1781]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; c.f.
<bibRefCitation id="7A1CB6A083257C034D72F973FD71F973" box="[379,687,1757,1782]" pageId="4" pageNumber="199" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
; fig. 1; Trivedi
<emphasis id="2CF9174383257C034F6CF970FC41F970" box="[869,927,1757,1781]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press; figs. 2C, 3A), the front is slightly concave medially (
<figureCitation id="86B6D7D483257C034D64F8AFFE18F89F" box="[365,454,1793,1818]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="4" pageNumber="199">Fig. 2A</figureCitation>
) (vs. front almost flat in
<taxonomicName id="D98DB0D283257C034EEAF8ACFCB4F89F" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[739,874,1794,1818]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034EEAF8ACFCB4F89F" box="[739,874,1794,1818]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; c.f.
<bibRefCitation id="7A1CB6A083257C034FACF8AFFB08F89F" box="[933,1238,1793,1818]" pageId="4" pageNumber="199" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
; fig. 1; Trivedi
<emphasis id="2CF9174383257C034981F8ACFF6AF8B8" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press; figs. 2C, 3A), there is no well-marked cervical groove on the dorsal surface of the carapace (
<figureCitation id="86B6D7D483257C034935F88BFA4AF8BB" box="[1340,1428,1829,1854]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="4" pageNumber="199">Fig. 2A</figureCitation>
) (vs. well-marked cervical grooves present on carapace in
<taxonomicName id="D98DB0D283257C034F37F8E4FC17F8E7" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[830,969,1866,1890]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034F37F8E4FC17F8E7" box="[830,969,1866,1890]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; c.f.
<bibRefCitation id="7A1CB6A083257C034802F8E7FA9BF8E7" box="[1035,1349,1865,1890]" pageId="4" pageNumber="199" refId="ref4236" refString="Hornell, J. &amp; Southwell, T. (1909) Description of a new species of Pinnotheres from Placuna placenta, with a note on the genus. In: Hornell, J. (Ed.), Report to the Government of Baroda on the Marine Zoology of Okhamandal in Kattiawar. Part 1. Richard Clay &amp; Sons Ltd., Bungay, Suffolk, pp. 99 - 103." type="book chapter">Hornell &amp; Southwell 1909</bibRefCitation>
; fig. 1; Trivedi
<emphasis id="2CF9174383257C034CE6F8C0FEF9F800" box="[239,295,1901,1925]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press; figs. 2C, 3A), the dactyli of left P2, P3 are glabrous (
<figureCitation id="86B6D7D483257C034FE2F8C3FBB0F800" box="[1003,1134,1901,1926]" captionStart="FIGURE 3" captionStartId="4.[151,250,1269,1291]" captionTargetBox="[288,1255,362,1232]" captionTargetId="figure@4.[269,1317,341,1248]" captionTargetPageId="4" captionText="FIGURE 3. Arcotheres shahi n. sp., holotype, female (CL 6.13mm, CW 6.77 mm) (ZL-AR-CR-97). A- D, left P2P5 dorsal view; A- C, right P2P4 outer view. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). E H, left P2-P5, dorsal view." httpUri="https://zenodo.org/record/1287091/files/figure.png" pageId="4" pageNumber="199">Figs. 3A, B</figureCitation>
) (vs. dactyli of left P2P5 setose on dorsal and ventral margin in
<taxonomicName id="D98DB0D283257C034E5BF83CFD07F82F" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" box="[594,729,1938,1962]" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C034E5BF83CFD07F82F" box="[594,729,1938,1962]" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; cf. Trivedi
<emphasis id="2CF9174383257C034F60F83CFC7DF82F" box="[873,931,1938,1962]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press; fig. 4AD), and the G1 is strongly curved and its dorsal margin is glabrous (
<figureCitation id="86B6D7D483257C034E66F81BFD16F84B" box="[623,712,1973,1998]" captionStart="FIGURE 2" captionStartId="3.[151,250,1553,1575]" captionTargetBox="[338,1264,642,1506]" captionTargetId="figure@3.[305,1282,625,1532]" captionTargetPageId="3" captionText="FIGURE 2. Arcotheres shahi n. sp., paratype, male (CL 3.43 mm, CW 3.30 mm) (ZL-AR-CR-98). A, habitus, dorsal view; B, right chela outer view; C, pleon and telson; D, G1, abdominal view; E, G2, abdominal view." httpUri="https://zenodo.org/record/1287089/files/figure.png" pageId="4" pageNumber="199">Fig. 2D</figureCitation>
) (vs. G1 gently curved with fringe setae on dorsal margin in
<taxonomicName id="D98DB0D283257C03498DF818FF20F877" ID-CoL="67PXW" baseAuthorityName="Hornell &amp; Southwell" baseAuthorityYear="1909" class="Malacostraca" family="Pinnotheridae" genus="Arcotheres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="199" phylum="Arthropoda" rank="species" species="placunae">
<emphasis id="2CF9174383257C03498DF818FF20F877" italics="true" pageId="4" pageNumber="199">A. placunae</emphasis>
</taxonomicName>
; cf. Trivedi
<emphasis id="2CF9174383257C034D81F874FE1FF877" box="[392,449,2010,2034]" italics="true" pageId="4" pageNumber="199">et al.</emphasis>
in press; fig. 3D).
</paragraph>
</subSubSection>
</treatment>
</document>