treatments-xml/data/1D/6F/B7/1D6FB737FF93FFA9FF2FF9FF5129FF7C.xml

<document id="AF6D77087216A2258EFAD5E29967577E" ID-CLB-Dataset="46415" ID-DOI="10.5281/zenodo.204428" ID-GBIF-Dataset="b27a74a0-2042-4720-839a-c5176cb48a38" ID-ISSN="1175-5326" ID-Zenodo-Dep="204428" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460385671311" checkinUser="plazi" docAuthor="Bellis, Glenn &amp; Dyce, Alan" docDate="2011" docId="1D6FB737FF93FFA9FF2FF9FF5129FF7C" docLanguage="en" docName="zt03014p058.pdf" docOrigin="Zootaxa 3014" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Culicoides pseudostigmaticus Tokunaga" docType="treatment" docVersion="11" lastPageNumber="44" masterDocId="E156CF4FFF97FFA0FFB8FFEA5414FFA8" masterDocTitle="Marksomyia, a new subgenus of Culicoides Latreille (Diptera: Ceratopogonidae) from the Australasian biogeographic region with descriptions of two new species" masterLastPageNumber="38" masterPageNumber="35" pageNumber="39" updateTime="1698297491458" updateUser="ExternalLinkService">
<mods:mods id="1C513AFD9B64CE26B054B1F8854A9025" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7A12CC478B73705ED55128C5BC8AC1D6">
<mods:title id="D8D77FC5FB723683B4B31CCCC953F011">Marksomyia, a new subgenus of Culicoides Latreille (Diptera: Ceratopogonidae) from the Australasian biogeographic region with descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name id="2351669FC48993589EADDD82EA95949D" type="personal">
<mods:role id="AF4D86E1969589CDC9B1111DC9E10F1F">
<mods:roleTerm id="222E924EA4FCCB705375E598A2BBE367">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EB559C8E6A3DCE99FFF4C93A1CBF0EF0">Bellis, Glenn</mods:namePart>
</mods:name>
<mods:name id="FFD9A1FF4E22EADC7D64FEF539168C6B" type="personal">
<mods:role id="D427C8B9CDD0FCBB195DBF687B0C723D">
<mods:roleTerm id="C96D25A1C0078BAD60A43B63E2CFF740">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65D87793734CEFD7185AC12E68A00F4B">Dyce, Alan</mods:namePart>
</mods:name>
<mods:typeOfResource id="B80EBAFAAEE77F2F5C8779F666023F3A">text</mods:typeOfResource>
<mods:relatedItem id="377CB06912D75BCF753DB125C1ACBE7F" type="host">
<mods:titleInfo id="4C4A7102EE2417AF20CB5B81C252FA78">
<mods:title id="6B1BF3C3664F041FA6B4A05323821615">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="59E3F705B5A4ADA304767B0A672D0E87">
<mods:date id="C47D19C459C4B716EA4727FC6599C3FD">2011</mods:date>
<mods:detail id="C0D7108046A901350B4AF34BCCCE43B5" type="volume">
<mods:number id="43C1C9C995E9CDF479FB2D0997C756DB">3014</mods:number>
</mods:detail>
<mods:extent id="D0CF06CBB51B593A66C47A2C84EB8800" unit="page">
<mods:start id="E0D3645DC97C32F09BF875E13AA68728">35</mods:start>
<mods:end id="93D61183DA4E807F7FB39B4C181812AE">38</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="44AB7858569A2E1C394EBBD6A7375182">journal article</mods:classification>
<mods:identifier id="7D68EEBBB62D028AD56BDA8444E1D986" type="CLB-Dataset">46415</mods:identifier>
<mods:identifier id="9D6E1C5B49127A32E8CD2108DAB1730D" type="DOI">10.5281/zenodo.204428</mods:identifier>
<mods:identifier id="3AD623E8C3E548B13F7DC775759608AE" type="GBIF-Dataset">b27a74a0-2042-4720-839a-c5176cb48a38</mods:identifier>
<mods:identifier id="EA54C57EBAA462429AEB33D81B87190C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A1FBA09E070078A05F0F4B1681CA6E2F" type="Zenodo-Dep">204428</mods:identifier>
</mods:mods>
<treatment id="1D6FB737FF93FFA9FF2FF9FF5129FF7C" ID-DOI="http://doi.org/10.5281/zenodo.5673950" ID-GBIF-Taxon="119415367" ID-Zenodo-Dep="5673950" LSID="urn:lsid:plazi:treatment:1D6FB737FF93FFA9FF2FF9FF5129FF7C" httpUri="http://treatment.plazi.org/id/1D6FB737FF93FFA9FF2FF9FF5129FF7C" lastPageId="9" lastPageNumber="44" pageId="4" pageNumber="39">
<subSubSection id="DDDC55AAFF93FFA4FF2FF9FF5625F9F8" pageId="4" pageNumber="39" type="nomenclature">
<paragraph id="95790621FF93FFA4FF2FF9FF566AF987" blockId="4.[151,638,1557,1616]" box="[151,638,1557,1583]" pageId="4" pageNumber="39">
<heading id="CE31B14DFF93FFA4FF2FF9FF566AF987" bold="true" box="[151,638,1557,1583]" fontSize="11" level="1" pageId="4" pageNumber="39" reason="1">
<taxonomicName id="52C67DA2FF93FFA4FF2FF9FF566AF987" authority="Tokunaga" authorityName="Tokunaga" box="[151,638,1557,1583]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF93FFA4FF2FF9FF566AF987" bold="true" box="[151,638,1557,1583]" pageId="4" pageNumber="39">
<emphasis id="A7B2DA33FF93FFA4FF2FF9FF55EEF987" bold="true" box="[151,506,1557,1583]" italics="true" pageId="4" pageNumber="39">Culicoides pseudostigmaticus</emphasis>
Tokunaga
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="95790621FF93FFA4FF2FF9D25625F9F8" blockId="4.[151,638,1557,1616]" box="[151,561,1592,1616]" pageId="4" pageNumber="39">
(
<figureCitation id="0DFD1AA4FF93FFA4FF27F9D254F3F9F8" box="[159,231,1592,1616]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1399,1422]" captionTargetBox="[187,1398,218,1321]" captionTargetId="figure@4.[151,1436,193,1378]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Eye separation of female Culicoides (Marksomyia) species. 1. C. marksi Lee &amp; Reye. 2. C. pseudostigmaticus Tokunaga. 3. C. zentae Bellis &amp; Dyce sp. nov. 4. C. kayi Bellis &amp; Dyce sp. nov. 5. C. parvimaculatus Lee &amp; Reye. 6. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204429/files/figure.png" pageId="4" pageNumber="39">Figs 2</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FF4BF9D25517F9F8" box="[243,259,1592,1616]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[195,1309,231,1843]" captionTargetId="figure@5.[151,1436,193,1871]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Female Culicoides (Marksomyia) species. 7. C. marksi Lee &amp; Reye, mandibular teeth and palp. 8. C. pseudostigmaticus Tokunaga, mandibular teeth and palp. 9. C. zentae Bellis &amp; Dyce sp. nov., mandibular teeth and palp. 10. C. kayi Bellis &amp; Dyce sp. nov., mandibular teeth and palp. 11. C. parvimaculatus Lee &amp; Reye, mandibular teeth and palp. 12. C. dycei Lee &amp; Reye, mandibular teeth and palp." httpUri="https://zenodo.org/record/204430/files/figure.png" pageId="4" pageNumber="39">8</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FEB7F9D25539F9F8" box="[271,301,1592,1616]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,264,1104,1127]" captionTargetBox="[202,1435,214,1036]" captionTargetId="figure@6.[151,1436,193,1083]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Antennae of female Culicoides (Marksomyia) species. 13. C. marksi Lee &amp; Reye, antennal segment 6 and antenna. 14. C. pseudostigmaticus Tokunaga. 15. C. zentae Bellis &amp; Dyce sp. nov. 16. C. kayi Bellis &amp; Dyce sp. nov. 17. C. parvimaculatus Lee &amp; Reye. 18. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204431/files/figure.png" pageId="4" pageNumber="39">14</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FE81F9D2554CF9F8" box="[313,344,1592,1616]" captionStart="FIGURES 19 – 24" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[187,1404,215,1824]" captionTargetId="figure@7.[151,1436,193,1871]" captionTargetPageId="7" captionText="FIGURES 19 – 24. Female Culicoides (Marksomyia) species 19. C. marksi Lee &amp; Reye, legs and tibial comb. 20. C. pseudostigmaticus Tokunaga, legs and tibial comb. 21. C. zentae Bellis &amp; Dyce sp. nov., legs and tibial comb. 22. C. kayi Bellis &amp; Dyce sp. nov., legs and tibial comb. 23. C. parvimaculatus Lee &amp; Reye, legs and tibial comb. 24. C. dycei Lee &amp; Reye, legs and tibial comb." httpUri="https://zenodo.org/record/204432/files/figure.png" pageId="4" pageNumber="39">20</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FEDBF9D25596F9F8" box="[355,386,1592,1616]" captionStart="FIGURES 25 – 32" captionStartId="8.[151,264,1616,1639]" captionTargetBox="[160,1418,200,1568]" captionTargetId="figure@8.[151,1436,178,1608]" captionTargetPageId="8" captionText="FIGURES 25 – 32. Wings of female Culicoides (Marksomyia) species. 25. C. marksi Lee &amp; Reye. 26. C. marksi Lee &amp; Reye, rare aberrant wing pattern. 27. C. pseudostigmaticus Tokunaga. 28. C. zentae Bellis &amp; Dyce sp. nov. 29. C. kayi Bellis &amp; Dyce sp. nov. 30. C. parvimaculatus Lee &amp; Reye. 31.. C. dycei Lee &amp; Reye 32. C. dycei Lee &amp; Reye, rare aberrant wing pattern." httpUri="https://zenodo.org/record/204433/files/figure.png" pageId="4" pageNumber="39">27</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FE35F9D255BFF9F8" box="[397,427,1592,1616]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1892,1915]" captionTargetBox="[186,1399,217,1834]" captionTargetId="figure@10.[151,1436,193,1871]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Female Culicoides (Marksomyia) species. 33. C. marksi Lee &amp; Reye, spermathecae and genital sclerotisation. 34. C. pseudostigmaticus Tokunaga, spermathecae and genital sclerotisation. 35. C. zentae Bellis &amp; Dyce sp. nov., spermathecae and genital sclerotisation. 36. C. kayi Bellis &amp; Dyce sp. nov., spermathecae and genital sclerotisation. 37. C. parvimaculatus Lee &amp; Reye, spermathecae and genital sclerotisation. 38. C. dycei Lee &amp; Reye, spermathecae and genital sclerotisation." httpUri="https://zenodo.org/record/204434/files/figure.png" pageId="4" pageNumber="39">34</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FE0FF9D255C1F9F8" box="[439,469,1592,1616]" captionStart="FIGURES 39 – 44" captionStartId="11.[151,264,1151,1174]" captionTargetBox="[193,1391,223,1085]" captionTargetId="figure@11.[151,1436,193,1130]" captionTargetPageId="11" captionText="FIGURES 39 – 44. Antennae of male Culicoides (Marksomyia) species. Plume aristae are not shown although attachment bases are indicated where present. 39. C. marksi Lee &amp; Reye. 40. C. pseudostigmaticus Tokunaga. 41. C. zentae Bellis &amp; Dyce sp. nov. 42. C. kayi Bellis &amp; Dyce sp. nov. 43. C. parvimaculatus Lee &amp; Reye. 44. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204435/files/figure.png" pageId="4" pageNumber="39">40</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FE59F9D255EBF9F8" box="[481,511,1592,1616]" captionStart="FIGURES 45 – 50" captionStartId="13.[151,264,1876,1899]" captionTargetBox="[299,1291,193,1847]" captionTargetId="figure@13.[298,1292,193,1847]" captionTargetPageId="13" captionText="FIGURES 45 – 50. Male Culicoides (Marksomyia) species 45. C. marksi Lee &amp; Reye, hypopygium, parameres and aedeagus. 46. C. pseudostigmaticus Tokunaga, hypopygium, parameres and aedeagus. 47. C. zentae Bellis &amp; Dyce sp. nov., hypopygium, parameres and aedeagus. 48. C. kayi Bellis &amp; Dyce sp. nov., hypopygium, parameres and aedeagus. 49. C. parvimaculatus Lee &amp; Reye, hypopygium, parameres and aedeagus. 50. C. dycei Lee &amp; Reye, hypopygium, parameres and aedeagus." httpUri="https://zenodo.org/record/204436/files/figure.png" pageId="4" pageNumber="39">46</figureCitation>
, 
<figureCitation id="0DFD1AA4FF93FFA4FDB3F9D2563CF9F8" box="[523,552,1592,1616]" captionStart="FIGURES 51 – 56" captionStartId="14.[151,264,1155,1178]" captionTargetBox="[151,1425,200,1083]" captionTargetId="figure@14.[151,1436,193,1134]" captionTargetPageId="14" captionText="FIGURES 51 – 56. Wings of male Culicoides (Marksomyia) species. 51. C. marksi Lee &amp; Reye. 52. C. pseudostigmaticus Tokunaga. 53. C. zentae Bellis &amp; Dyce sp. nov. 54. C. kayi Bellis &amp; Dyce sp. nov. 55. C. parvimaculatus Lee &amp; Reye. 56. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204437/files/figure.png" pageId="4" pageNumber="39">52</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF93FFA4FF2FF991568FF939" box="[151,667,1659,1681]" pageId="4" pageNumber="39" type="reference_group">
<paragraph id="95790621FF93FFA4FF2FF991568FF939" blockId="4.[151,667,1659,1681]" box="[151,667,1659,1681]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="B5D6210FFF93FFA4FF2FF991568FF939" box="[151,667,1659,1681]" pageId="4" pageNumber="39">
<taxonomicName id="52C67DA2FF93FFA4FF2FF9915683F939" authority="Tokunaga 1959:234" authorityName="Tokunaga" authorityPageNumber="234" authorityYear="1959" box="[151,663,1659,1681]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF93FFA4FF2FF99155D0F939" box="[151,452,1659,1681]" italics="true" pageId="4" pageNumber="39">Culicoides pseudostigmaticus</emphasis>
<treatmentCitation id="14672030FF93FFA4FE73F9915683F939" author="Tokunaga" box="[459,663,1659,1681]" page="234" pageId="4" pageNumber="39" year="1959">
<bibRefCitation id="F1577BD0FF93FFA4FE73F991567EF939" author="Tokunaga" box="[459,618,1659,1681]" pageId="4" pageNumber="39" refString="Tokunaga, M. (1959) New Guinea biting midges (Diptera: Ceratopogonidae). Pacific Insects, 1, 177 - 314." type="journal article" year="1959">Tokunaga 1959</bibRefCitation>
:234
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF93FFA4FF2FF9565715F8F0" pageId="4" pageNumber="39" type="materials_examined">
<paragraph id="95790621FF93FFA4FF2FF9565715F8F0" blockId="4.[151,1436,1723,2014]" pageId="4" pageNumber="39">
<emphasis id="A7B2DA33FF93FFA4FF2FF956562EF97C" bold="true" box="[151,570,1723,1748]" pageId="4" pageNumber="39">
Non-type material examined. 
<collectingCountry id="EDD146B1FF93FFA4FE43F9515621F97C" box="[507,565,1723,1748]" name="Papua New Guinea" pageId="4" pageNumber="39">PNG</collectingCountry>
,
</emphasis>
DPI Stn Urimo, NG, 
<date id="E17820E1FF93FFA4FC95F95157D8F97B" box="[813,972,1723,1747]" pageId="4" pageNumber="39" value="1977-12-02" valueMax="1977-12-03" valueMin="1977-12-02">2–3.Dec.1977</date>
, B. Kadeu (
<specimenCount id="83C0CDA8FF93FFA4FBEBF95150ABF97B" box="[1107,1215,1723,1747]" pageId="4" pageNumber="39" type="female">3 females</specimenCount>
<collectionCode id="F3D79EE4FF93FFA4FB7FF9515104F97B" LSID="urn:lsid:biocol.org:col:32981" box="[1223,1296,1723,1747]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="4" pageNumber="39">ANIC</collectionCode>
); East Sepik Prov., Wirul Mission via Weewak, 
<geoCoordinate id="F0F260E6FF93FFA4FDFEF93656D7F95C" box="[582,707,1756,1780]" direction="south" orientation="latitude" pageId="4" pageNumber="39" precision="1" value="-3.5782">3°34.692'S</geoCoordinate>
, 
<geoCoordinate id="F0F260E6FF93FFA4FD6FF9365764F95C" box="[727,880,1756,1780]" direction="east" orientation="longitude" pageId="4" pageNumber="39" precision="1" value="143.64612">143º38.767'E</geoCoordinate>
, 
<date id="E17820E1FF93FFA4FC3BF9365004F95C" box="[899,1040,1756,1780]" pageId="4" pageNumber="39" value="2007-10-22">22.Oct.2007</date>
, LT, A.Rice (
<specimenCount id="83C0CDA8FF93FFA4FB78F9365100F95C" box="[1216,1300,1756,1780]" pageId="4" pageNumber="39" type="male">1 male</specimenCount>
, 
<specimenCount id="83C0CDA8FF93FFA4FA90F9365188F95C" box="[1320,1436,1756,1780]" pageId="4" pageNumber="39" type="female">4 females</specimenCount>
AQISNT); Balamuk, 
<geoCoordinate id="F0F260E6FF93FFA4FE32F91455E3F8BE" box="[394,503,1790,1814]" direction="south" orientation="latitude" pageId="4" pageNumber="39" precision="9" value="-8.905666">8º54.34'S</geoCoordinate>
, 
<geoCoordinate id="F0F260E6FF93FFA4FDBBF914568FF8BE" box="[515,667,1790,1814]" direction="east" orientation="longitude" pageId="4" pageNumber="39" precision="1" value="141.2764">141º16.584'E</geoCoordinate>
, 
<date id="E17820E1FF93FFA4FD1FF9145756F8BE" box="[679,834,1790,1814]" pageId="4" pageNumber="39" value="2007-10-03" valueMax="2007-10-07" valueMin="2007-10-03">3–7.Oct.2007</date>
, LT, J.Schmidt (
<specimenCount id="83C0CDA8FF93FFA4FC45F9145049F8BE" box="[1021,1117,1790,1814]" pageId="4" pageNumber="39" type="female">1 female</specimenCount>
AQISNT); 
<emphasis id="A7B2DA33FF93FFA4FB5CF917514BF8BE" bold="true" box="[1252,1375,1789,1814]" pageId="4" pageNumber="39">
<collectingCountry id="EDD146B1FF93FFA4FB5CF917514FF8BE" box="[1252,1371,1789,1814]" name="Indonesia" pageId="4" pageNumber="39">Indonesia</collectingCountry>
,
</emphasis>
Irian Jaya (
<collectingCountry id="EDD146B1FF93FFA4FF60F8F55536F89F" box="[216,290,1823,1847]" name="Papua New Guinea" pageId="4" pageNumber="39">Papua</collectingCountry>
), Jayapura, 
<date id="E17820E1FF93FFA4FE10F8F55621F89F" box="[424,565,1823,1847]" pageId="4" pageNumber="39" value="1989-10-23">23.Oct.1989</date>
, Sukarsih (
<specimenCount id="83C0CDA8FF93FFA4FD0EF8F55702F89F" box="[694,790,1823,1847]" pageId="4" pageNumber="39" type="female">1 female</specimenCount>
BALITVET); Jayapura, 
<date id="E17820E1FF93FFA4FB89F8F550A2F89F" box="[1073,1206,1823,1847]" pageId="4" pageNumber="39" value="1989-11-06">6.Nov.1989</date>
, Sukarsih, 
<specimenCount id="83C0CDA8FF93FFA4FA8AF8F5518CF89F" box="[1330,1432,1823,1847]" pageId="4" pageNumber="39" type="female">1 female</specimenCount>
; Jayapura, 
<date id="E17820E1FF93FFA4FEB1F8AA5583F8F0" box="[265,407,1856,1880]" pageId="4" pageNumber="39" value="1989-09-25">25.Sep.1989</date>
, Sukarsih (
<specimenCount id="83C0CDA8FF93FFA4FDAEF8AA5674F8F0" box="[534,608,1856,1880]" pageId="4" pageNumber="39" type="male">1 male</specimenCount>
BALITVET).
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF93FFA4FF7FF88B54D3F876" pageId="4" pageNumber="39" type="diagnosis">
<paragraph id="95790621FF93FFA4FF7FF88B54D3F876" blockId="4.[151,1436,1723,2014]" pageId="4" pageNumber="39">
<emphasis id="A7B2DA33FF93FFA4FF7FF88B5555F8D2" bold="true" box="[199,321,1889,1914]" pageId="4" pageNumber="39">Diagnosis.</emphasis>
Wing with single pale spot in cell M4. Female with SCo distribution 3, 11–15, six SCh on 4–7, four to five SCh on 8–10, single SCh on 11–14, SCh absent from 15; spermathecae subspherical with proximal half nonsclerotised. Male with apical half of parameres slightly curved laterad, ventral membrane of ninth sternite spiculate.
</paragraph>
</subSubSection>
<caption id="C1B956A9FF92FFA5FF2FF88E5639F87E" httpUri="https://zenodo.org/record/204430/files/figure.png" pageId="5" pageNumber="40" targetBox="[195,1309,231,1843]" targetPageId="5">
<paragraph id="95790621FF92FFA5FF2FF88E5639F87E" blockId="5.[151,1436,1892,2006]" pageId="5" pageNumber="40">
<emphasis id="A7B2DA33FF92FFA5FF2FF88E555CF8D2" bold="true" box="[151,328,1892,1915]" pageId="5" pageNumber="40">FIGURES 7–12.</emphasis>
Female 
<taxonomicName id="52C67DA2FF92FFA5FE19F88F56B1F8D2" authority="Marksomyia" authorityName="Marksomyia" box="[417,677,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF92FFA5FE19F88F5619F8D3" box="[417,525,1893,1915]" italics="true" pageId="5" pageNumber="40">Culicoides</emphasis>
(
<emphasis id="A7B2DA33FF92FFA5FDA5F88F568FF8D3" box="[541,667,1893,1915]" italics="true" pageId="5" pageNumber="40">Marksomyia</emphasis>
)
</taxonomicName>
species. 7. 
<taxonomicName id="52C67DA2FF92FFA5FCA5F88F501CF8D2" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[797,1032,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF92FFA5FCA5F88F5795F8D3" box="[797,897,1893,1915]" italics="true" pageId="5" pageNumber="40">C. marksi</emphasis>
Lee &amp; Reye
</taxonomicName>
, mandibular teeth and palp. 8. 
<taxonomicName id="52C67DA2FF92FFA5FAFEF88F559BF831" authority="Tokunaga" authorityName="Tokunaga" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF92FFA5FAFEF88F5530F831" italics="true" pageId="5" pageNumber="40">C. pseudostigmaticus</emphasis>
Tokunaga
</taxonomicName>
, mandibular teeth and palp. 9. 
<taxonomicName id="52C67DA2FF92FFA5FD7DF86E57ADF831" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[709,953,1923,1945]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF92FFA5FD7DF86E5737F831" box="[709,803,1924,1945]" italics="true" pageId="5" pageNumber="40">C. zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF92FFA5FC07F86E501BF831" bold="true" box="[959,1039,1924,1945]" pageId="5" pageNumber="40">
<taxonomicNameLabel id="BC816748FF92FFA5FC07F86E501BF831" box="[959,1039,1924,1945]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, mandibular teeth and palp. 10. 
<taxonomicName id="52C67DA2FF92FFA5FAECF86E553CF810" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="kayi" status="sp. nov.">
<emphasis id="A7B2DA33FF92FFA5FAECF86E518FF831" box="[1364,1435,1923,1945]" italics="true" pageId="5" pageNumber="40">C. kayi</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF92FFA5FE96F848556BF81F" bold="true" box="[302,383,1954,1975]" pageId="5" pageNumber="40">
<taxonomicNameLabel id="BC816748FF92FFA5FE96F848556BF81F" box="[302,383,1954,1975]" pageId="5" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, mandibular teeth and palp. 11. 
<taxonomicName id="52C67DA2FF92FFA5FD7DF8495017F810" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[709,1027,1954,1976]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF92FFA5FD7DF8495794F810" box="[709,896,1954,1976]" italics="true" pageId="5" pageNumber="40">C. parvimaculatus</emphasis>
Lee &amp; Reye
</taxonomicName>
, mandibular teeth and palp. 12. 
<taxonomicName id="52C67DA2FF92FFA5FAF1F8495507F87E" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF92FFA5FAF1F8495188F810" box="[1353,1436,1954,1976]" italics="true" pageId="5" pageNumber="40">C. dycei</emphasis>
Lee &amp; Reye
</taxonomicName>
, mandibular teeth and palp.
</paragraph>
</caption>
<caption id="C1B956A9FF91FFA6FF2FFBBA56D6FB0C" httpUri="https://zenodo.org/record/204431/files/figure.png" pageId="6" pageNumber="41" targetBox="[202,1435,214,1036]" targetPageId="6">
<paragraph id="95790621FF91FFA6FF2FFBBA56D6FB0C" blockId="6.[151,1436,1104,1188]" pageId="6" pageNumber="41">
<emphasis id="A7B2DA33FF91FFA6FF2FFBBA5541FBCE" bold="true" box="[151,341,1104,1127]" pageId="6" pageNumber="41">FIGURES 13–18.</emphasis>
Antennae of female 
<taxonomicName id="52C67DA2FF91FFA6FD95FBBB5724FBCF" authority="Marksomyia" authorityName="Marksomyia" box="[557,816,1105,1127]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF91FFA6FD95FBBB568DFBCF" box="[557,665,1105,1127]" italics="true" pageId="6" pageNumber="41">Culicoides</emphasis>
(
<emphasis id="A7B2DA33FF91FFA6FD11FBBB5733FBCF" box="[681,807,1105,1127]" italics="true" pageId="6" pageNumber="41">Marksomyia</emphasis>
)
</taxonomicName>
species. 13. 
<taxonomicName id="52C67DA2FF91FFA6FC0DFBB8508BFBCF" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[949,1183,1105,1127]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF91FFA6FC0DFBB8500DFBCF" box="[949,1049,1105,1127]" italics="true" pageId="6" pageNumber="41">C. marksi</emphasis>
Lee &amp; Reye
</taxonomicName>
, antennal segment 6 and antenna. 14. 
<taxonomicName id="52C67DA2FF91FFA6FEA3FB9A5673FB2D" authority="Tokunaga." authorityName="Tokunaga." box="[283,615,1135,1157]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF91FFA6FEA3FB9A55E0FB2D" box="[283,500,1135,1157]" italics="true" pageId="6" pageNumber="41">C. pseudostigmaticus</emphasis>
Tokunaga.
</taxonomicName>
15. 
<taxonomicName id="52C67DA2FF91FFA6FD2DFB9A5784FB2D" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[661,912,1135,1158]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF91FFA6FD2DFB9A56E0FB2E" box="[661,756,1136,1158]" italics="true" pageId="6" pageNumber="41">C. zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF91FFA6FC2FFB9A57FDFB2D" bold="true" box="[919,1001,1136,1157]" pageId="6" pageNumber="41">
<taxonomicNameLabel id="BC816748FF91FFA6FC2FFB9A57FDFB2D" box="[919,1001,1136,1157]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
16. 
<taxonomicName id="52C67DA2FF91FFA6FBA0FB9A50E8FB2D" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[1048,1276,1135,1157]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="kayi" status="sp. nov.">
<emphasis id="A7B2DA33FF91FFA6FBA0FB9A5075FB2D" box="[1048,1121,1135,1157]" italics="true" pageId="6" pageNumber="41">C. kayi</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF91FFA6FABCFB9A5141FB2D" bold="true" box="[1284,1365,1136,1157]" pageId="6" pageNumber="41">
<taxonomicNameLabel id="BC816748FF91FFA6FABCFB9A5141FB2D" box="[1284,1365,1136,1157]" pageId="6" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
17. 
<taxonomicName id="52C67DA2FF91FFA6FA3CFB9A55A8FB0C" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF91FFA6FA3CFB9A5520FB0C" italics="true" pageId="6" pageNumber="41">C. parvimaculatus</emphasis>
Lee &amp; Reye.
</taxonomicName>
18. 
<taxonomicName id="52C67DA2FF91FFA6FE5FFB6556D6FB0C" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." box="[487,706,1166,1188]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF91FFA6FE5FFB65562FFB0C" box="[487,571,1166,1188]" italics="true" pageId="6" pageNumber="41">C. dycei</emphasis>
Lee &amp; Reye.
</taxonomicName>
</paragraph>
</caption>
<subSubSection id="DDDC55AAFF91FFA8FF7FFB2455DCF941" lastPageId="8" lastPageNumber="43" pageId="6" pageNumber="41" type="description">
<paragraph id="95790621FF91FFA6FF7FFB245725FAE2" blockId="6.[151,1437,1230,2016]" pageId="6" pageNumber="41">
<emphasis id="A7B2DA33FF91FFA6FF7FFB245560FB4E" bold="true" box="[199,372,1230,1254]" pageId="6" pageNumber="41">Female. Head.</emphasis>
Eyes bare except for a weak band of interfacetal hairs medially, separated by a distance of at least one facet (
<figureCitation id="0DFD1AA4FF91FFA6FEFCFB055596FAAF" box="[324,386,1263,1287]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1399,1422]" captionTargetBox="[187,1398,218,1321]" captionTargetId="figure@4.[151,1436,193,1378]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Eye separation of female Culicoides (Marksomyia) species. 1. C. marksi Lee &amp; Reye. 2. C. pseudostigmaticus Tokunaga. 3. C. zentae Bellis &amp; Dyce sp. nov. 4. C. kayi Bellis &amp; Dyce sp. nov. 5. C. parvimaculatus Lee &amp; Reye. 6. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204429/files/figure.png" pageId="6" pageNumber="41">Fig 2</figureCitation>
), proboscis short. Palpus pale brown with five segments (fig 8), segment 3 expanding apically then abruptly narrowed beyond an irregular, shallow sensory pit with several protruding capitate sensilla. Antennomeres (fig 14) 4–10 barrel shaped, 11–15 cylindrical, short.
</paragraph>
<paragraph id="95790621FF91FFA6FF7DFABE57DDFA1B" blockId="6.[151,1437,1230,2016]" pageId="6" pageNumber="41">
<emphasis id="A7B2DA33FF91FFA6FF7DFABE5530FAC4" bold="true" box="[197,292,1364,1388]" pageId="6" pageNumber="41">Thorax.</emphasis>
Legs (fig 20) dark brown with dark knees, fore &amp; mid femora pale basally and with conspicuous apical pale band, hind femora dark brown with less conspicuous apical pale band; all tibiae with distinct basal pale band. Wing (fig 27) strongly patterned with a single pale spot in cell M4.
</paragraph>
<paragraph id="95790621FF91FFA6FF7DFA5555CEFA53" blockId="6.[151,1437,1230,2016]" pageId="6" pageNumber="41">
<emphasis id="A7B2DA33FF91FFA6FF7DFA555555FA70" bold="true" box="[197,321,1471,1496]" pageId="6" pageNumber="41">Abdomen.</emphasis>
Three developed, ovoid spermathecae partially to almost completely sclerotised, ducts short; sclerotised ring tapered (
<figureCitation id="0DFD1AA4FF91FFA6FEC7FA0955D9FA53" box="[383,461,1507,1531]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1892,1915]" captionTargetBox="[186,1399,217,1834]" captionTargetId="figure@10.[151,1436,193,1871]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Female Culicoides (Marksomyia) species. 33. C. marksi Lee &amp; Reye, spermathecae and genital sclerotisation. 34. C. pseudostigmaticus Tokunaga, spermathecae and genital sclerotisation. 35. C. zentae Bellis &amp; Dyce sp. nov., spermathecae and genital sclerotisation. 36. C. kayi Bellis &amp; Dyce sp. nov., spermathecae and genital sclerotisation. 37. C. parvimaculatus Lee &amp; Reye, spermathecae and genital sclerotisation. 38. C. dycei Lee &amp; Reye, spermathecae and genital sclerotisation." httpUri="https://zenodo.org/record/204434/files/figure.png" pageId="6" pageNumber="41">Fig 34</figureCitation>
).
</paragraph>
<paragraph id="95790621FF91FFA6FF7FF9E257C8F94F" blockId="6.[151,1437,1230,2016]" pageId="6" pageNumber="41">
<emphasis id="A7B2DA33FF91FFA6FF7FF9E2554DF988" bold="true" box="[199,345,1544,1568]" pageId="6" pageNumber="41">Male. Head.</emphasis>
Eyes bare except for a weak band of interfacetal hairs medially. Palpus similar to female with a shallow pit on segment 3. Antenna (
<figureCitation id="0DFD1AA4FF91FFA6FD91F9C25661F9E8" box="[553,629,1576,1600]" captionStart="FIGURES 39 – 44" captionStartId="11.[151,264,1151,1174]" captionTargetBox="[193,1391,223,1085]" captionTargetId="figure@11.[151,1436,193,1130]" captionTargetPageId="11" captionText="FIGURES 39 – 44. Antennae of male Culicoides (Marksomyia) species. Plume aristae are not shown although attachment bases are indicated where present. 39. C. marksi Lee &amp; Reye. 40. C. pseudostigmaticus Tokunaga. 41. C. zentae Bellis &amp; Dyce sp. nov. 42. C. kayi Bellis &amp; Dyce sp. nov. 43. C. parvimaculatus Lee &amp; Reye. 44. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204435/files/figure.png" pageId="6" pageNumber="41">Fig 40</figureCitation>
) of specimens examined partially feminised with all antennomeres fully independent and lacking the oblique row of plume verticils present in males of other species of 
<emphasis id="A7B2DA33FF91FFA6FB04F9A0515CF9CA" box="[1212,1352,1610,1634]" italics="true" pageId="6" pageNumber="41">Marksomyia</emphasis>
; antennomeres 3–10 broader than antennomeres 11–15 with gradual increase in length from basal to distal antennomeres. SCh of two distinct 
<typeStatus id="4A7DB883FF91FFA6FEC4F96455AEF90E" box="[380,442,1678,1702]" pageId="6" pageNumber="41">types</typeStatus>
, 3–10 with basal whorl of five to eight very long SCh resembling plume aristae, 11–14 with basal whorl of two to four short SCh resembling those present on distal antennomeres of males of other species of 
<emphasis id="A7B2DA33FF91FFA6FF50F9255560F94F" box="[232,372,1743,1767]" italics="true" pageId="6" pageNumber="41">Marksomyia</emphasis>
. Non-plume SCh on 3 reaching to antennomere 7 or 8.
</paragraph>
<paragraph id="95790621FF91FFA6FF7DF91A55C9F848" blockId="6.[151,1437,1230,2016]" pageId="6" pageNumber="41">
<emphasis id="A7B2DA33FF91FFA6FF7DF91A552EF8A1" bold="true" box="[197,314,1776,1801]" pageId="6" pageNumber="41">Genitalia.</emphasis>
(fig 46) Ninth tergite with caudal margin convex. Ninth sternite with moderately deep, caudomedial excavation, ventral membrane with a continuous narrow band of spicules across its anterior margin. Gonocoxite short and broad with dorsal root long and simple; ventral root reduced to a short, stout point. Gonostylus slender, distally curving gently to a pointed apex. Aedeagus with shoulders thicker than basal arms and almost perpendicular to distal process; distal process short with a rounded expansion. Parameres weakly joined, with straight basal arms angled sharply to weakly swollen stem lacking ventral lobe narrowing gradually to simple sharp-tipped distal portion curved lateroventrad.
</paragraph>
<caption id="C1B956A9FF90FFA7FF2FF88E5505F87E" httpUri="https://zenodo.org/record/204432/files/figure.png" pageId="7" pageNumber="42" targetBox="[187,1404,215,1824]" targetPageId="7">
<paragraph id="95790621FF90FFA7FF2FF88E5505F87E" blockId="7.[151,1436,1892,2006]" pageId="7" pageNumber="42">
<emphasis id="A7B2DA33FF90FFA7FF2FF88E5542F8D2" bold="true" box="[151,342,1892,1915]" pageId="7" pageNumber="42">FIGURES 19–24.</emphasis>
Female 
<taxonomicName id="52C67DA2FF90FFA7FE0BF88F56A3F8D2" authority="Marksomyia" authorityName="Marksomyia" box="[435,695,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF90FFA7FE0BF88F560BF8D3" box="[435,543,1893,1915]" italics="true" pageId="7" pageNumber="42">Culicoides</emphasis>
(
<emphasis id="A7B2DA33FF90FFA7FD88F88F56BAF8D3" box="[560,686,1893,1915]" italics="true" pageId="7" pageNumber="42">Marksomyia</emphasis>
)
</taxonomicName>
species 19. 
<taxonomicName id="52C67DA2FF90FFA7FC83F88F503FF8D2" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[827,1067,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF90FFA7FC83F88F57B4F8D3" box="[827,928,1893,1915]" italics="true" pageId="7" pageNumber="42">C. marksi</emphasis>
Lee &amp; Reye
</taxonomicName>
, legs and tibial comb. 20. 
<taxonomicName id="52C67DA2FF90FFA7FAFDF88F5584F831" authority="Tokunaga" authorityName="Tokunaga" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF90FFA7FAFDF88F5530F831" italics="true" pageId="7" pageNumber="42">C. pseudostigmaticus</emphasis>
Tokunaga
</taxonomicName>
, legs and tibial comb. 21. 
<taxonomicName id="52C67DA2FF90FFA7FD23F86E5785F831" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[667,913,1923,1945]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF90FFA7FD23F86E56EEF831" box="[667,762,1924,1945]" italics="true" pageId="7" pageNumber="42">C. zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF90FFA7FC21F86E57FDF831" bold="true" box="[921,1001,1924,1945]" pageId="7" pageNumber="42">
<taxonomicNameLabel id="BC816748FF90FFA7FC21F86E57FDF831" box="[921,1001,1924,1945]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, legs and tibial comb. 22. 
<taxonomicName id="52C67DA2FF90FFA7FB40F86E54D8F810" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="kayi" status="sp. nov.">
<emphasis id="A7B2DA33FF90FFA7FB40F86E5154F831" box="[1272,1344,1923,1945]" italics="true" pageId="7" pageNumber="42">C. kayi</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF90FFA7FF6AF8485536F81F" bold="true" box="[210,290,1954,1975]" pageId="7" pageNumber="42">
<taxonomicNameLabel id="BC816748FF90FFA7FF6AF8485536F81F" box="[210,290,1954,1975]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, legs and tibial comb. 23. 
<taxonomicName id="52C67DA2FF90FFA7FD92F8495773F810" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[554,871,1954,1976]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF90FFA7FD92F84956F0F810" box="[554,740,1954,1976]" italics="true" pageId="7" pageNumber="42">C. parvimaculatus</emphasis>
Lee &amp; Reye
</taxonomicName>
, legs and tibial comb. 24. 
<taxonomicName id="52C67DA2FF90FFA7FBD4F8495154F810" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[1132,1344,1954,1976]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF90FFA7FBD4F84950AAF810" box="[1132,1214,1954,1976]" italics="true" pageId="7" pageNumber="42">C. dycei</emphasis>
Lee &amp; Reye
</taxonomicName>
, legs and tibial comb.
</paragraph>
</caption>
<caption id="C1B956A9FF9FFFA8FF2FF9BA5169F90C" httpUri="https://zenodo.org/record/204433/files/figure.png" pageId="8" pageNumber="43" targetBox="[160,1418,200,1568]" targetPageId="8">
<paragraph id="95790621FF9FFFA8FF2FF9BA5169F90C" blockId="8.[151,1436,1616,1700]" pageId="8" pageNumber="43">
<emphasis id="A7B2DA33FF9FFFA8FF2FF9BA5541F9CE" bold="true" box="[151,341,1616,1639]" pageId="8" pageNumber="43">FIGURES 25–32.</emphasis>
Wings of female 
<taxonomicName id="52C67DA2FF9FFFA8FDB4F9BB571BF9CF" authority="Marksomyia" authorityName="Marksomyia" box="[524,783,1617,1639]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF9FFFA8FDB4F9BB566CF9CF" box="[524,632,1617,1639]" italics="true" pageId="8" pageNumber="43">Culicoides</emphasis>
(
<emphasis id="A7B2DA33FF9FFFA8FD30F9BB5712F9CF" box="[648,774,1617,1639]" italics="true" pageId="8" pageNumber="43">Marksomyia</emphasis>
)
</taxonomicName>
species. 25. 
<taxonomicName id="52C67DA2FF9FFFA8FC2CF9B85096F9CF" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." box="[916,1154,1617,1639]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF9FFFA8FC2CF9B857E3F9CF" box="[916,1015,1617,1639]" italics="true" pageId="8" pageNumber="43">C. marksi</emphasis>
Lee &amp; Reye.
</taxonomicName>
26. 
<taxonomicName id="52C67DA2FF9FFFA8FB17F9B8518CF9CF" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[1199,1432,1617,1639]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF9FFFA8FB17F9B85106F9CF" box="[1199,1298,1617,1639]" italics="true" pageId="8" pageNumber="43">C. marksi</emphasis>
Lee &amp; Reye
</taxonomicName>
, rare aberrant wing pattern. 27. 
<taxonomicName id="52C67DA2FF9FFFA8FE77F99A570DF92D" authority="Tokunaga." authorityName="Tokunaga." box="[463,793,1647,1669]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF9FFFA8FE77F99A56BCF92D" box="[463,680,1647,1669]" italics="true" pageId="8" pageNumber="43">C. pseudostigmaticus</emphasis>
Tokunaga.
</taxonomicName>
28. 
<taxonomicName id="52C67DA2FF9FFFA8FCFCF99A502EF92D" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[836,1082,1647,1670]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF9FFFA8FCFCF99A57B6F92E" box="[836,930,1648,1670]" italics="true" pageId="8" pageNumber="43">C. zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF9FFFA8FBF8F99A5084F92D" bold="true" box="[1088,1168,1648,1669]" pageId="8" pageNumber="43">
<taxonomicNameLabel id="BC816748FF9FFFA8FBF8F99A5084F92D" box="[1088,1168,1648,1669]" pageId="8" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
29. 
<taxonomicName id="52C67DA2FF9FFFA8FB05F99A5188F92D" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[1213,1436,1647,1669]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="kayi" status="sp. nov.">
<emphasis id="A7B2DA33FF9FFFA8FB05F99A5110F92D" box="[1213,1284,1647,1669]" italics="true" pageId="8" pageNumber="43">C. kayi</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF9FFFA8FF2FF96554F3F90C" bold="true" box="[151,231,1679,1700]" pageId="8" pageNumber="43">
<taxonomicNameLabel id="BC816748FF9FFFA8FF2FF96554F3F90C" box="[151,231,1679,1700]" pageId="8" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
30. 
<taxonomicName id="52C67DA2FF9FFFA8FEACF9655643F90C" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." box="[276,599,1678,1700]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF9FFFA8FEACF96555DBF90C" box="[276,463,1678,1700]" italics="true" pageId="8" pageNumber="43">C. parvimaculatus</emphasis>
Lee &amp; Reye.
</taxonomicName>
31.. 
<taxonomicName id="52C67DA2FF9FFFA8FD37F9655776F90C" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[655,866,1678,1700]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF9FFFA8FD37F96556F6F90C" box="[655,738,1678,1700]" italics="true" pageId="8" pageNumber="43">C. dycei</emphasis>
Lee &amp; Reye
</taxonomicName>
32. 
<taxonomicName id="52C67DA2FF9FFFA8FC37F9655071F90C" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[911,1125,1678,1700]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF9FFFA8FC37F96557F6F90C" box="[911,994,1678,1700]" italics="true" pageId="8" pageNumber="43">C. dycei</emphasis>
Lee &amp; Reye
</taxonomicName>
, rare aberrant wing pattern.
</paragraph>
</caption>
<paragraph id="95790621FF9FFFA8FF7FF93B55DCF941" blockId="8.[151,1436,1745,2036]" box="[199,456,1745,1769]" pageId="8" pageNumber="43">
<emphasis id="A7B2DA33FF9FFFA8FF7FF93B5546F941" bold="true" box="[199,338,1745,1769]" pageId="8" pageNumber="43">Immatures.</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9FFFA8FF7FF91855E8F8A2" box="[199,508,1778,1803]" pageId="8" pageNumber="43" type="distribution">
<paragraph id="95790621FF9FFFA8FF7FF91855E8F8A2" blockId="8.[151,1436,1745,2036]" box="[199,508,1778,1803]" pageId="8" pageNumber="43">
<emphasis id="A7B2DA33FF9FFFA8FF7FF9185574F8A3" bold="true" box="[199,352,1778,1803]" pageId="8" pageNumber="43">Distribution.</emphasis>
New 
<collectingCountry id="EDD146B1FF9FFFA8FE1BF91855ECF8A2" box="[419,504,1778,1802]" name="Guinea-Bissau" pageId="8" pageNumber="43">Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9FFFA8FF7FF8F955B0F883" box="[199,420,1811,1836]" pageId="8" pageNumber="43" type="biology_ecology">
<paragraph id="95790621FF9FFFA8FF7FF8F955B0F883" blockId="8.[151,1436,1745,2036]" box="[199,420,1811,1836]" pageId="8" pageNumber="43">
<emphasis id="A7B2DA33FF9FFFA8FF7FF8F95533F884" bold="true" box="[199,295,1811,1836]" pageId="8" pageNumber="43">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9FFFA9FF7FF8DF5129FF7C" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43" type="discussion">
<paragraph id="95790621FF9FFFA8FF7FF8DF50B0F85B" blockId="8.[151,1436,1745,2036]" pageId="8" pageNumber="43">
<emphasis id="A7B2DA33FF9FFFA8FF7FF8DF552FF8E5" bold="true" box="[199,315,1845,1869]" pageId="8" pageNumber="43">Remarks.</emphasis>
While feminisation of the male antenna is not unknown within the genus 
<taxonomicName id="52C67DA2FF9FFFA8FBCCF8DF5195F8E5" authority="Dyce 1996" authorityName="Dyce" authorityYear="1996" box="[1140,1409,1845,1870]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF9FFFA8FBCCF8DF50EFF8E5" box="[1140,1275,1845,1869]" italics="true" pageId="8" pageNumber="43">Culicoides (</emphasis>
<bibRefCitation id="F1577BD0FF9FFFA8FB44F8DC516DF8E5" author="Dyce" box="[1276,1401,1845,1870]" pageId="8" pageNumber="43" refString="Dyce A. L. (1996) Description of the male of Culicoides agas Wirth and Hubert (Diptera: Ceratopogonidae). Memoirs of the Entomological Society of Washington, 18, 113 - 118." type="journal article" year="1996">Dyce 1996</bibRefCitation>
)
</taxonomicName>
, it is possible that the male specimens of 
<taxonomicName id="52C67DA2FF9FFFA8FDFCF8BD5727F8C6" box="[580,819,1878,1902]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF9FFFA8FDFCF8BD5727F8C6" box="[580,819,1878,1902]" italics="true" pageId="8" pageNumber="43">C. pseudostigmaticus</emphasis>
</taxonomicName>
examined in this study are in fact intersexes. This condition is observed occasionally in field collected material and is often caused by infestation with mermithid parasites (Poinar &amp; Sarto i Monteys 2008). No such parasites were observed in the specimens examined herein. These specimens are the only males known for this species and for this reason they were described herein although we acknowledge that they may not represent the normal morphology of the male of this species.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7DFF7D5129FF7C" blockId="9.[151,1436,151,212]" pageId="9" pageNumber="44">
Male and female specimens of 
<taxonomicName id="52C67DA2FF9EFFA9FDA7FF72571AFF18" box="[543,782,152,176]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF9EFFA9FDA7FF72571AFF18" box="[543,782,152,176]" italics="true" pageId="9" pageNumber="44">C. pseudostigmaticus</emphasis>
</taxonomicName>
can be distinguished from all other 
<emphasis id="A7B2DA33FF9EFFA9FB25FF72513DFF18" box="[1181,1321,152,176]" italics="true" pageId="9" pageNumber="44">Marksomyia</emphasis>
excepting 
<taxonomicName id="52C67DA2FF9EFFA9FF2FFF5754E6FF7C" box="[151,242,188,212]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF9EFFA9FF2FFF5754E6FF7C" box="[151,242,188,212]" italics="true" pageId="9" pageNumber="44">C. dycei</emphasis>
</taxonomicName>
by the single pale spot in cell M4. See comments under 
<taxonomicName id="52C67DA2FF9EFFA9FCD4FF5757DCFF7C" box="[876,968,188,212]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF9EFFA9FCD4FF5757DCFF7C" box="[876,968,188,212]" italics="true" pageId="9" pageNumber="44">C. dycei</emphasis>
</taxonomicName>
for other distinguishing features.
</paragraph>
</subSubSection>
</treatment>
</document>