treatments-xml/data/BA/49/24/BA492440124EFFF0FCD9FC7FFCEFA8D6.xml

<document id="C346D67BA771FF4DE41E476CEA289401" ID-CLB-Dataset="53759" ID-DOI="10.6620/ZS.2018.57-18" ID-GBIF-Dataset="cff4ab26-6fdc-462b-aaa0-b8e8f87afd71" ID-PMC="PMC6624643" ID-PubMed="31966258" ID-Zenodo-Dep="8064335" IM.bibliography_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687182456079" checkinUser="valdenar" docAuthor="Yoshida 1, Takahiro, Karner, Michael &amp; Hirowatari, Toshiya" docDate="2018" docId="BA492440124EFFF0FCD9FC7FFCEFA8D6" docLanguage="en" docName="ZoolStud.57.18.1-18.pdf" docOrigin="Zoological studies 57 (18)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Psammoecus taiwanensis Ng &amp; Lin 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="6" masterDocId="46705C38124FFFF5FFA6FFC2FF9FA960" masterDocTitle="A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="2" updateTime="1687352464173" updateUser="ExternalLinkService">
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<mods:title id="B611B1222537FDA99A54251003B19B8B">A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="8EAB028D628A142AAEDA50A20E3AAFD1">Yoshida 1, Takahiro</mods:namePart>
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<mods:namePart id="853D49533F1E7CE19996343C0EC55D87">Karner, Michael</mods:namePart>
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<mods:namePart id="CB632E558EABD6C448CB3FF789BF1DF1">Hirowatari, Toshiya</mods:namePart>
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<mods:title id="2D9A0C47FFC675A6833C32C73E07CFA7">Zoological studies</mods:title>
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<mods:date id="B17BEE85FDC2115EDF2AF215C387A413">2018</mods:date>
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<treatment id="BA492440124EFFF0FCD9FC7FFCEFA8D6" ID-DOI="http://doi.org/10.5281/zenodo.8055788" ID-Zenodo-Dep="8055788" LSID="urn:lsid:plazi:treatment:BA492440124EFFF0FCD9FC7FFCEFA8D6" httpUri="http://treatment.plazi.org/id/BA492440124EFFF0FCD9FC7FFCEFA8D6" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
<subSubSection id="7AFAC6DD124EFFF4FCD9FC7FFAACAAB6" box="[895,1331,957,982]" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="325F9556124EFFF4FCD9FC7FFAACAAB6" blockId="1.[826,1400,957,1067]" box="[895,1331,957,982]" pageId="1" pageNumber="2">
<heading id="6917223A124EFFF4FCD9FC7FFAACAAB6" bold="true" box="[895,1331,957,982]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
<emphasis id="00944944124EFFF4FCD9FC7FFAACAAB6" bold="true" box="[895,1331,957,982]" pageId="1" pageNumber="2">
<taxonomicName id="F5E0EED5124EFFF4FCD9FC7FFB56AAB6" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" box="[895,1225,957,982]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="00944944124EFFF4FCD9FC7FFB56AAB6" bold="true" box="[895,1225,957,982]" italics="true" pageId="1" pageNumber="2">Psammoecus taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F124EFFF4FB77FC7FFAACAAB6" box="[1233,1331,957,982]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="7AFAC6DD124EFFF0FC60FC1FFCEFA8D6" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2" type="description">
<paragraph id="325F9556124EFFF4FC60FC1FFB32AD4B" blockId="1.[826,1400,957,1067]" pageId="1" pageNumber="2">
(
<figureCitation id="AADB89D3124EFFF4FC6BFC1FFBABAA96" box="[973,1076,989,1014]" captionStart="Fig" captionStartId="2.[174,208,1844,1864]" captionTargetBox="[174,1451,308,1818]" captionTargetId="figure-67@2.[174,1451,308,1818]" captionTargetPageId="2" captionText="Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="1" pageNumber="2">Figs. 1A</figureCitation>
, 
<figureCitation id="AADB89D3124EFFF4FB99FC1FFBCEAA96" box="[1087,1105,989,1014]" captionStart="Fig" captionStartId="3.[136,170,1954,1974]" captionTargetBox="[326,1224,1377,1928]" captionTargetId="figure-537@3.[326,1224,1377,1928]" captionTargetPageId="3" captionText="Fig. 2. Psammoecus taiwanensis sp. nov. in dorsal view (A) and ventral view (B). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064339" httpUri="https://zenodo.org/record/8064339/files/figure.png" pageId="1" pageNumber="2">2</figureCitation>
, 
<figureCitation id="AADB89D3124EFFF4FBFBFC1FFBF3AA96" box="[1117,1132,989,1014]" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="1" pageNumber="2">3</figureCitation>
and 
<figureCitation id="AADB89D3124EFFF4FB01FC1EFB29AA96" box="[1191,1206,988,1014]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="1" pageNumber="2">4</figureCitation>
A-C) 
<uri id="46719954124EFFF4FC9CFC3AFB32AD4B" pageId="1" pageNumber="2">
urn:lsid:zoobank.org:act: 
<uuid id="4646AF83124EFFF4FB9FFC3AFB32AD4B" pageId="1" pageNumber="2">549E3797-8024-4C89-A13C-547166490EAA</uuid>
</uri>
</paragraph>
<paragraph id="325F9556124EFFF4FCCFFB92FA1AAEC9" blockId="1.[813,1414,1104,1962]" pageId="1" pageNumber="2">
<emphasis id="00944944124EFFF4FCCFFB92FB9CAD09" box="[873,1027,1104,1129]" italics="true" pageId="1" pageNumber="2">Type series</emphasis>
: 
<materialsCitation id="82889F0B124EFFF4FBBCFB92FA1DADC9" collectingDate="2014-07-02" collectionCode="NMNS" collectorName="T. Yoshida" county="Ren-ai Township" location="Sungkang" municipality="Jing Ying Village" pageId="1" pageNumber="2" specimenCount="1" typeStatus="holotype">
<typeStatus id="ED5B2BF4124EFFF4FBBCFB92FB0BAD09" box="[1050,1172,1104,1129]" pageId="1" pageNumber="2">Holotype</typeStatus>
: male, 
<location id="373FC38D124EFFF4FB5FFB92FA1EAD0A" LSID="urn:lsid:plazi:treatment:BA492440124EFFF0FCD9FC7FFCEFA8D6:373FC38D124EFFF4FB5FFB92FA1EAD0A" box="[1273,1409,1104,1130]" county="Ren-ai Township" municipality="Jing Ying Village" name="Sungkang" pageId="1" pageNumber="2">Sungkang</location>
, 
<collectingMunicipality id="D23B0F2C124EFFF4FC8BFBB2FC64ADE9" box="[813,1019,1136,1161]" pageId="1" pageNumber="2">Jing Ying Village</collectingMunicipality>
, 
<collectingCounty id="DB3EEDDA124EFFF4FBAEFBB2FB4AADE9" box="[1032,1237,1136,1161]" pageId="1" pageNumber="2">Ren-ai Township</collectingCounty>
, 
<date id="465EB396124EFFF4FB44FBB2FAFBADE9" box="[1250,1380,1136,1161]" pageId="1" pageNumber="2" value="2014-07-02">
<collectingDate id="561A4A7E124EFFF4FB44FBB2FAFBADE9" box="[1250,1380,1136,1161]" pageId="1" pageNumber="2" value="2014-07-02">2-VII-2014</collectingDate>
</date>
, 
<collectorName id="9F15F080124EFFF4FAD7FBB2FC10ADC9" pageId="1" pageNumber="2">T. Yoshida</collectorName>
leg. (
<collectionCode id="54F10D93124EFFF4FC7FFB52FBAEADCA" box="[985,1073,1168,1194]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="1" pageNumber="2">NMNS</collectionCode>
, male genitalia illustrated)
</materialsCitation>
. 
<typeStatus id="ED5B2BF4124EFFF4FC8BFB72FC35ADA9" box="[813,938,1200,1225]" pageId="1" pageNumber="2">Paratypes</typeStatus>
: 
<collectingCountry id="4AF7D5C6124EFFF4FC10FB72FB80ADA9" box="[950,1055,1200,1225]" name="Taiwan" pageId="1" pageNumber="2">TAIWAN</collectingCountry>
: [
<collectingRegion id="F0245BB4124EFFF4FB97FB72FB6DADAA" box="[1073,1266,1200,1226]" country="Taiwan" name="New Taipei" pageId="1" pageNumber="2">New Taipei City</collectingRegion>
] 
<specimenCount id="24E65EDF124EFFF4FAA7FB72FAF5ADA9" box="[1281,1386,1200,1225]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
&amp; 
<specimenCount id="24E65EDF124EFFF4FC8BFB12FCF5AD89" box="[813,874,1232,1257]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, Wulai District, 
<date id="465EB396124EFFF4FBBAFB12FB0BAD89" box="[1052,1172,1232,1257]" pageId="1" pageNumber="2" value="1970-06-04">4-VI-1970</date>
, Y. Hori leg. (EUMJ). [
<collectingRegion id="F0245BB4124EFFF4FC94FB32FC49AC6A" box="[818,982,1264,1290]" country="Taiwan" name="Yilan" pageId="1" pageNumber="2">Yilan County</collectingRegion>
] 2 exs, Fushan Botanical Garden, Yuanshan Township, 
<date id="465EB396124EFFF4FBE0FAD2FA99AC49" box="[1094,1286,1296,1321]" pageId="1" pageNumber="2" value="2000-09-25" valueMax="2000-09-27" valueMin="2000-09-25">25-27-IX-2000</date>
, L. Papp, L. Peregovits &amp; L. Ronkay leg., at light (HNHM; studied by MK); 2 exs, Cueifong Lake, Nan-ao Township, alt. 
<quantity id="F51838B3124EFFF4FC7DFAB2FBA5ACE9" box="[987,1082,1392,1417]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="1" pageNumber="2" unit="m" value="1950.0">1950 m</quantity>
, 
<date id="465EB396124EFFF4FBEEFAB2FB5FACE9" box="[1096,1216,1392,1417]" pageId="1" pageNumber="2" value="2004-04-01">1-IV-2004</date>
, T. Kurihara leg. (ELKU). [
<collectingRegion id="F0245BB4124EFFF4FC02FA52FBF4ACCA" box="[932,1131,1424,1450]" country="Taiwan" name="Nantou" pageId="1" pageNumber="2">Nantou County</collectingRegion>
] 
<specimenCount id="24E65EDF124EFFF4FBDAFA52FB5EACC9" box="[1148,1217,1424,1449]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, Gaofeng, 
<date id="465EB396124EFFF4FAF5FA52FC16ACA9" pageId="1" pageNumber="2" value="2009-05-07" valueMax="2009-05-08" valueMin="2009-05-07">7-8- V-2009</date>
, J. Aoki leg. (ELKU); 
<specimenCount id="24E65EDF124EFFF4FB06FA72FA96ACA9" box="[1184,1289,1456,1481]" pageId="1" pageNumber="2" type="male">3 males</specimenCount>
, 
<specimenCount id="24E65EDF124EFFF4FABEFA72FA1AACA9" box="[1304,1413,1456,1481]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
&amp; 2 exs, Tun-yuan-trail, Jing Ying Village, Ren-ai Township, 
<date id="465EB396124EFFF4FC77FA32FBCAAF69" box="[977,1109,1520,1545]" pageId="1" pageNumber="2" value="2014-07-01">1-VII-2014</date>
, T. Yoshida leg. (ELKU); 3 exs, same locality, 
<date id="465EB396124EFFF4FB9EF9D2FB20AF49" box="[1080,1215,1552,1577]" pageId="1" pageNumber="2" value="2014-07-06">6-VII-2014</date>
, T. Yoshida leg. (ELKU); 10 exs, same data as 
<typeStatus id="ED5B2BF4124EFFF4FB3AF9F2FA9AAF29" box="[1180,1285,1584,1609]" pageId="1" pageNumber="2">holotype</typeStatus>
. (ELKU); 
<specimenCount id="24E65EDF124EFFF4FAD1F9F2FCF3AF09" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
, Tsiufeng, Ren-ai Township, 
<date id="465EB396124EFFF4FB6DF992FAC2AF09" box="[1227,1373,1616,1641]" pageId="1" pageNumber="2" value="1966-07-10">10-VII-1966</date>
, H. Kamiya leg. (KUM). [
<collectingRegion id="F0245BB4124EFFF4FB82F9B2FB4FAFEA" box="[1060,1232,1648,1674]" country="Taiwan" name="Chiayi" pageId="1" pageNumber="2">Chiayi County</collectingRegion>
] 
<specimenCount id="24E65EDF124EFFF4FB7BF9B2FA83AFE9" box="[1245,1308,1648,1673]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, Alishan, Alishan Township, 
<date id="465EB396124EFFF4FBB5F952FB01AFC9" box="[1043,1182,1680,1705]" pageId="1" pageNumber="2" value="1968-08-01">1-VIII-1968</date>
, K. Y. leg. (ELKU); 
<specimenCount id="24E65EDF124EFFF4FC8BF972FCF2AFA9" box="[813,877,1712,1737]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, Fenchihu, Zhuqi Township, 
<date id="465EB396124EFFF4FB6AF972FAC1AFA9" box="[1228,1374,1712,1737]" pageId="1" pageNumber="2" value="1966-07-22">22-VII-1966</date>
, H. Kamiya leg. (KUM); 2 exs, same locality, 
<date id="465EB396124EFFF4FA95F912FCF2AE69" pageId="1" pageNumber="2" value="1966-07-23">23-VII-1966</date>
, H. Kamiya leg. (KUM); 
<specimenCount id="24E65EDF124EFFF4FB33F932FB4AAE69" box="[1173,1237,1776,1801]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, same locality, 
<date id="465EB396124EFFF4FC8BF8D2FC5EAE49" box="[813,961,1808,1833]" pageId="1" pageNumber="2" value="1966-07-24">24-VII-1966</date>
, H. Kamiya leg. (KUM); 
<specimenCount id="24E65EDF124EFFF4FB54F8D2FAABAE49" box="[1266,1332,1808,1833]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, same locality, 
<date id="465EB396124EFFF4FC31F8F2FBB8AE29" box="[919,1063,1840,1865]" pageId="1" pageNumber="2" value="1968-06-22">22-VI-1968</date>
, M. Tomokuni leg. (EUMJ). [
<collectingRegion id="F0245BB4124EFFF4FC94F892FC75AE0A" box="[818,1002,1872,1898]" country="Taiwan" name="Kaohsiung" pageId="1" pageNumber="2">Kaohsiung City</collectingRegion>
] 
<specimenCount id="24E65EDF124EFFF4FC51F892FBAAAE09" box="[1015,1077,1872,1897]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, Shanping, Liouguei District, 
<date id="465EB396124EFFF4FC8BF8B2FC72AEE9" box="[813,1005,1904,1929]" pageId="1" pageNumber="2" value="1980-03-20" valueMax="1980-03-21" valueMin="1980-03-20">20-21-III-1980</date>
, T. Niisato leg. (EUMJ); 
<specimenCount id="24E65EDF124EFFF4FA9CF8B2FA1FAEE9" box="[1338,1408,1904,1929]" pageId="1" pageNumber="2" type="generic">1 ex.</specimenCount>
, Mt. Su-shan (Shi Shan) near Liouguei, 
<collectingRegion id="F0245BB4124EFFF4FABBF852FA1AAEC9" box="[1309,1413,1936,1961]" country="Taiwan" name="Taoyuan" pageId="1" pageNumber="2">Taoyuan</collectingRegion>
</paragraph>
<footnote id="51FB8958124EFFF4FF2EF81FFE07AE8F" box="[136,408,2013,2031]" pageId="1" pageNumber="2">
<paragraph id="325F9556124EFFF4FF2EF81FFE07AE8F" blockId="1.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="1" pageNumber="2">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="669FC5DE124DFFF7FF08F8F6FCAFAEE0" ID-DOI="http://doi.org/10.5281/zenodo.8064337" ID-Zenodo-Dep="8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="2" pageNumber="3" startId="2.[174,208,1844,1864]" targetBox="[174,1451,308,1818]" targetPageId="2" targetType="figure">
<paragraph id="325F9556124DFFF7FF08F8F6FCAFAEE0" blockId="2.[174,1451,1844,1920]" pageId="2" pageNumber="3">
<emphasis id="00944944124DFFF7FF08F8F6FF73AE28" bold="true" box="[174,236,1844,1864]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
<taxonomicName id="F5E0EED5124DFFF7FF51F8F6FE59AE28" box="[247,454,1844,1864]" pageId="2" pageNumber="3">
<emphasis id="00944944124DFFF7FF51F8F6FEE7AE28" box="[247,376,1844,1864]" italics="true" pageId="2" pageNumber="3">Psammoecus</emphasis>
species
</taxonomicName>
of Taiwan, habitus. (A) 
<taxonomicName id="F5E0EED5124DFFF7FD05F8F6FCB3AE28" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" box="[675,812,1844,1864]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="00944944124DFFF7FD05F8F6FD2EAE28" box="[675,689,1844,1864]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FD1BF8F6FCB3AE28" box="[701,812,1844,1864]" italics="true" pageId="2" pageNumber="3">taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F124DFFF7FC94F8F6FCE3AE28" box="[818,892,1844,1864]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(B) 
<taxonomicName id="F5E0EED5124DFFF7FC05F8F6FBEEAE28" authority="Grouvelle" authorityName="Grouvelle" box="[931,1137,1844,1864]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="00944944124DFFF7FC05F8F6FC2EAE28" box="[931,945,1844,1864]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FC1BF8F6FB8EAE28" box="[957,1041,1844,1864]" italics="true" pageId="2" pageNumber="3">delicatus</emphasis>
Grouvelle
</taxonomicName>
(C) 
<taxonomicName id="F5E0EED5124DFFF7FB3DF8F6FAF1AE28" authority="Grouvelle" authorityName=", Grouvelle" authorityYear="1912" box="[1179,1390,1844,1864]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="00944944124DFFF7FB3DF8F6FB36AE28" box="[1179,1193,1844,1864]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FB13F8F6FA91AE28" box="[1205,1294,1844,1864]" italics="true" pageId="2" pageNumber="3">harmandi</emphasis>
Grouvelle
</taxonomicName>
(D) 
<taxonomicName id="F5E0EED5124DFFF7FA31F8F6FEFEAE04" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1883" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="00944944124DFFF7FA31F8F6FA3AAE28" box="[1431,1445,1844,1864]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FF08F892FE9FAE04" box="[174,256,1872,1892]" italics="true" pageId="2" pageNumber="3">dentatus</emphasis>
Grouvelle
</taxonomicName>
(E) 
<taxonomicName id="F5E0EED5124DFFF7FE2AF892FD0AAE04" authority="Motschulsky" authorityName="Motschulsky" authorityYear="1858" box="[396,661,1872,1892]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="00944944124DFFF7FE2AF892FE05AE04" box="[396,410,1872,1892]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FE01F892FD85AE04" box="[423,538,1872,1892]" italics="true" pageId="2" pageNumber="3">trimaculatus</emphasis>
Motschulsky
</taxonomicName>
(F) 
<taxonomicName id="F5E0EED5124DFFF7FD18F892FCE2AE04" authority="Reitter" authorityName="Reitter" authorityYear="1874" box="[702,893,1872,1892]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="triguttatus">
<emphasis id="00944944124DFFF7FD18F892FD53AE04" box="[702,716,1872,1892]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FD7FF892FCA8AE04" box="[729,823,1872,1892]" italics="true" pageId="2" pageNumber="3">triguttatus</emphasis>
Reitter
</taxonomicName>
(G) 
<taxonomicName id="F5E0EED5124DFFF7FC0FF892FA82AE04" authority="Yoshida and Hirowatari" authorityName="Yoshida and Hirowatari" authorityYear="2014" box="[937,1309,1872,1892]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="labyrinthicus">
<emphasis id="00944944124DFFF7FC0FF892FC28AE04" box="[937,951,1872,1892]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FC62F892FBA4AE04" box="[964,1083,1872,1892]" italics="true" pageId="2" pageNumber="3">labyrinthicus</emphasis>
Yoshida and Hirowatari
</taxonomicName>
(H) 
<taxonomicName id="F5E0EED5124DFFF7FAEEF892FE96AEE0" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1892" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="simonis">
<emphasis id="00944944124DFFF7FAEEF892FAC9AE04" box="[1352,1366,1872,1892]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FAC5F892FA34AE04" box="[1379,1451,1872,1892]" italics="true" pageId="2" pageNumber="3">simonis</emphasis>
Grouvelle
</taxonomicName>
(I) 
<taxonomicName id="F5E0EED5124DFFF7FE8FF8AEFD3BAEE0" authority="Yoshida and Hirowatari. Scale" authorityName="Yoshida and Hirowatari. Scale" authorityYear="2013" box="[297,676,1900,1920]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hiranoi">
<emphasis id="00944944124DFFF7FE8FF8AEFEA8AEE0" box="[297,311,1900,1920]" italics="true" pageId="2" pageNumber="3">P</emphasis>
. 
<emphasis id="00944944124DFFF7FEE5F8AEFE1CAEE0" box="[323,387,1900,1920]" italics="true" pageId="2" pageNumber="3">hiranoi</emphasis>
Yoshida and Hirowatari. Scale
</taxonomicName>
bar = 1.0 mm.
</paragraph>
</caption>
<footnote id="51FB8958124DFFF7FB3AF81FFA34AE8F" box="[1180,1451,2013,2031]" pageId="2" pageNumber="3">
<paragraph id="325F9556124DFFF7FB3AF81FFA34AE8F" blockId="2.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="2" pageNumber="3">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="325F9556124CFFF6FF2EFF3FFDF8A876" blockId="3.[136,737,253,1334]" box="[136,615,253,278]" pageId="3" pageNumber="4">
District, 
<date id="465EB396124CFFF6FF4EFF3FFEF0A876" box="[232,367,253,278]" pageId="3" pageNumber="4" value="1986-04-25">25-IV-1986</date>
, K. Baba leg. (KUM).
</paragraph>
<paragraph id="325F9556124CFFF6FF63FEDFFD11A836" blockId="3.[136,737,253,1334]" pageId="3" pageNumber="4">
<emphasis id="00944944124CFFF6FF63FEDFFEDEA856" box="[197,321,285,310]" italics="true" pageId="3" pageNumber="4">Etymology</emphasis>
: The specific name is derived from 
<collectingCountry id="4AF7D5C6124CFFF6FF2EFEFFFF40A836" box="[136,223,317,342]" name="Taiwan" pageId="3" pageNumber="4">Taiwan</collectingCountry>
, the 
<typeStatus id="ED5B2BF4124CFFF6FEB0FEFFFED7A836" box="[278,328,317,342]" pageId="3" pageNumber="4">type</typeStatus>
locality of the new species.
</paragraph>
<paragraph id="325F9556124CFFF6FF63FE9FFF44ABB6" blockId="3.[136,737,253,1334]" pageId="3" pageNumber="4">
<emphasis id="00944944124CFFF6FF63FE9FFED2A816" box="[197,333,349,374]" italics="true" pageId="3" pageNumber="4">Diagnosis</emphasis>
: This species resembles 
<taxonomicName id="F5E0EED5124CFFF6FD63FE9FFE38A8F6" authority="Grouvelle, 1882" authorityName="Grouvelle" authorityYear="1882" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="00944944124CFFF6FD63FE9FFD48A816" box="[709,727,349,374]" italics="true" pageId="3" pageNumber="4">P</emphasis>
. 
<emphasis id="00944944124CFFF6FF2EFEBFFF49A8F6" box="[136,214,381,406]" italics="true" pageId="3" pageNumber="4">piceus</emphasis>
Grouvelle, 1882
</taxonomicName>
; it is distinguished by the distal portions of parameres bearing dense short pubescence, in comparison with distal portions of parameres bearing only two apical setae in 
<taxonomicName id="F5E0EED5124CFFF6FD6EFE1FFF49AB76" authorityName="Grouvelle" authorityYear="1882" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="piceus">
<emphasis id="00944944124CFFF6FD6EFE1FFD45A896" box="[712,730,477,502]" italics="true" pageId="3" pageNumber="4">P</emphasis>
. 
<emphasis id="00944944124CFFF6FF2EFE3FFF49AB76" box="[136,214,509,534]" italics="true" pageId="3" pageNumber="4">piceus</emphasis>
</taxonomicName>
. This species also resembles 
<taxonomicName id="F5E0EED5124CFFF6FDEBFE3FFD7EAB76" authorityName=", Grouvelle" authorityYear="1912" box="[589,737,509,534]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="00944944124CFFF6FDEBFE3FFDC0AB76" box="[589,607,509,534]" italics="true" pageId="3" pageNumber="4">P</emphasis>
. 
<emphasis id="00944944124CFFF6FDD7FE3FFD7EAB76" box="[625,737,509,534]" italics="true" pageId="3" pageNumber="4">harmandi</emphasis>
</taxonomicName>
but is easily distinguished by the presence of a row of long erect setae along each lateral elytral margin, the expanded anterior angles of pronotum and the stick-shaped and elongate parameres (cone-shaped and stout in 
<taxonomicName id="F5E0EED5124CFFF6FE7EFD5FFDF0ABD6" authorityName=", Grouvelle" authorityYear="1912" box="[472,623,669,694]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="00944944124CFFF6FE7EFD5FFE75ABD6" box="[472,490,669,694]" italics="true" pageId="3" pageNumber="4">P</emphasis>
. 
<emphasis id="00944944124CFFF6FE5BFD5FFDF0ABD6" box="[509,623,669,694]" italics="true" pageId="3" pageNumber="4">harmandi</emphasis>
</taxonomicName>
) (
<figureCitation id="AADB89D3124CFFF6FD2CFD5FFD7FABD6" box="[650,736,669,694]" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
and 
<figureCitation id="AADB89D3124CFFF6FF1AFD7FFF50ABB6" box="[188,207,701,726]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
).
</paragraph>
<paragraph id="325F9556124CFFF6FF63FD1FFF29AA76" blockId="3.[136,737,253,1334]" pageId="3" pageNumber="4">
<emphasis id="00944944124CFFF6FF63FD1FFED4AB96" box="[197,331,733,758]" italics="true" pageId="3" pageNumber="4">Description</emphasis>
: Body length: 
<quantity id="F51838B3124CFFF6FE58FD1FFD39AB96" box="[510,678,733,758]" metricMagnitude="-3" metricUnit="m" metricValue="3.13" metricValueMax="3.43" metricValueMin="2.83" pageId="3" pageNumber="4" unit="mm" value="3.13" valueMax="3.43" valueMin="2.83">2.83-3.43 mm</quantity>
(
<emphasis id="00944944124CFFF6FD1FFD1FFD57AB96" box="[697,712,733,758]" italics="true" pageId="3" pageNumber="4">n</emphasis>
= 20).
</paragraph>
<paragraph id="325F9556124CFFF6FF63FCDFFE3EAD56" blockId="3.[136,737,253,1334]" pageId="3" pageNumber="4">
<emphasis id="00944944124CFFF6FF63FCDFFECFAA56" box="[197,336,797,822]" italics="true" pageId="3" pageNumber="4">Coloration</emphasis>
: (
<figureCitation id="AADB89D3124CFFF6FEC8FCDFFE7EAA56" box="[366,481,797,822]" captionStart="Fig" captionStartId="2.[174,208,1844,1864]" captionTargetBox="[174,1451,308,1818]" captionTargetId="figure-67@2.[174,1451,308,1818]" captionTargetPageId="2" captionText="Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="3" pageNumber="4">Figs. 1A</figureCitation>
and 
<figureCitation id="AADB89D3124CFFF6FD8BFCDFFDDDAA56" box="[557,578,797,822]" captionStart="Fig" captionStartId="3.[136,170,1954,1974]" captionTargetBox="[326,1224,1377,1928]" captionTargetId="figure-537@3.[326,1224,1377,1928]" captionTargetPageId="3" captionText="Fig. 2. Psammoecus taiwanensis sp. nov. in dorsal view (A) and ventral view (B). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064339" httpUri="https://zenodo.org/record/8064339/files/figure.png" pageId="3" pageNumber="4">2</figureCitation>
). Head and pronotum dark brown. Elytra somewhat lighter colored, with a quadrate black macula on each elytron at middle, connected by narrow horizontal band at posterior 1/3, round maculae at posterior 1/4, connected to lateral margins, darkened around humeri and end of elytra. Antennae dark brown, 7th and/or 8th to 10th antennomere darker, 11th (apex) yellowish-brown.
</paragraph>
<paragraph id="325F9556124CFFF6FF63FBFFFC4FA816" blockId="3.[136,737,253,1334]" lastBlockId="3.[813,1415,253,1334]" pageId="3" pageNumber="4">
Head: (
<figureCitation id="AADB89D3124CFFF6FE87FBFFFE49AD36" box="[289,470,1085,1110]" captionStart="Fig" captionStartId="3.[136,170,1954,1974]" captionTargetBox="[326,1224,1377,1928]" captionTargetId="figure-537@3.[326,1224,1377,1928]" captionTargetPageId="3" captionText="Fig. 2. Psammoecus taiwanensis sp. nov. in dorsal view (A) and ventral view (B). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064339" httpUri="https://zenodo.org/record/8064339/files/figure.png" pageId="3" pageNumber="4">Fig. 2A and B</figureCitation>
). Wide, HW/HL 1.28- 1.43; IE/HL 0.86-0.98. Temples somewhat enlarged behind eyes, narrowed gradually toward basis. Eyes of moderate size, slightly prominent. Dorsal surface with coarse punctation. Antennae long; antennomeres with pubescence of moderate to large length; distal portion of 7th to 10th and entire 11th antennomeres with short pubescence, very dense on 8th to 11th antennomeres; approximate ratios of antennomere lengths of 
<typeStatus id="ED5B2BF4124CFFF6FB4AFEDFFAC4A856" box="[1260,1371,285,310]" pageId="3" pageNumber="4">holotype</typeStatus>
as follows: 2.6: 1.0: 1.2: 1.3: 1.3: 1.5: 1.4: 1.2: 1.1: 1.0: 1.7.
</paragraph>
<paragraph id="325F9556124CFFF6FCCFFEBFFBC5AB96" blockId="3.[813,1415,253,1334]" pageId="3" pageNumber="4">
Pronotum: (
<figureCitation id="AADB89D3124CFFF6FC57FEBFFBEAA8F6" box="[1009,1141,381,406]" captionStart="Fig" captionStartId="3.[136,170,1954,1974]" captionTargetBox="[326,1224,1377,1928]" captionTargetId="figure-537@3.[326,1224,1377,1928]" captionTargetPageId="3" captionText="Fig. 2. Psammoecus taiwanensis sp. nov. in dorsal view (A) and ventral view (B). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064339" httpUri="https://zenodo.org/record/8064339/files/figure.png" pageId="3" pageNumber="4">Figs. 2A, B</figureCitation>
and 
<figureCitation id="AADB89D3124CFFF6FB09FEBFFB4BA8F6" box="[1199,1236,381,406]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="3" pageNumber="4">4B</figureCitation>
). Subquadrate, widest near anterior margin, straightly narrowed toward base, lateral margins slightly flattened; lateral areas of basis somewhat flattened, PW/ PL 1.18-1.27. Punctation on pronotal disk as on vertex; pubescence composed of numerous setae of moderate length, very long setae on teeth of lateral margins and anterior and posterior angles (
<figureCitation id="AADB89D3124CFFF6FC92FDBFFC0CABF6" box="[820,915,637,662]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="3" pageNumber="4">Fig. 4B</figureCitation>
). Anterior angle with two small teeth and few small tubercles; lateral margin with some very small teeth, anterior tooth largest; posterior angle with small tooth (
<figureCitation id="AADB89D3124CFFF6FC55FD1FFBD1AB96" box="[1011,1102,733,758]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="3" pageNumber="4">Fig. 4B</figureCitation>
).
</paragraph>
<paragraph id="325F9556124CFFF6FCCFFD3FFBDAAAB7" blockId="3.[813,1415,253,1334]" pageId="3" pageNumber="4">
Elytra: (
<figureCitation id="AADB89D3124CFFF6FC77FD3FFBDFAA76" box="[977,1088,765,790]" captionStart="Fig" captionStartId="3.[136,170,1954,1974]" captionTargetBox="[326,1224,1377,1928]" captionTargetId="figure-537@3.[326,1224,1377,1928]" captionTargetPageId="3" captionText="Fig. 2. Psammoecus taiwanensis sp. nov. in dorsal view (A) and ventral view (B). Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064339" httpUri="https://zenodo.org/record/8064339/files/figure.png" pageId="3" pageNumber="4">Figs. 2A</figureCitation>
and 
<figureCitation id="AADB89D3124CFFF6FB21FD3FFB2DAA77" box="[1159,1202,765,791]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="3" pageNumber="4">4C</figureCitation>
). Elongate-oval, EW/BL 0.40-0.45. Rows of punctures wider than interstices. Pubescence composed of numerous semi-erect setae of medium length, some very long, erect setae along each lateral margin, longer toward humeral swelling, with some long setae on lateral margin (
<figureCitation id="AADB89D3124CFFF6FC7BFC7FFBA6AAB7" box="[989,1081,957,983]" captionStart="Fig" captionStartId="5.[136,170,1925,1945]" captionTargetBox="[136,1413,723,1900]" captionTargetId="figure-215@5.[136,1413,723,1900]" captionTargetPageId="5" captionText="Fig. 4. SEM images of Psammoecus taiwanensis sp. nov. (A-C) and Psammoecus harmandi Grouvelle (D-F). A and D habitus B and E pronotum C and F anterolateral elytron." figureDoi="http://doi.org/10.5281/zenodo.8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="3" pageNumber="4">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="325F9556124CFFF0FCCFFC1FFDC9ABF6" blockId="3.[813,1415,253,1334]" lastBlockId="5.[136,738,253,662]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
Male genitalia: (
<figureCitation id="AADB89D3124CFFF6FB9EFC1FFB17AA96" box="[1080,1160,989,1014]" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). Spiculum gastrale (
<figureCitation id="AADB89D3124CFFF6FC92FC3FFC0DAD76" box="[820,914,1021,1046]" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
) with long and very thin strut, diverging widely around apex; branches slightly widened, connected by membrane; lateral sclerites elongate, curved inwards around apices, with short and thin strut. Parameres (
<figureCitation id="AADB89D3124CFFF6FBA3FBBFFBC0ADF6" box="[1029,1119,1149,1174]" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
) stick-shaped, elongate, depressed at base, protruding dorsally at inner basal angles, narrowed at basal 1/3 to 2/3, dorsally densely punctate, punctures of various size on basal 1/3, with a short seta on each inner margin of basal 1/3, dorsally densely with short setae on distal 1/2 or 1/3 to apices, with a long seta and a very long seta on each apex. Phallobase (
<figureCitation id="AADB89D3124AFFF0FD14FEDFFF35A836" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="5" pageNumber="6">Fig. 3B</figureCitation>
) long, upper layer deeply and widely incised at distal 1/3, with incision covered by a membrane, lower layer thin, extending toward base. Penis (
<figureCitation id="AADB89D3124AFFF0FD15FEBFFF35A8D6" captionStart="Fig" captionStartId="4.[174,208,1926,1946]" captionTargetBox="[223,1417,256,1893]" captionTargetId="figure-6@4.[178,1448,229,1906]" captionTargetPageId="4" captionText="Fig. 3. Male genital organs of Psammoecus taiwanensis sp. nov., holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="5" pageNumber="6">Fig. 3B</figureCitation>
) long, sub-parallel, gradually narrowed around apex, with rounded protrusion at apex, finely punctate near apex. Internal sac recurved around apex, with a thin ringed structure and a thin apical strut, densely armed with numerous spines on apical half near space of the ringed structure, with two spinous parallel armed lines around middle, with dense, minute spines near middle.
</paragraph>
<caption id="669FC5DE124CFFF6FF2EF860FBCFAED6" ID-DOI="http://doi.org/10.5281/zenodo.8064339" ID-Zenodo-Dep="8064339" box="[136,1104,1953,1974]" httpUri="https://zenodo.org/record/8064339/files/figure.png" pageId="3" pageNumber="4" startId="3.[136,170,1954,1974]" targetBox="[326,1224,1377,1928]" targetPageId="3" targetType="figure">
<paragraph id="325F9556124CFFF6FF2EF860FBCFAED6" blockId="3.[136,1104,1953,2031]" box="[136,1104,1953,1974]" pageId="3" pageNumber="4">
<emphasis id="00944944124CFFF6FF2EF860FF59AED6" bold="true" box="[136,198,1954,1974]" pageId="3" pageNumber="4">Fig. 2.</emphasis>
<taxonomicName id="F5E0EED5124CFFF6FF74F860FE57AED6" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" box="[210,456,1954,1974]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="00944944124CFFF6FF74F860FE57AED6" box="[210,456,1954,1974]" italics="true" pageId="3" pageNumber="4">Psammoecus taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F124CFFF6FE68F860FD86AED6" box="[462,537,1954,1974]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
in dorsal view (A) and ventral view (B). Scale bar = 1.0 mm.
</paragraph>
</caption>
<footnote id="51FB8958124CFFF6FF2EF81FFE07AE8F" box="[136,408,2013,2031]" pageId="3" pageNumber="4">
<paragraph id="325F9556124CFFF6FF2EF81FFE07AE8F" blockId="3.[136,1104,1953,2031]" box="[136,408,2013,2031]" pageId="3" pageNumber="4">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="669FC5DE124BFFF1FF08F844FF63AED6" ID-DOI="http://doi.org/10.5281/zenodo.8064341" ID-Zenodo-Dep="8064341" httpUri="https://zenodo.org/record/8064341/files/figure.png" pageId="4" pageNumber="5" startId="4.[174,208,1926,1946]" targetBox="[223,1417,256,1893]" targetPageId="4" targetType="figure">
<paragraph id="325F9556124BFFF1FF08F844FF63AED6" blockId="4.[174,1451,1925,1974]" pageId="4" pageNumber="5">
<emphasis id="00944944124BFFF1FF08F844FF72AEFA" bold="true" box="[174,237,1926,1946]" pageId="4" pageNumber="5">Fig. 3.</emphasis>
Male genital organs of 
<taxonomicName id="F5E0EED5124BFFF1FE7CF844FD4DAEFA" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" box="[474,722,1926,1946]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="00944944124BFFF1FE7CF844FD4DAEFA" box="[474,722,1926,1946]" italics="true" pageId="4" pageNumber="5">Psammoecus taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F124BFFF1FD7CF844FCB6AEFA" box="[730,809,1926,1946]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm.
</paragraph>
</caption>
<footnote id="51FB8958124BFFF1FB3AF81FFA34AE8F" box="[1180,1451,2013,2031]" pageId="4" pageNumber="5">
<paragraph id="325F9556124BFFF1FB3AF81FFA34AE8F" blockId="4.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="4" pageNumber="5">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="325F9556124AFFF0FCCFFF3FFBC7A876" blockId="5.[813,1413,253,438]" box="[873,1112,253,278]" pageId="5" pageNumber="6">
<emphasis id="00944944124AFFF0FCCFFF3FFC71A876" box="[873,1006,253,278]" italics="true" pageId="5" pageNumber="6">Distribution</emphasis>
: 
<collectingCountry id="4AF7D5C6124AFFF0FC5BFF3FFBCBA876" box="[1021,1108,253,278]" name="Taiwan" pageId="5" pageNumber="6">Taiwan</collectingCountry>
.
</paragraph>
<paragraph id="325F9556124AFFF0FCCFFEDFFCEFA8D6" blockId="5.[813,1413,253,438]" pageId="5" pageNumber="6">
<emphasis id="00944944124AFFF0FCCFFEDFFC49A856" box="[873,982,285,310]" italics="true" pageId="5" pageNumber="6">Remarks</emphasis>
: This new species is distributed in high altitudes of 
<collectingCountry id="4AF7D5C6124AFFF0FC5FFEFFFBD2A836" box="[1017,1101,317,342]" name="Taiwan" pageId="5" pageNumber="6">Taiwan</collectingCountry>
and occurs sympatrically with 
<taxonomicName id="F5E0EED5124AFFF0FCCEFE9FFC61A816" authorityName=", Grouvelle" authorityYear="1912" box="[872,1022,349,374]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="00944944124AFFF0FCCEFE9FFCE5A816" box="[872,890,349,374]" italics="true" pageId="5" pageNumber="6">P</emphasis>
. 
<emphasis id="00944944124AFFF0FC2AFE9FFC61A816" box="[908,1022,349,374]" italics="true" pageId="5" pageNumber="6">harmandi</emphasis>
</taxonomicName>
. On Tun-yuan-trail, the species was collected from dead leaves of broad-leaved trees.
</paragraph>
</subSubSection>
</treatment>
</document>