treatments-xml/data/BA/49/24/BA492440124AFFFCFCF9FE1FFF74AC96.xml

<document id="1688F48B487871249E5A2F283447596F" ID-CLB-Dataset="53759" ID-DOI="10.6620/ZS.2018.57-18" ID-GBIF-Dataset="cff4ab26-6fdc-462b-aaa0-b8e8f87afd71" ID-PMC="PMC6624643" ID-PubMed="31966258" ID-Zenodo-Dep="8064335" IM.bibliography_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1687182456079" checkinUser="valdenar" docAuthor="Yoshida 1, Takahiro, Karner, Michael &amp; Hirowatari, Toshiya" docDate="2018" docId="BA492440124AFFFCFCF9FE1FFF74AC96" docLanguage="en" docName="ZoolStud.57.18.1-18.pdf" docOrigin="Zoological studies 57 (18)" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2015-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2015-.journal_article" docStyleVersion="1" docTitle="Psammoecus delicatus Grouvelle 1908" docType="treatment" docVersion="4" lastPageNumber="10" masterDocId="46705C38124FFFF5FFA6FFC2FF9FA960" masterDocTitle="A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="18" masterPageNumber="1" pageNumber="6" updateTime="1687352464173" updateUser="ExternalLinkService">
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<mods:title id="5BC7A99C54F2CC185EF6D4197E3937B1">A Revision of Taiwanese Species in the Genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
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<mods:namePart id="99F28DA126CE998B704BBABB8430AEC0">Yoshida 1, Takahiro</mods:namePart>
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<mods:namePart id="42741B00CCA82CA75C9CB658E140BC7C">Karner, Michael</mods:namePart>
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<subSubSection id="7AFAC6DD124AFFF0FCF9FE1FFACCA896" box="[863,1363,476,502]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="325F9556124AFFF0FCF9FE1FFACCA896" blockId="5.[863,1363,476,534]" box="[863,1363,476,502]" pageId="5" pageNumber="6">
<heading id="6917223A124AFFF0FCF9FE1FFACCA896" bold="true" box="[863,1363,476,502]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="6" reason="2">
<taxonomicName id="F5E0EED5124AFFF0FCF9FE1FFACCA896" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[863,1363,476,502]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="00944944124AFFF0FCF9FE1FFACCA896" bold="true" box="[863,1363,476,502]" pageId="5" pageNumber="6">
<emphasis id="00944944124AFFF0FCF9FE1FFB18A896" bold="true" box="[863,1159,477,502]" italics="true" pageId="5" pageNumber="6">Psammoecus delicatus</emphasis>
<bibRefCitation id="5671E8A7124AFFF0FB28FE1EFACCA896" author="Grouvelle A." box="[1166,1363,476,502]" pageId="5" pageNumber="6" pagination="315 - 495" refId="ref12782" refString="Grouvelle A. 1908. Coleopteres de la region Indienne. Rhysodidae, Trogositidae, Nitidulidae, Colydiidae, Cucujidae. Annales de la Societe Entomologique de France 77: 315 - 495." type="journal article" year="1908">Grouvelle, 1908</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="7AFAC6DD124AFFF0FC4DFE3FFB58AB76" box="[1003,1223,509,534]" pageId="5" pageNumber="6" type="description">
<paragraph id="325F9556124AFFF0FC4DFE3FFB58AB76" blockId="5.[863,1363,476,534]" box="[1003,1223,509,534]" pageId="5" pageNumber="6">
(
<figureCitation id="AADB89D3124AFFF0FC54FE3FFBC7AB76" box="[1010,1112,509,534]" captionStart="Fig" captionStartId="2.[174,208,1844,1864]" captionTargetBox="[174,1451,308,1818]" captionTargetId="figure-67@2.[174,1451,308,1818]" captionTargetPageId="2" captionText="Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="5" pageNumber="6">Figs. 1B</figureCitation>
, 
<figureCitation id="AADB89D3124AFFF0FBC3FE3FFBEBAB76" box="[1125,1140,509,534]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
and 
<figureCitation id="AADB89D3124AFFF0FB09FE3FFB20AB76" box="[1199,1215,509,534]" captionStart="Fig" captionStartId="8.[174,209,1925,1945]" captionTargetBox="[196,1418,255,1898]" captionTargetId="figure-6@8.[177,1448,240,1911]" captionTargetPageId="8" captionText="Fig. 6. Male genital organs of Psammoecus delicatus Grouvelle, paralectotype of Psammoecus x-notatus Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="5" pageNumber="6">6</figureCitation>
)
</paragraph>
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<treatmentCitationGroup id="12F0B278124AFFF0FC8BFDFAFBF7ABE1" pageId="5" pageNumber="6">
<treatmentCitation id="B341B347124AFFF0FC8BFDFAFB41AB2C" author="Grouvelle A." box="[813,1246,567,588]" page="477" pageId="5" pageNumber="6" year="1908">
<taxonomicName id="F5E0EED5124AFFF0FC8BFDFAFB41AB2C" authority="Grouvelle, 1908: 477" authorityName="Grouvelle" authorityPageNumber="477" authorityYear="1908" box="[813,1246,567,588]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="00944944124AFFF0FC8BFDFAFB96AB2C" box="[813,1033,568,588]" italics="true" pageId="5" pageNumber="6">Psammoecus delicatus</emphasis>
<bibRefCitation id="5671E8A7124AFFF0FBB7FDF5FB41AB2C" author="Grouvelle A." box="[1041,1246,567,588]" pageId="5" pageNumber="6" pagination="315 - 495" refId="ref12782" refString="Grouvelle A. 1908. Coleopteres de la region Indienne. Rhysodidae, Trogositidae, Nitidulidae, Colydiidae, Cucujidae. Annales de la Societe Entomologique de France 77: 315 - 495." type="journal article" year="1908">Grouvelle, 1908: 477</bibRefCitation>
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. 
<typeStatus id="ED5B2BF4124AFFF0FB41FDFAFA8AAB2C" box="[1255,1301,568,588]" pageId="5" pageNumber="6">Type</typeStatus>
locality: 
<collectingCountry id="4AF7D5C6124AFFF0FACDFDF5FC00AB06" name="Sri Lanka" pageId="5" pageNumber="6">Sri Lanka</collectingCountry>
; 
<typeStatus id="ED5B2BF4124AFFF0FC01FD90FC4AAB06" box="[935,981,594,614]" pageId="5" pageNumber="6">Type</typeStatus>
examined. - 
<treatmentCitation id="B341B347124AFFF0FBF4FD90FB51AB06" author="Pal TK" box="[1106,1230,594,614]" page="21" pageId="5" pageNumber="6" year="1985">
<bibRefCitation id="5671E8A7124AFFF0FBF4FD90FB51AB06" author="Pal TK" box="[1106,1230,594,614]" pageId="5" pageNumber="6" pagination="1 - 54" refId="ref13613" refString="Pal TK. 1985. A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae). Records of the Zoological Survey of India 71: 1 - 54." type="journal article" year="1985">Pal 1985: 21</bibRefCitation>
</treatmentCitation>
, fig. 6. - 
<treatmentCitation id="B341B347124AFFF0FAB9FD90FB8DABE1" author="Sen Gupta T &amp; Pal TK" page="172" pageId="5" pageNumber="6" year="1996">
<bibRefCitation id="5671E8A7124AFFF0FAB9FD90FB8DABE1" author="Sen Gupta T &amp; Pal TK" pageId="5" pageNumber="6" refId="ref13823" refString="Sen Gupta T, Pal TK. 1996. Fauna of India and the Adjacent Countries. Calvicornia [sic]: Coleoptera. Family Silvanidae. Zoological Survey of India, Calcutta." type="book" year="1996">Sen Gupta and Pal 1996: 172</bibRefCitation>
</treatmentCitation>
, fig. 243.
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</paragraph>
<paragraph id="325F9556124AFFF3FC8BFD4AFE94A800" blockId="5.[813,1414,567,668]" lastBlockId="6.[174,776,252,459]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<treatmentCitationGroup id="12F0B278124AFFF3FC8BFD4AFE94A800" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<taxonomicName id="F5E0EED5124AFFF0FC8BFD4AFB54ABFC" ID-CoL="6YB5" authorityName="Latreille" authorityYear="1829" box="[813,1227,647,668]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="x-notatus">
<emphasis id="00944944124AFFF0FC8BFD4AFB83ABFC" box="[813,1052,648,668]" italics="true" pageId="5" pageNumber="6">Psammoecus x-notatus</emphasis>
<treatmentCitation id="B341B347124AFFF0FB82FD45FB6CABFC" author="Grouvelle A." box="[1060,1267,647,668]" page="84" pageId="5" pageNumber="6" year="1912">
<bibRefCitation id="5671E8A7124AFFF0FB82FD45FB6CABFC" author="Grouvelle A." box="[1060,1267,647,668]" pageId="5" pageNumber="6" pagination="81 - 94" refId="ref12816" refString="Grouvelle A. 1912. Psammoecus nouveaux du Musee de Leide. Notes from the Leyden Museum 34 (2): 81 - 94." type="journal article" year="1912">Grouvelle, 1912: 84</bibRefCitation>
</treatmentCitation>
</taxonomicName>
. Type locality: 
<collectingCountry id="4AF7D5C61249FFF3FF45FF3EFEB1A870" box="[227,302,252,272]" name="Taiwan" pageId="6" pageNumber="7">Taiwan</collectingCountry>
: Akau (= 
<collectingRegion id="F0245BB41249FFF3FE32FF3EFDBDA870" box="[404,546,252,272]" country="Taiwan" name="Pingtung" pageId="6" pageNumber="7">Pingtung City</collectingRegion>
, 
<collectingRegion id="F0245BB41249FFF3FD88FF3EFD41A870" box="[558,734,252,272]" country="Taiwan" name="Pingtung" pageId="6" pageNumber="7">Pingtung County</collectingRegion>
) for 
<typeStatus id="ED5B2BF41249FFF3FF45FED5FEA7A84B" box="[227,312,279,299]" pageId="6" pageNumber="7">lectotype</typeStatus>
(
<specimenCount id="24E65EDF1249FFF3FEE3FED5FE7BA84B" box="[325,484,279,299]" pageId="6" pageNumber="7" type="generic" typeStatus="paralectotypes">3 paralectotypes</specimenCount>
were collected from Kosempo (= Jiaxian District, 
<collectingRegion id="F0245BB41249FFF3FE33FEF0FDB5A825" box="[405,554,305,326]" country="Taiwan" name="Kaohsiung" pageId="6" pageNumber="7">Kaohsiung City</collectingRegion>
)); Type examined. syn. nov.
</treatmentCitationGroup>
</paragraph>
<caption id="669FC5DE124AFFF0FF2EF847FE61AED5" ID-DOI="http://doi.org/10.5281/zenodo.8064343" ID-Zenodo-Dep="8064343" httpUri="https://zenodo.org/record/8064343/files/figure.png" pageId="5" pageNumber="6" startId="5.[136,170,1925,1945]" targetBox="[136,1413,723,1900]" targetPageId="5" targetType="figure">
<paragraph id="325F9556124AFFF0FF2EF847FE61AED5" blockId="5.[136,1413,1925,2031]" pageId="5" pageNumber="6">
<emphasis id="00944944124AFFF0FF2EF847FF59AEF9" bold="true" box="[136,198,1925,1945]" pageId="5" pageNumber="6">Fig. 4.</emphasis>
SEM images of 
<taxonomicName id="F5E0EED5124AFFF0FECFF847FDC0AEF9" authority="Ng &amp; Lin, 2023" authorityName="Ng &amp; Lin" authorityYear="2023" box="[361,607,1925,1945]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="taiwanensis" status="sp. nov.">
<emphasis id="00944944124AFFF0FECFF847FDC0AEF9" box="[361,607,1925,1945]" italics="true" pageId="5" pageNumber="6">Psammoecus taiwanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F124AFFF0FDC0F847FD2EAEF9" box="[614,689,1925,1945]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
(A-C) and 
<taxonomicName id="F5E0EED5124AFFF0FCBEF847FBC6AEF9" authority="Grouvelle" authorityName=", Grouvelle" authorityYear="1912" box="[792,1113,1925,1945]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="harmandi">
<emphasis id="00944944124AFFF0FCBEF847FC67AEF9" box="[792,1016,1925,1945]" italics="true" pageId="5" pageNumber="6">Psammoecus harmandi</emphasis>
Grouvelle
</taxonomicName>
(D-F). A and D habitus B and E pronotum C and F anterolateral elytron.
</paragraph>
</caption>
<footnote id="51FB8958124AFFF0FF2EF81FFE07AE8F" box="[136,408,2013,2031]" pageId="5" pageNumber="6">
<paragraph id="325F9556124AFFF0FF2EF81FFE07AE8F" blockId="5.[136,1413,1925,2031]" box="[136,408,2013,2031]" pageId="5" pageNumber="6">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="325F95561249FFF3FF08FEA5FDE1A8F6" blockId="6.[174,776,252,459]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="12F0B2781249FFF3FF08FEA5FDE1A8F6" pageId="6" pageNumber="7">
<treatmentCitation id="B341B3471249FFF3FF08FEA5FDF2A81B" author="Grouvelle A." box="[174,621,359,379]" page="18" pageId="6" pageNumber="7" year="1919">
<taxonomicName id="F5E0EED51249FFF3FF08FEA5FDF2A81B" authority="Grouvelle, 1919: 18" authorityName="Grouvelle" authorityPageNumber="18" authorityYear="1919" box="[174,621,359,379]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitidior">
<emphasis id="009449441249FFF3FF08FEA5FE14A81B" box="[174,395,359,379]" italics="true" pageId="6" pageNumber="7">Psammoecus nitidior</emphasis>
<bibRefCitation id="5671E8A71249FFF3FE30FEA5FDF2A81B" author="Grouvelle A." box="[406,621,359,379]" pageId="6" pageNumber="7" refId="ref12842" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus Latr. In: Memoires Entomologiques. Etudes sur les coleopteres. II. Societe Entomologiques de France, Paris." type="book" year="1919">Grouvelle, 1919: 18</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="ED5B2BF41249FFF3FDDCFEA5FD33A81B" box="[634,684,359,379]" pageId="6" pageNumber="7">Type</typeStatus>
locality: 
<collectingCountry id="4AF7D5C61249FFF3FF45FE40FEDFA8F6" box="[227,320,386,406]" name="Indonesia" pageId="6" pageNumber="7">Indonesia</collectingCountry>
(
<collectingRegion id="F0245BB41249FFF3FEEDFE40FEE1A8F6" box="[331,382,386,406]" country="Indonesia" name="Jawa Timur" pageId="6" pageNumber="7">Java</collectingRegion>
); 
<typeStatus id="ED5B2BF41249FFF3FE2AFE40FE25A8F6" box="[396,442,386,406]" pageId="6" pageNumber="7">Type</typeStatus>
examined. syn. nov.
</treatmentCitationGroup>
</paragraph>
<paragraph id="325F95561249FFF3FF08FE5EFDDEA8AB" blockId="6.[174,776,252,459]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="12F0B2781249FFF3FF08FE5EFDDEA8AB" pageId="6" pageNumber="7">
<treatmentCitation id="B341B3471249FFF3FF08FE5EFDF1A8D0" author="Grouvelle A." box="[174,622,412,432]" page="28" pageId="6" pageNumber="7" year="1919">
<taxonomicName id="F5E0EED51249FFF3FF08FE5EFDF1A8D0" authority="Grouvelle, 1919: 28" authorityName="Grouvelle" authorityPageNumber="28" authorityYear="1919" box="[174,622,412,432]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="009449441249FFF3FF08FE5EFE12A8D0" box="[174,397,412,432]" italics="true" pageId="6" pageNumber="7">Psammoecus raffrayi</emphasis>
<bibRefCitation id="5671E8A71249FFF3FE31FE5EFDF1A8D0" author="Grouvelle A." box="[407,622,412,432]" pageId="6" pageNumber="7" refId="ref12842" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus Latr. In: Memoires Entomologiques. Etudes sur les coleopteres. II. Societe Entomologiques de France, Paris." type="book" year="1919">Grouvelle, 1919: 28</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. 
<typeStatus id="ED5B2BF41249FFF3FDDDFE5EFD32A8D0" box="[635,685,412,432]" pageId="6" pageNumber="7">Type</typeStatus>
locality: 
<collectingCountry id="4AF7D5C61249FFF3FF45FE75FED7A8AB" box="[227,328,439,459]" name="Singapore" pageId="6" pageNumber="7">Singapore</collectingCountry>
; 
<typeStatus id="ED5B2BF41249FFF3FEF6FE75FEE1A8AB" box="[336,382,439,459]" pageId="6" pageNumber="7">Type</typeStatus>
examined. syn. nov.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="7AFAC6DD1249FFFCFF4CFE30FF74AC96" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="325F95561249FFF3FF4CFE30FEDDAF4B" blockId="6.[174,776,498,1963]" pageId="6" pageNumber="7">
<emphasis id="009449441249FFF3FF4CFE30FE13AB6B" box="[234,396,498,523]" italics="true" pageId="6" pageNumber="7">Type series</emphasis>
: 
<materialsCitation id="82889F0B1249FFF3FE03FE30FDDEABEB" collectionCode="MNHN" country="Sri Lanka" location="Grouvelle's" pageId="6" pageNumber="7" specimenCount="1" typeStatus="lectotype">
<typeStatus id="ED5B2BF41249FFF3FE03FE30FDB4AB6B" box="[421,555,498,523]" pageId="6" pageNumber="7">Lectotype</typeStatus>
: male, “ 
<collectingCountry id="4AF7D5C61249FFF3FD0EFE30FC9BAB6B" box="[680,772,498,524]" name="Sri Lanka" pageId="6" pageNumber="7">Ceylan</collectingCountry>
| Horn.”; “Type” [red label]; “Museum Paris | Collection | Grouvelle”; “ 
<taxonomicName id="F5E0EED51249FFF3FE7DFDF0FDE0AB2B" authorityName="Latreille" authorityYear="1829" box="[475,639,562,587]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="009449441249FFF3FE7DFDF0FDE0AB2B" box="[475,639,562,587]" italics="true" pageId="6" pageNumber="7">Psammoecus</emphasis>
</taxonomicName>
| 
<taxonomicName id="F5E0EED51249FFF3FD3DFDF0FC98AB2B" box="[667,775,562,587]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441249FFF3FD3DFDF0FC98AB2B" box="[667,775,562,587]" italics="true" pageId="6" pageNumber="7">delicatus</emphasis>
</taxonomicName>
| ty. | Grouv” [
<location id="373FC38D1249FFF3FEFDFD90FE78AB0C" LSID="urn:lsid:plazi:treatment:BA492440124AFFFCFCF9FE1FFF74AC96:373FC38D1249FFF3FEFDFD90FE78AB0C" box="[347,487,594,620]" country="Sri Lanka" name="Grouvelle's" pageId="6" pageNumber="7">Grouvelle’s</location>
hand] (
<collectionCode id="54F10D931249FFF3FDEEFD90FD3FAB0B" box="[584,672,594,619]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by MK in 2015, here designated)
</materialsCitation>
. 
<typeStatus id="ED5B2BF41249FFF3FDE8FDB0FC9CABEB" box="[590,771,626,651]" pageId="6" pageNumber="7">Paralectotypes</typeStatus>
: 
<specimenCount id="24E65EDF1249FFF3FF08FD50FEADABCB" box="[174,306,658,683]" pageId="6" pageNumber="7" type="female">4 females</specimenCount>
, “Sumatra | Palembang”; “ 
<taxonomicName id="F5E0EED51249FFF3FD2DFD50FD63ABCB" box="[651,764,658,683]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441249FFF3FD2DFD50FD63ABCB" box="[651,764,658,683]" italics="true" pageId="6" pageNumber="7">delicatus</emphasis>
</taxonomicName>
” [printed label] (
<collectionCode id="54F10D931249FFF3FED6FD70FE54ABAB" box="[368,459,690,715]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by MK in 2015, deposited in the collection of Grouvelle and Horn). 
<materialsCitation id="82889F0B1249FFF3FEAFFD30FC9CAA2B" collectionCode="MNHN" country="Sri Lanka" pageId="6" pageNumber="7" specimenCount="1" typeStatus="lectotype">
<typeStatus id="ED5B2BF41249FFF3FEAFFD30FE1AAA6B" box="[265,389,754,779]" pageId="6" pageNumber="7">Lectotype</typeStatus>
of 
<taxonomicName id="F5E0EED51249FFF3FE15FD30FDB1AA6B" authorityName="Grouvelle" authorityYear="1919" box="[435,558,754,779]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitidior" status="syn. nov.">
<emphasis id="009449441249FFF3FE15FD30FE5AAA6B" box="[435,453,754,779]" italics="true" pageId="6" pageNumber="7">P</emphasis>
. 
<emphasis id="009449441249FFF3FE7FFD30FDB1AA6B" box="[473,558,754,779]" italics="true" pageId="6" pageNumber="7">nitidior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F1249FFF3FD9CFD30FD2DAA6B" box="[570,690,754,779]" pageId="6" pageNumber="7">syn. nov.</taxonomicNameLabel>
: male, “Java”; “Type” [red label]; “ 
<taxonomicName id="F5E0EED51249FFF3FDAAFCD0FDFEAA4B" authorityName="Grouvelle" authorityYear="1919" box="[524,609,786,811]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitidior">
<emphasis id="009449441249FFF3FDAAFCD0FDFEAA4B" box="[524,609,786,811]" italics="true" pageId="6" pageNumber="7">nitidior</emphasis>
</taxonomicName>
| ty. Grouv”. (
<collectionCode id="54F10D931249FFF3FF12FCF0FE95AA2B" box="[180,266,818,843]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by TY in 2017, here designated)
</materialsCitation>
. 
<typeStatus id="ED5B2BF41249FFF3FF08FC90FEC1AA0B" box="[174,350,850,875]" pageId="6" pageNumber="7">Paralectotypes</typeStatus>
of 
<taxonomicName id="F5E0EED51249FFF3FE23FC90FE6AAA0B" authorityName="Grouvelle" authorityYear="1919" box="[389,501,850,875]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nitidior" status="syn. nov.">
<emphasis id="009449441249FFF3FE23FC90FE08AA0B" box="[389,407,850,875]" italics="true" pageId="6" pageNumber="7">P</emphasis>
. 
<emphasis id="009449441249FFF3FE00FC90FE6AAA0B" box="[422,501,850,875]" italics="true" pageId="6" pageNumber="7">nitidior</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F1249FFF3FE5BFC90FDF4AA0B" box="[509,619,850,875]" pageId="6" pageNumber="7">syn. nov.</taxonomicNameLabel>
: 6 exs, same label data as the 
<typeStatus id="ED5B2BF41249FFF3FED8FCB0FE72AAEB" box="[382,493,882,907]" pageId="6" pageNumber="7">lectotype</typeStatus>
. (
<collectionCode id="54F10D931249FFF3FDA6FCB0FDC9AAEB" box="[512,598,882,907]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by TY in 2017, here designated). 
<materialsCitation id="82889F0B1249FFF3FE53FC50FE55AD6B" collectionCode="MNHN" collectorName="A. Grouv" country="Singapore" location="Singapore" pageId="6" pageNumber="7" specimenCount="1" typeStatus="lectotype">
<typeStatus id="ED5B2BF41249FFF3FE53FC50FDF6AACB" box="[501,617,914,939]" pageId="6" pageNumber="7">Lectotype</typeStatus>
of 
<taxonomicName id="F5E0EED51249FFF3FD34FC50FC99AACB" authorityName="Grouvelle" authorityYear="1919" box="[658,774,914,939]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="raffrayi" status="syn. nov.">
<emphasis id="009449441249FFF3FD34FC50FD3BAACB" box="[658,676,914,939]" italics="true" pageId="6" pageNumber="7">P</emphasis>
. 
<emphasis id="009449441249FFF3FD13FC50FC99AACB" box="[693,774,914,939]" italics="true" pageId="6" pageNumber="7">raffrayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1BA7F43F1249FFF3FF08FC70FE80AAAB" box="[174,287,946,971]" pageId="6" pageNumber="7">syn. nov.</taxonomicNameLabel>
: female, “ 
<collectingCountry id="4AF7D5C61249FFF3FE3EFC70FD86AAAC" box="[408,537,946,972]" name="Singapore" pageId="6" pageNumber="7">Singapore</collectingCountry>
”; “Type” [red label]; “ 
<emphasis id="009449441249FFF3FF11FC10FE8CAA8B" box="[183,275,978,1003]" italics="true" pageId="6" pageNumber="7">Raffrayi</emphasis>
| ty. 
<collectorName id="9F15F0801249FFF3FEF6FC10FE5FAA8C" box="[336,448,978,1004]" pageId="6" pageNumber="7">A. Grouv</collectorName>
”. (
<collectionCode id="54F10D931249FFF3FE7BFC10FDACAA8B" box="[477,563,978,1003]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by TY in 2017, here designated)
</materialsCitation>
. 
<materialsCitation id="82889F0B1249FFF3FE71FC30FC9CADAB" collectionCode="RMNH" country="Taiwan" location="Grouvelle's" pageId="6" pageNumber="7" specimenCount="1" typeStatus="lectotype">
<typeStatus id="ED5B2BF41249FFF3FE71FC30FDD4AD6B" box="[471,587,1010,1035]" pageId="6" pageNumber="7">Lectotype</typeStatus>
of 
<emphasis id="009449441249FFF3FDD0FC30FD17AD6B" box="[630,648,1010,1035]" italics="true" pageId="6" pageNumber="7">P</emphasis>
. 
<emphasis id="009449441249FFF3FD3FFC30FC98AD6B" box="[665,775,1010,1035]" italics="true" pageId="6" pageNumber="7">x-notatus</emphasis>
syn. nov.: male, “H. | Sauter. | Akau | 
<collectingCountry id="4AF7D5C61249FFF3FD2CFBD0FD69AD4B" box="[650,758,1042,1067]" name="Taiwan" pageId="6" pageNumber="7">Formosa</collectingCountry>
| 1-10.12.07 [round label]”; “ 
<taxonomicName id="F5E0EED51249FFF3FDA7FBF0FD27AD2B" authorityName="Boudier" authorityYear="1834" box="[513,696,1074,1099]" class="Insecta" family="Silvanidae" genus="Psammoechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="009449441249FFF3FDA7FBF0FD27AD2B" box="[513,696,1074,1099]" italics="true" pageId="6" pageNumber="7">Psammoechus</emphasis>
</taxonomicName>
[sic] | 
<emphasis id="009449441249FFF3FF08FB90FE84AD0B" box="[174,283,1106,1131]" italics="true" pageId="6" pageNumber="7">x-nigrum</emphasis>
| type. Grouv.” [not 
<location id="373FC38D1249FFF3FD87FB90FD32AD0C" LSID="urn:lsid:plazi:treatment:BA492440124AFFFCFCF9FE1FFF74AC96:373FC38D1249FFF3FD87FB90FD32AD0C" box="[545,685,1106,1132]" country="Taiwan" name="Grouvelle's" pageId="6" pageNumber="7">Grouvelle’s</location>
hand]; “Museum Leiden. | 
<emphasis id="009449441249FFF3FE0CFBB0FDB7ADEB" box="[426,552,1138,1163]" italics="true" pageId="6" pageNumber="7">Psammoe</emphasis>
| 
<emphasis id="009449441249FFF3FDE1FBB0FDEDADEB" box="[583,626,1138,1163]" italics="true" pageId="6" pageNumber="7">cus</emphasis>
| 
<emphasis id="009449441249FFF3FD36FBB0FC98ADEB" box="[656,775,1138,1163]" italics="true" pageId="6" pageNumber="7">x notatus</emphasis>
| Det: Grouv”; “type” [blue label]; “Cat. No | 5” (
<collectionCode id="54F10D931249FFF3FF12FB70FE95ADAB" box="[180,266,1202,1227]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
; studied by MK in 2014, here designated)
</materialsCitation>
. 
<typeStatus id="ED5B2BF41249FFF3FF08FB10FEF8AD8B" box="[174,359,1234,1259]" pageId="6" pageNumber="7">Paralectotypes</typeStatus>
of 
<emphasis id="009449441249FFF3FE32FB10FE39AD8B" box="[404,422,1234,1259]" italics="true" pageId="6" pageNumber="7">P</emphasis>
. 
<emphasis id="009449441249FFF3FE1FFB10FDB3AD8B" box="[441,556,1234,1259]" italics="true" pageId="6" pageNumber="7">x-notatus</emphasis>
syn. nov.: 2 exs, same label data as the 
<typeStatus id="ED5B2BF41249FFF3FE6AFB30FDA4AC6B" box="[460,571,1266,1291]" pageId="6" pageNumber="7">lectotype</typeStatus>
. (
<collectionCode id="54F10D931249FFF3FDE8FB30FD3BAC6B" box="[590,676,1266,1291]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by TY in 2017, here designated); 
<specimenCount id="24E65EDF1249FFF3FDF5FAD0FD5CAC4B" box="[595,707,1298,1323]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
, “H. | Sauter. | Kosempo | 
<collectingCountry id="4AF7D5C61249FFF3FE1EFAF0FDB9AC2B" box="[440,550,1330,1355]" name="Taiwan" pageId="6" pageNumber="7">Formosa</collectingCountry>
| Jan. 08” [round label]; “ 
<taxonomicName id="F5E0EED51249FFF3FEBEFA90FE56AC0B" authorityName="Latreille" authorityYear="1829" box="[280,457,1362,1387]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="009449441249FFF3FEBEFA90FE56AC0B" box="[280,457,1362,1387]" italics="true" pageId="6" pageNumber="7">Psammoecus</emphasis>
</taxonomicName>
| 
<emphasis id="009449441249FFF3FE4DFA90FDC0AC0B" box="[491,607,1362,1387]" italics="true" pageId="6" pageNumber="7">x nigrum</emphasis>
| G. Grouv” [Grouvelle’s hand]; “type”; “Museum Leiden. | 
<taxonomicName id="F5E0EED51249FFF3FF08FA50FECFACCB" authorityName="Latreille" authorityYear="1829" box="[174,336,1426,1451]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="009449441249FFF3FF08FA50FECFACCB" box="[174,336,1426,1451]" italics="true" pageId="6" pageNumber="7">Psammoecus</emphasis>
</taxonomicName>
| 
<emphasis id="009449441249FFF3FEC0FA50FE4DACCB" box="[358,466,1426,1451]" italics="true" pageId="6" pageNumber="7">x notatus</emphasis>
| Det: Grouv”; “type” [blue label]; “Type” [red label] (
<collectionCode id="54F10D931249FFF3FE7CFA70FDAFACAB" box="[474,560,1458,1483]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
; studied by MK in 2014, here designated); 
<specimenCount id="24E65EDF1249FFF3FE78FA10FDACAC8B" box="[478,563,1490,1515]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
and 
<specimenCount id="24E65EDF1249FFF3FDD3FA10FD29AC8B" box="[629,694,1490,1515]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same label data. (
<collectionCode id="54F10D931249FFF3FEE4FA30FE04AF6B" box="[322,411,1522,1547]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
; studied by TY in 2017, here designated).
</paragraph>
<paragraph id="325F95561249FFF3FF4CF9F0FAA5AED7" blockId="6.[174,776,498,1963]" lastBlockId="6.[850,1452,253,1975]" pageId="6" pageNumber="7">
<emphasis id="009449441249FFF3FF4CF9F0FD90AF2B" box="[234,527,1586,1611]" italics="true" pageId="6" pageNumber="7">Specimens examined</emphasis>
: 
<collectingCountry id="4AF7D5C61249FFF3FD80F9F0FD04AF2B" box="[550,667,1586,1611]" name="Taiwan" pageId="6" pageNumber="7">TAIWAN</collectingCountry>
: [
<collectingRegion id="F0245BB41249FFF3FD15F9F0FE97AF0C" country="Taiwan" name="Miaoli" pageId="6" pageNumber="7">Miaoli County</collectingRegion>
] 
<specimenCount id="24E65EDF1249FFF3FEBFF990FEF0AF0B" box="[281,367,1618,1643]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FD85F990FDECAF0C" box="[547,627,1618,1644]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 3235- 1443), Sanyi, 
<date id="465EB3961249FFF3FEC5F9B0FD82AFEB" box="[355,541,1650,1675]" pageId="6" pageNumber="7" value="1999-06-14" valueMax="1999-06-16" valueMin="1999-06-14">14-16-VI-1999</date>
, C. S. Lin &amp; W. T. Yang leg., UV Light. (
<collectionCode id="54F10D931249FFF3FE02F950FE65AFCC" box="[420,506,1682,1708]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
). [
<collectingRegion id="F0245BB41249FFF3FDB7F950FD2CAFCC" box="[529,691,1682,1708]" country="Taiwan" name="Taichung" pageId="6" pageNumber="7">Taichung City</collectingRegion>
] 2 exs, Wufeng, Wufeng Disctrict, 
<date id="465EB3961249FFF3FE58F970FD15AFAB" box="[510,650,1714,1739]" pageId="6" pageNumber="7" value="1992-04-24">24-IV-1992</date>
, Smetana leg. (
<collectionCode id="54F10D931249FFF3FF4EF910FEDFAF8C" box="[232,320,1746,1772]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="7" type="Museum">MHNG</collectionCode>
; studied by MK); 
<specimenCount id="24E65EDF1249FFF3FDA9F910FDD1AF8B" box="[527,590,1746,1771]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Fengyuan, 
<date id="465EB3961249FFF3FD46F910FE8DAE6B" pageId="6" pageNumber="7" value="1990-04-22">22- IV-1990</date>
, Smetana leg. (
<collectionCode id="54F10D931249FFF3FE7EF930FDADAE6C" box="[472,562,1778,1804]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="6" pageNumber="7" type="Museum">MHNG</collectionCode>
; studied by MK). [
<collectingRegion id="F0245BB41249FFF3FF15F8D0FEF3AE4C" box="[179,364,1810,1836]" country="Taiwan" name="Nantou" pageId="6" pageNumber="7">Nantou County</collectingRegion>
] 
<specimenCount id="24E65EDF1249FFF3FEDDF8D0FE51AE4B" box="[379,462,1810,1835]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FDDFF8D0FD57AE4C" box="[633,712,1810,1836]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 1602-131), Chushan (= Zhushan Township), 
<date id="465EB3961249FFF3FD70F8F0FE8AAE0B" pageId="6" pageNumber="7" value="1993-11-08" valueMax="1993-11-09" valueMin="1993-11-08">8-9- XI-1993</date>
, C. S. Lin &amp; W. T. Yang leg., UV Light. (
<collectionCode id="54F10D931249FFF3FF13F8B0FE8FAEEC" box="[181,272,1906,1932]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FE81F8B0FEF8AEEB" box="[295,359,1906,1931]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FD86F8B0FDEEAEEC" box="[544,625,1906,1932]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 3513- 284), Lienhuachih, Yuchi Township, 
<date id="465EB3961249FFF3FD35F850FC5CA876" box="[659,963,253,1963]" pageId="6" pageNumber="7" value="1998-07-06" valueMax="1998-08-12" valueMin="1998-07-06">6-VII-12- VIII-1998</date>
, C. S. Lin &amp; W. T. Yang leg., Malaise Trap (
<collectionCode id="54F10D931249FFF3FCFFFEDFFC2DA857" box="[857,946,285,311]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FC6EFEDFFB95A856" box="[968,1034,285,310]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, NW Yuchi, Yuchi Township, 
<date id="465EB3961249FFF3FADEFEDFFC0AA836" pageId="6" pageNumber="7" value="2009-10-08">8-X-2009</date>
, Dányi &amp; Lazányi leg. (
<collectionCode id="54F10D931249FFF3FB66FEFFFA85A836" box="[1216,1306,317,342]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
: studied by MK); 
<specimenCount id="24E65EDF1249FFF3FC31FE9FFC49A816" box="[919,982,349,374]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FB2CFE9FFB45A817" box="[1162,1242,349,375]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 1582-328), Fenghuangku, Lugu Township, 
<date id="465EB3961249FFF3FB74FEBFFA1CA8F6" box="[1234,1411,381,406]" pageId="6" pageNumber="7" value="1993-09-20" valueMax="1993-09-22" valueMin="1993-09-20">20-22-IX-1993</date>
, C. S. Lin &amp; M. L. Chan leg., UV Light (
<collectionCode id="54F10D931249FFF3FB54FE5FFAD6A8D7" box="[1266,1353,413,439]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FAFDFE5FFA34A8D6" box="[1371,1451,413,438]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(Catalog No. 1337-5), Shih-tzu-tou, 
<date id="465EB3961249FFF3FB51FE7FFAF0A8B6" box="[1271,1391,445,470]" pageId="6" pageNumber="7" value="1987-09-06">6-IX-1987</date>
, I. C. Hsu leg., Sweeping Net. (
<collectionCode id="54F10D931249FFF3FB2FFE1FFB7FA897" box="[1161,1248,477,503]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
). [
<collectingRegion id="F0245BB41249FFF3FB5FFE1FFA39A897" box="[1273,1446,477,503]" country="Taiwan" name="Chiayi" pageId="6" pageNumber="7">Chiayi County</collectingRegion>
] 
<specimenCount id="24E65EDF1249FFF3FCF4FE3FFC05AB76" box="[850,922,509,534]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Chuchi, Zhuqi Township, 
<date id="465EB3961249FFF3FAA0FE3FFA39AB76" box="[1286,1446,509,534]" pageId="6" pageNumber="7" value="1966-07-20">20-VII-1966</date>
, H. Kamiya leg. (
<collectionCode id="54F10D931249FFF3FB85FDDFFBF6AB56" box="[1059,1129,541,566]" pageId="6" pageNumber="7">KUM</collectionCode>
). [
<collectingRegion id="F0245BB41249FFF3FB20FDDFFAD3AB57" box="[1158,1356,541,567]" country="Taiwan" name="Kaohsiung" pageId="6" pageNumber="7">Kaohsiung City</collectingRegion>
] 7 exs, Kosempo (= Jiaxian District), 
<date id="465EB3961249FFF3FB08FDFFFB62AB36" box="[1198,1277,573,598]" pageId="6" pageNumber="7" value="1908-01">I-1908</date>
, H. Sauter leg. (
<collectionCode id="54F10D931249FFF3FCFFFD9FFC34AB16" box="[857,939,605,630]" pageId="6" pageNumber="7">ZMHB</collectionCode>
, 
<collectionCode id="54F10D931249FFF3FC10FD9FFB95AB16" box="[950,1034,605,630]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
; studied by MK); 
<specimenCount id="24E65EDF1249FFF3FB71FD9FFAB7AB16" box="[1239,1320,605,630]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
and 4 exs, same label data. (
<collectionCode id="54F10D931249FFF3FB8DFDBFFBEEABF7" box="[1067,1137,637,663]" pageId="6" pageNumber="7">SDEI</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FB20FDBFFB5AABF6" box="[1158,1221,637,662]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Mt. Ta Yuen Shan near Liouguei, 
<date id="465EB3961249FFF3FBABFD5FFB15ABD6" box="[1037,1162,669,694]" pageId="6" pageNumber="7" value="1989-06-05">5-VI-1989</date>
, K. Baba leg. (
<collectionCode id="54F10D931249FFF3FAE5FD5FFA18ABD6" box="[1347,1415,669,694]" pageId="6" pageNumber="7">KUM</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FA3AFD5FFCE5ABB6" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Liu Kui (= Liouguei), 
<date id="465EB3961249FFF3FB24FD7FFA95ABB6" box="[1154,1290,701,726]" pageId="6" pageNumber="7" value="1989-08-02">2-VIII-1989</date>
, K. Baba leg. (
<collectionCode id="54F10D931249FFF3FCFCFD1FFC01AB96" box="[858,926,733,758]" pageId="6" pageNumber="7">KUM</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FC15FD1FFC6BAB96" box="[947,1012,733,758]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, same locality, 
<date id="465EB3961249FFF3FB14FD1FFAA4AB96" box="[1202,1339,733,758]" pageId="6" pageNumber="7" value="1986-03-30">30-III-1986</date>
, K. Baba leg. (
<collectionCode id="54F10D931249FFF3FC37FD3FFC7BAA76" box="[913,996,765,790]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="6" pageNumber="7">EUMJ</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FC5CFD3FFBA2AA76" box="[1018,1085,765,790]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Shanping Forest Recreation Area, near Liukuei, 
<date id="465EB3961249FFF3FB9EFCDFFB79AA56" box="[1080,1254,797,822]" pageId="6" pageNumber="7" value="2009-11-19" valueMax="2009-11-21" valueMin="2009-11-19">19-21-XI-2009</date>
, Ronkay &amp; Merkl leg. (
<collectionCode id="54F10D931249FFF3FC32FCFFFC70AA36" box="[916,1007,829,854]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
; studied by MK). [
<collectingRegion id="F0245BB41249FFF3FB7BFCFFFA3AAA37" box="[1245,1445,829,855]" country="Taiwan" name="Taitung" pageId="6" pageNumber="7">Taitung County</collectingRegion>
] 8 exs, Pilam (= 
<collectingRegion id="F0245BB41249FFF3FBBBFC9FFBE1AA16" box="[1053,1150,861,886]" country="Taiwan" name="Taitung" pageId="6" pageNumber="7">Taitung</collectingRegion>
), 
<date id="465EB3961249FFF3FB35FC9FFB71AA16" box="[1171,1262,861,886]" pageId="6" pageNumber="7" value="1908-02">II-1908</date>
, H. Sauter leg. (
<collectionCode id="54F10D931249FFF3FCFFFCBFFC30AAF6" box="[857,943,893,918]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
; studied by MK). [
<collectingRegion id="F0245BB41249FFF3FB27FCBFFAD4AAF7" box="[1153,1355,893,919]" country="Taiwan" name="Pingtung" pageId="6" pageNumber="7">Pingtung County</collectingRegion>
] 11 exs, Akau (= 
<collectingRegion id="F0245BB41249FFF3FC18FC5FFBF7AAD7" box="[958,1128,925,951]" country="Taiwan" name="Pingtung" pageId="6" pageNumber="7">Pingtung City</collectingRegion>
), 
<date id="465EB3961249FFF3FBDBFC5FFAB6AAD6" box="[1149,1321,925,950]" pageId="6" pageNumber="7" value="1907-12-01" valueMax="1907-12-10" valueMin="1907-12-01">1-10-XII-1907</date>
, H. Sauter leg. (
<collectionCode id="54F10D931249FFF3FC2BFC7FFC7CAAB6" box="[909,995,957,982]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="6" pageNumber="7">HNHM</collectionCode>
; studied by MK); 7 exs, same locality, 
<date id="465EB3961249FFF3FCF4FC1FFC61AA96" box="[850,1022,989,1014]" pageId="6" pageNumber="7" value="1907-12-01" valueMax="1907-12-10" valueMin="1907-12-01">1-10-XII-1907</date>
, H. Sauter leg. (
<collectionCode id="54F10D931249FFF3FB6CFC1FFA81AA96" box="[1226,1310,989,1014]" pageId="6" pageNumber="7">ZMHB</collectionCode>
; studied by MK); 
<specimenCount id="24E65EDF1249FFF3FC34FC3FFC70AD76" box="[914,1007,1021,1046]" pageId="6" pageNumber="7" type="male">2 males</specimenCount>
and 8 exs, same locality, 
<date id="465EB3961249FFF3FAB8FC3FFA19AD76" box="[1310,1414,1021,1046]" pageId="6" pageNumber="7" value="1907-12">XII-1907</date>
, H. Sauter leg. (
<collectionCode id="54F10D931249FFF3FC42FBDFFBB5AD57" box="[996,1066,1053,1079]" pageId="6" pageNumber="7">SDEI</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FB9BFBDFFBE7AD56" box="[1085,1144,1053,1078]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FAB9FBDFFAF1AD57" box="[1311,1390,1053,1079]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6920-303), Nanjenshan, Manzhou Township, 
<date id="465EB3961249FFF3FACBFBFFFC2DAD16" pageId="6" pageNumber="7" value="2009-04-01" valueMax="2009-04-19" valueMin="2009-04-01">1-19- IV-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., SLAM Trap. (
<collectionCode id="54F10D931249FFF3FCFFFBBFFC2BADF7" box="[857,948,1149,1175]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FC6DFBBFFB94ADF6" box="[971,1035,1149,1174]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FB62FBBFFA8AADF7" box="[1220,1301,1149,1175]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6920- 82), same locality &amp; same period, M. L. Jeng &amp; T. R. Chen leg., Flight Interception Trap. (
<collectionCode id="54F10D931249FFF3FA96FB7FFA17ADB7" box="[1328,1416,1213,1239]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 2 exs (Catalog No. 
<collectionCode id="54F10D931249FFF3FB86FB1FFBF0AD97" box="[1056,1135,1245,1271]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6469-40 &amp; 48), same locality, 
<date id="465EB3961249FFF3FC14FB3FFBE3AC76" box="[946,1148,1277,1302]" pageId="6" pageNumber="7" value="2009-06-21" valueMax="2009-07-04" valueMin="2009-06-21">21-VI-4-VII-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Flight Interception Trap. (
<collectionCode id="54F10D931249FFF3FB6DFADFFABBAC57" box="[1227,1316,1309,1335]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FA9CFADFFA10AC56" box="[1338,1423,1309,1334]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
&amp; 
<specimenCount id="24E65EDF1249FFF3FCF4FAFFFC0FAC36" box="[850,912,1341,1366]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FB9BFAFFFB13AC37" box="[1085,1164,1341,1367]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6469-486 &amp; 503), same locality, 
<date id="465EB3961249FFF3FBA0FA9FFB2BAC16" box="[1030,1204,1373,1398]" pageId="6" pageNumber="7" value="2009-07-04" valueMax="2009-07-14" valueMin="2009-07-04">4-14-VII-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Flight Interception Trap. (
<collectionCode id="54F10D931249FFF3FAA3FABFFAC3ACF7" box="[1285,1372,1405,1431]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FAC9FABFFA33ACF6" box="[1391,1452,1405,1430]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FC56FA5FFBA0ACD7" box="[1008,1087,1437,1463]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6674-322), same locality, 
<date id="465EB3961249FFF3FCF4FA7FFBB9ACB6" box="[850,1062,1469,1494]" pageId="6" pageNumber="7" value="2009-09-27" valueMax="2009-10-12" valueMin="2009-09-27">27-IX-12-X-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Flight Interception Trap. (
<collectionCode id="54F10D931249FFF3FB26FA1FFB48AC97" box="[1152,1239,1501,1527]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FB4CFA1FFAA2AC96" box="[1258,1341,1501,1526]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FC20FA3FFC4AAF77" box="[902,981,1533,1559]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6632-57), Manchou (= Manzhou) Lite Chulaoshushan, 
<date id="465EB3961249FFF3FBC1F9DFFAC6AF56" box="[1127,1369,1565,1590]" pageId="6" pageNumber="7" value="2009-08-30" valueMax="2009-09-14" valueMin="2009-08-30">30-VIII-14-IX-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Malaise Trap. (
<collectionCode id="54F10D931249FFF3FAE1F9FFFA3EAF37" box="[1351,1441,1597,1623]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FCF4F99FFC0FAF16" box="[850,912,1629,1654]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FBE7F99FFB0FAF17" box="[1089,1168,1629,1655]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6579-378), same locality, 
<date id="465EB3961249FFF3FC11F9BFFBF4AFF6" box="[951,1131,1661,1686]" pageId="6" pageNumber="7" value="2009-09-12" valueMax="2009-09-26" valueMin="2009-09-12">12-26-IX-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Malaise Trap. (
<collectionCode id="54F10D931249FFF3FBE0F95FFB01AFD7" box="[1094,1182,1693,1719]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FB15F95FFA99AFD6" box="[1203,1286,1693,1718]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(Catalog No. 
<collectionCode id="54F10D931249FFF3FCF4F97FFC3EAFB7" box="[850,929,1725,1751]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6525-85), Manchou (= Manzhou) Lite No: 169 Hsien Rd. 
<quantity id="F51838B31249FFF3FBE6F91FFB07AF96" box="[1088,1176,1757,1782]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="7" unit="km" value="3.7">3.7 km</quantity>
, 
<date id="465EB3961249FFF3FB03F91FFAC0AF96" box="[1189,1375,1757,1782]" pageId="6" pageNumber="7" value="2009-04-19" valueMax="2009-05-01" valueMin="2009-04-19">19-IV-1-V-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Flight Interception Trap. (
<collectionCode id="54F10D931249FFF3FCFFF8DFFC2CAE57" box="[857,947,1821,1847]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FC6FF8DFFB93AE56" box="[969,1036,1821,1846]" pageId="6" pageNumber="7" type="generic">1 ex.</specimenCount>
, Sichongxi, Checheng Township, 
<date id="465EB3961249FFF3FCF4F8FFFC22AE36" box="[850,957,1853,1878]" pageId="6" pageNumber="7" value="1986-01-01">1-I-1986</date>
, K. Baba leg. (
<collectionCode id="54F10D931249FFF3FBD0F8FFFB57AE36" box="[1142,1224,1853,1878]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="6" pageNumber="7">EUMJ</collectionCode>
); 
<specimenCount id="24E65EDF1249FFF3FB7BF8FFFAA9AE36" box="[1245,1334,1853,1878]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
. (Catalog No. 
<collectionCode id="54F10D931249FFF3FC20F89FFC4AAE17" box="[902,981,1885,1911]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
ENT 6632-57), Manchou (= Manzhou) Lite Chulaoshushan, 
<date id="465EB3961249FFF3FBF5F8BFFA82AEF6" box="[1107,1309,1917,1942]" pageId="6" pageNumber="7" value="2009-10-30" valueMax="2009-09-14" valueMin="2009-10-30">30-X-14-IX-2009</date>
, M. L. Jeng &amp; T. R. Chen leg., Malaise Trap. (
<collectionCode id="54F10D931249FFF3FB7EF85FFAB0AED7" box="[1240,1327,1949,1975]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="7">NMNS</collectionCode>
).
</paragraph>
<footnote id="51FB89581249FFF3FB3AF81FFA34AE8F" box="[1180,1451,2013,2031]" pageId="6" pageNumber="7">
<paragraph id="325F95561249FFF3FB3AF81FFA34AE8F" blockId="6.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="6" pageNumber="7">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="325F95561248FFF2FF63FF3FFE9DAB56" blockId="7.[136,738,253,1174]" pageId="7" pageNumber="8">
<emphasis id="009449441248FFF2FF63FF3FFEA5A876" box="[197,314,253,278]" italics="true" pageId="7" pageNumber="8">Diagnosis</emphasis>
: This species differs from the other Taiwanese species by the very long and slender teeth on lateral margins of its pronotum, the pattern of these teeth, the pubescence composed of moderate and long setae on head, pronotum and elytra, and its large, strongly protruding eyes. 
<taxonomicName id="F5E0EED51248FFF2FF2EFE7FFE03A8B6" box="[136,412,445,470]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441248FFF2FF2EFE7FFE03A8B6" box="[136,412,445,470]" italics="true" pageId="7" pageNumber="8">Psammoecus delicatus</emphasis>
</taxonomicName>
is related to 
<taxonomicName id="F5E0EED51248FFF2FD99FE7FFE9EA896" authority="Pal, 1985" authorityName="Pal" authorityYear="1985" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="complexus">
<emphasis id="009449441248FFF2FD99FE7FFDCEA8B6" box="[575,593,445,470]" italics="true" pageId="7" pageNumber="8">P</emphasis>
. 
<emphasis id="009449441248FFF2FDC4FE7FFD7EA8B6" box="[610,737,445,470]" italics="true" pageId="7" pageNumber="8">complexus</emphasis>
<bibRefCitation id="5671E8A71248FFF2FF2EFE1FFE9EA896" author="Pal TK" box="[136,257,477,502]" pageId="7" pageNumber="8" pagination="1 - 54" refId="ref13613" refString="Pal TK. 1985. A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae). Records of the Zoological Survey of India 71: 1 - 54." type="journal article" year="1985">Pal, 1985</bibRefCitation>
</taxonomicName>
, but can easily be distinguished by the different shape of the lateral pronotal teeth (see 
<bibRefCitation id="5671E8A71248FFF2FF2EFDDFFF68AB56" author="Pal TK" box="[136,247,541,566]" pageId="7" pageNumber="8" pagination="1 - 54" refId="ref13613" refString="Pal TK. 1985. A revision of Indian Psammoecus Latreille (Coleoptera: Silvanidae). Records of the Zoological Survey of India 71: 1 - 54." type="journal article" year="1985">Pal 1985</bibRefCitation>
).
</paragraph>
<paragraph id="325F95561248FFF2FF63FDFFFF29AB16" blockId="7.[136,738,253,1174]" pageId="7" pageNumber="8">
<emphasis id="009449441248FFF2FF63FDFFFED4AB36" box="[197,331,573,598]" italics="true" pageId="7" pageNumber="8">Description</emphasis>
: Body length: 
<quantity id="F51838B31248FFF2FE58FDFFFD39AB36" box="[510,678,573,598]" metricMagnitude="-3" metricUnit="m" metricValue="2.6799999999999997" metricValueMax="2.98" metricValueMin="2.38" pageId="7" pageNumber="8" unit="mm" value="2.6799999999999997" valueMax="2.98" valueMin="2.38">2.38-2.98 mm</quantity>
(
<emphasis id="009449441248FFF2FD1FFDFFFD57AB36" box="[697,712,573,598]" italics="true" pageId="7" pageNumber="8">n</emphasis>
= 16).
</paragraph>
<paragraph id="325F95561248FFF2FF63FDBFFD20AAD7" blockId="7.[136,738,253,1174]" pageId="7" pageNumber="8">
<emphasis id="009449441248FFF2FF63FDBFFED0ABF6" box="[197,335,637,662]" italics="true" pageId="7" pageNumber="8">Coloration</emphasis>
: (
<figureCitation id="AADB89D31248FFF2FEC8FDBFFE7DABF6" box="[366,482,637,662]" captionStart="Fig" captionStartId="2.[174,208,1844,1864]" captionTargetBox="[174,1451,308,1818]" captionTargetId="figure-67@2.[174,1451,308,1818]" captionTargetPageId="2" captionText="Fig. 1. Psammoecus species of Taiwan, habitus. (A) P. taiwanensis sp. nov. (B) P. delicatus Grouvelle (C) P. harmandi Grouvelle (D) P. dentatus Grouvelle (E) P. trimaculatus Motschulsky (F) P. triguttatus Reitter (G) P. labyrinthicus Yoshida and Hirowatari (H) P. simonis Grouvelle (I) P. hiranoi Yoshida and Hirowatari. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064337" httpUri="https://zenodo.org/record/8064337/files/figure.png" pageId="7" pageNumber="8">Figs. 1B</figureCitation>
and 
<figureCitation id="AADB89D31248FFF2FD8BFDBFFDDDABF6" box="[557,578,637,662]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">5</figureCitation>
). Head and pronotum yellowish-brown. Elytra yellowish-brown with variable x-shaped blackish-brown maculae covering posterior 2/3 (
<figureCitation id="AADB89D31248FFF2FE0CFD1FFDC2AB96" box="[426,605,733,758]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">Fig. 5A and B</figureCitation>
); posterior half of maculae in lighter color specimen obscure or absent; sometimes posterior 1/3 of elytra darkened (
<figureCitation id="AADB89D31248FFF2FEB4FCFFFEE6AA37" box="[274,377,829,855]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">Fig. 5C</figureCitation>
). Antennae yellowish-brown basally, 7th to 10th antennomere sometimes darkened, 11th antennomere lighter colored, rarely also 10th antennomere lighter colored (
<figureCitation id="AADB89D31248FFF2FDF1FC5FFD2CAAD7" box="[599,691,925,951]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">Fig. 5C</figureCitation>
).
</paragraph>
<paragraph id="325F95561248FFF2FF63FC7FFAEFA816" blockId="7.[136,738,253,1174]" lastBlockId="7.[813,1414,253,1174]" pageId="7" pageNumber="8">
Head: (
<figureCitation id="AADB89D31248FFF2FE8EFC7FFEE1AAB6" box="[296,382,957,982]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
). Wide, HW/HL 1.34-1.47; IE/HL 0.79-0.89. Temples gradually narrowed toward bases. Eyes large, well prominent. Dorsal surface punctation sparse and weak, pubescence composed of moderate and long setae. Antennae long; antennomeres with moderate and long, erect setae; distal portion of 7th to 10th and entire 11th antennomere covered with short pubescence, dense on 8th to 11th; approximate ratios of antennomere lengths of 
<typeStatus id="ED5B2BF41248FFF2FBF5FEFFFB21A836" box="[1107,1214,317,342]" pageId="7" pageNumber="8">lectotype</typeStatus>
as follows: 3.0: 1.0: 1.3: 1.4: 1.4: 1.5: 1.2: 1.2: 1.2: 1.2: 2.4.
</paragraph>
<paragraph id="325F95561248FFF2FCCFFEBFFC0BAA76" blockId="7.[813,1414,253,1174]" pageId="7" pageNumber="8">
Pronotum: (
<figureCitation id="AADB89D31248FFF2FC5BFEBFFBD4A8F6" box="[1021,1099,381,406]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
). Wide, point of greatest width variable, between anterior 1/3 and middle, distinctly narrowed toward base; lateral areas slightly flattened, PW/PL 1.32-1.42. Punctation on dorsal surface denser and slightly stronger than on vertex. Pubescence composed of numerous moderate and fewer long setae, long seta on each tooth and tubercle on anterior, lateral and posterior margins and anterior and posterior angles. Anterior angle with a few large tubercles; lateral margin with some dorso-laterally bent short to long teeth; posterior angle with a very small tooth and a small tubercle.
</paragraph>
<paragraph id="325F95561248FFF2FCCFFCDFFB58AD76" blockId="7.[813,1414,253,1174]" pageId="7" pageNumber="8">
Elytra: (
<figureCitation id="AADB89D31248FFF2FC73FCDFFBB4AA56" box="[981,1067,797,822]" captionStart="Fig" captionStartId="7.[137,172,1897,1917]" captionTargetBox="[136,1413,1235,1871]" captionTargetId="figure-449@7.[136,1413,1235,1871]" captionTargetPageId="7" captionText="Fig. 5. Habitus of Psammoecus spp. synonymized with Psammoecus delicatus Grouvelle. (A) Psammoecus x-notatus Grouvelle, paralectotype, male (B) Psammoecus nitidior Grouvelle, lectotype, male (C) Psammoecus raffrayi Grouvelle, lectotype, female. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
). Oval, EW/BL 0.39-0.44. Rows of punctures slightly wider than interstices. Pubescence composed of many semi-erect moderate setae, most of interstices with rows of long erect setae, very long erect setae in a row along lateral margins, setae longer toward humeral swelling, anterior setae on small tubercles, with some long setae on lateral margin.
</paragraph>
<paragraph id="325F95561248FFFCFCCFFBDFFEBAAA36" blockId="7.[813,1414,253,1174]" lastBlockId="9.[136,738,253,1526]" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
Male genitalia: (
<figureCitation id="AADB89D31248FFF2FB9EFBDFFB18AD56" box="[1080,1159,1053,1078]" captionStart="Fig" captionStartId="8.[174,209,1925,1945]" captionTargetBox="[196,1418,255,1898]" captionTargetId="figure-6@8.[177,1448,240,1911]" captionTargetPageId="8" captionText="Fig. 6. Male genital organs of Psammoecus delicatus Grouvelle, paralectotype of Psammoecus x-notatus Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
). Spiculum gastrale (
<figureCitation id="AADB89D31248FFF2FC92FBFFFC14AD36" box="[820,907,1085,1110]" captionStart="Fig" captionStartId="8.[174,209,1925,1945]" captionTargetBox="[196,1418,255,1898]" captionTargetId="figure-6@8.[177,1448,240,1911]" captionTargetPageId="8" captionText="Fig. 6. Male genital organs of Psammoecus delicatus Grouvelle, paralectotype of Psammoecus x-notatus Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="7" pageNumber="8">Fig. 6A</figureCitation>
) with short and thick strut, enlarged around base, diverging widely near middle; branches obscure and thin, connected by membrane; lateral sclerites elongate triangular, enlarged inwards near anterior 1/3. Parameres (
<figureCitation id="AADB89D31246FFFCFE5BFEDFFDC6A856" box="[509,601,285,310]" captionStart="Fig" captionStartId="8.[174,209,1925,1945]" captionTargetBox="[196,1418,255,1898]" captionTargetId="figure-6@8.[177,1448,240,1911]" captionTargetPageId="8" captionText="Fig. 6. Male genital organs of Psammoecus delicatus Grouvelle, paralectotype of Psammoecus x-notatus Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="9" pageNumber="10">Fig. 6B</figureCitation>
) small and stout, fused basally, rounded at apices, apex sometimes somewhat angular, dorsally with some small punctures around middle of inner margins, densely covered with setae of various length near apices, dorsally with some short, sparse setae near inner margins of apical 1/3, with a long apicolateral seta. Phallobase (
<figureCitation id="AADB89D31246FFFCFE09FE3FFD98AB76" box="[431,519,509,534]" captionStart="Fig" captionStartId="8.[174,209,1925,1945]" captionTargetBox="[196,1418,255,1898]" captionTargetId="figure-6@8.[177,1448,240,1911]" captionTargetPageId="8" captionText="Fig. 6. Male genital organs of Psammoecus delicatus Grouvelle, paralectotype of Psammoecus x-notatus Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="9" pageNumber="10">Fig. 6B</figureCitation>
) long, subparallel, basal half of upper layer diverging, the branches widening distally, with membrane between bases of the branches, lower layer narrow, exposed around base. Penis (
<figureCitation id="AADB89D31246FFFCFE97FDBFFE0DABF7" box="[305,402,637,663]" captionStart="Fig" captionStartId="8.[174,209,1925,1945]" captionTargetBox="[196,1418,255,1898]" captionTargetId="figure-6@8.[177,1448,240,1911]" captionTargetPageId="8" captionText="Fig. 6. Male genital organs of Psammoecus delicatus Grouvelle, paralectotype of Psammoecus x-notatus Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="9" pageNumber="10">Fig. 6C</figureCitation>
) long, gradually narrowed toward apex, somewhat rounded at apex, with fine dense punctation around apex. Internal sac with very long and thin structure partly ringed around apex, with a pair of small spines near middle (sometimes absent), with a pair of large spines around base.
</paragraph>
<caption id="669FC5DE1248FFF2FF2FF8ABFE91AED5" ID-DOI="http://doi.org/10.5281/zenodo.8064347" ID-Zenodo-Dep="8064347" httpUri="https://zenodo.org/record/8064347/files/figure.png" pageId="7" pageNumber="8" startId="7.[137,172,1897,1917]" targetBox="[136,1413,1235,1871]" targetPageId="7" targetType="figure">
<paragraph id="325F95561248FFF2FF2FF8ABFE91AED5" blockId="7.[136,1415,1897,1973]" pageId="7" pageNumber="8">
<emphasis id="009449441248FFF2FF2FF8ABFF55AE1D" bold="true" box="[137,202,1897,1917]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
Habitus of 
<taxonomicName id="F5E0EED51248FFF2FEE0F8ABFE63AE1D" box="[326,508,1897,1917]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="009449441248FFF2FEE0F8ABFE56AE1D" box="[326,457,1897,1917]" italics="true" pageId="7" pageNumber="8">Psammoecus</emphasis>
spp.
</taxonomicName>
synonymized with 
<taxonomicName id="F5E0EED51248FFF2FD1FF8ABFB97AE1D" authority="Grouvelle." authorityName="Grouvelle." box="[697,1032,1897,1917]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441248FFF2FD1FF8ABFC04AE1D" box="[697,923,1897,1917]" italics="true" pageId="7" pageNumber="8">Psammoecus delicatus</emphasis>
Grouvelle.
</taxonomicName>
(A) 
<emphasis id="009449441248FFF2FB92F8ABFA85AE1D" box="[1076,1306,1897,1917]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="F5E0EED51248FFF2FB92F8ABFB28AE1D" authorityName="Latreille" authorityYear="1829" box="[1076,1207,1897,1917]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
x-notatus
</emphasis>
Grouvelle, paralectotype, male (B) 
<taxonomicName id="F5E0EED51248FFF2FEC9F847FD01AEF9" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1919" box="[367,670,1925,1945]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nitidior">
<emphasis id="009449441248FFF2FEC9F847FDA8AEF9" box="[367,567,1925,1945]" italics="true" pageId="7" pageNumber="8">Psammoecus nitidior</emphasis>
Grouvelle
</taxonomicName>
, lectotype, male (C) 
<taxonomicName id="F5E0EED51248FFF2FCC3F847FB09AEF9" authority="Grouvelle" authorityName="Grouvelle" authorityYear="1919" box="[869,1174,1925,1945]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="009449441248FFF2FCC3F847FBB0AEF9" box="[869,1071,1925,1945]" italics="true" pageId="7" pageNumber="8">Psammoecus raffrayi</emphasis>
Grouvelle
</taxonomicName>
, lectotype, female. Scale bar = 1.0 mm.
</paragraph>
</caption>
<footnote id="51FB89581248FFF2FF2EF81FFE07AE8F" box="[136,408,2013,2031]" pageId="7" pageNumber="8">
<paragraph id="325F95561248FFF2FF2EF81FFE07AE8F" blockId="7.[136,408,2013,2031]" box="[136,408,2013,2031]" pageId="7" pageNumber="8">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<caption id="669FC5DE1247FFFDFF08F847FD0DAED5" ID-DOI="http://doi.org/10.5281/zenodo.8064349" ID-Zenodo-Dep="8064349" httpUri="https://zenodo.org/record/8064349/files/figure.png" pageId="8" pageNumber="9" startId="8.[174,209,1925,1945]" targetBox="[196,1418,255,1898]" targetPageId="8" targetType="figure">
<paragraph id="325F95561247FFFDFF08F847FD0DAED5" blockId="8.[174,1451,1925,1973]" pageId="8" pageNumber="9">
<emphasis id="009449441247FFFDFF08F847FF71AEF9" bold="true" box="[174,238,1925,1945]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
Male genital organs of 
<taxonomicName id="F5E0EED51247FFFDFE44F847FCB4AEF9" authority="Grouvelle" authorityName="Grouvelle" box="[482,811,1925,1945]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441247FFFDFE44F847FD5CAEF9" box="[482,707,1925,1945]" italics="true" pageId="8" pageNumber="9">Psammoecus delicatus</emphasis>
Grouvelle
</taxonomicName>
, paralectotype of 
<emphasis id="009449441247FFFDFC7CF847FB21AEF9" box="[986,1214,1925,1945]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="F5E0EED51247FFFDFC7CF847FBC2AEF9" authorityName="Latreille" authorityYear="1829" box="[986,1117,1925,1945]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
x-notatus
</emphasis>
Grouvelle. (A) spiculum gastrale (B) tegmen (C) penis. Scale bar = 0.2 mm.
</paragraph>
</caption>
<footnote id="51FB89581247FFFDFB3AF81FFA34AE8F" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">
<paragraph id="325F95561247FFFDFB3AF81FFA34AE8F" blockId="8.[1180,1451,2013,2031]" box="[1180,1451,2013,2031]" pageId="8" pageNumber="9">© 2018 Academia Sinica, Taiwan</paragraph>
</footnote>
<paragraph id="325F95561246FFFCFF63FC9FFE13AAF6" blockId="9.[136,738,253,1526]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFF63FC9FFEFAAA16" box="[197,357,861,886]" italics="true" pageId="9" pageNumber="10">Distribution</emphasis>
: 
<collectingCountry id="4AF7D5C61246FFFCFEDBFC9FFE43AA16" box="[381,476,861,886]" name="Taiwan" pageId="9" pageNumber="10">Taiwan</collectingCountry>
; 
<collectingCountry id="4AF7D5C61246FFFCFE52FC9FFDA7AA16" box="[500,568,861,886]" name="India" pageId="9" pageNumber="10">India</collectingCountry>
; 
<collectingCountry id="4AF7D5C61246FFFCFDF7FC9FFD48AA16" box="[593,727,861,887]" name="Sri Lanka" pageId="9" pageNumber="10">Sri Lanka</collectingCountry>
; 
<collectingCountry id="4AF7D5C61246FFFCFF2EFCBFFE99AAF7" box="[136,262,893,919]" name="Singapore" pageId="9" pageNumber="10">Singapore</collectingCountry>
; 
<collectingCountry id="4AF7D5C61246FFFCFEB6FCBFFE17AAF6" box="[272,392,893,918]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
.
</paragraph>
<paragraph id="325F95561246FFFCFF63FC5FFEDBAC76" blockId="9.[136,738,253,1526]" pageId="9" pageNumber="10">
<emphasis id="009449441246FFFCFF63FC5FFEA0AAD6" box="[197,319,925,950]" italics="true" pageId="9" pageNumber="10">Remarks</emphasis>
: The external morphological characters as well as the male genital structures of 
<emphasis id="009449441246FFFCFF2EFC1FFF05AA96" box="[136,154,989,1014]" italics="true" pageId="9" pageNumber="10">P</emphasis>
. 
<emphasis id="009449441246FFFCFF0AFC1FFE84AA96" box="[172,283,989,1014]" italics="true" pageId="9" pageNumber="10">x-notatus</emphasis>
and 
<taxonomicName id="F5E0EED51246FFFCFEFAFC1FFE4FAA96" authorityName="Grouvelle" authorityYear="1919" box="[348,464,989,1014]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nitidior">
<emphasis id="009449441246FFFCFEFAFC1FFEF1AA96" box="[348,366,989,1014]" italics="true" pageId="9" pageNumber="10">P</emphasis>
. 
<emphasis id="009449441246FFFCFE26FC1FFE4FAA96" box="[384,464,989,1014]" italics="true" pageId="9" pageNumber="10">nitidior</emphasis>
</taxonomicName>
type specimens show the typical morphology of 
<taxonomicName id="F5E0EED51246FFFCFE64FC3FFDCCAD76" box="[450,595,1021,1046]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441246FFFCFE64FC3FFE4BAD76" box="[450,468,1021,1046]" italics="true" pageId="9" pageNumber="10">P</emphasis>
. 
<emphasis id="009449441246FFFCFE40FC3FFDCCAD76" box="[486,595,1021,1046]" italics="true" pageId="9" pageNumber="10">delicatus</emphasis>
</taxonomicName>
<emphasis id="009449441246FFFCFDF5FC3FFDC8AD76" box="[595,599,1021,1046]" italics="true" pageId="9" pageNumber="10">.</emphasis>
Hence, we conclude that these species are junior synonyms of 
<taxonomicName id="F5E0EED51246FFFCFF0CFBFFFEA4AD36" box="[170,315,1085,1110]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441246FFFCFF0CFBFFFF23AD36" box="[170,188,1085,1110]" italics="true" pageId="9" pageNumber="10">P</emphasis>
. 
<emphasis id="009449441246FFFCFF68FBFFFEA4AD36" box="[206,315,1085,1110]" italics="true" pageId="9" pageNumber="10">delicatus</emphasis>
</taxonomicName>
. Although Grouvelle’s description of 
<taxonomicName id="F5E0EED51246FFFCFF0FFB9FFEBEAD16" authorityName="Grouvelle" authorityYear="1919" box="[169,289,1117,1142]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="009449441246FFFCFF0FFB9FFF24AD16" box="[169,187,1117,1142]" italics="true" pageId="9" pageNumber="10">P</emphasis>
. 
<emphasis id="009449441246FFFCFF6BFB9FFEBEAD16" box="[205,289,1117,1142]" italics="true" pageId="9" pageNumber="10">raffrayi</emphasis>
</taxonomicName>
suggests that at least 
<specimenCount id="24E65EDF1246FFFCFDE7FB9FFD7EAD16" box="[577,737,1117,1142]" pageId="9" pageNumber="10" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
should be present, only 
<specimenCount id="24E65EDF1246FFFCFE6FFBBFFDC5ADF6" box="[457,602,1149,1174]" pageId="9" pageNumber="10" type="female">one female</specimenCount>
specimen could be found and studied. Since 
<taxonomicName id="F5E0EED51246FFFCFD9EFB5FFD5CADD6" box="[568,707,1181,1206]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="delicatus">
<emphasis id="009449441246FFFCFD9EFB5FFD5CADD6" box="[568,707,1181,1206]" italics="true" pageId="9" pageNumber="10">P. delicatus</emphasis>
</taxonomicName>
is well characterized by its external morphological characters, we have no doubt that 
<taxonomicName id="F5E0EED51246FFFCFD91FB1FFD35AD96" authorityName="Grouvelle" authorityYear="1919" box="[567,682,1245,1270]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="raffrayi">
<emphasis id="009449441246FFFCFD91FB1FFD35AD96" box="[567,682,1245,1270]" italics="true" pageId="9" pageNumber="10">P. raffrayi</emphasis>
</taxonomicName>
is a junior synonym.
</paragraph>
<paragraph id="325F95561246FFFCFF63FADFFF74AC96" blockId="9.[136,738,253,1526]" pageId="9" pageNumber="10">
The unusual morphology of the spiculum gastrale is remarkable compared to that of other 
<taxonomicName id="F5E0EED51246FFFCFF2EFA9FFEB5AC16" authorityName="Latreille" authorityYear="1829" box="[136,298,1373,1398]" class="Insecta" family="Silvanidae" genus="Psammoecus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="009449441246FFFCFF2EFA9FFEB5AC16" box="[136,298,1373,1398]" italics="true" pageId="9" pageNumber="10">Psammoecus</emphasis>
</taxonomicName>
. This may indicate that this species belongs to a distinct species group. Studies of extensive material from the Oriental region revealed numerous closely related yet undescribed species.
</paragraph>
</subSubSection>
</treatment>
</document>