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<document id="3292A6EAD89AC6253CC2F62244579E3B" ID-CLB-Dataset="21035" ID-DOI="10.5281/zenodo.4004389" ID-GBIF-Dataset="03168ca4-19e8-4ef5-a000-0e23e0c1a0d2" ID-ISSN="0253-116X" ID-Zenodo-Dep="4004389" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598542413075" checkinUser="valdenar" docAuthor="Smetana, Tephens &amp; Shavrin, A." docDate="2018" docId="8948DC625D63FFA3FF36FB76FEF2A51A" docLanguage="en" docName="Linzer.biol.Beitr.50.1.0825-0836.pdf.imf" docOrigin="Linzer biologische Beiträge 50 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Quedius (Raphirus) aedilis Smetana &amp; Shavrin, 2018, nov. sp." docType="treatment" docVersion="7" lastPageNumber="834" masterDocId="7571A41A5D6BFFAAFF84FFFEFFCBA633" masterDocTitle="Contribution to the knowledge of the genus Quedius S, 1829 of Siberia and Russian Far East (Coleoptera: Staphylinidae: Staphylinini: Quediina)" masterLastPageNumber="836" masterPageNumber="825" pageNumber="833" updateTime="1698845623269" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="C07A896C4C0CE4578785A84D217D0152">Contribution to the knowledge of the genus Quedius S, 1829 of Siberia and Russian Far East (Coleoptera: Staphylinidae: Staphylinini: Quediina)</mods:title>
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<mods:name id="2412CB737051B64D45436953D60885D0" type="personal">
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<mods:namePart id="4DABAAB8151C14B6CA7AA1F082752DF3">Smetana, Tephens</mods:namePart>
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<mods:namePart id="50F8ECB11FB946B455DE65BDFC49A1BD">Shavrin, A.</mods:namePart>
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<mods:title id="057A8B28A65BCB346CB848E8443516E3">Linzer biologische Beiträge</mods:title>
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<mods:part id="4E2D6C83D26DD507403670BABF6745DA">
<mods:date id="73240210F4DD37BE85E5333833BA3EB1">2018</mods:date>
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<mods:number id="B59F8C52FEDA5CF9CD4052E1F842507D">2018-07-27</mods:number>
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<mods:number id="F6DF97305A3F940AF4F8B91425C7931C">50</mods:number>
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<mods:classification id="3571431A1C4C6D2F2A24E08D0D46AD0C">journal article</mods:classification>
<mods:identifier id="2ECF877D39FE611E6BBDAE14C73C8381" type="CLB-Dataset">21035</mods:identifier>
<mods:identifier id="31A039CCF29AFA2435450EF064F361DE" type="DOI">10.5281/zenodo.4004389</mods:identifier>
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<mods:identifier id="E57D57E8213A8AABDD5A284B4E5EBAEB" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="701340ED86271FB67A7F67034DA731A8" type="Zenodo-Dep">4004389</mods:identifier>
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<treatment id="8948DC625D63FFA3FF36FB76FEF2A51A" ID-DOI="http://doi.org/10.5281/zenodo.4329299" ID-GBIF-Taxon="167376785" ID-Zenodo-Dep="4329299" LSID="urn:lsid:plazi:treatment:8948DC625D63FFA3FF36FB76FEF2A51A" httpUri="http://treatment.plazi.org/id/8948DC625D63FFA3FF36FB76FEF2A51A" lastPageId="9" lastPageNumber="834" pageId="8" pageNumber="833">
<subSubSection id="49FB3EFF5D63FFA2FF36FB76FE1FA2EF" pageId="8" pageNumber="833" type="nomenclature">
<paragraph id="015E6D745D63FFA2FF36FB76FCDBA2AD" blockId="8.[178,784,1160,1182]" box="[178,784,1160,1182]" pageId="8" pageNumber="833">
<heading id="5A16DA185D63FFA2FF36FB76FCDBA2AD" box="[178,784,1160,1182]" fontSize="8" level="2" pageId="8" pageNumber="833" reason="2">
<emphasis id="3395B1665D63FFA2FF36FB76FCDBA2AD" bold="true" box="[178,784,1160,1182]" pageId="8" pageNumber="833">
<taxonomicName id="C6E116F75D63FFA2FF36FB76FE07A2AD" ID-CoL="4R2ZH" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[178,460,1160,1182]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="833" phylum="Arthropoda" rank="species" species="aedilis" subGenus="Raphirus">
<emphasis id="3395B1665D63FFA2FF36FB76FEC2A2AD" bold="true" box="[178,265,1160,1182]" italics="true" pageId="8" pageNumber="833">Quedius</emphasis>
(
<emphasis id="3395B1665D63FFA2FE9CFB76FEB2A2AD" bold="true" box="[280,377,1160,1182]" italics="true" pageId="8" pageNumber="833">Raphirus</emphasis>
)
<emphasis id="3395B1665D63FFA2FE0CFB76FE07A2AD" bold="true" box="[392,460,1160,1182]" italics="true" pageId="8" pageNumber="833">aedilis</emphasis>
</taxonomicName>
SMETANA
<taxonomicNameLabel id="28A60C1D5D63FFA2FDBBFB77FD59A2AD" box="[575,658,1161,1182]" pageId="8" pageNumber="833" rank="species">nov. sp.</taxonomicNameLabel>
(
<figureCitation id="99DA71F15D63FFA2FD1AFB76FCC2A2AD" box="[670,777,1160,1182]" captionStart="Figs 12-21" captionStartId="4.[178,218,1414,1436]" captionTargetBox="[188,1067,221,1405]" captionTargetId="figure@4.[178,1085,212,1405]" captionTargetPageId="4" captionText="Figs 12-21: 12-16. Quedius ryvkini nov.sp. (12) apical portion of male sternite 8; (13) tergite 10 of male genital segment; (14) sternite 9 of male genital segment; (15) aedoeagus, ventral view; (16) apical portion of underside of paramere with sensory peg setae. 17-21. Quedius aedilis nov.sp. (17) apical portion of male sternite 8; (18) tergite 10 of male genital segment; (19) sternite 9 of male genital segment; (20) aedoeagus, ventral view; (21) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004393" httpUri="https://zenodo.org/record/4004393/files/figure.png" pageId="8" pageNumber="833">Figs 18-21</figureCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="015E6D745D63FFA2FF36FB51FE1FA2EF" blockId="8.[178,1086,1198,1372]" pageId="8" pageNumber="833">
T y p e l o c a l i t y. ROSSIA: Primorskyi krai, Sikhote-Alinskyi zapovednik, Kabanyi kliuch. 45˚ 07’N 135˚ 52’E,
<specimenCount id="17E7A6FD5D63FFA2FE12FB38FE1FA2EF" box="[406,468,1222,1244]" pageId="8" pageNumber="833" type="generic">
<quantity id="C619C0915D63FFA2FE12FB38FE1BA2EF" box="[406,464,1222,1244]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="833" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="49FB3EFF5D63FFA3FF36FB19FEF2A51A" lastPageId="9" lastPageNumber="834" pageId="8" pageNumber="833" type="description">
<paragraph id="015E6D745D63FFA2FF36FB19FDF9A36F" blockId="8.[178,1086,1198,1372]" pageId="8" pageNumber="833">
<materialsCitation id="B18967295D63FFA2FF36FB19FDF9A36F" ID-GBIF-Occurrence="2852847307" collectingDate="2015-04-27" collectorName="D. E. Shcherbakov &amp; D. E. Shcherbakov." determinerName="A. B. Ryvkin" elevation="500" location="Sikhote-Alinskiy Nature Reserve" pageId="8" pageNumber="833" specimenCount="501" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
T y p e m a t e r i a l.
<typeStatus id="DE5AD3D65D63FFA2FE28FB18FE36A2CF" box="[428,509,1254,1276]" pageId="8" pageNumber="833" type="holotype">Holotype</typeStatus>
(
<specimenCount id="17E7A6FD5D63FFA2FD8AFB19FDEBA2CE" box="[526,544,1255,1277]" pageId="8" pageNumber="833" type="male">♂</specimenCount>
), ROSSIA: &quot;Primorskyi krai, Sikhote-Alinskyi zapovednik, Kabaniy kliuch. 45˚ 07’N 135˚ 52’E,
<specimenCount id="17E7A6FD5D63FFA2FDA3FB00FDAEA327" box="[551,613,1278,1300]" pageId="8" pageNumber="833" type="generic">
<quantity id="C619C0915D63FFA2FDA3FB00FDA9A327" box="[551,610,1278,1300]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="833" unit="m" value="500.0">
<elevation id="8ACC8A475D63FFA2FDA3FB00FDA9A327" box="[551,610,1278,1300]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="833" unit="m" value="500.0">500 m</elevation>
</quantity>
.
</specimenCount>
zheltye tarepky. 24-
<date id="755F4BB45D63FFA2FC93FB00FCB7A327" box="[791,892,1278,1300]" pageId="8" pageNumber="833" value="2015-04-27">
<collectingDate id="651BB25C5D63FFA2FC93FB00FCB7A327" box="[791,892,1278,1300]" pageId="8" pageNumber="833" value="2015-04-27">27.04.2015</collectingDate>
</date>
.
<collectorName id="AC1408A25D63FFA2FC00FB01FBE0A327" box="[900,1067,1278,1300]" pageId="8" pageNumber="833">D. E. Shcherbakov</collectorName>
&quot; / &quot;
<taxonomicName id="C6E116F75D63FFA2FF6CFAE8FE5CA31F" authority="" box="[232,407,1302,1324]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="833" phylum="Arthropoda" rank="subGenus" subGenus="Raphirus">
<emphasis id="3395B1665D63FFA2FF6CFAE8FEFAA318" box="[232,305,1302,1323]" italics="true" pageId="8" pageNumber="833">Quedius</emphasis>
(
<emphasis id="3395B1665D63FFA2FEC4FAE8FE5BA318" box="[320,400,1302,1323]" italics="true" pageId="8" pageNumber="833">Raphirus</emphasis>
)
</taxonomicName>
sp. nov. 2
<determinerName id="F48281B05D63FFA2FD85FAE9FDB0A31F" box="[513,635,1302,1324]" pageId="8" pageNumber="833">A. B. Ryvkin</determinerName>
det. 2015&quot;. [
<collectingRegion id="C325A3965D63FFA2FD76FAE8FC56A31F" box="[754,925,1302,1324]" country="Russia" name="Primorskiy" pageId="8" pageNumber="833">Maritime Province</collectingRegion>
,
<location id="043E3BAF5D63FFA2FC2CFAE8FEA3A377" LSID="urn:lsid:plazi:treatment:8948DC625D63FFA3FF36FB76FEF2A51A:043E3BAF5D63FFA2FC2CFAE8FEA3A377" name="Sikhote-Alinskiy Nature Reserve" pageId="8" pageNumber="833" stateProvince="Primorskiy">Sikhote-Alinskiy Nature Reserve</location>
, Kabaniy source, 45˚ 07’N 135˚ 52’E,
<quantity id="C619C0915D63FFA2FD3BFAD0FD33A377" box="[703,760,1326,1348]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="833" unit="m" value="500.0">
<elevation id="8ACC8A475D63FFA2FD3BFAD0FD33A377" box="[703,760,1326,1348]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="833" unit="m" value="500.0">500 m</elevation>
</quantity>
,
<collectingMethod id="D8A015635D63FFA2FC85FAD0FC5FA377" box="[769,916,1326,1348]" pageId="8" pageNumber="833">yellow pan traps</collectingMethod>
, 24-
<date id="755F4BB45D63FFA2FC3DFAD0FBD4A377" box="[953,1055,1326,1348]" pageId="8" pageNumber="833" value="2015-04-27">
<collectingDate id="651BB25C5D63FFA2FC3DFAD0FBD4A377" box="[953,1055,1326,1348]" pageId="8" pageNumber="833" value="2015-04-27">27.04.2015</collectingDate>
</date>
.
<collectorName id="AC1408A25D63FFA2FBA3FAD1FEB9A36F" pageId="8" pageNumber="833">D. E. Shcherbakov.</collectorName>
] Deposited in NSMT.
</materialsCitation>
</paragraph>
<paragraph id="015E6D745D63FFA2FF36FA95FCF5A3AD" blockId="8.[178,1085,1387,1438]" pageId="8" pageNumber="833">
D i a g n o s i s. A species in all characers similar to
<taxonomicName id="C6E116F75D63FFA2FD59FA95FC89A3B3" authority="Smetana &amp; Shavrin, 2018" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[733,834,1387,1408]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="833" phylum="Arthropoda" rank="species" species="ryvkini" status="sp. nov.">
<emphasis id="3395B1665D63FFA2FD59FA95FC89A3B3" box="[733,834,1387,1408]" italics="true" pageId="8" pageNumber="833">Q. ryvkini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="28A60C1D5D63FFA2FCCDFA95FC5EA3B3" box="[841,917,1387,1408]" pageId="8" pageNumber="833" rank="species">nov.sp.</taxonomicNameLabel>
, but different by darker body coloration and by the differently shaped aedoeagus.
</paragraph>
<paragraph id="015E6D745D63FFA2FF36FA50FD23A028" blockId="8.[178,1086,1453,1563]" pageId="8" pageNumber="833">D e s c r i p t i o n. Head black, pronotum uniformly piceous, elytra piceous, abdomen piceous-black to black with apical margins of tergites more or less narrowly paler. Mandibles rufotestaceous, palpi, antennae and legs uniformly pale testaceous but first three antennal segments vaguely, indefinitely darkened.</paragraph>
<paragraph id="015E6D745D63FFA3FF36F9D5FE31A7E8" blockId="8.[178,1085,1579,1601]" lastBlockId="9.[178,1085,219,475]" lastPageId="9" lastPageNumber="834" pageId="8" pageNumber="833">
Male. First four segments of front tarsus similar to those of
<taxonomicName id="C6E116F75D63FFA2FCA1F9D5FC47A073" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[805,908,1579,1600]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="833" phylum="Arthropoda" rank="species" species="ryvkini">
<emphasis id="3395B1665D63FFA2FCA1F9D5FCFCA073" box="[805,823,1579,1600]" italics="true" pageId="8" pageNumber="833">Q</emphasis>
.
<emphasis id="3395B1665D63FFA2FCC2F9D5FC47A073" box="[838,908,1579,1600]" italics="true" pageId="8" pageNumber="833">ryvkini</emphasis>
</taxonomicName>
, but less dilated, but segment two as wide as apex of tibia, as in
<taxonomicName id="C6E116F75D62FFA3FD2AFF25FCDDA6C3" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[686,790,219,240]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="834" phylum="Arthropoda" rank="species" species="ryvkini">
<emphasis id="3395B1665D62FFA3FD2AFF25FD0BA6C3" box="[686,704,219,240]" italics="true" pageId="9" pageNumber="834">Q</emphasis>
.
<emphasis id="3395B1665D62FFA3FD54FF25FCDDA6C3" box="[720,790,219,240]" italics="true" pageId="9" pageNumber="834">ryvkini</emphasis>
</taxonomicName>
. Sternite 8 with four strong setae on right side and three on left side; with medioapical emargination similar, but more acute and less deep (
<figureCitation id="99DA71F15D62FFA3FE47FEEBFDDBA71F" box="[451,528,277,300]" captionStart="Figs 12-21" captionStartId="4.[178,218,1414,1436]" captionTargetBox="[188,1067,221,1405]" captionTargetId="figure@4.[178,1085,212,1405]" captionTargetPageId="4" captionText="Figs 12-21: 12-16. Quedius ryvkini nov.sp. (12) apical portion of male sternite 8; (13) tergite 10 of male genital segment; (14) sternite 9 of male genital segment; (15) aedoeagus, ventral view; (16) apical portion of underside of paramere with sensory peg setae. 17-21. Quedius aedilis nov.sp. (17) apical portion of male sternite 8; (18) tergite 10 of male genital segment; (19) sternite 9 of male genital segment; (20) aedoeagus, ventral view; (21) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004393" httpUri="https://zenodo.org/record/4004393/files/figure.png" pageId="9" pageNumber="834">Fig. 17</figureCitation>
). Both tergite 10 and sternite 9 of genital segment not appreciably different from those of
<taxonomicName id="C6E116F75D62FFA3FD98FECDFD4AA77B" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[540,641,307,328]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="834" phylum="Arthropoda" rank="species" species="ryvkini">
<emphasis id="3395B1665D62FFA3FD98FECDFDE5A77B" box="[540,558,307,328]" italics="true" pageId="9" pageNumber="834">Q</emphasis>
.
<emphasis id="3395B1665D62FFA3FDBFFECDFD4AA77B" box="[571,641,307,328]" italics="true" pageId="9" pageNumber="834">ryvkini</emphasis>
</taxonomicName>
(
<figureCitation id="99DA71F15D62FFA3FD0BFECDFCCEA77A" box="[655,773,307,329]" captionStart="Figs 12-21" captionStartId="4.[178,218,1414,1436]" captionTargetBox="[188,1067,221,1405]" captionTargetId="figure@4.[178,1085,212,1405]" captionTargetPageId="4" captionText="Figs 12-21: 12-16. Quedius ryvkini nov.sp. (12) apical portion of male sternite 8; (13) tergite 10 of male genital segment; (14) sternite 9 of male genital segment; (15) aedoeagus, ventral view; (16) apical portion of underside of paramere with sensory peg setae. 17-21. Quedius aedilis nov.sp. (17) apical portion of male sternite 8; (18) tergite 10 of male genital segment; (19) sternite 9 of male genital segment; (20) aedoeagus, ventral view; (21) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004393" httpUri="https://zenodo.org/record/4004393/files/figure.png" pageId="9" pageNumber="834">Figs 18, 19</figureCitation>
). Aedoeagus (
<figureCitation id="99DA71F15D62FFA3FC12FECDFBC2A77B" box="[918,1033,307,329]" captionStart="Figs 12-21" captionStartId="4.[178,218,1414,1436]" captionTargetBox="[188,1067,221,1405]" captionTargetId="figure@4.[178,1085,212,1405]" captionTargetPageId="4" captionText="Figs 12-21: 12-16. Quedius ryvkini nov.sp. (12) apical portion of male sternite 8; (13) tergite 10 of male genital segment; (14) sternite 9 of male genital segment; (15) aedoeagus, ventral view; (16) apical portion of underside of paramere with sensory peg setae. 17-21. Quedius aedilis nov.sp. (17) apical portion of male sternite 8; (18) tergite 10 of male genital segment; (19) sternite 9 of male genital segment; (20) aedoeagus, ventral view; (21) apical portion of underside of paramere with sensory peg setae." figureDoi="http://doi.org/10.5281/zenodo.4004393" httpUri="https://zenodo.org/record/4004393/files/figure.png" pageId="9" pageNumber="834">Figs 20, 21</figureCitation>
with median lobe similar in shape to that of
<taxonomicName id="C6E116F75D62FFA3FDE6FEAEFD00A756" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[610,715,336,357]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="834" phylum="Arthropoda" rank="species" species="ryvkini">
<emphasis id="3395B1665D62FFA3FDE6FEAEFDBFA756" box="[610,628,336,357]" italics="true" pageId="9" pageNumber="834">Q</emphasis>
.
<emphasis id="3395B1665D62FFA3FD01FEAEFD00A756" box="[645,715,336,357]" italics="true" pageId="9" pageNumber="834">ryvkini</emphasis>
</taxonomicName>
but narrower; paramere similar in shape, but markedly narrower, with subacute apex not reaching apex of median lobe; four apical setae, median pair markedly longer than lateral pair and two long setae at each lateral margin close to apex; sensory peg setae on underside less numerous, forming a narrow field on apical portion.
</paragraph>
<paragraph id="015E6D745D62FFA3FF36FE15FEA2A432" blockId="9.[178,361,491,513]" box="[178,361,491,513]" pageId="9" pageNumber="834">Female unknown.</paragraph>
<paragraph id="015E6D745D62FFA3FF36FDEEFE9EA416" blockId="9.[178,341,528,550]" box="[178,341,528,550]" pageId="9" pageNumber="834">
Length
<quantity id="C619C0915D62FFA3FE84FDEEFE9BA416" box="[256,336,528,550]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="9" pageNumber="834" unit="mm" value="7.5">7.5 mm</quantity>
.
</paragraph>
<paragraph id="015E6D745D62FFA3FF36FDC8FDCBA45B" blockId="9.[178,1085,565,617]" pageId="9" pageNumber="834">
E t y m o l o g y. The specific epithet is the Latin masculine noun
<emphasis id="3395B1665D62FFA3FC06FDCBFC0DA479" box="[898,966,565,586]" italics="true" pageId="9" pageNumber="834">aedilis</emphasis>
, -
<emphasis id="3395B1665D62FFA3FC5AFDC8FC24A478" box="[990,1007,566,587]" italics="true" pageId="9" pageNumber="834">is</emphasis>
, m (an aedile, official in ancient Rome).
</paragraph>
<paragraph id="015E6D745D62FFA3FF36FD86FF2EA4FB" blockId="9.[178,1085,632,712]" pageId="9" pageNumber="834">
G e o g r a p h i c a l d i s t r i b u t i o n.
<taxonomicName id="C6E116F75D62FFA3FD14FD86FCFAA4BE" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[656,817,632,653]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="834" phylum="Arthropoda" rank="species" species="aedilis">
<emphasis id="3395B1665D62FFA3FD14FD86FCFAA4BE" box="[656,817,632,653]" italics="true" pageId="9" pageNumber="834">Quedius aedilis</emphasis>
</taxonomicName>
is at present known only from the
<typeStatus id="DE5AD3D65D62FFA3FE94FD68FEF7A498" box="[272,316,662,683]" pageId="9" pageNumber="834">type</typeStatus>
locality in Sikhote-Alin range in the
<collectingRegion id="C325A3965D62FFA3FD3FFD6BFCB1A498" box="[699,890,661,683]" country="Russia" name="Primorskiy" pageId="9" pageNumber="834">Maritime Province</collectingRegion>
of the Russian Far East.
</paragraph>
<paragraph id="015E6D745D62FFA3FF36FD27FEF2A51A" blockId="9.[178,1085,728,809]" pageId="9" pageNumber="834">
B i o n o m i c s.The
<typeStatus id="DE5AD3D65D62FFA3FE18FD26FE3EA4DD" box="[412,501,728,750]" pageId="9" pageNumber="834" type="holotype">holotype</typeStatus>
was taken from a yellow pan trap, but it is not known in what habitat the traps were set. It is apparent that
<taxonomicName id="C6E116F75D62FFA3FD49FD08FCFEA539" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[717,821,757,779]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="834" phylum="Arthropoda" rank="species" species="ryvkini">
<emphasis id="3395B1665D62FFA3FD49FD08FD14A538" box="[717,735,758,779]" italics="true" pageId="9" pageNumber="834">Q</emphasis>
.
<emphasis id="3395B1665D62FFA3FD6BFD0BFCFEA539" box="[751,821,757,778]" italics="true" pageId="9" pageNumber="834">ryvkini</emphasis>
</taxonomicName>
and
<taxonomicName id="C6E116F75D62FFA3FCEAFD08FC1EA539" authorityName="Smetana &amp; Shavrin" authorityYear="2018" box="[878,981,757,779]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="834" phylum="Arthropoda" rank="species" species="aedilis">
<emphasis id="3395B1665D62FFA3FCEAFD08FC1EA539" box="[878,981,757,779]" italics="true" pageId="9" pageNumber="834">Q. aedilis</emphasis>
</taxonomicName>
share the same habitat.
</paragraph>
</subSubSection>
</treatment>
</document>
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