treatments-xml/data/80/77/87/807787F6B82DFFCE49A1D755FCEE3BED.xml

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<mods:title id="0DE64E932AA48341DA5A270EF9314113">Revision of the Oriental tiger beetle genus Heptodonta Hope, 1838 (Coleoptera: Cicindelidae)</mods:title>
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<mods:namePart id="0E5E62C866885CEE2F5447290512F88A">Görn, Sebastian</mods:namePart>
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<paragraph id="086136E0B82DFFCB49A1D755FD203E64" blockId="19.[151,648,1086,1149]" box="[151,648,1086,1113]" pageId="19" pageNumber="20">
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<taxonomicName id="CFDE4D63B82DFFCB49A1D755FD203E64" ID-CoL="3KX7X" authority="(Schaum, 1862)" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[151,648,1086,1113]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B82DFFCB49A1D755FD203E64" bold="true" box="[151,648,1086,1113]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49A1D755FE173E65" bold="true" box="[151,447,1086,1113]" italics="true" pageId="19" pageNumber="20">Heptodonta melanopyga</emphasis>
(
<bibRefCitation id="6C4F4B11B82DFFCB48E6D755FD283E64" author="Schaum, H." box="[464,640,1086,1112]" pageId="19" pageNumber="20" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum, 1862</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="40C4656BB82DFFCB49A1D708FECA3E41" box="[151,354,1122,1149]" pageId="19" pageNumber="20" type="description">
<paragraph id="086136E0B82DFFCB49A1D708FECA3E41" blockId="19.[151,648,1086,1149]" box="[151,354,1122,1149]" pageId="19" pageNumber="20">
<figureCitation id="90E52A65B82DFFCB49A1D708FF4B3E41" box="[151,227,1123,1149]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1827,1851]" captionTargetBox="[177,1392,191,1791]" captionTargetId="figure-15@5.[177,1410,181,1803]" captionTargetPageId="5" captionText="FIGURES 1–7. 1–2—anterior elytra: 1—Heptodonta melanopyga (Schaum, 1862), ♀, Philippines, Luzon, Banaue (PSCH); 2—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). 3–4—pronotum: 3—H. arrowi W. Horn, 1900, ♂, Myanmar, Magway Region, Pyon Village (SGCH); 4—Heptodonta horii sp. nov., ♂, holotype, Myanmar, Bago Re- gion, Chanthakwin (JWCW); 5–7—head: 5—Heptodonta tempesta sp. nov., ♂, holotype, India, Meghalaya, Balpakram NP (JWCW); 6—Heptodonta pulchella (Hope, 1831), ♂, Nepal, Bagmati, Sollenta-Dhading (JWCW); 7—Heptodonta schuelei sp. nov., ♂, holotype, India, Meghalaya, Cherrapunjee (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578783" httpUri="https://zenodo.org/record/4578783/files/figure.png" pageId="19" pageNumber="20">Figs. 1</figureCitation>
, 
<figureCitation id="90E52A65B82DFFCB49C7D708FEB83E41" box="[241,272,1123,1149]" captionStart="FIGURES 44–47" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[205,1333,189,1867]" captionTargetId="figure-15@17.[197,1389,181,1887]" captionTargetPageId="17" captionText="FIGURES 44–47. 44–47—habitus: 44–45—Heptodonta nigrosericea (W. Horn, 1930): 44—♀, lectotype, Philippines, [Min- danao], Columbogan (SDEI); 45—♂, holotype of syn. H. lumawigi Wiesner, 1980, Philippines, Canuawua (JWCW). 46—H. melanopyga (Schaum, 1862), ♂, lectotype, Philippines, Luzon (MNB). 47—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578799" httpUri="https://zenodo.org/record/4578799/files/figure.png" pageId="19" pageNumber="20">46</figureCitation>
, 
<figureCitation id="90E52A65B82DFFCB482DD709FECA3E41" box="[283,354,1122,1149]" captionStart="FIGURES 48–54" captionStartId="20.[151,264,1924,1948]" captionTargetBox="[152,1435,184,1871]" captionTargetId="figure-20@20.[151,1435,181,1900]" captionTargetPageId="20" captionText="FIGURES 48–54. Heptodonta melanopyga (Schaum, 1862), Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="19" pageNumber="20">48–54</figureCitation>
</paragraph>
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<subSubSection id="40C4656BB82DFFCB49A1D7CCFCAB3F06" pageId="19" pageNumber="20" type="reference_group">
<paragraph id="086136E0B82DFFCB49A1D7CCFD743E83" blockId="19.[151,859,1191,1338]" box="[151,732,1191,1215]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="28CE11CEB82DFFCB49A1D7CCFD743E83" box="[151,732,1191,1215]" pageId="19" pageNumber="20">
<treatmentCitation id="897F10F1B82DFFCB49A1D7CCFD703E83" author="Schaum, H." box="[151,728,1191,1215]" page="173" pageId="19" pageNumber="20" year="1862">
<taxonomicName id="CFDE4D63B82DFFCB49A1D7CCFD703E83" authority="Schaum, 1862: 173" authorityName="Schaum" authorityPageNumber="173" authorityYear="1862" box="[151,728,1191,1215]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga" subGenus="Heptadonta">
<emphasis id="3AAAEAF2B82DFFCB49A1D7CCFF523E83" box="[151,250,1191,1215]" italics="true" pageId="19" pageNumber="20">Cicindela</emphasis>
(
<emphasis id="3AAAEAF2B82DFFCB483ED7CCFE283E83" box="[264,384,1191,1215]" italics="true" pageId="19" pageNumber="20">Heptadonta</emphasis>
) 
<emphasis id="3AAAEAF2B82DFFCB48B9D7CCFDA33E83" box="[399,523,1191,1215]" italics="true" pageId="19" pageNumber="20">melanopyga</emphasis>
<bibRefCitation id="6C4F4B11B82DFFCB4B27D7CCFD703E83" author="Schaum, H." box="[529,728,1191,1215]" pageId="19" pageNumber="20" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum, 1862: 173</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="086136E0B82DFFCB49A1D7ADFDC03EE2" blockId="19.[151,859,1191,1338]" box="[151,616,1222,1246]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="28CE11CEB82DFFCB49A1D7ADFDC03EE2" box="[151,616,1222,1246]" pageId="19" pageNumber="20">
<taxonomicName id="CFDE4D63B82DFFCB49A1D7ADFE393EE2" ID-CoL="3KX7X" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[151,401,1222,1246]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B82DFFCB49A1D7ADFE393EE2" box="[151,401,1222,1246]" italics="true" pageId="19" pageNumber="20">Heptodonta melanopyga</emphasis>
</taxonomicName>
. 
<treatmentCitation id="897F10F1B82DFFCB48A8D7ADFDCC3EE2" author="Chaudoir, M." box="[414,612,1222,1246]" page="20" pageId="19" pageNumber="20" year="1865">
<bibRefCitation id="6C4F4B11B82DFFCB48A8D7ADFDCC3EE2" author="Chaudoir, M." box="[414,612,1222,1246]" pageId="19" pageNumber="20" refId="ref38256" refString="Chaudoir, M. (1865) Catalogue de la collection de Cicindeletes de M. le Baron Chaudoir. Nys, Bruxelles, 64 pp. https: // doi. org / 10.5962 / bhl. title. 134840" type="book" year="1865">Chaudoir, 1865: 20</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="086136E0B82DFFCB49A1D78FFDDF3EC1" blockId="19.[151,859,1191,1338]" box="[151,631,1252,1277]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="28CE11CEB82DFFCB49A1D78FFDDF3EC1" box="[151,631,1252,1277]" pageId="19" pageNumber="20">
<taxonomicName id="CFDE4D63B82DFFCB49A1D78FFE393EC0" authority="Fleutiaux, 1892" box="[151,401,1252,1276]" class="Insecta" family="Carabidae" genus="Heptadonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B82DFFCB49A1D78FFE393EC0" box="[151,401,1252,1276]" italics="true" pageId="19" pageNumber="20">Heptadonta melanopyga</emphasis>
</taxonomicName>
. 
<treatmentCitation id="897F10F1B82DFFCB48A8D78FFDDB3EC1" author="Fleutiaux, E." box="[414,627,1252,1277]" page="128" pageId="19" pageNumber="20" year="1892">
<bibRefCitation id="6C4F4B11B82DFFCB48A8D78FFDDB3EC1" author="Fleutiaux, E." box="[414,627,1252,1277]" pageId="19" pageNumber="20" refId="ref38476" refString="Fleutiaux, E. (1892) Catalogue systematique des Cicindelidae. Imprimerie H. Vaillant-Carmanne, Liege, 186 pp. https: // doi. org / 10.5962 / bhl. title. 48634" type="book" year="1892">Fleutiaux, 1892: 128</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="086136E0B82DFFCB49A1D668FCF33F27" blockId="19.[151,859,1191,1338]" box="[151,859,1283,1307]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="28CE11CEB82DFFCB49A1D668FCF33F27" box="[151,859,1283,1307]" pageId="19" pageNumber="20">
<taxonomicName id="CFDE4D63B82DFFCB49A1D668FDD73F27" ID-CoL="8LSDX" authorityName="Agassiz" authorityYear="1846" box="[151,639,1283,1307]" class="Insecta" family="Carabidae" genus="Odontochila" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="3AAAEAF2B82DFFCB49A1D668FEBE3F27" box="[151,278,1283,1307]" italics="true" pageId="19" pageNumber="20">Odontochila</emphasis>
(“group” 
<emphasis id="3AAAEAF2B82DFFCB484BD668FE5D3F27" box="[381,501,1283,1307]" italics="true" pageId="19" pageNumber="20">Heptodonta</emphasis>
) 
<emphasis id="3AAAEAF2B82DFFCB4B35D668FDD73F27" box="[515,639,1283,1307]" italics="true" pageId="19" pageNumber="20">melanopyga</emphasis>
</taxonomicName>
. W. 
<treatmentCitation id="897F10F1B82DFFCB4B9BD66FFCFF3F27" author="Horn, W." box="[685,855,1283,1307]" page="203" pageId="19" pageNumber="20" year="1910">
<bibRefCitation id="6C4F4B11B82DFFCB4B9BD66FFCFF3F27" author="Horn, W." box="[685,855,1283,1307]" pageId="19" pageNumber="20" pagination="105 - 208" refId="ref39211" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, Fascicule 82 B, pp. 105 - 208, pls. 6 - 15." type="journal article" year="1910">Horn, 1910: 203</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="086136E0B82DFFCB49A1D649FCAB3F06" blockId="19.[151,859,1191,1338]" box="[151,771,1314,1338]" pageId="19" pageNumber="20">
<treatmentCitationGroup id="28CE11CEB82DFFCB49A1D649FCAB3F06" box="[151,771,1314,1338]" pageId="19" pageNumber="20">
<taxonomicName id="CFDE4D63B82DFFCB49A1D649FD8F3F06" ID-CoL="8LSDX" authority="W. Horn, 1926" box="[151,551,1314,1338]" class="Insecta" family="Carabidae" genus="Odontochila" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga" subGenus="Heptodonta">
<emphasis id="3AAAEAF2B82DFFCB49A1D649FEBE3F06" box="[151,278,1314,1338]" italics="true" pageId="19" pageNumber="20">Odontochila</emphasis>
(
<emphasis id="3AAAEAF2B82DFFCB4813D649FE353F06" box="[293,413,1314,1338]" italics="true" pageId="19" pageNumber="20">Heptodonta</emphasis>
) 
<emphasis id="3AAAEAF2B82DFFCB489DD649FD8F3F06" box="[427,551,1314,1338]" italics="true" pageId="19" pageNumber="20">melanopyga</emphasis>
</taxonomicName>
. W. 
<treatmentCitation id="897F10F1B82DFFCB4B63D649FD573F06" author="Horn, W." box="[597,767,1314,1338]" page="123" pageId="19" pageNumber="20" year="1926">
<bibRefCitation id="6C4F4B11B82DFFCB4B63D649FD573F06" author="Horn, W." box="[597,767,1314,1338]" pageId="19" pageNumber="20" pagination="1 - 345" refId="ref39316" refString="Horn, W. (1926) Carabidae, Cicindelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, 86, pp. 1 - 345." type="journal article" year="1926">Horn, 1926: 123</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82DFFCB49A1D608FC0D3CA1" pageId="19" pageNumber="20" type="materials_examined">
<paragraph id="086136E0B82DFFCB49A1D608FDB03F41" blockId="19.[151,1437,1378,1982]" box="[151,536,1378,1405]" pageId="19" pageNumber="20">
<materialsCitation id="B8B63CBDB82DFFCB49A1D608FDBC3F41" ID-GBIF-Occurrence="3045076617" box="[151,532,1378,1405]" country="Philippines" location="Luzon" pageId="19" pageNumber="20" specimenCount="1" typeStatus="holotype">
<emphasis id="3AAAEAF2B82DFFCB49A1D608FE9E3F40" bold="true" box="[151,310,1378,1405]" pageId="19" pageNumber="20">
<typeStatus id="D7658842B82DFFCB49A1D608FF7A3F41" box="[151,210,1379,1405]" pageId="19" pageNumber="20">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="70C97670B82DFFCB480AD608FE173F41" box="[316,447,1379,1405]" name="Philippines" pageId="19" pageNumber="20">Philippines</collectingCountry>
: 
<location id="0D01603BB82DFFCB48FFD608FDBC3F41" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82DFFCB48FFD608FDBC3F41" box="[457,532,1379,1405]" country="Philippines" name="Luzon" pageId="19" pageNumber="20">Luzon</location>
</materialsCitation>
.
</paragraph>
<paragraph id="086136E0B82DFFCB49F1D6ECFC0D3CA1" blockId="19.[151,1437,1378,1982]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D6ECFE5D3F9D" bold="true" box="[199,501,1414,1441]" pageId="19" pageNumber="20">
Type material. 
<typeStatus id="D7658842B82DFFCB4849D6ECFE5D3F9D" box="[383,501,1415,1441]" pageId="19" pageNumber="20" type="lectotype">Lectotype</typeStatus>
</emphasis>
(designated here): 
<specimenCount id="1ED8FD69B82DFFCB4BE6D6ECFD4D3F9D" box="[720,741,1415,1441]" pageId="19" pageNumber="20" type="male">♂</specimenCount>
in MNB, labelled: “42450” [printed] // “Hist.-Coll. (Coleop- tera) / Nr. 42450 / 
<taxonomicName id="CFDE4D63B82DFFCB4855D6C0FDF63FF9" authorityName="Schaum" authorityYear="1862" box="[355,606,1451,1477]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga">Cicindela melanopyga</taxonomicName>
/ Schaum* / Luzon, Coll. Schaum / Zool. Mus. 
<collectingRegion id="CA1AF802B82DFFCB4D44D6C0FB133FF9" box="[1138,1211,1451,1477]" country="Germany" name="Berlin" pageId="19" pageNumber="20">Berlin</collectingRegion>
” [yellow with black frame, printed] // “melanopyga / Schaum.* / Luzon. (Semper)” [yellow with black frame, handwritten]. 
<emphasis id="3AAAEAF2B82DFFCB4C17D6A4FF623C31" bold="true" pageId="19" pageNumber="20">
<typeStatus id="D7658842B82DFFCB4C17D6A4FF623C31" pageId="19" pageNumber="20" type="paralectotype">Paralectotype</typeStatus>
</emphasis>
: 
<specimenCount id="1ED8FD69B82DFFCB49EFD698FEAA3C31" box="[217,258,1523,1549]" pageId="19" pageNumber="20" type="female">1 ♀</specimenCount>
in MNB: “Hist.-Coll. (
<taxonomicName id="CFDE4D63B82DFFCB4B32D698FD2B3C32" authorityName="Linnaeus" authorityYear="1758" box="[516,643,1523,1550]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
) / Nr. 42450 / 
<taxonomicName id="CFDE4D63B82DFFCB4A15D698FBB63C31" authorityName="Schaum" authorityYear="1862" box="[803,1054,1523,1549]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="melanopyga">Cicindela melanopyga</taxonomicName>
/ Schaum* / Luzon, Coll. Schaum / Zool. Mus. 
<collectingRegion id="CA1AF802B82DFFCB4807D57CFED23C0D" box="[305,378,1559,1585]" country="Germany" name="Berlin" pageId="19" pageNumber="20">Berlin</collectingRegion>
” [yellow with black frame, printed]. All type specimens labelled: “ 
<typeStatus id="D7658842B82DFFCB4DB5D57CFAAC3C0D" box="[1155,1284,1559,1585]" pageId="19" pageNumber="20" type="syntype">SYNTYPE</typeStatus>
/ 
<taxonomicName id="CFDE4D63B82DFFCB4C29D57CFA223C0D" authorityName="Linne" authorityYear="1758" box="[1311,1418,1559,1585]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Cicindela</taxonomicName>
/ melanopyga / 
<bibRefCitation id="6C4F4B11B82DFFCB480FD550FE733C69" author="Schaum, H." box="[313,475,1595,1621]" pageId="19" pageNumber="20" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum, 1862</bibRefCitation>
/ labelled by MFNB 2018” [red, printed] // “ 
<typeStatus id="D7658842B82DFFCB4AE9D550FB2A3C69" box="[991,1154,1595,1621]" pageId="19" pageNumber="20" type="lectotype">LECTOTYPE</typeStatus>
(or 
<typeStatus id="D7658842B82DFFCB4D82D550FA343C69" box="[1204,1436,1595,1621]" pageId="19" pageNumber="20" type="paralectotype">PARALECTOTYPE</typeStatus>
respectively) / 
<taxonomicName id="CFDE4D63B82DFFCB480AD534FE0F3C45" authorityName="Linne" authorityYear="1758" box="[316,423,1631,1657]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Cicindela</taxonomicName>
/ melanopyga / 
<bibRefCitation id="6C4F4B11B82DFFCB4B6ED534FD503C45" author="Schaum, H." box="[600,760,1631,1657]" pageId="19" pageNumber="20" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum, 1862</bibRefCitation>
/ design. Sebastian Görn 2020” [red, printed] // “ 
<taxonomicName id="CFDE4D63B82DFFCB4C20D534FA343C45" authorityName="Analis. Hope" authorityYear="1838" box="[1302,1436,1631,1657]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Heptodonta</taxonomicName>
/ melanopyga / (
<bibRefCitation id="6C4F4B11B82DFFCB487AD5E8FE473CA2" author="Schaum, H." box="[332,495,1667,1694]" pageId="19" pageNumber="20" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum, 1862</bibRefCitation>
) / det. Sebastian Görn 2020” [printed].
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82DFFCB49F1D5CDFE5B3DA5" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="086136E0B82DFFCB49F1D5CDFE5B3DA5" blockId="19.[151,1437,1378,1982]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D5CDFEEA3CFC" bold="true" box="[199,322,1702,1728]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
Distinguishable from 
<taxonomicName id="CFDE4D63B82DFFCB4B08D5CCFD0C3CFD" baseAuthorityName="Fabricius" baseAuthorityYear="1801" box="[574,676,1703,1729]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="analis">
<emphasis id="3AAAEAF2B82DFFCB4B08D5CCFD0C3CFD" box="[574,676,1703,1729]" italics="true" pageId="19" pageNumber="20">H. analis</emphasis>
</taxonomicName>
, 
<taxonomicName id="CFDE4D63B82DFFCB4B85D5CCFC953CFD" box="[691,829,1703,1729]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="abasileia" status="sp. nov.">
<emphasis id="3AAAEAF2B82DFFCB4B85D5CCFC953CFD" box="[691,829,1703,1729]" italics="true" pageId="19" pageNumber="20">H. abasileia</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B82DFFCB4A72D5CCFC363CFD" bold="true" box="[836,926,1703,1729]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="21995789B82DFFCB4A72D5CCFC363CFD" box="[836,926,1703,1729]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="CFDE4D63B82DFFCB4A9AD5CCFB833CFD" box="[940,1067,1703,1729]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="halensis" status="sp. nov.">
<emphasis id="3AAAEAF2B82DFFCB4A9AD5CCFB833CFD" box="[940,1067,1703,1729]" italics="true" pageId="19" pageNumber="20">H. halensis</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B82DFFCB4D04D5CCFB243CFD" bold="true" box="[1074,1164,1703,1729]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="21995789B82DFFCB4D04D5CCFB243CFD" box="[1074,1164,1703,1729]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and 
<taxonomicName id="CFDE4D63B82DFFCB4DFCD5CCFAD13CFC" baseAuthorityName="W. Horn" baseAuthorityYear="1930" box="[1226,1401,1703,1728]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrosericea">
<emphasis id="3AAAEAF2B82DFFCB4DFCD5CCFAD13CFC" box="[1226,1401,1703,1728]" italics="true" pageId="19" pageNumber="20">H. nigrosericea</emphasis>
</taxonomicName>
by lacking mediolateral and anteroapical elytral impressions. In contrast to continental species, with shiny lateroapical lustre, due to deeper apical impression, round aedeagal tip, short rather rectangular labrum in males, and triangular, usually less elongated labrum in females. Distinguishable from the similar 
<taxonomicName id="CFDE4D63B82DFFCB4ACCD478FB1D3D11" authorityName="Cassola" authorityYear="2000" box="[1018,1205,1811,1837]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mindoroensis">
<emphasis id="3AAAEAF2B82DFFCB4ACCD478FB1D3D11" box="[1018,1205,1811,1837]" italics="true" pageId="19" pageNumber="20">H. mindoroensis</emphasis>
</taxonomicName>
, by slightly dilated, more robust elytra, irregular, more fused elytral punctation (
<figureCitation id="90E52A65B82DFFCB4A0BD45CFCD73D6D" box="[829,895,1847,1873]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1827,1851]" captionTargetBox="[177,1392,191,1791]" captionTargetId="figure-15@5.[177,1410,181,1803]" captionTargetPageId="5" captionText="FIGURES 1–7. 1–2—anterior elytra: 1—Heptodonta melanopyga (Schaum, 1862), ♀, Philippines, Luzon, Banaue (PSCH); 2—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). 3–4—pronotum: 3—H. arrowi W. Horn, 1900, ♂, Myanmar, Magway Region, Pyon Village (SGCH); 4—Heptodonta horii sp. nov., ♂, holotype, Myanmar, Bago Re- gion, Chanthakwin (JWCW); 5–7—head: 5—Heptodonta tempesta sp. nov., ♂, holotype, India, Meghalaya, Balpakram NP (JWCW); 6—Heptodonta pulchella (Hope, 1831), ♂, Nepal, Bagmati, Sollenta-Dhading (JWCW); 7—Heptodonta schuelei sp. nov., ♂, holotype, India, Meghalaya, Cherrapunjee (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578783" httpUri="https://zenodo.org/record/4578783/files/figure.png" pageId="19" pageNumber="20">Fig. 1</figureCitation>
), and rather orthogonal aedeagus. In contrast to 
<taxonomicName id="CFDE4D63B82DFFCB49A1D430FEB13D48" box="[151,281,1883,1908]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="3AAAEAF2B82DFFCB49A1D430FEB13D48" box="[151,281,1883,1908]" italics="true" pageId="19" pageNumber="20">H. wiesneri</emphasis>
</taxonomicName>
<emphasis id="3AAAEAF2B82DFFCB4817D430FED33D49" bold="true" box="[289,379,1883,1909]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="21995789B82DFFCB4817D430FED33D49" box="[289,379,1883,1909]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, pronotum with continuously converging lateral margin and relatively coarse transverse striation, aedeagus more elongated.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82DFFCD49F1D4C9FC9E3C81" lastPageId="21" lastPageNumber="22" pageId="19" pageNumber="20" type="description">
<paragraph id="086136E0B82DFFCB49F1D4C9FACB3D81" blockId="19.[151,1437,1378,1982]" box="[199,1379,1954,1982]" pageId="19" pageNumber="20">
<emphasis id="3AAAEAF2B82DFFCB49F1D4C9FEDD3D80" bold="true" box="[199,373,1954,1980]" pageId="19" pageNumber="20">Redescription.</emphasis>
Body size: Length 
<quantity id="CF269B05B82DFFCB4B66D4C9FD5B3D81" box="[592,755,1954,1981]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" metricValueMax="1.35" metricValueMin="1.03" pageId="19" pageNumber="20" unit="mm" value="11.9" valueMax="13.5" valueMin="10.3">10.3–13.5 mm</quantity>
(without labrum), width 
<quantity id="CF269B05B82DFFCB4D3BD4C8FB3C3D81" box="[1037,1172,1955,1981]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="4.1" metricValueMin="3.2" pageId="19" pageNumber="20" unit="mm" value="3.65" valueMax="4.1" valueMin="3.2">3.2–4.1 mm</quantity>
(
<figureCitation id="90E52A65B82DFFCB4D95D4C8FAA93D81" box="[1187,1281,1955,1981]" captionStart="FIGURES 44–47" captionStartId="17.[151,264,1908,1932]" captionTargetBox="[205,1333,189,1867]" captionTargetId="figure-15@17.[197,1389,181,1887]" captionTargetPageId="17" captionText="FIGURES 44–47. 44–47—habitus: 44–45—Heptodonta nigrosericea (W. Horn, 1930): 44—♀, lectotype, Philippines, [Min- danao], Columbogan (SDEI); 45—♂, holotype of syn. H. lumawigi Wiesner, 1980, Philippines, Canuawua (JWCW). 46—H. melanopyga (Schaum, 1862), ♂, lectotype, Philippines, Luzon (MNB). 47—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578799" httpUri="https://zenodo.org/record/4578799/files/figure.png" pageId="19" pageNumber="20">Figs. 46</figureCitation>
, 
<figureCitation id="90E52A65B82DFFCB4C3AD4C8FAFF3D81" box="[1292,1367,1955,1981]" captionStart="FIGURES 48–54" captionStartId="20.[151,264,1924,1948]" captionTargetBox="[152,1435,184,1871]" captionTargetId="figure-20@20.[151,1435,181,1900]" captionTargetPageId="20" captionText="FIGURES 48–54. Heptodonta melanopyga (Schaum, 1862), Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="19" pageNumber="20">48–49</figureCitation>
).
</paragraph>
<caption id="5CA16668B82AFFCC49A1D4EFFBE33DD8" ID-DOI="http://doi.org/10.5281/zenodo.4578804" ID-Zenodo-Dep="4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,264,1924,1948]" targetBox="[152,1435,184,1871]" targetPageId="20">
<paragraph id="086136E0B82AFFCC49A1D4EFFBE33DD8" blockId="20.[151,1437,1924,2020]" pageId="20" pageNumber="21">
<emphasis id="3AAAEAF2B82AFFCC49A1D4EFFEFB3DA0" bold="true" box="[151,339,1924,1948]" pageId="20" pageNumber="21">FIGURES 48–54.</emphasis>
<taxonomicName id="CFDE4D63B82AFFCC486FD4EFFD523DA0" authority="(Schaum, 1862)" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[345,762,1924,1948]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B82AFFCC486FD4EFFDFA3DA0" box="[345,594,1924,1948]" italics="true" pageId="20" pageNumber="21">Heptodonta melanopyga</emphasis>
(
<bibRefCitation id="6C4F4B11B82AFFCC4B68D4EFFD5B3DA0" author="Schaum, H." box="[606,755,1924,1948]" pageId="20" pageNumber="21" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum, 1862</bibRefCitation>
)
</taxonomicName>
, Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm.
</paragraph>
</caption>
<paragraph id="086136E0B82BFFCD49F1D3FCFA2F3895" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
Dorsal surface of head copper or bronze with metallic green areas on vertex and lateral occiput, or metallic bluish-green with iridescent yellowish-bronze areas on frons, vertex, and occiput, plus blue areas on vertex, temples blue. Frons transversely to irregularly grooved, crumpled wrinkles in transition to vertex. Vertex median with parallel longitudinal striation, that converging in transition to the orbital plates. Orbital plates extensively smooth, with fine, convex to parallel converging striae and two setae on each side. Occiput with irregular transverse grooves, anteromedian irregularly rugose. Genae glabrous and shallowly grooved, violet-blue posterior and yellowish-green anterior. Clypeus glabrous, iridescent bronze and/or green, often with two dull dark-blue frontal impressions. Labrum testaceous, with four setae, five apical teeth, and one lateral tooth on each side, three median teeth acute, third apical teeth often reduced, shape variable, male labrum rectangular and short (
<quantity id="CF269B05B82BFFCD4D32D2DCFB003BED" box="[1028,1192,439,465]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" metricValueMax="7.0" metricValueMin="4.8" pageId="21" pageNumber="22" unit="mm" value="0.59" valueMax="0.7" valueMin="0.48">0.48–0.70 mm</quantity>
long, 1.28–1.73 wide mm, 
<figureCitation id="90E52A65B82BFFCD49E6D2B0FE8A3BC9" box="[208,290,474,501]" captionStart="FIGURES 48–54" captionStartId="20.[151,264,1924,1948]" captionTargetBox="[152,1435,184,1871]" captionTargetId="figure-20@20.[151,1435,181,1900]" captionTargetPageId="20" captionText="FIGURES 48–54. Heptodonta melanopyga (Schaum, 1862), Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="21" pageNumber="22">Fig. 50</figureCitation>
); female labrum antero-triangular or three median teeth protruding (
<quantity id="CF269B05B82BFFCD4D25D2B1FB1E3BC9" box="[1043,1206,474,501]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" metricValueMax="10.5" metricValueMin="8.299999999999999" pageId="21" pageNumber="22" unit="mm" value="0.94" valueMax="1.05" valueMin="0.83">0.83–1.05 mm</quantity>
long, 
<quantity id="CF269B05B82BFFCD4DCDD2B0FA353BC9" box="[1275,1437,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="1.515" metricValueMax="1.63" metricValueMin="1.4" pageId="21" pageNumber="22" unit="mm" value="1.515" valueMax="1.63" valueMin="1.4">1.40–1.63 mm</quantity>
wide, 
<figureCitation id="90E52A65B82BFFCD49EAD294FE873825" box="[220,303,510,537]" captionStart="FIGURES 48–54" captionStartId="20.[151,264,1924,1948]" captionTargetBox="[152,1435,184,1871]" captionTargetId="figure-20@20.[151,1435,181,1900]" captionTargetPageId="20" captionText="FIGURES 48–54. Heptodonta melanopyga (Schaum, 1862), Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="21" pageNumber="22">Fig. 51</figureCitation>
). Mandibles testaceous, with black teeth. Labial and maxillary palpi testaceous, terminal palpomeres apically to entirely black, pre-terminal maxillary palpomere sometimes apically dark testaceous in females. Antennae slender, extending back over the middle of the elytra, metallic black, scape, pedicel, and antennomeres 3–4 dorsally rarely with a gentle violet lustre, pedicel testaceous at base, antennomeres 5–6 sometimes lightened, scape with a single apical seta, antennomeres 3–4 with few scattered setae, antennomeres 5–11 finely and evenly pubescent.
</paragraph>
<paragraph id="086136E0B82BFFCD49F1D1D8FCF839F5" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
Thorax entirely glabrous. Pronotum most frequently copper or bronze with green lateral margin, but also green or bluish-green with bronze disc and blue lateral margin, strong median line usually dull blue with green margin, pronotal shape variable, usually slightly longer in small specimens and slightly wider in large specimens (
<quantity id="CF269B05B82BFFCD4C1CD190FF6B3905" metricMagnitude="-3" metricUnit="m" metricValue="2.365" metricValueMax="2.63" metricValueMin="2.1" pageId="21" pageNumber="22" unit="mm" value="2.365" valueMax="2.63" valueMin="2.1">2.10–2.63 mm</quantity>
long, 
<quantity id="CF269B05B82BFFCD4831D074FE013905" box="[263,425,799,825]" metricMagnitude="-3" metricUnit="m" metricValue="2.405" metricValueMax="2.78" metricValueMin="2.03" pageId="21" pageNumber="22" unit="mm" value="2.405" valueMax="2.78" valueMin="2.03">2.03–2.78 mm</quantity>
wide), pronounced transverse grooves generally irregular on disc, distinct anterior and posterior sulci, anterior lobe wider than posterior, lateral margins of the median lobe moderately to distinctly converging to the base. Episterna violet, pro- and mesepisternum sometimes and metepisternum frequently covered with iridescent bluish-green to yellowish-green lustre. Prosternum, posterior mesosternum, metasternum, and epimera iridescent bluish-green, yellowish-green, or yellowish-bronze.
</paragraph>
<paragraph id="086136E0B82BFFCD49F1D0B8FBE43E9D" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
Elytra elongate, length 
<quantity id="CF269B05B82BFFCD48FED0B8FDF839D1" box="[456,592,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.4" metricValueMin="6.1" pageId="21" pageNumber="22" unit="mm" value="7.25" valueMax="8.4" valueMin="6.1">6.1–8.4 mm</quantity>
, slightly dilated laterally to almost parallel, maximal width posterior, bronze with green lateral colouration or green with bronze lustre along the sutures and blue lateral colouration, lateral margin violet, apex green to violet, discal and juxtahumeral impression moderate or slightly pronounced, impression on posterior declivity moderate or lacking, apical impression deep, basodiscal convexity and posterior gibbosity moderate, punctures irregularly distributed and decreasing in size to apex (
<figureCitation id="90E52A65B82BFFCD4AD9D708FB9A3E41" box="[1007,1074,1123,1149]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1827,1851]" captionTargetBox="[177,1392,191,1791]" captionTargetId="figure-15@5.[177,1410,181,1803]" captionTargetPageId="5" captionText="FIGURES 1–7. 1–2—anterior elytra: 1—Heptodonta melanopyga (Schaum, 1862), ♀, Philippines, Luzon, Banaue (PSCH); 2—H. mindoroensis Cassola, 2000, ♂, Philippines, Mindoro, Mt. Halcon (JWCW). 3–4—pronotum: 3—H. arrowi W. Horn, 1900, ♂, Myanmar, Magway Region, Pyon Village (SGCH); 4—Heptodonta horii sp. nov., ♂, holotype, Myanmar, Bago Re- gion, Chanthakwin (JWCW); 5–7—head: 5—Heptodonta tempesta sp. nov., ♂, holotype, India, Meghalaya, Balpakram NP (JWCW); 6—Heptodonta pulchella (Hope, 1831), ♂, Nepal, Bagmati, Sollenta-Dhading (JWCW); 7—Heptodonta schuelei sp. nov., ♂, holotype, India, Meghalaya, Cherrapunjee (JWCW). Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4578783" httpUri="https://zenodo.org/record/4578783/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
), punctures irregularly fused to grooves running more transverse anterior and more converging to the apex posterior.
</paragraph>
<paragraph id="086136E0B82BFFCD49F1D7C0FECB3F45" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">Coxae dark testaceous with iridescent bluish-green to yellowish-bronze lustre, pro- and mesocoxae anteriorly and metacoxae laterally densely covered with long white setae, mesocoxae posteriorly with few white setae. Trochanters and femora testaceous. Generally, distal femoral metallic black colouration expanded dorsally on profemur, dorsally and posteriorly on mesofemur, and over all sides on metafemur, covering usually one-fifth to one-third of the femora. Tibiae and tarsi metallic black. Proximoapical tibiae testaceous or at least slightly lightened. Claws rufous-testaceous.</paragraph>
<paragraph id="086136E0B82BFFCD49F1D6E8FDA23C35" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
Abdominal sternites glabrous, apart from few long setae at posterior margins, some minute single setae on sternites 4–5 of females, and in males some minute setae on posterior margins of sternites 4–5 and on posterolateral margins of sternites 2–5, dark-brown with iridescent violet-blue to yellowish-green reflections; anal sternite posterior without reflections (
<figureCitation id="90E52A65B82BFFCD489FD684FE553C35" box="[425,509,1518,1545]" captionStart="FIGURES 48–54" captionStartId="20.[151,264,1924,1948]" captionTargetBox="[152,1435,184,1871]" captionTargetId="figure-20@20.[151,1435,181,1900]" captionTargetPageId="20" captionText="FIGURES 48–54. Heptodonta melanopyga (Schaum, 1862), Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="21" pageNumber="22">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="086136E0B82BFFCD49F1D578FE5D3CA5" blockId="21.[151,1437,151,2013]" pageId="21" pageNumber="22">
Aedeagus angled, length 
<quantity id="CF269B05B82BFFCD48DCD578FD3A3C11" box="[490,658,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="2.19" metricValueMax="2.3" metricValueMin="2.08" pageId="21" pageNumber="22" unit="mm" value="2.19" valueMax="2.3" valueMin="2.08">2.08–2.30 mm</quantity>
, ventral side straight or convex, apex ventrally convex and thus tip pointed rather dorsally, dorsoapically slightly ascending over approximately a third of the aedeagus, dorsomedian convexity moderately and aedeagal base steeply sloping. Inner sac in left lateral aspect median with spoon-shaped diagonal sclerite (
<figureCitation id="90E52A65B82BFFCD4869D514FE413CA5" box="[351,489,1662,1689]" captionStart="FIGURES 48–54" captionStartId="20.[151,264,1924,1948]" captionTargetBox="[152,1435,184,1871]" captionTargetId="figure-20@20.[151,1435,181,1900]" captionTargetPageId="20" captionText="FIGURES 48–54. Heptodonta melanopyga (Schaum, 1862), Philippines, Luzon, Banaue (PSCH): 48, 52–54—♂: 48—habitus, 52—lateral body, 53—aedeagus in left lateral aspect, 54—aedeagus in right lateral aspect; 49, 51—♀: 49—habitus, 51—head; 50—head, ♂, lectotype, Philippines, Luzon (MNB). Scales: 48–52 = 1 mm, 53–54 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4578804" httpUri="https://zenodo.org/record/4578804/files/figure.png" pageId="21" pageNumber="22">Figs. 53–54</figureCitation>
).
</paragraph>
<paragraph id="086136E0B82BFFCD49F1D5C9FC9E3C81" blockId="21.[151,1437,151,2013]" box="[199,822,1698,1725]" pageId="21" pageNumber="22">
<emphasis id="3AAAEAF2B82BFFCD49F1D5C9FEE53C80" bold="true" box="[199,333,1698,1724]" pageId="21" pageNumber="22">Variability.</emphasis>
Variable in pronotal shape and colouration.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB82BFFCE49F1D5ADFDD13AC5" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="distribution">
<paragraph id="086136E0B82BFFCD49F1D5ADFA393CDE" blockId="21.[151,1437,151,2013]" box="[199,1425,1734,1762]" pageId="21" pageNumber="22">
<emphasis id="3AAAEAF2B82BFFCD49F1D5ADFEC93CDC" bold="true" box="[199,353,1734,1760]" pageId="21" pageNumber="22">Distribution.</emphasis>
Northern and central 
<emphasis id="3AAAEAF2B82BFFCD4B60D5ADFCA13CDC" bold="true" box="[598,777,1734,1760]" pageId="21" pageNumber="22">
<collectingCountry id="70C97670B82BFFCD4B60D5ADFCA13CDC" box="[598,777,1734,1760]" name="Philippines" pageId="21" pageNumber="22">PHILIPPINES</collectingCountry>
</emphasis>
(Batan, Luzon, Marindugue, Mindoro, Negros, Sibuyan).
</paragraph>
<paragraph id="086136E0B82BFFCE49F1D581FDD13AC5" blockId="21.[151,1437,151,2013]" lastBlockId="22.[151,1436,150,465]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="3AAAEAF2B82BFFCD49F1D581FD913D38" bold="true" box="[199,569,1770,1796]" pageId="21" pageNumber="22">
<collectingRegion id="CA1AF802B82BFFCD49F1D581FEEF3D38" box="[199,327,1770,1796]" pageId="21" pageNumber="22">Additional</collectingRegion>
material examined.
</emphasis>
<materialsCitation id="B8B63CBDB82BFFCD4B74D580FA913D39" ID-GBIF-Occurrence="3045076488" box="[578,1337,1770,1797]" collectingDate="1983-09-03" collectionCode="ZSM" country="Philippines" location="Basco" pageId="21" pageNumber="22" specimenCount="136" stateProvince="Batan">
<specimenCount id="1ED8FD69B82BFFCD4B74D580FD453D39" box="[578,749,1771,1797]" pageId="21" pageNumber="22" type="generic">136 specimens</specimenCount>
. 
<emphasis id="3AAAEAF2B82BFFCD4BCCD581FBB83D39" bold="true" box="[762,1040,1770,1797]" pageId="21" pageNumber="22">
<collectingCountry id="70C97670B82BFFCD4BCCD581FC053D38" box="[762,941,1770,1796]" name="Philippines" pageId="21" pageNumber="22">PHILIPPINES</collectingCountry>
: 
<collectingRegion id="CA1AF802B82BFFCD4AF6D580FBA43D39" box="[960,1036,1771,1797]" pageId="21" pageNumber="22">Batan</collectingRegion>
:
</emphasis>
<location id="0D01603BB82BFFCD4D2CD580FBC93D39" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4D2CD580FBC93D39" box="[1050,1121,1771,1797]" country="Philippines" name="Basco" pageId="21" pageNumber="22" stateProvince="Batan">Basco</location>
, 
<date id="7C601020B82BFFCD4D59D580FB483D39" box="[1135,1248,1771,1797]" pageId="21" pageNumber="22" value="1983-09-03">
<collectingDate id="6C24E9C8B82BFFCD4D59D580FB483D39" box="[1135,1248,1771,1797]" pageId="21" pageNumber="22" value="1983-09-03">3.IX.1983</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4DC7D580FA983D39" box="[1265,1328,1771,1797]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="21" pageNumber="22" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4C71D580FD803D15" ID-GBIF-Occurrence="3045076310" collectingDate="1986-05-27" collectionCode="JWCW" country="Philippines" isTailEnumeration="true" location="Mt. Province" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Luzon">
<emphasis id="3AAAEAF2B82BFFCD4C71D580FA343D39" bold="true" box="[1351,1436,1771,1797]" pageId="21" pageNumber="22">
<collectingRegion id="CA1AF802B82BFFCD4C71D580FA303D39" box="[1351,1432,1771,1797]" pageId="21" pageNumber="22">Luzon</collectingRegion>
:
</emphasis>
<location id="0D01603BB82BFFCD49A1D464FE993D15" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD49A1D464FE993D15" box="[151,305,1807,1833]" country="Philippines" name="Mt. Province" pageId="21" pageNumber="22" stateProvince="Luzon">Mt. Province</location>
, 
<date id="7C601020B82BFFCD4877D464FE1C3D15" box="[321,436,1807,1833]" pageId="21" pageNumber="22" value="1986-05-27">
<collectingDate id="6C24E9C8B82BFFCD4877D464FE1C3D15" box="[321,436,1807,1833]" pageId="21" pageNumber="22" value="1986-05-27">27.V.1986</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD48F1D464FD883D15" box="[455,544,1807,1833]" pageId="21" pageNumber="22">JWCW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4B00D464FC353D15" ID-GBIF-Occurrence="3045076484" box="[566,925,1807,1833]" collectionCode="JWCW" country="Philippines" location="Abra Province" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Abra">
<collectingRegion id="CA1AF802B82BFFCD4B00D464FD483D15" box="[566,736,1807,1833]" country="Philippines" name="Abra" pageId="21" pageNumber="22">Abra Province</collectingRegion>
, 1978 (
<collectionCode id="6ECFAE25B82BFFCD4A0AD464FC3D3D15" box="[828,917,1807,1833]" pageId="21" pageNumber="22">JWCW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4A9AD464FAFE3D15" ID-GBIF-Occurrence="3045076349" box="[940,1366,1807,1833]" collectingDate="2012-06" collectionCode="JWCW" country="Philippines" location="Isabella Province" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Isabela">
<collectingRegion id="CA1AF802B82BFFCD4A9AD464FBDD3D15" box="[940,1141,1807,1833]" country="Philippines" name="Isabela" pageId="21" pageNumber="22">Isabella Province</collectingRegion>
, 
<date id="7C601020B82BFFCD4DB3D464FB4A3D15" box="[1157,1250,1807,1833]" pageId="21" pageNumber="22" value="2012-06">
<collectingDate id="6C24E9C8B82BFFCD4DB3D464FB4A3D15" box="[1157,1250,1807,1833]" pageId="21" pageNumber="22" value="2012-06">VI.2012</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4DC3D464FAE63D15" box="[1269,1358,1807,1833]" pageId="21" pageNumber="22">JWCW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4C50D464FE6D3D71" ID-GBIF-Occurrence="3045076593" collectionCode="SNMF, MNB, SDEI" country="Philippines" location="Imugan" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Luzon">
<location id="0D01603BB82BFFCD4C50D464FF173D71" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4C50D464FF173D71" country="Philippines" name="Imugan" pageId="21" pageNumber="22" stateProvince="Luzon">Imugan</location>
(
<collectionCode id="6ECFAE25B82BFFCD49E6D458FEB73D71" box="[208,287,1843,1869]" pageId="21" pageNumber="22">SNMF</collectionCode>
, 
<collectionCode id="6ECFAE25B82BFFCD481DD458FEC63D71" box="[299,366,1843,1869]" pageId="21" pageNumber="22">MNB</collectionCode>
, 
<collectionCode id="6ECFAE25B82BFFCD484CD458FE153D71" box="[378,445,1843,1869]" pageId="21" pageNumber="22">SDEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD48E4D458FBA73D71" ID-GBIF-Occurrence="3045076322" box="[466,1039,1843,1869]" collectingDate="2016-08" collectionCode="PSCH" country="Philippines" location="North Kayapa" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Nueva Vizcaya">
<location id="0D01603BB82BFFCD48E4D458FDDC3D71" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD48E4D458FDDC3D71" box="[466,628,1843,1869]" country="Philippines" name="North Kayapa" pageId="21" pageNumber="22" stateProvince="Nueva Vizcaya">North Kayapa</location>
, 
<collectingRegion id="CA1AF802B82BFFCD4BB7D458FC873D71" box="[641,815,1843,1869]" country="Philippines" name="Nueva Vizcaya" pageId="21" pageNumber="22">Nueva Vizcaya</collectingRegion>
, 
<date id="7C601020B82BFFCD4A0DD458FC023D71" box="[827,938,1843,1869]" pageId="21" pageNumber="22" value="2016-08">
<collectingDate id="6C24E9C8B82BFFCD4A0DD458FC023D71" box="[827,938,1843,1869]" pageId="21" pageNumber="22" value="2016-08">VIII.2016</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4A8DD458FBAF3D71" box="[955,1031,1843,1869]" pageId="21" pageNumber="22">PSCH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4D2AD458FF473D4D" ID-GBIF-Occurrence="3045076524" collectingDate="2016-07" collectionCode="PSCH, SGCH" country="Philippines" location="Banaue" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Ifugao">
<collectingRegion id="CA1AF802B82BFFCD4D2AD458FBC03D71" box="[1052,1128,1843,1869]" country="Philippines" name="Ifugao" pageId="21" pageNumber="22">Ifugao</collectingRegion>
, 
<location id="0D01603BB82BFFCD4D42D458FB633D71" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4D42D458FB633D71" box="[1140,1227,1843,1869]" country="Philippines" name="Banaue" pageId="21" pageNumber="22" stateProvince="Ifugao">Banaue</location>
, 
<date id="7C601020B82BFFCD4DE1D458FA953D71" box="[1239,1341,1843,1869]" pageId="21" pageNumber="22" value="2016-07">
<collectingDate id="6C24E9C8B82BFFCD4DE1D458FA953D71" box="[1239,1341,1843,1869]" pageId="21" pageNumber="22" value="2016-07">VII.2016</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4C78D458FA303D71" box="[1358,1432,1843,1869]" pageId="21" pageNumber="22">PSCH</collectionCode>
, 
<collectionCode id="6ECFAE25B82BFFCD49A1D43CFF4F3D4D" box="[151,231,1879,1905]" pageId="21" pageNumber="22">SGCH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD49CDD43CFD383D4D" ID-GBIF-Occurrence="3045076392" box="[251,656,1879,1905]" collectingDate="2016-07" collectionCode="PSCH" country="Philippines" location="Subic" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Zambales">
<collectingRegion id="CA1AF802B82BFFCD49CDD43CFEC33D4D" box="[251,363,1879,1905]" country="Philippines" name="Zambales" pageId="21" pageNumber="22">Zambales</collectingRegion>
, 
<location id="0D01603BB82BFFCD4841D43CFE123D4D" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4841D43CFE123D4D" box="[375,442,1879,1905]" country="Philippines" name="Subic" pageId="21" pageNumber="22" stateProvince="Zambales">Subic</location>
, 
<date id="7C601020B82BFFCD48F0D43CFD833D4D" box="[454,555,1879,1905]" pageId="21" pageNumber="22" value="2016-07">
<collectingDate id="6C24E9C8B82BFFCD48F0D43CFD833D4D" box="[454,555,1879,1905]" pageId="21" pageNumber="22" value="2016-07">VII.2016</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4B0AD43CFD203D4D" box="[572,648,1879,1905]" pageId="21" pageNumber="22">PSCH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4BABD43CFC7F3D4D" ID-GBIF-Occurrence="3045076312" box="[669,983,1879,1905]" collectingDate="2016-07" collectionCode="PSCH" country="Philippines" location="Luisiana" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Luzon">
<location id="0D01603BB82BFFCD4BABD43CFCA83D4D" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4BABD43CFCA83D4D" box="[669,768,1879,1905]" country="Philippines" name="Luisiana" pageId="21" pageNumber="22" stateProvince="Luzon">Luisiana</location>
, 
<date id="7C601020B82BFFCD4A3AD43CFCDA3D4D" box="[780,882,1879,1905]" pageId="21" pageNumber="22" value="2016-07">
<collectingDate id="6C24E9C8B82BFFCD4A3AD43CFCDA3D4D" box="[780,882,1879,1905]" pageId="21" pageNumber="22" value="2016-07">VII.2016</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4AB4D43CFC663D4D" box="[898,974,1879,1905]" pageId="21" pageNumber="22">PSCH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4AD5D43CFAEF3D4D" ID-GBIF-Occurrence="3045076497" box="[995,1351,1879,1905]" collectingDate="1990-06" collectionCode="PSCH" country="Philippines" isTailEnumeration="true" location="Luisiana" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Mt. Province">
<collectingRegion id="CA1AF802B82BFFCD4AD5D43CFBD13D4D" box="[995,1145,1879,1905]" inferredBySuffix="true" pageId="21" pageNumber="22">Mt. Province</collectingRegion>
, 
<date id="7C601020B82BFFCD4DB3D43CFB4A3D4D" box="[1157,1250,1879,1905]" pageId="21" pageNumber="22" value="1990-06">
<collectingDate id="6C24E9C8B82BFFCD4DB3D43CFB4A3D4D" box="[1157,1250,1879,1905]" pageId="21" pageNumber="22" value="1990-06">VI.1990</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4DC4D43CFA963D4D" box="[1266,1342,1879,1905]" pageId="21" pageNumber="22">PSCH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4C65D43CFD0A3DA9" ID-GBIF-Occurrence="3045076547" collectingDate="2004-06" collectionCode="CKCE" country="Philippines" location="Bayombong" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Nueva Vizcaya">
<collectingRegion id="CA1AF802B82BFFCD4C65D43CFF5C3DA9" country="Philippines" name="Nueva Vizcaya" pageId="21" pageNumber="22">Nueva Vizcaya</collectingRegion>
, near 
<location id="0D01603BB82BFFCD4809D410FE633DA9" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4809D410FE633DA9" box="[319,459,1915,1941]" country="Philippines" name="Bayombong" pageId="21" pageNumber="22" stateProvince="Nueva Vizcaya">Bayombong</location>
, 
<date id="7C601020B82BFFCD48EFD410FD9E3DA9" box="[473,566,1915,1941]" pageId="21" pageNumber="22" value="2004-06">
<collectingDate id="6C24E9C8B82BFFCD48EFD410FD9E3DA9" box="[473,566,1915,1941]" pageId="21" pageNumber="22" value="2004-06">VI.2004</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4B7FD410FD323DA9" box="[585,666,1915,1941]" pageId="21" pageNumber="22">CKCE</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4B99D410FB223DA9" ID-GBIF-Occurrence="3045076318" box="[687,1162,1915,1941]" collectingDate="2014-11" collectionCode="JWCW" country="Philippines" location="Sierra Madre" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Aurora">
<collectingRegion id="CA1AF802B82BFFCD4B99D410FCA93DA9" box="[687,769,1915,1941]" country="Philippines" name="Aurora" pageId="21" pageNumber="22">Aurora</collectingRegion>
, 
<location id="0D01603BB82BFFCD4A27D410FC033DA9" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4A27D410FC033DA9" box="[785,939,1915,1941]" country="Philippines" name="Sierra Madre" pageId="21" pageNumber="22" stateProvince="Aurora">Sierra Madre</location>
, 
<date id="7C601020B82BFFCD4A8CD410FBBF3DA9" box="[954,1047,1915,1941]" pageId="21" pageNumber="22" value="2014-11">
<collectingDate id="6C24E9C8B82BFFCD4A8CD410FBBF3DA9" box="[954,1047,1915,1941]" pageId="21" pageNumber="22" value="2014-11">XI.2014</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD4D1FD410FB2A3DA9" box="[1065,1154,1915,1941]" pageId="21" pageNumber="22">JWCW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB82BFFCD4DAFD410FF453D85" ID-GBIF-Occurrence="3045076507" collectingDate="1914-06" collectionCode="MNB" country="Philippines" location="Mt. Banahao" pageId="21" pageNumber="22" specimenCount="1" stateProvince="Luzon">
<location id="0D01603BB82BFFCD4DAFD410FA993DA9" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4DAFD410FA993DA9" box="[1177,1329,1915,1941]" country="Philippines" name="Mt. Banahao" pageId="21" pageNumber="22" stateProvince="Luzon">Mt. Banahao</location>
, 
<date id="7C601020B82BFFCD4C76D410FA343DA9" box="[1344,1436,1915,1941]" pageId="21" pageNumber="22" value="1914-06">
<collectingDate id="6C24E9C8B82BFFCD4C76D410FA343DA9" box="[1344,1436,1915,1941]" pageId="21" pageNumber="22" value="1914-06">VI.1914</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B82BFFCD49A9D4F4FF4C3D85" box="[159,228,1951,1977]" pageId="21" pageNumber="22">MNB</collectionCode>
)
</materialsCitation>
; 
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<location id="0D01603BB82BFFCD49CFD4F4FEEB3D85" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD49CFD4F4FEEB3D85" box="[249,323,1951,1977]" country="Philippines" name="Limay" pageId="21" pageNumber="22" stateProvince="Luzon">Limay</location>
, 
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<location id="0D01603BB82BFFCD4B28D4F4FDD03D85" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4B28D4F4FDD03D85" box="[542,632,1951,1977]" country="Philippines" name="Bilucao" pageId="21" pageNumber="22" stateProvince="Laguna">Bilucao</location>
, 
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(
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<location id="0D01603BB82BFFCD4A50D4F4FC543D85" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4A50D4F4FC543D85" box="[870,1020,1951,1977]" country="Philippines" name="Mt. Makiling" pageId="21" pageNumber="22" stateProvince="Luzon">Mt. Makiling</location>
(
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; 
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<location id="0D01603BB82BFFCD4D52D4F4FB753D85" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4D52D4F4FB753D85" box="[1124,1245,1951,1977]" country="Philippines" name="Mt. Limay" pageId="21" pageNumber="22" stateProvince="Luzon">Mt. Limay</location>
(
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)
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<collectingRegion id="CA1AF802B82BFFCD4C73D4F4FA343D85" box="[1349,1436,1951,1977]" country="Philippines" name="Quezon" pageId="21" pageNumber="22">Quezon</collectingRegion>
<location id="0D01603BB82BFFCD49A1D4A8FF653DE1" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD49A1D4A8FF653DE1" box="[151,205,1987,2013]" country="Philippines" name="Park" pageId="21" pageNumber="22" stateProvince="Quezon">Park</location>
, 
<date id="7C601020B82BFFCD49ECD4A8FEE43DE1" box="[218,332,1987,2013]" pageId="21" pageNumber="22" value="1932-06-01">
<collectingDate id="6C24E9C8B82BFFCD49ECD4A8FEE43DE1" box="[218,332,1987,2013]" pageId="21" pageNumber="22" value="1932-06-01">1.VI.1932</collectingDate>
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(
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)
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<collectingMunicipality id="E805AC9AB82BFFCD4883D4A8FDBB3DE1" box="[437,531,1987,2013]" pageId="21" pageNumber="22">Tayabas</collectingMunicipality>
, 
<location id="0D01603BB82BFFCD4B16D4A8FDD53DE1" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4B16D4A8FDD53DE1" box="[544,637,1987,2013]" country="Philippines" municipality="Tayabas" name="Malinao" pageId="21" pageNumber="22" stateProvince="Luzon">Malinao</location>
(
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, 
<collectingRegion id="CA1AF802B82BFFCD4A7ED4A8FC0D3DE1" box="[840,933,1987,2013]" country="Philippines" name="Benguet" pageId="21" pageNumber="22">Benguet</collectingRegion>
(
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<collectingRegion id="CA1AF802B82BFFCD4D39D4A8FBCE3DE1" box="[1039,1126,1987,2013]" country="Philippines" name="Quezon" pageId="21" pageNumber="22">Quezon</collectingRegion>
<collectingMunicipality id="E805AC9AB82BFFCD4D59D4A8FB0D3DE1" box="[1135,1189,1987,2013]" pageId="21" pageNumber="22">Park</collectingMunicipality>
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<location id="0D01603BB82BFFCD4D84D4A8FAB83DE1" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB82BFFCD4D84D4A8FAB83DE1" box="[1202,1296,1987,2013]" country="Philippines" municipality="Park" name="Tabayas" pageId="21" pageNumber="22" stateProvince="Quezon">Tabayas</location>
, 
<date id="7C601020B82BFFCD4C2BD4A8FA343DE1" box="[1309,1436,1987,2013]" pageId="21" pageNumber="22" value="1931-06-16">
<collectingDate id="6C24E9C8B82BFFCD4C2BD4A8FA343DE1" box="[1309,1436,1987,2013]" pageId="21" pageNumber="22" value="1931-06-16">16.VI.1931</collectingDate>
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(
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)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE49CBD3FDFCA93A8D" ID-GBIF-Occurrence="3045076548" box="[253,769,150,177]" collectingDate="1984-09" collectionCode="ZSM" country="Philippines" location="Marindugue" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Marindugue">
<emphasis id="3AAAEAF2B828FFCE49CBD3FDFE343A8C" bold="true" box="[253,412,150,176]" pageId="22" pageNumber="23">
<collectingRegion id="CA1AF802B828FFCE49CBD3FDFE3F3A8C" box="[253,407,150,176]" pageId="22" pageNumber="23">Marindugue</collectingRegion>
:
</emphasis>
<location id="0D01603BB828FFCE489CD3FCFD9E3A8D" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE489CD3FCFD9E3A8D" box="[426,566,151,177]" country="Philippines" name="Marindugue" pageId="22" pageNumber="23" stateProvince="Marindugue">Marindugue</location>
, 
<date id="7C601020B828FFCE4B71D3FCFD0C3A8D" box="[583,676,151,177]" pageId="22" pageNumber="23" value="1984-09">
<collectingDate id="6C24E9C8B828FFCE4B71D3FCFD0C3A8D" box="[583,676,151,177]" pageId="22" pageNumber="23" value="1984-09">IX.1984</collectingDate>
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(
<collectionCode id="6ECFAE25B828FFCE4B8FD3FCFD503A8D" box="[697,760,151,177]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="22" pageNumber="23" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE4A25D3FDFBCD3A8D" ID-GBIF-Occurrence="3045076601" box="[787,1125,150,177]" collectionCode="SNMF" country="Philippines" location="Mindoro" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Mindoro">
<emphasis id="3AAAEAF2B828FFCE4A25D3FDFC2D3A8C" bold="true" box="[787,901,150,176]" pageId="22" pageNumber="23">
<collectingRegion id="CA1AF802B828FFCE4A25D3FDFC293A8C" box="[787,897,150,176]" pageId="22" pageNumber="23">Mindoro</collectingRegion>
:
</emphasis>
<location id="0D01603BB828FFCE4AA5D3FCFC5D3A8D" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE4AA5D3FCFC5D3A8D" box="[915,1013,151,177]" country="Philippines" name="Mindoro" pageId="22" pageNumber="23" stateProvince="Mindoro">Mindoro</location>
(
<collectionCode id="6ECFAE25B828FFCE4D3DD3FCFBF53A8D" box="[1035,1117,151,177]" pageId="22" pageNumber="23">SNMF</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE4D41D3FCFEC73AE9" ID-GBIF-Occurrence="3045076481" collectingDate="1985-07" collectionCode="JWCW" country="Philippines" location="Negros oriental" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Negros Oriental">
<emphasis id="3AAAEAF2B828FFCE4D41D3FCFB7C3A8D" bold="true" box="[1143,1236,151,177]" pageId="22" pageNumber="23">Negros:</emphasis>
<collectingRegion id="CA1AF802B828FFCE4DD7D3FCFA303A8D" box="[1249,1432,151,177]" country="Philippines" name="Negros Oriental" pageId="22" pageNumber="23">Negros oriental</collectingRegion>
, 
<date id="7C601020B828FFCE49A1D3D1FF553AE9" box="[151,253,186,213]" pageId="22" pageNumber="23" value="1985-07">
<collectingDate id="6C24E9C8B828FFCE49A1D3D1FF553AE9" box="[151,253,186,213]" pageId="22" pageNumber="23" value="1985-07">VII.1985</collectingDate>
</date>
(
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)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE484BD3D0FCBA3AE9" ID-GBIF-Occurrence="3045076579" box="[381,786,187,213]" collectingDate="1991-05-28" collectingDateMax="1991-06-30" collectingDateMin="1991-05-28" collectionCode="SMNS" country="Philippines" location="Murcia" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Negros">
<location id="0D01603BB828FFCE484BD3D0FE793AE9" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE484BD3D0FE793AE9" box="[381,465,187,213]" country="Philippines" name="Murcia" pageId="22" pageNumber="23" stateProvince="Negros">Murcia</location>
, 
<date id="7C601020B828FFCE48E8D3D0FD0E3AE9" box="[478,678,187,213]" pageId="22" pageNumber="23" value="1991-05-28" valueMax="1991-06-30" valueMin="1991-05-28">
<collectingDate id="6C24E9C8B828FFCE48E8D3D0FD0E3AE9" box="[478,678,187,213]" pageId="22" pageNumber="23" value="1991-05-28" valueMax="1991-06-30" valueMin="1991-05-28">28.V.–30.VI.1991</collectingDate>
</date>
(
<collectionCode id="6ECFAE25B828FFCE4B8ED3D0FCA23AE9" box="[696,778,187,213]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="22" pageNumber="23">SMNS</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE4A16D3D0FB813AE9" ID-GBIF-Occurrence="3045076486" box="[800,1065,187,213]" collectingDate="1986-05" collectionCode="ZSM" country="Philippines" location="Negros" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Negros">
<location id="0D01603BB828FFCE4A16D3D0FCDC3AE9" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE4A16D3D0FCDC3AE9" box="[800,884,187,213]" country="Philippines" name="Negros" pageId="22" pageNumber="23" stateProvince="Negros">Negros</location>
, 
<date id="7C601020B828FFCE4AB7D3D0FC793AE9" box="[897,977,187,213]" pageId="22" pageNumber="23" value="1986-05">
<collectingDate id="6C24E9C8B828FFCE4AB7D3D0FC793AE9" box="[897,977,187,213]" pageId="22" pageNumber="23" value="1986-05">V.1986</collectingDate>
</date>
(
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)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE4D01D3D1FEA83AC5" ID-GBIF-Occurrence="3045076408" collectingDate="2015-04" collectionCode="JWCW" country="Philippines" location="Mt. Guiting" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Sibuyan">
<emphasis id="3AAAEAF2B828FFCE4D01D3D1FB0B3AE8" bold="true" box="[1079,1187,186,212]" pageId="22" pageNumber="23">
<collectingRegion id="CA1AF802B828FFCE4D01D3D1FB353AE8" box="[1079,1181,186,212]" pageId="22" pageNumber="23">Sibuyan</collectingRegion>
:
</emphasis>
<location id="0D01603BB828FFCE4D9AD3D0FA9D3AE9" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE4D9AD3D0FA9D3AE9" box="[1196,1333,187,213]" country="Philippines" name="Mt. Guiting" pageId="22" pageNumber="23" stateProvince="Sibuyan">Mt. Guiting</location>
, 
<date id="7C601020B828FFCE4C75D3D1FA343AE9" box="[1347,1436,186,213]" pageId="22" pageNumber="23" value="2015-04">
<collectingDate id="6C24E9C8B828FFCE4C75D3D1FA343AE9" box="[1347,1436,186,213]" pageId="22" pageNumber="23" value="2015-04">IV.2015</collectingDate>
</date>
(
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)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE483DD3B4FE1A3AC5" ID-GBIF-Occurrence="3045076460" box="[267,434,223,249]" collectionCode="ZSM" country="Philippines" location="Espana" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Sibuyan">
<location id="0D01603BB828FFCE483DD3B4FEF43AC5" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE483DD3B4FEF43AC5" box="[267,348,223,249]" country="Philippines" name="Espana" pageId="22" pageNumber="23" stateProvince="Sibuyan">Espana</location>
(
<collectionCode id="6ECFAE25B828FFCE485DD3B4FE023AC5" box="[363,426,223,249]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="22" pageNumber="23" type="Museum">ZSM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B8B63CBDB828FFCE4888D3B4FDDD3AC5" ID-GBIF-Occurrence="3045076463" box="[446,629,223,249]" collectionCode="SDEI" country="Philippines" location="Sibuyan" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Sibuyan">
<location id="0D01603BB828FFCE4888D3B4FDB23AC5" LSID="urn:lsid:plazi:treatment:807787F6B82DFFCE49A1D755FCEE3BED:0D01603BB828FFCE4888D3B4FDB23AC5" box="[446,538,223,249]" country="Philippines" name="Sibuyan" pageId="22" pageNumber="23" stateProvince="Sibuyan">Sibuyan</location>
(
<collectionCode id="6ECFAE25B828FFCE4B1FD3B4FDC53AC5" box="[553,621,223,249]" pageId="22" pageNumber="23">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB828FFCE49F1D269FA993B21" box="[199,1329,258,285]" pageId="22" pageNumber="23" type="biology_ecology">
<paragraph id="086136E0B828FFCE49F1D269FA993B21" blockId="22.[151,1436,150,465]" box="[199,1329,258,285]" pageId="22" pageNumber="23">
<emphasis id="3AAAEAF2B828FFCE49F1D269FE8F3B20" bold="true" box="[199,295,258,284]" pageId="22" pageNumber="23">Biology.</emphasis>
According to 
<bibRefCitation id="6C4F4B11B828FFCE48F0D268FD053B21" author="Medina, M. N. D. &amp; Cabras, A. A. &amp; Ibanez, J. &amp; Opiso, G. &amp; Villanueva, R. J. T." box="[454,685,259,285]" pageId="22" pageNumber="23" pagination="37 - 45" refId="ref39775" refString="Medina, M. N. D., Cabras, A. A., Ibanez, J., Opiso, G. &amp; Villanueva, R. J. T. (2020) Annotated list of tiger beetles (Coleoptera, Cicindelidae) in Calanasan, Apayao Province, Luzon, Philippines. Check List, 16 (1), 37 - 45. https: // doi. org / 10.15560 / 16.1.37" type="journal article" year="2020">
Medina 
<emphasis id="3AAAEAF2B828FFCE4B15D268FDF33B21" box="[547,603,259,285]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
(2020)
</bibRefCitation>
“on well-exposed sandy ground near the irrigation dam.”
</paragraph>
</subSubSection>
<subSubSection id="40C4656BB828FFCE49F1D24CFCEE3BED" pageId="22" pageNumber="23" type="discussion">
<paragraph id="086136E0B828FFCE49F1D24CFCEE3BED" blockId="22.[151,1436,150,465]" pageId="22" pageNumber="23">
<emphasis id="3AAAEAF2B828FFCE49F1D24CFE933B7D" bold="true" box="[199,315,295,321]" pageId="22" pageNumber="23">Remarks.</emphasis>
Common in the northern part of the 
<collectingCountry id="70C97670B828FFCE4BD8D24CFCC43B7D" box="[750,876,295,321]" name="Philippines" pageId="22" pageNumber="23">Philippines</collectingCountry>
(
<bibRefCitation id="6C4F4B11B828FFCE4A4BD24CFBB63B7D" author="Schaum, H." box="[893,1054,295,321]" pageId="22" pageNumber="23" pagination="172 - 184" refId="ref40935" refString="Schaum, H. (1862) Die Cicindeliden der philippinischen Inseln. Berliner Entomologische Zeitschrift, 6, 172 - 184." type="journal article" year="1862">Schaum 1862</bibRefCitation>
; 
<bibRefCitation id="6C4F4B11B828FFCE4D1BD24CFB723B7D" author="Chaudoir, M." box="[1069,1242,294,321]" pageId="22" pageNumber="23" refId="ref38256" refString="Chaudoir, M. (1865) Catalogue de la collection de Cicindeletes de M. le Baron Chaudoir. Nys, Bruxelles, 64 pp. https: // doi. org / 10.5962 / bhl. title. 134840" type="book" year="1865">Chaudoir 1865</bibRefCitation>
; 
<bibRefCitation id="6C4F4B11B828FFCE4DDFD24CFA3F3B7D" author="Fleutiaux, E." box="[1257,1431,295,321]" pageId="22" pageNumber="23" refId="ref38476" refString="Fleutiaux, E. (1892) Catalogue systematique des Cicindelidae. Imprimerie H. Vaillant-Carmanne, Liege, 186 pp. https: // doi. org / 10.5962 / bhl. title. 48634" type="book" year="1892">Fleutiaux 1892</bibRefCitation>
; 
<bibRefCitation id="6C4F4B11B828FFCE49A1D220FEBC3B59" author="Horn, W." box="[151,276,331,357]" pageId="22" pageNumber="23" pagination="105 - 208" refId="ref39211" refString="Horn, W. (1910) Coleoptera Adephaga, Fam. Carabidae, Subfam. Cicindelinae. In: Wytsman, P., Genera Insectorum, Fascicule 82 B, pp. 105 - 208, pls. 6 - 15." type="journal article" year="1910">Horn 1910</bibRefCitation>
, 
<bibRefCitation id="6C4F4B11B828FFCE4816D220FEF33B59" author="Horn, W." box="[288,347,331,357]" pageId="22" pageNumber="23" pagination="1 - 345" refId="ref39316" refString="Horn, W. (1926) Carabidae, Cicindelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, 86, pp. 1 - 345." type="journal article" year="1926">1926</bibRefCitation>
; 
<bibRefCitation id="6C4F4B11B828FFCE485ED220FDAF3B59" author="Wiesner, J." box="[360,519,331,357]" pageId="22" pageNumber="23" pagination="119 - 127" refId="ref41195" refString="Wiesner, J. (1980) Beitrage zur Kenntnis der philippinischen Cicindelidae. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 119 - 127." type="journal article" year="1980">Wiesner 1980</bibRefCitation>
, 
<bibRefCitation id="6C4F4B11B828FFCE4B25D220FDE63B59" author="Wiesner, J." box="[531,590,331,357]" pageId="22" pageNumber="23" refId="ref41340" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt, Checklist of the Tiger Beetles of the World, 27. Beitrag zur Kenntnis der Cicindelidae. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">1992</bibRefCitation>
; 
<bibRefCitation id="6C4F4B11B828FFCE4B6DD220FD5F3B59" author="Cassola, F." box="[603,759,331,357]" pageId="22" pageNumber="23" pagination="491 - 509" refId="ref37970" refString="Cassola, F. (2000) Studies on tiger beetles 102. The Cicindelidae collected by Roland A. Muller in the Philippine Islands, with description of three new species (Coleoptera: Cicindelidae). Zoologische Mededelingen, 73, 491 - 509." type="journal article" year="2000">Cassola 2000</bibRefCitation>
; Cabras 
<emphasis id="3AAAEAF2B828FFCE4A6FD220FC3A3B59" box="[857,914,331,357]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
2016; 
<bibRefCitation id="6C4F4B11B828FFCE4AD4D220FBD53B59" author="Medina, M. N. D. &amp; Cabras, A. A. &amp; Ibanez, J. &amp; Opiso, G. &amp; Villanueva, R. J. T." box="[994,1149,331,357]" pageId="22" pageNumber="23" pagination="37 - 45" refId="ref39775" refString="Medina, M. N. D., Cabras, A. A., Ibanez, J., Opiso, G. &amp; Villanueva, R. J. T. (2020) Annotated list of tiger beetles (Coleoptera, Cicindelidae) in Calanasan, Apayao Province, Luzon, Philippines. Check List, 16 (1), 37 - 45. https: // doi. org / 10.15560 / 16.1.37" type="journal article" year="2020">Medina 2020</bibRefCitation>
). The co-occurrence with 
<taxonomicName id="CFDE4D63B828FFCE49A1D204FEF83BB5" authorityName="Cassola" authorityYear="2000" box="[151,336,367,393]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="mindoroensis">
<emphasis id="3AAAEAF2B828FFCE49A1D204FEF83BB5" box="[151,336,367,393]" italics="true" pageId="22" pageNumber="23">H. mindoroensis</emphasis>
</taxonomicName>
on the island Mindoro is unclear; both specimens are old material determined by W. Horn. Also, the existence of 
<taxonomicName id="CFDE4D63B828FFCE487BD2F8FE503B91" baseAuthorityName="Schaum" baseAuthorityYear="1862" box="[333,504,403,429]" class="Insecta" family="Carabidae" genus="Heptodonta" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="melanopyga">
<emphasis id="3AAAEAF2B828FFCE487BD2F8FE503B91" box="[333,504,403,429]" italics="true" pageId="22" pageNumber="23">H. melanopyga</emphasis>
</taxonomicName>
on Mindanao needs to be verified, since the only evidence for this occurrence is an entry in the list of Philippine tiger beetles by 
<bibRefCitation id="6C4F4B11B828FFCE4BA7D2DCFCEA3BED" author="Wiesner, J." box="[657,834,439,465]" pageId="22" pageNumber="23" pagination="119 - 127" refId="ref41195" refString="Wiesner, J. (1980) Beitrage zur Kenntnis der philippinischen Cicindelidae. Mitteilungen der Munchner Entomologischen Gesellschaft, 70, 119 - 127." type="journal article" year="1980">Wiesner (1980)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>