334 lines
34 KiB
XML
334 lines
34 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1046.57052" ID-GBIF-Dataset="348e46a4-8819-4e3c-b4b9-b5e88bee10d7" ID-PMC="PMC8238928" ID-Pensoft-Pub="1313-2970-1046-1" ID-Pensoft-UUID="DBC42C98E4DA5074B86525CF2FB2FA64" ID-PubMed="34239337" ID-ZooBank="66E61C4EFAFE45DE9145DB38199BDEC3" ModsDocID="1313-2970-1046-1" checkinTime="1624307367557" checkinUser="pensoft" docAuthor="Jordaens, Kurt, Goergen, Georg, Skevington, Jeffrey H., Kelso, Scott & Meyer, Marc De" docDate="2021" docId="579D924128DE5AFF974E995F3D727CA1" docLanguage="en" docName="ZooKeys 1046: 1-141" docOrigin="ZooKeys 1046" docPubDate="2021-06-21" docSource="http://dx.doi.org/10.3897/zookeys.1046.57052" docTitle="Mesembrius ingratus" docType="treatment" docVersion="4" id="DBC42C98E4DA5074B86525CF2FB2FA64" lastPageNumber="1" masterDocId="DBC42C98E4DA5074B86525CF2FB2FA64" masterDocTitle="Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data" masterLastPageNumber="141" masterPageNumber="1" pageNumber="1" updateTime="1668150539886" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Revision of the Afrotropical species of the hover fly genus Mesembrius Rondani (Diptera, Syrphidae) using morphological and molecular data</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jordaens, Kurt</mods:namePart>
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<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
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<mods:nameIdentifier type="email">kurt.jordaens@africamuseum.be</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Goergen, Georg</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4496-0495</mods:nameIdentifier>
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<mods:affiliation>International Institute for Tropical Agriculture (IITA), Biodiversity Centre, 08 BP 0932 Tri Postal, Cotonou, Benin</mods:affiliation>
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<mods:namePart>Skevington, Jeffrey H.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1445-9870</mods:nameIdentifier>
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<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:namePart>Kelso, Scott</mods:namePart>
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<mods:affiliation>Canadian National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, K. W. Neatby Building, 960 Carling Avenue, Ottawa, ON K 1 A 0 C 6, Canada</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Meyer, Marc De</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0755-2898</mods:nameIdentifier>
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<mods:affiliation>Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B- 3080 Tervuren, Belgium</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-21</mods:number>
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<mods:detail type="volume">
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<mods:number>1046</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>141</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1046.57052</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1046.57052</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1046-1</mods:identifier>
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<mods:identifier type="ZooBank">66E61C4EFAFE45DE9145DB38199BDEC3</mods:identifier>
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<treatment ID-GBIF-Taxon="183114810" LSID="urn:lsid:plazi:treatment:579D924128DE5AFF974E995F3D727CA1" httpUri="http://treatment.plazi.org/id/579D924128DE5AFF974E995F3D727CA1" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="579D9241-28DE-5AFF-974E-995F3D727CA1" authority="(Loew, 1858)" baseAuthorityName="Loew" baseAuthorityYear="1858" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">Mesembrius ingratus (Loew, 1858)</taxonomicName>
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<figureCitation captionStart="Figures 11, 12" captionStartId="F6" captionText="Figures 11, 12. Mesembrius spp., habitus, lateral view 11 M. ingratus (Loew) (♂) 12 M. longipilosus sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures11-12" httpUri="https://binary.pensoft.net/fig/557467" pageId="0" pageNumber="1">Figs 11</figureCitation>
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<figureCitation captionStart="Figures 52–57" captionStartId="F25" captionText="Figures 52 - 57. Mesembrius spp., head, frontal view 52 M. copelandi sp. nov. (♂) 53 M. cyanipennis (Bezzi) (♂) 54 M. ingratus (Loew) (♂) 55 M. longipilosus sp. nov. (♂) 56 M. madagascariensis Keiser (♂) 57 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures52-57" httpUri="https://binary.pensoft.net/fig/557486" pageId="0" pageNumber="1">, 54</figureCitation>
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<figureCitation captionStart="Figures 89–94" captionStartId="F32" captionText="Figures 89 - 94. Mesembrius spp., abdomen, dorsal view 89 M. copelandi sp. nov. (♂) 90 M. cyanipennis (Bezzi) (♂) 91 M. ingratus (Loew) (♂) 92 M. longipilosus sp. nov. (♂) 93 M. madagascariensis Keiser (♂) 94 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures89-94" httpUri="https://binary.pensoft.net/fig/557493" pageId="0" pageNumber="1">, 91</figureCitation>
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<figureCitation captionStart="Figures 131–134" captionStartId="F40" captionText="Figures 131 - 134. Mesembrius spp., right wing 131 M. copelandi sp. nov. (♂) 132 M. cyanipennis (Bezzi) (♂) 133 M. ingratus (Loew) (♂) 134 M. longipilosus sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures131-134" httpUri="https://binary.pensoft.net/fig/557501" pageId="0" pageNumber="1">, 133</figureCitation>
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<figureCitation captionStart="Figures 155–158" captionStartId="F46" captionText="Figures 155 - 158. Mesembrius, proleg, dorsal view 155 M. sulcus sp. nov. (♂) 156 M. tibialis sp. nov. (♂) 157 M. arcuatus sp. nov. (♂) 158 M. ingratus (Loew) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures155-158" pageId="0" pageNumber="1">, 158</figureCitation>
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<figureCitation captionStart="Figures 189–192" captionStartId="F54" captionText="Figures 189 - 192. Mesembrius spp., metaleg, posterior view 189 M. tarsatus (Bigot) (♂) 190 M. ingratus (Loew) (♂) 191 M. arcuatus sp. nov. (♂) 192 M. nigriceps Curran (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures189-192" httpUri="https://binary.pensoft.net/fig/557514" pageId="0" pageNumber="1">, 190</figureCitation>
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<figureCitation captionStart="Figures 205–216" captionStartId="F58" captionText="Figures 205 - 216. Mesembrius spp., male genitalia, lateral view 205 M. arcuatus sp. nov. 206 M. caffer (Loew) (nominal morph) 207 M. caffer (Loew) (spined morph) 208 M. ctenifer Hull syn. nov. 209 M. capensis (Macquart) 210 M. chapini Curran 211 M. copelandi sp. nov. 212 M. cyanipennis (Bezzi) 213 M. ingratus (Loew) 214 M. longipilosus sp. nov. 215 M. madagascariensis Keiser 216 M. minor (Bezzi)." figureDoi="10.3897/zookeys.1046.57052.figures205-216" httpUri="https://binary.pensoft.net/fig/557518" pageId="0" pageNumber="1">, 213</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Loew" authorityYear="1858" class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">Helophilus ingratus</taxonomicName>
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Loew, 1858: 380.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Loew" authorityYear="1858" class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">Helophilus ingratus</taxonomicName>
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-
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<bibRefCitation author="Loew, H" journalOrPublisher="Abhandlungen der Naturwissenschaftlichen Verein zu Sachsen und Thueringen in Halle" pageId="0" pageNumber="1" pagination="57 - 402" publicationUrl="https://www.biodiversitylibrary.org/page/8687188" refId="B39" refString="Loew, H, 1860. Die Dipteren-Fauna Suedafrika's. Erste Abtheilung. Abhandlungen der Naturwissenschaftlichen Verein zu Sachsen und Thueringen in Halle 2 (1858-1861): 57 - 402, https://www.biodiversitylibrary.org/page/8687188" title="Die Dipteren-Fauna Suedafrika's. Erste Abtheilung." url="https://www.biodiversitylibrary.org/page/8687188" volume="2" year="1860">Loew (1860)</bibRefCitation>
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: 386 -
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<bibRefCitation author="Herve-Bazin, J" journalOrPublisher="Revue Zoologique Africaine" pageId="0" pageNumber="1" pagination="279 - 298" publicationUrl="https://www.biodiversitylibrary.org/page/32991575" refId="B25" refString="Herve-Bazin, J, 1914a. Syrphidae (Dipt.) recueillis au Congo belge par le Dr J. Bequaert IV. Revue Zoologique Africaine 3: 279 - 298, https://www.biodiversitylibrary.org/page/32991575" title="Syrphidae (Dipt.) recueillis au Congo belge par le Dr J. Bequaert IV." url="https://www.biodiversitylibrary.org/page/32991575" volume="3" year="1914 a">
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<normalizedToken originalValue="Hervé-Bazin">Herve-Bazin</normalizedToken>
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(1914a)
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</bibRefCitation>
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: 297.
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||
</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Loew" authorityYear="1858" class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus (Mesembrius) ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus" subGenus="Mesembrius">Helophilus (Mesembrius) ingratus</taxonomicName>
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-
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<bibRefCitation author="Bezzi, M" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/10933663" refId="B3" refString="Bezzi, M, 1915. The Syrphidae of the Ethiopian Region Based on Material in the collection of the British Museum (Natural History), with Descriptions of new Genera and Species. British Museum (Natural History), London, https://www.biodiversitylibrary.org/page/10933663" title="The Syrphidae of the Ethiopian Region Based on Material in the collection of the British Museum (Natural History), with Descriptions of new Genera and Species." url="https://www.biodiversitylibrary.org/page/10933663" year="1915">Bezzi (1915)</bibRefCitation>
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: 97.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName class="Insecta" family="Syrphidae" genus="Tubifera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tubifera ingrata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingrata">Tubifera ingrata</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.5147" author="Kertesz, K" journalOrPublisher="Verhandlungen der Naturforschenden Gesellschaft in Basel" pageId="0" pageNumber="1" refId="B35" refString="Kertesz, K, 1910. Catalogus Dipterorum hucusque descriptorum VII, Budapestini, 467 pp. https://doi.org/10.5962/bhl.title.5147" title="Catalogus Dipterorum hucusque descriptorum VII, Budapestini, 467 pp." url="https://doi.org/10.5962/bhl.title.5147" year="1910">
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<normalizedToken originalValue="Kertész">Kertesz</normalizedToken>
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(1910)
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</bibRefCitation>
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: 254.
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||
</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Loew" baseAuthorityYear="1858" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">Mesembrius ingratus</taxonomicName>
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-
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<bibRefCitation author="Curran, CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="33 - 89" publicationUrl="http://digitallibrary.amnh.org/handle/2246/1640?show=full" refId="B10" refString="Curran, CH, 1927. Diptera of the American Museum Congo Expedition. Part I. - Bibionidae, Bombyliidae, Dolichopodidae, Syrphidae, Trypaneidae. Bulletin of the American Museum of Natural History 57: 33 - 89, http://digitallibrary.amnh.org/handle/2246/1640?show=full" title="Diptera of the American Museum Congo Expedition. Part I. - Bibionidae, Bombyliidae, Dolichopodidae, Syrphidae, Trypaneidae." url="http://digitallibrary.amnh.org/handle/2246/1640?show=full" volume="57" year="1927">Curran (1927)</bibRefCitation>
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: 60 -
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<bibRefCitation author="Szilady, Z" journalOrPublisher="Annales Historico - Naturales Musei Nationalis Hungarici" pageId="0" pageNumber="1" pagination="91 - 101" publicationUrl="http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_1942_Vol_35_z_91.pdf" refId="B52" refString="Szilady, Z, 1942. Ostafrikanische Syrphiden und Bombyliden (Dipt.) im Ungarischen Nationalmuseum. Annales Historico - Naturales Musei Nationalis Hungarici 35: 91 - 101, http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_1942_Vol_35_z_91.pdf" title="Ostafrikanische Syrphiden und Bombyliden (Dipt.) im Ungarischen Nationalmuseum." url="http://publication.nhmus.hu/pdf/annHNHM/Annals_HNHM_1942_Vol_35_z_91.pdf" volume="35" year="1942">
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<normalizedToken originalValue="Szilády">Szilady</normalizedToken>
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(1942)
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</bibRefCitation>
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: 92 -
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<bibRefCitation author="Smith, KGV" editor="Crosskey, RW" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="1" pagination="488 - 510" refId="B49" refString="Smith, KGV, Vockeroth, JR, 1980. 38. Family Syrphidae. In: Crosskey, RW, Ed., Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London: 488 - 510" title="38. Family Syrphidae." volumeTitle="Catalogue of the Diptera of the Afrotropical Region." year="1980">Smith and Vockeroth (1980)</bibRefCitation>
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: 504.
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName baseAuthorityName="Loew" baseAuthorityYear="1858" class="Insecta" family="Syrphidae" genus="Mesembrius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mesembrius ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesembrius ingratus</emphasis>
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</taxonomicName>
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males are holoptic, have a loose black apical pile brush on the profemur, a densely yellow pilose scutum with vague longitudinal vittae, a tuft of black pile on the posterior side of the probasitarsus, a slender metatibia and a posterior deep depression in the proximal half of the metatibia which ventrally extends into a deep groove. It can be distinguished from any other species by the apical pile brush of the profemur which is loose and yellowish with some black pile interspersed (black dorsally, yellow ventrally in
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<taxonomicName lsidName="M. arcuatus" pageId="0" pageNumber="1" rank="species" species="arcuatus">
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<emphasis italics="true" pageId="0" pageNumber="1">M. arcuatus</emphasis>
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</taxonomicName>
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sp. nov.; black in
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<taxonomicName lsidName="M. tarsatus" pageId="0" pageNumber="1" rank="species" species="tarsatus">
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<emphasis italics="true" pageId="0" pageNumber="1">M. tarsatus</emphasis>
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</taxonomicName>
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) and by the deep groove on the metatibia (strongly compressed in
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<taxonomicName lsidName="M. arcuatus" pageId="0" pageNumber="1" rank="species" species="arcuatus">
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<emphasis italics="true" pageId="0" pageNumber="1">M. arcuatus</emphasis>
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</taxonomicName>
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sp. nov.; with a rounded swelling in
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<taxonomicName lsidName="M. tarsatus" pageId="0" pageNumber="1" rank="species" species="tarsatus">
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<emphasis italics="true" pageId="0" pageNumber="1">M. tarsatus</emphasis>
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</taxonomicName>
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). The female is unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Examined material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Loew" authorityYear="1858" class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">
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<emphasis italics="true" pageId="0" pageNumber="1">Helophilus ingratus</emphasis>
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</taxonomicName>
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Loew:
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<typeStatus>Holotype</typeStatus>
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, male, "
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<taxonomicName class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Helophilus</taxonomicName>
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//
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<taxonomicName authorityName="Loew" authorityYear="1858" class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">Helophilus ingratus</taxonomicName>
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"
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<normalizedToken originalValue="“209”">"209"</normalizedToken>
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<normalizedToken originalValue="“209”">"209"</normalizedToken>
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"
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<typeStatus>HOLOTYPUS</typeStatus>
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//
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<taxonomicName authorityName="Loew" authorityYear="1858" class="Insecta" family="Syrphidae" genus="Helophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helophilus ingratus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ingratus">Helophilus ingratus</taxonomicName>
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// Loew, 1858" "design. Kassebeer 1993" "NHRS-BYWS // 000002619" [NRMS].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="Mount" country="Malawi" location="Mulanje" specimenCount="1" specimenCount-male="1" stateProvince="Mulanje">
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<collectingCountry name="Malawi">Malawi</collectingCountry>
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•
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
;
|
||
<collectorName>Mount</collectorName>
|
||
<collectingRegion country="Malawi" name="Mulanje">Mulanje</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<date value="1912-10-20">20 Oct 1912</date>
|
||
; S.A. Neave leg.; NHMUK.
|
||
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Senegal" location="Dakar" specimenCount="2" specimenCount-male="2" stateProvince="Dakar">
|
||
<collectingCountry name="Senegal">Senegal</collectingCountry>
|
||
•
|
||
<specimenCount type="male">2♂♂</specimenCount>
|
||
;
|
||
<collectingRegion country="Senegal" name="Dakar">Dakar</collectingRegion>
|
||
</materialsCitation>
|
||
;
|
||
<date value="1945-01-14">14 Jan 1945</date>
|
||
; collector unknown; RMNH.
|
||
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="South Africa" location="KwaZulu-Natal" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
|
||
<collectingCountry name="South Africa">South Africa</collectingCountry>
|
||
•
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
;
|
||
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation collectingDate="2010-12-13" collectingDateMax="2010-12-16" collectingDateMin="2010-12-13" collectorName="J. G. H. Londt" country="South Africa" location="Manguzi Forest Reserve" specimenCount="1">
|
||
<location LSID="urn:lsid:plazi:treatment:579D924128DE5AFF974E995F3D727CA1:17E98392E1709EFC7C022A769C44F036" country="South Africa" name="Manguzi Forest Reserve">Manguzi Forest Reserve</location>
|
||
;
|
||
<collectingDate value="2010-12-13" valueMax="2010-12-16" valueMin="2010-12-13">13-16 Dec 2010</collectingDate>
|
||
;
|
||
<collectorName>J.G.H. Londt</collectorName>
|
||
leg.; NMSA
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="South Africa" location="KwaZulu-Natal" specimenCount="1" specimenCount-male="1" stateProvince="KwaZulu-Natal">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
;
|
||
<collectingRegion country="South Africa" name="KwaZulu-Natal">KwaZulu-Natal</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation collectingDate="1988-02-02" collectorName="J. G. H. Londt" country="St. Lucia" location="Park Reserve" specimenCount="1">
|
||
<collectingCountry name="Saint Lucia">St. Lucia</collectingCountry>
|
||
<location LSID="urn:lsid:plazi:treatment:579D924128DE5AFF974E995F3D727CA1:E2C6D20476BA6313C5C10B0471D08EC2" country="St. Lucia" name="Park Reserve">Park Reserve</location>
|
||
;
|
||
<collectingDate value="1988-02-02">2 Feb 1988</collectingDate>
|
||
;
|
||
<collectorName>J.G.H. Londt</collectorName>
|
||
leg.; NMSA
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation collectingDate="1988-04-26" collectorName="J. G. H. Londt" country="St. Lucia" location="Ngoya Forest Reserve" municipality="KwaZulu-Natal" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount type="male">1♂</specimenCount>
|
||
;
|
||
<collectingMunicipality>KwaZulu-Natal</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:579D924128DE5AFF974E995F3D727CA1:0496F464FA971D9537ACFA99501EBC92" country="St. Lucia" municipality="KwaZulu-Natal" name="Ngoya Forest Reserve">Ngoya Forest Reserve</location>
|
||
;
|
||
<collectingDate value="1988-04-26">26 Apr 1988</collectingDate>
|
||
;
|
||
<collectorName>J.G.H. Londt</collectorName>
|
||
leg.; NMSA
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="description">
|
||
<paragraph pageId="0" pageNumber="1">Re-description male</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 11, 12" captionStartId="F6" captionText="Figures 11, 12. Mesembrius spp., habitus, lateral view 11 M. ingratus (Loew) (♂) 12 M. longipilosus sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures11-12" httpUri="https://binary.pensoft.net/fig/557467" pageId="0" pageNumber="1">11</figureCitation>
|
||
).
|
||
</emphasis>
|
||
Body length: 10.9-11.1 mm. Wing length: 8.0-8.4 mm.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 52–57" captionStartId="F25" captionText="Figures 52 - 57. Mesembrius spp., head, frontal view 52 M. copelandi sp. nov. (♂) 53 M. cyanipennis (Bezzi) (♂) 54 M. ingratus (Loew) (♂) 55 M. longipilosus sp. nov. (♂) 56 M. madagascariensis Keiser (♂) 57 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures52-57" httpUri="https://binary.pensoft.net/fig/557486" pageId="0" pageNumber="1">54</figureCitation>
|
||
). Eyes bare; holoptic, length of eye contiguity approx. 1/3 the length of ocellar triangle. Face white with dark medial vitta; white pilose; white pollinose. Vertical triangle with black pile in ventral half and at ocellar triangle; yellow pilose in dorsal half; yellow pollinose. Distance between lateral ocellus and eye margin 1/2 width of ocellus. Frontal triangle and gena white; white pilose; white pollinose. Frontal prominence shiny black; black pilose. Occiput black, but strongly white-grey pollinose; yellow-brown pilose, with a row of almost equally long black pile at dorsal eye margin. Antenna reddish-brown.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax.</emphasis>
|
||
Scutum dark brown with dorsally a pair of vague yellow vittae; yellow-brown pilose. Scutellum uniformly light yellow-brown; yellow pilose.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs.</emphasis>
|
||
All legs dark brown, except for pro- and mesotarsi which are yellow-brown. All femora and tibiae with long, loose yellow pile. Proleg (Fig.
|
||
<figureCitation captionStart="Figures 155–158" captionStartId="F46" captionText="Figures 155 - 158. Mesembrius, proleg, dorsal view 155 M. sulcus sp. nov. (♂) 156 M. tibialis sp. nov. (♂) 157 M. arcuatus sp. nov. (♂) 158 M. ingratus (Loew) (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures155-158" pageId="0" pageNumber="1">158</figureCitation>
|
||
): Femur with a loose, yellow apical pile brush interspersed with some long black pile; very short thick pile and longer thin black pile at proximal 1/4 ventrally. Tibia with long, black pile, except dorsally. Basitarsus black pilose dorsally, with tuft of black pile on posterior side, short orange pilose ventrally. Other tarsi black pilose dorsally, short orange pilose ventrally. Mesoleg: Femur long yellow pilose posteriorly and posterodorsally, except at distal end where pile is black; short black pilose anteriorly and anterodorsally; ventrally with longer black pile. Tibia long black and short yellow pilose. Tarsi short black pilose dorsally; short yellow pilose ventrally with a few thick black spines. Metaleg (Fig.
|
||
<figureCitation captionStart="Figures 189–192" captionStartId="F54" captionText="Figures 189 - 192. Mesembrius spp., metaleg, posterior view 189 M. tarsatus (Bigot) (♂) 190 M. ingratus (Loew) (♂) 191 M. arcuatus sp. nov. (♂) 192 M. nigriceps Curran (♂). Abbreviations: bt-basitarsus, fem-femur, tib-tibia." figureDoi="10.3897/zookeys.1046.57052.figures189-192" httpUri="https://binary.pensoft.net/fig/557514" pageId="0" pageNumber="1">190</figureCitation>
|
||
): Femur very slender; covered with long, thin yellow pile; shorter black pile on ventral 1/4 and at distal end. Tibia with a deep posterior depression in the proximal half which is extended as a groove on the ventral side, demarcated with short, dense black pile; proximal half dorsoventrally flattened; predominantly black pilose. Tarsi black pilose dorsally; orange pilose ventrally.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 131–134" captionStartId="F40" captionText="Figures 131 - 134. Mesembrius spp., right wing 131 M. copelandi sp. nov. (♂) 132 M. cyanipennis (Bezzi) (♂) 133 M. ingratus (Loew) (♂) 134 M. longipilosus sp. nov. (♂)." figureDoi="10.3897/zookeys.1046.57052.figures131-134" httpUri="https://binary.pensoft.net/fig/557501" pageId="0" pageNumber="1">133</figureCitation>
|
||
). Entire wing uniformly dense microtrichose.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 89–94" captionStartId="F32" captionText="Figures 89 - 94. Mesembrius spp., abdomen, dorsal view 89 M. copelandi sp. nov. (♂) 90 M. cyanipennis (Bezzi) (♂) 91 M. ingratus (Loew) (♂) 92 M. longipilosus sp. nov. (♂) 93 M. madagascariensis Keiser (♂) 94 M. minor (Bezzi) (♂)." figureDoi="10.3897/zookeys.1046.57052.figures89-94" httpUri="https://binary.pensoft.net/fig/557493" pageId="0" pageNumber="1">91</figureCitation>
|
||
). Tergite II with pair of very large, yellow triangular maculae; yellow pilose; black marking hourglass-shaped; posterior black marking equal in size or somewhat narrower than anterior black marking, but more vague because of medial white pollinosity; posterior black marking with black pile that posterolaterally extends into the yellow maculae. Tergite III with yellow fascia of variable size, occupying approx. the entire tergite; yellow pilose; with posterior, triangular black marking; black pilose; white pollinose. Tergite IV black; white pilose and white pollinose on anterior fascia, black and yellow pilose on remainder of tergite.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figures 205–216" captionStartId="F58" captionText="Figures 205 - 216. Mesembrius spp., male genitalia, lateral view 205 M. arcuatus sp. nov. 206 M. caffer (Loew) (nominal morph) 207 M. caffer (Loew) (spined morph) 208 M. ctenifer Hull syn. nov. 209 M. capensis (Macquart) 210 M. chapini Curran 211 M. copelandi sp. nov. 212 M. cyanipennis (Bezzi) 213 M. ingratus (Loew) 214 M. longipilosus sp. nov. 215 M. madagascariensis Keiser 216 M. minor (Bezzi)." figureDoi="10.3897/zookeys.1046.57052.figures205-216" httpUri="https://binary.pensoft.net/fig/557518" pageId="0" pageNumber="1">213</figureCitation>
|
||
). Epandrium: Dorsal lobe of surstylus somewhat elongated, broadly rounded, with short, black spines on almost entire surface; dorsally long yellow pilose. Ventral lobe of surstylus straight; bare.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="female">
|
||
<paragraph pageId="0" pageNumber="1">Female.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">Malawi, Senegal and South Africa.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="comments">
|
||
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation author="Curran, CH" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="1" pagination="33 - 89" publicationUrl="http://digitallibrary.amnh.org/handle/2246/1640?show=full" refId="B10" refString="Curran, CH, 1927. Diptera of the American Museum Congo Expedition. Part I. - Bibionidae, Bombyliidae, Dolichopodidae, Syrphidae, Trypaneidae. Bulletin of the American Museum of Natural History 57: 33 - 89, http://digitallibrary.amnh.org/handle/2246/1640?show=full" title="Diptera of the American Museum Congo Expedition. Part I. - Bibionidae, Bombyliidae, Dolichopodidae, Syrphidae, Trypaneidae." url="http://digitallibrary.amnh.org/handle/2246/1640?show=full" volume="57" year="1927">Curran (1927)</bibRefCitation>
|
||
cites several specimens from three localities in the Democratic Republic of the Congo. In a later publication,
|
||
<bibRefCitation author="Curran, CH" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="1" pagination="1 - 11" publicationUrl="http://digitallibrary.amnh.org/bitstream/handle/2246/2208/N1010.pdf?sequence=1" refId="B12" refString="Curran, CH, 1939. Records and descriptions of African Syrphidae III (Diptera). American Museum Novitates 1025: 1 - 11, http://digitallibrary.amnh.org/bitstream/handle/2246/2208/N1010.pdf?sequence=1" title="Records and descriptions of African Syrphidae III (Diptera)." url="http://digitallibrary.amnh.org/bitstream/handle/2246/2208/N1010.pdf?sequence=1" volume="1025" year="1939">Curran (1939)</bibRefCitation>
|
||
suggests that these specimens belong to
|
||
<taxonomicName lsidName="M. tarsatus." pageId="0" pageNumber="1" rank="species" species="tarsatus.">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. tarsatus.</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation author="Bezzi, M" journalOrPublisher="British Museum (Natural History), London" pageId="0" pageNumber="1" publicationUrl="https://www.biodiversitylibrary.org/page/10933663" refId="B3" refString="Bezzi, M, 1915. The Syrphidae of the Ethiopian Region Based on Material in the collection of the British Museum (Natural History), with Descriptions of new Genera and Species. British Museum (Natural History), London, https://www.biodiversitylibrary.org/page/10933663" title="The Syrphidae of the Ethiopian Region Based on Material in the collection of the British Museum (Natural History), with Descriptions of new Genera and Species." url="https://www.biodiversitylibrary.org/page/10933663" year="1915">Bezzi (1915)</bibRefCitation>
|
||
also mentions the species from Uganda and Durban in South Africa, but
|
||
<bibRefCitation author="Curran, CH" journalOrPublisher="American Museum Novitates" pageId="0" pageNumber="1" pagination="1 - 11" publicationUrl="http://digitallibrary.amnh.org/bitstream/handle/2246/2208/N1010.pdf?sequence=1" refId="B12" refString="Curran, CH, 1939. Records and descriptions of African Syrphidae III (Diptera). American Museum Novitates 1025: 1 - 11, http://digitallibrary.amnh.org/bitstream/handle/2246/2208/N1010.pdf?sequence=1" title="Records and descriptions of African Syrphidae III (Diptera)." url="http://digitallibrary.amnh.org/bitstream/handle/2246/2208/N1010.pdf?sequence=1" volume="1025" year="1939">Curran (1939)</bibRefCitation>
|
||
also considers these specimens as
|
||
<taxonomicName lsidName="M. tarsatus" pageId="0" pageNumber="1" rank="species" species="tarsatus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. tarsatus</emphasis>
|
||
</taxonomicName>
|
||
. We have not encountered females that could be associated with the male of
|
||
<taxonomicName lsidName="M. ingratus" pageId="0" pageNumber="1" rank="species" species="ingratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. ingratus</emphasis>
|
||
</taxonomicName>
|
||
. No DNA barcodes are available for
|
||
<taxonomicName lsidName="M. ingratus" pageId="0" pageNumber="1" rank="species" species="ingratus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">M. ingratus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |