treatments-xml/data/83/66/87/8366879DE935853FFF6FE80F0CF4F8BE.xml

<document id="7488EC6C283BDA25B70EABF46E79FCE8" ID-CLB-Dataset="38982" ID-DOI="10.11646/zootaxa.4116.1.1" ID-GBIF-Dataset="c41804b4-7a66-4a00-afc1-c7a31871329f" ID-ISSN="1175-5326" ID-Zenodo-Dep="257322" ID-ZooBank="22D15539-E49E-4D6C-BFCF-D4DBC72BA640" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464331980959" checkinUser="plazi" docAuthor="Mathis, Wayne N. &amp; Marinoni, Luciane" docDate="2016" docId="8366879DE935853FFF6FE80F0CF4F8BE" docLanguage="en" docName="zootaxa.4116.1..pdf.imf" docOrigin="Zootaxa 4116 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Neoephydra dasycephala Mathis &amp; Marinoni, 2016, sp. nov." docType="treatment" docVersion="17" lastPageNumber="43" masterDocId="7F5FFFE5E9118515FFF8E95A0E0AFFE9" masterDocTitle="Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States" masterLastPageNumber="110" masterPageNumber="1" pageNumber="37" updateTime="1698630767650" updateUser="plazi">
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<mods:title id="A750BC03473CE4E393A8DA139CB4A259">Revision of Ephydrini Zetterstedt (Diptera: Ephydridae) from the Americas south of the United States</mods:title>
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<mods:namePart id="10A9773C385C379FC559F2F23FA40C61">Mathis, Wayne N.</mods:namePart>
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<mods:namePart id="9A0CEC201A0B5E2322ADB343B8C03D5B">Marinoni, Luciane</mods:namePart>
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<mods:date id="3F6492BF7DB73C11C993C8A00740B3A4">2016</mods:date>
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<treatment id="8366879DE935853FFF6FE80F0CF4F8BE" ID-DOI="http://doi.org/10.5281/zenodo.6074084" ID-GBIF-Taxon="120731782" ID-Zenodo-Dep="6074084" LSID="urn:lsid:plazi:treatment:8366879DE935853FFF6FE80F0CF4F8BE" httpUri="http://treatment.plazi.org/id/8366879DE935853FFF6FE80F0CF4F8BE" lastPageId="42" lastPageNumber="43" pageId="36" pageNumber="37">
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<paragraph id="0B70368BE9358531FF6FE80F0C4DFE86" blockId="36.[151,583,341,400]" box="[151,583,341,367]" pageId="36" pageNumber="37">
<heading id="503881E7E9358531FF6FE80F0C4DFE86" bold="true" box="[151,583,341,367]" fontSize="11" level="1" pageId="36" pageNumber="37" reason="1">
<emphasis id="39BBEA99E9358531FF6FE80F0C4DFE86" bold="true" box="[151,583,341,367]" pageId="36" pageNumber="37">
9. 
<taxonomicName id="CCCF4D08E9358531FF4CE80F0FE8FE86" ID-CoL="8CZ3M" box="[180,482,341,367]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dasycephala" status="sp. nov.">
<emphasis id="39BBEA99E9358531FF4CE80F0FE8FE86" bold="true" box="[180,482,341,367]" italics="true" pageId="36" pageNumber="37">Neoephydra dasycephala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228857E2E9358531FE11E80F0C4DFE86" box="[489,583,341,367]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="43D56500E9358531FF6FE82D0F7BFE79" box="[151,369,375,400]" pageId="36" pageNumber="37" type="description">
<paragraph id="0B70368BE9358531FF6FE82D0F7BFE79" blockId="36.[151,583,341,400]" box="[151,369,375,400]" pageId="36" pageNumber="37">
<figureCitation id="93F42A0EE9358531FF6FE82D0F14FE79" box="[151,286,375,400]" captionStart="FIGURES 50 – 52" captionStartId="36.[151,264,1964,1987]" captionTargetBox="[153,1428,709,1935]" captionTargetId="figure@36.[151,1436,705,1944]" captionTargetPageId="36" captionText="FIGURES 50 – 52. Structures of male terminalia of Neoephydra dasycephala sp. nov. (50) tergite 5, epandrium, cerci, surstyli, lateral view; (51) same, posterior view; (52) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399424/files/figure.png" pageId="36" pageNumber="37">Figs. 50–52</figureCitation>
, Map 9
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9358530FF6FE8E50D6EFE44" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="0B70368BE9358531FF6FE8E50B27FD64" blockId="36.[151,1436,447,653]" pageId="36" pageNumber="37">
<emphasis id="39BBEA99E9358531FF6FE8E50F1BFE31" bold="true" box="[151,273,447,472]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
Specimens of this species could easily be confused with those of either congener of this species group but may be distinguished by the following characters: larger, body length up to 5.00 mm; legs mostly grayish green, densely microtomentose, but with ventral surface of at least hindtibia somewhat rufous; medial portion of scutum brown, mostly microtomentose, subshiny vittae less distinct, as in specimens of 
<emphasis id="39BBEA99E9358531FB74EB770B20FDAC" box="[1164,1322,556,581]" italics="true" pageId="36" pageNumber="37">
<taxonomicName id="CCCF4D08E9358531FB74EB770B29FDAC" box="[1164,1315,556,581]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="mallonota">N. mallonota</taxonomicName>
;
</emphasis>
and male terminalia with distinctive conformation. Moderately large shore flies, body length 4.30–5.00 mm (averaging 
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); grayish blue or grayish green to brown dorsally, progressively paler, grayer toward ventral surface.
</paragraph>
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<paragraph id="0B70368BE9358531FF6FEEF60B5EF80B" blockId="36.[151,1436,1964,2018]" pageId="36" pageNumber="37">
<emphasis id="39BBEA99E9358531FF6FEEF60F5EF82B" bold="true" box="[151,340,1964,1987]" pageId="36" pageNumber="37">FIGURES 50–52.</emphasis>
Structures of male terminalia of 
<taxonomicName id="CCCF4D08E9358531FD5BEEF70DA9F82A" box="[675,931,1965,1987]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="dasycephala" status="sp. nov.">
<emphasis id="39BBEA99E9358531FD5BEEF70DA9F82A" box="[675,931,1965,1987]" italics="true" pageId="36" pageNumber="37">Neoephydra dasycephala</emphasis>
</taxonomicName>
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<emphasis id="39BBEA99E9358531FC53EEF40DF6F82A" bold="true" box="[939,1020,1966,1987]" pageId="36" pageNumber="37">sp. nov.</emphasis>
</taxonomicNameLabel>
(50) tergite 5, epandrium, cerci, surstyli, lateral view; (51) same, posterior view; (52) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view.
</paragraph>
</caption>
<paragraph id="0B70368BE9348530FF3FE9CD0D6EFE44" blockId="37.[151,1437,151,861]" pageId="37" pageNumber="38">
<emphasis id="39BBEA99E9348530FF3FE9CD0F07FF59" box="[199,269,151,176]" italics="true" pageId="37" pageNumber="38">Head:</emphasis>
Head ratio 0.57–0.60; frontal ratio 0.47–0.52; mesofrons with dark greenish blue, metallic luster, more or less densely pilose; ocellar triangle and lateral margins of frons mostly concolorous, microtomentose, brownish gray to charcoal gray, more charcoal colored toward anterior margins; area between fronto-orbital seta insertions and eye less microtomentose, grayish green, slightly subshiny, bare. Antenna unicolorous, microtomentose, blackish brown to black; aristal length equal to combined length of pedicel and basal flagellomere. Facial ratio 0.76-0.81; face densely microtomentose, dark brown dorsally, gradually paler ventrally, grayish tan to nearly white. Eye ratio 0.96–0.99; gena-to-eye ratio 0.62–0.66; gena very high, coloration pale tannish white anteriorly, darker posteriorly with some faint grayish green or grayish blue tinges.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9348533FF3FE8ED0F96FEF4" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="0B70368BE9348530FF3FE8ED0F19FCB4" blockId="37.[151,1437,151,861]" pageId="37" pageNumber="38">
<emphasis id="39BBEA99E9348530FF3FE8ED0F2BFE39" box="[199,289,439,464]" italics="true" pageId="37" pageNumber="38">Thorax:</emphasis>
Mostly microtomentose; scutum brown medially with paler, grayish blue margins, but with some faint, subshiny vittae. Anterior margin with 2 distinct, subshiny brown vittae which become grayer and duller from transverse suture backward; area laterad and mediad of subshiny anterior vittae darker brown, subshiny, and fused posteriorly; gray microtomentose lateral margins particularly evident from humeral callus posteriorly through notopleural region, more bluish and slightly subshiny posteriorly; dorsum of scutellum concolorous with posteriormost portion of scutum, subshiny with metallic brownish blue luster; lateral and apical margins grayish blue, duller; anepisternum brown along dorsal margin, gradually grayer toward ventral margin; other pleural areas mostly grayish tan, unicolorous. Wing mostly hyaline or with faint, grayish brown coloration; costal vein ratio 0.39–0.41; M vein ratio 0.89–0.92. Coxae grayish white to silvery; femora generally grayish green, densely microtomentose, concolorous; tibiae concolorous with femora but some specimens with rufous coloration on ventral surface, especially on hindleg; tarsi grayish black in freshly emerged specimens, paler, rufous in older specimens.
</paragraph>
<paragraph id="0B70368BE9348530FF6FEE1E0D14F8B3" blockId="37.[151,798,1860,1882]" box="[151,798,1860,1882]" pageId="37" pageNumber="38">
<emphasis id="39BBEA99E9348530FF6FEE1E0EE0F8B0" bold="true" box="[151,234,1860,1881]" pageId="37" pageNumber="38">MAP 9.</emphasis>
Distribution map for 
<taxonomicName id="CCCF4D08E9348530FE3EEE1E0CCDF8B3" box="[454,711,1860,1882]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="dasycephala" status="sp. nov.">
<emphasis id="39BBEA99E9348530FE3EEE1E0CCDF8B3" box="[454,711,1860,1882]" italics="true" pageId="37" pageNumber="38">Neoephydra dasycephala</emphasis>
</taxonomicName>
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<emphasis id="39BBEA99E9348530FD35EE1F0D14F8B3" bold="true" box="[717,798,1861,1882]" pageId="37" pageNumber="38">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
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<emphasis id="39BBEA99E9348530FF3FEEDD0F30F849" box="[199,314,1927,1952]" italics="true" pageId="37" pageNumber="38">Abdomen:</emphasis>
Unicolorous, microtomentose to subshiny, grayish green or grayish blue, often with some faint golden, metallic luster when viewed from posteroblique angle. Tergite 5 (
<figureCitation id="93F42A0EE9348530FC0DEEF10A40F82D" box="[1013,1098,1963,1988]" captionStart="FIGURES 50 – 52" captionStartId="36.[151,264,1964,1987]" captionTargetBox="[153,1428,709,1935]" captionTargetId="figure@36.[151,1436,705,1944]" captionTargetPageId="36" captionText="FIGURES 50 – 52. Structures of male terminalia of Neoephydra dasycephala sp. nov. (50) tergite 5, epandrium, cerci, surstyli, lateral view; (51) same, posterior view; (52) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399424/files/figure.png" pageId="37" pageNumber="38">Fig. 50</figureCitation>
) of male forming equilateral triangle. Male terminalia (
<figureCitation id="93F42A0EE9348530FE39EE950C46F801" box="[449,588,1999,2024]" captionStart="FIGURES 50 – 52" captionStartId="36.[151,264,1964,1987]" captionTargetBox="[153,1428,709,1935]" captionTargetId="figure@36.[151,1436,705,1944]" captionTargetPageId="36" captionText="FIGURES 50 – 52. Structures of male terminalia of Neoephydra dasycephala sp. nov. (50) tergite 5, epandrium, cerci, surstyli, lateral view; (51) same, posterior view; (52) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399424/files/figure.png" pageId="37" pageNumber="38">Figs. 50–51</figureCitation>
): margins of epandrium from posterior view enlarged toward middle, with lateral prominences at juncture of surstyli with ventral margin of epandrium; posterior surstylar processes closely opposed basally, divergent apically; 2 other pairs of lateral, surstylar processes, basal pair with convergent orientation; aedeagus long, slender, crescent-shaped, tapered gradually to apex; phallapodeme well developed, almost as wide as long.
</paragraph>
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<subSubSection id="43D56500E9378533FF3FE87D0D05FDF1" pageId="38" pageNumber="39" type="materials_examined">
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<typeStatus id="D4748829E9378533FF3FE87D0F08FEA9" box="[199,258,295,320]" pageId="38" pageNumber="39">Type</typeStatus>
material.
</emphasis>
The 
<typeStatus id="D4748829E9378533FE41E87D0C11FEA9" box="[441,539,295,320]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
male is labeled “ 
<date id="7F71104BE9378533FD13E8720D60FEA9" box="[747,874,295,320]" pageId="38" pageNumber="39" value="1965-10-18">18 X 1965</date>
[
<date id="7F71104BE9378533FC84E8720A19FEA9" box="[892,1043,295,320]" pageId="38" pageNumber="39" value="1965-10-18">18 Oct 1965</date>
] Pusi [
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, 
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], 
<collectionCode id="6DDEAE4EE9378533FAA8E8720B97FEA9" box="[1360,1437,296,320]" pageId="38" pageNumber="39">PUNO</collectionCode>
<collectingCountry id="73D8761BE9378533FF6FE8170ED5FE8D" box="[151,223,333,356]" name="Peru" pageId="38" pageNumber="39">PERU</collectingCountry>
250 Coll. J. C. Hitchcock, Jr./♂/ 
<typeStatus id="D4748829E9378533FD97E8160D0CFE8D" box="[623,774,332,356]" pageId="38" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
♂ 
<taxonomicName id="CCCF4D08E9378533FCCDE8160AA5FE8C" authority="Mathis" authorityName="Mathis" box="[821,1199,332,357]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="dasycephala">Neoephydra dasycephala Mathis</taxonomicName>
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[red].” The 
<typeStatus id="D4748829E9378533FF6FE8350EF3FE61" box="[151,249,367,392]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
is directly pinned, is in excellent condition, and is deposited in the 
<collectionCode id="6DDEAE4EE9378533FBFAE82A0A5CFE61" box="[1026,1110,368,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="39">USNM</collectionCode>
. The allotype female and 10 
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(
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, 3♀; 
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) bear the same locality label data as the 
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. Other 
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are as follows: 
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<collectingCountry id="73D8761BE9378533FF6FE8E20F38FE26" box="[151,306,440,463]" name="Argentina" pageId="38" pageNumber="39">ARGENTINA</collectingCountry>
. Jujuy:
</emphasis>
Cangrejillos (S La Quiaca; 
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, 
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; 
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), 
<date id="7F71104BE9378533FBF8E8E20A87FE39" box="[1024,1165,439,464]" pageId="38" pageNumber="39" value="1968-10-28">28 Oct 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E9378533FAD8E8E20B4FFE39" box="[1312,1349,440,464]" pageId="38" pageNumber="39" type="generic">1♂</specimenCount>
; 
<collectionCode id="6DDEAE4EE9378533FAA9E8E20B85FE39" box="[1361,1423,440,464]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="38" pageNumber="39">CNC</collectionCode>
); Santa Catalina (
<geoCoordinate id="6EFB504CE9378533FEB1E8860FA9FE1C" box="[329,419,476,501]" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="-22.083334">22°05'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE9378533FE57E8860C1FFE1C" box="[431,533,476,501]" direction="west" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="-66.3">66°18'W</geoCoordinate>
; 
<quantity id="CC379B6EE9378533FDD8E8860C72FE1D" box="[544,632,476,500]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="38" pageNumber="39" unit="m" value="3700.0">3700 m</quantity>
), 
<date id="7F71104BE9378533FD74E8860D10FE1C" box="[652,794,476,501]" pageId="38" pageNumber="39" value="1968-10-25">25 Oct 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E9378533FC52E8860DC5FE1D" box="[938,975,476,500]" pageId="38" pageNumber="39" type="generic">1♂</specimenCount>
; 
<collectionCode id="6DDEAE4EE9378533FC23E8860A13FE1D" box="[987,1049,476,500]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="38" pageNumber="39">CNC</collectionCode>
).
</paragraph>
<paragraph id="0B70368BE9378533FF3FE8A50D05FDF1" blockId="38.[151,1437,151,861]" box="[199,783,511,536]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FE8A50F6FFDF1" bold="true" box="[199,357,511,536]" pageId="38" pageNumber="39">
<typeStatus id="D4748829E9378533FF3FE8A50F08FDF1" box="[199,258,511,536]" pageId="38" pageNumber="39">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="73D8761BE9378533FE94EB5A0FA9FDFE" box="[364,419,512,535]" name="Peru" pageId="38" pageNumber="39">Peru</collectingCountry>
. Puno: Pusi (
<geoCoordinate id="6EFB504CE9378533FDCEE8A50C9AFDF1" box="[566,656,511,536]" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="925" value="-15.433333">15°26'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE9378533FD64E8A50D09FDF1" box="[668,771,511,536]" direction="west" orientation="longitude" pageId="38" pageNumber="39" precision="925" value="-69.933334">69°56'W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9378533FF3FEB7E0B43FDD4" box="[199,1353,548,573]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="0B70368BE9378533FF3FEB7E0B43FDD4" blockId="38.[151,1437,151,861]" box="[199,1353,548,573]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEB7E0F53FDD4" bold="true" box="[199,345,548,573]" pageId="38" pageNumber="39">Distribution</emphasis>
(Map 9). 
<emphasis id="39BBEA99E9378533FE31EB7E0C5DFDD4" box="[457,599,548,573]" italics="true" pageId="38" pageNumber="39">Neotropical:</emphasis>
<collectingCountry id="73D8761BE9378533FDA6EB7E0CC7FDD4" box="[606,717,548,573]" name="Argentina" pageId="38" pageNumber="39">Argentina</collectingCountry>
(Jujuy), 
<collectingCountry id="73D8761BE9378533FCCAEB7F0D6FFDD5" box="[818,869,549,572]" name="Peru" pageId="38" pageNumber="39">Peru</collectingCountry>
(Puno), between 15°–23°S and 65°–70°W.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9378533FF3FEB1D0FF8FD41" pageId="38" pageNumber="39" type="etymology">
<paragraph id="0B70368BE9378533FF3FEB1D0FF8FD41" blockId="38.[151,1437,151,861]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEB1D0F44FD89" bold="true" box="[199,334,583,608]" pageId="38" pageNumber="39">Etymology.</emphasis>
The specific epithet, 
<emphasis id="39BBEA99E9378533FDBDEB1D0CD2FD89" box="[581,728,583,608]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="CCCF4D08E9378533FDBDEB1D0CDEFD89" box="[581,724,583,608]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="dasycephala">dasycephala</taxonomicName>
,
</emphasis>
is of Greek derivation and is the combination of the adjective 
<emphasis id="39BBEA99E9378533FF6FEB360EDAFD6C" box="[151,208,620,645]" italics="true" pageId="38" pageNumber="39">dasy,</emphasis>
meaning hairy or shaggy, and the noun 
<emphasis id="39BBEA99E9378533FD69EB360CFAFD6C" box="[657,752,620,645]" italics="true" pageId="38" pageNumber="39">cephala,</emphasis>
meaning head, in allusion to the dense pilosity over much of the body, particularly the head.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9378533FF3FEBEE0F75FC02" pageId="38" pageNumber="39" type="discussion">
<paragraph id="0B70368BE9378533FF3FEBEE0B64FCB5" blockId="38.[151,1437,151,861]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEBEE0F31FD24" bold="true" box="[199,315,692,717]" pageId="38" pageNumber="39">Remarks.</emphasis>
Although we used colorational characters in the key to distinguish this species from 
<emphasis id="39BBEA99E9378533FAFAEBEF0B96FD24" box="[1282,1436,692,717]" italics="true" pageId="38" pageNumber="39">
<taxonomicName id="CCCF4D08E9378533FAFAEBEF0B9DFD24" box="[1282,1431,692,717]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mallonota">N. mallonota</taxonomicName>
,
</emphasis>
they may be difficult to interpret without reference to accurately determined material. As a further complication, facial coloration often varies considerably intraspecifically, although trends are evident for each species, as indicated in the key and descriptions. When identification of a specimen is doubtful, structures of the male terminalia should be examined. Unassociated female specimens will perhaps not be identifiable in some cases.
</paragraph>
<paragraph id="0B70368BE9378533FF6FEAEA0F75FC02" blockId="38.[151,567,943,1003]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF6FEAEA0FDCFC20" bold="true" box="[151,470,943,970]" pageId="38" pageNumber="39">
10. 
<taxonomicName id="CCCF4D08E9378533FF3BEAF50FDCFC20" box="[195,470,943,969]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mallonota" status="sp. nov.">
<emphasis id="39BBEA99E9378533FF3BEAF50FDCFC20" bold="true" box="[195,470,943,969]" italics="true" pageId="38" pageNumber="39">Neoephydra mallonota</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="228857E2E9378533FE25EAEA0C3DFC20" box="[477,567,944,969]" pageId="38" pageNumber="39" rank="species">
<emphasis id="39BBEA99E9378533FE25EAEA0C3DFC20" bold="true" box="[477,567,944,969]" pageId="38" pageNumber="39">sp. nov.</emphasis>
</taxonomicNameLabel>
<figureCitation id="93F42A0EE9378533FF6FEA880F14FC02" box="[151,286,978,1003]" captionStart="FIGURES 53 – 57" captionStartId="39.[151,264,1891,1914]" captionTargetBox="[153,1436,345,1860]" captionTargetId="figure@39.[151,1436,337,1870]" captionTargetPageId="39" captionText="FIGURES 53 – 57. Head, thorax, and structures of male terminalia of Neoephydra mallonota sp. nov. (53) head, lateroblique view; (54) mesonotum, dorsal view; (55) tergite 5, epandrium, cerci, surstyli, lateral view; (56) same, posterior view; (57) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399425/files/figure.png" pageId="38" pageNumber="39">Figs. 53–57</figureCitation>
, Map 10
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9378533FF6FED400A6BFA56" pageId="38" pageNumber="39" type="diagnosis">
<paragraph id="0B70368BE9378533FF6FED400DEBFB76" blockId="38.[151,1437,1050,2011]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF6FED400F1BFBDA" bold="true" box="[151,273,1050,1075]" pageId="38" pageNumber="39">Diagnosis.</emphasis>
Specimens of 
<taxonomicName id="CCCF4D08E9378533FE4FED460C40FBDA" box="[439,586,1050,1075]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="mallonota">
<emphasis id="39BBEA99E9378533FE4FED460C40FBDA" box="[439,586,1050,1075]" italics="true" pageId="38" pageNumber="39">N. mallonota</emphasis>
</taxonomicName>
may be distinguished from those of similar congeners of this species group by their generally darker appearance, especially the darker scutum and legs, and by the distinctive conformation of the male terminalia. Moderately large shore flies, body length 4.00– 
<quantity id="CC379B6EE9378533FC6EED390DF1FB93" box="[918,1019,1123,1147]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="38" pageNumber="39" unit="mm" value="4.3">4.30 mm</quantity>
(averaging 
<quantity id="CC379B6EE9378533FB79ED380AE0FB92" box="[1153,1258,1122,1147]" metricMagnitude="-3" metricUnit="m" metricValue="4.16" pageId="38" pageNumber="39" unit="mm" value="4.16">4.16 mm</quantity>
); with brown to dull greenish blue coloration dorsally, becoming paler and grayer ventrally.
</paragraph>
<paragraph id="0B70368BE9378533FF3FEDF00A6BFA56" blockId="38.[151,1437,1050,2011]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEDF00F0EFB2A" box="[199,260,1194,1219]" italics="true" pageId="38" pageNumber="39">Head</emphasis>
(
<figureCitation id="93F42A0EE9378533FEECEDF00F60FB2A" box="[276,362,1194,1219]" captionStart="FIGURES 53 – 57" captionStartId="39.[151,264,1891,1914]" captionTargetBox="[153,1436,345,1860]" captionTargetId="figure@39.[151,1436,337,1870]" captionTargetPageId="39" captionText="FIGURES 53 – 57. Head, thorax, and structures of male terminalia of Neoephydra mallonota sp. nov. (53) head, lateroblique view; (54) mesonotum, dorsal view; (55) tergite 5, epandrium, cerci, surstyli, lateral view; (56) same, posterior view; (57) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399425/files/figure.png" pageId="38" pageNumber="39">Fig. 53</figureCitation>
): Head ratio 0.59–0.61; frontal ratio 0.43–0.46; mesofrons with dark greenish blue to blackish blue metallic luster, densely pilose (best seen in profile); ocellar triangle and parafrons mostly concolorous, microtomentose; parafrons generally duller and more charcoal colored toward anterior margin but with a slightly subshiny, greenish blue vitta just mesad of larger fronto-orbital setae. Antenna microtomentose, mostly unicolorous, dark brown to black or with basal segments grayer. Facial ratio 0.81–0.83; face mostly dull, microtomentose, unicolorous, yellowish gray to grayish brown, frequently slightly paler in color ventrally. Eye ratio 0.98–1.02; gena-to-eye ratio 0.61–0.67; gena very wide, grayish white to silvery anteriorly, darker, more olivaceous to brown posteriorly, with some metallic although faint bluish green tinges.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E9378533FF3FEC900B43F85E" pageId="38" pageNumber="39" type="description">
<paragraph id="0B70368BE9378533FF3FEC900C78F936" blockId="38.[151,1437,1050,2011]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEC900F12FA0A" box="[199,280,1482,1507]" italics="true" pageId="38" pageNumber="39">Thorax</emphasis>
(
<figureCitation id="93F42A0EE9378533FED4EC900F8FFA0A" box="[300,389,1482,1507]" captionStart="FIGURES 53 – 57" captionStartId="39.[151,264,1891,1914]" captionTargetBox="[153,1436,345,1860]" captionTargetId="figure@39.[151,1436,337,1870]" captionTargetPageId="39" captionText="FIGURES 53 – 57. Head, thorax, and structures of male terminalia of Neoephydra mallonota sp. nov. (53) head, lateroblique view; (54) mesonotum, dorsal view; (55) tergite 5, epandrium, cerci, surstyli, lateral view; (56) same, posterior view; (57) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399425/files/figure.png" pageId="38" pageNumber="39">Fig. 54</figureCitation>
): Microtomentose to subshiny; scutum mostly brownish to brownish blue but with faint, subshiny, darker vittae, 1 medial and 2 lateral pairs; anterior portion of scutum and lateral margins more microtomentose, becoming distinctly shiny posteriorly; lateral margins from postpronotum posteriorly through notopleural area duller, grayish blue to blue, darker, shinier posteriorly; dorsum of scutellum shiny, nearly concolorous with shiny mesofrons. Pleural areas mostly olivaceous gray, paler and grayer ventrally; anepisternum more brownish. Wing hyaline to very slightly infumate, pale brown; costal vein ratio 0.35–0.39; M vein ratio 0.85– 0.88. Legs mostly unicolorous, densely microtomentose, greenish gray, gradually more charcoal colored apically; tibiae and tarsi unicolorous, charcoal gray.
</paragraph>
<paragraph id="0B70368BE9378533FF3FEFB00B43F85E" blockId="38.[151,1437,1050,2011]" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEFB00F30F8EA" box="[199,314,1770,1795]" italics="true" pageId="38" pageNumber="39">Abdomen:</emphasis>
Dorsal surface mostly unicolorous, grayish blue to bluish brown, subshiny to shiny with some metallic, bluish green to greenish blue luster; anteromedial portion of each segment tinged with some brownish coloration, duller; tergite 5 of male wider basally than long. Male terminalia (
<figureCitation id="93F42A0EE9378533FBD2EE680AB3F8A2" box="[1066,1209,1842,1867]" captionStart="FIGURES 53 – 57" captionStartId="39.[151,264,1891,1914]" captionTargetBox="[153,1436,345,1860]" captionTargetId="figure@39.[151,1436,337,1870]" captionTargetPageId="39" captionText="FIGURES 53 – 57. Head, thorax, and structures of male terminalia of Neoephydra mallonota sp. nov. (53) head, lateroblique view; (54) mesonotum, dorsal view; (55) tergite 5, epandrium, cerci, surstyli, lateral view; (56) same, posterior view; (57) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399425/files/figure.png" pageId="38" pageNumber="39">Figs. 55–57</figureCitation>
): lateral margins of epandrium more or less parallel, lacking any widening where surstyli attach; posterior surstylar processes closely opposed basally, continued parallel to each other to apex; 1 additional pair of lateral processes, each with apical setae; internal genitalic structures generally reduced in size; aedeagus very slender, much longer than wide.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E937853DFF3FEE980D19FB61" lastPageId="40" lastPageNumber="41" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="0B70368BE9378532FF3FEE980DDAFEF5" blockId="38.[151,1437,1050,2011]" lastBlockId="39.[151,1437,151,285]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="39BBEA99E9378533FF3FEE980F7DF832" bold="true" box="[199,375,1986,2011]" pageId="38" pageNumber="39">
<typeStatus id="D4748829E9378533FF3FEE980F08F832" box="[199,258,1986,2011]" pageId="38" pageNumber="39">Type</typeStatus>
material.
</emphasis>
The 
<typeStatus id="D4748829E9378533FE4CEE980C1CF832" box="[436,534,1986,2011]" pageId="38" pageNumber="39" type="holotype">holotype</typeStatus>
male is labeled “ 
<collectingCountry id="73D8761BE9378533FD27EE980D3CF832" box="[735,822,1986,2011]" name="Bolivia" pageId="38" pageNumber="39">Bolivia</collectingCountry>
, La Paz, 
<date id="7F71104BE9378533FC5BEE990A38F832" box="[931,1074,1986,2011]" pageId="38" pageNumber="39" value="1969-05-02">2 May 1969</date>
, P &amp; P Spangler [handwritten]/ 
<typeStatus id="D4748829E9368532FF6FE9C20F21FF59" box="[151,299,152,176]" pageId="39" pageNumber="40" type="holotype">HOLOTYPE</typeStatus>
♂ 
<taxonomicName id="CCCF4D08E9368532FEB5E9CD0CA8FF59" authority="Mathis" authorityName="Mathis" box="[333,674,151,176]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mallonota">Neoephydra mallonota Mathis</taxonomicName>
<collectionCode id="6DDEAE4EE9368532FD51E9C20CF0FF59" box="[681,762,152,176]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40">USNM</collectionCode>
[red].” The 
<typeStatus id="D4748829E9368532FC7CE9CD0DECFF59" box="[900,998,151,176]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
is double mounted (minuten in a block of polyporus), is in excellent condition, and is deposited in the 
<collectionCode id="6DDEAE4EE9368532FCA2E9E60DA1FF3D" box="[858,939,188,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40">USNM</collectionCode>
(
<date id="7F71104BE9368532FC42E9E60A1EFF3D" box="[954,1044,188,212]" pageId="39" pageNumber="40" value="12-10-02">100212</date>
). Four 
<typeStatus id="D4748829E9368532FB9BE9E70AC4FF3C" box="[1123,1230,189,213]" pageId="39" pageNumber="40" type="paratype">paratypes</typeStatus>
(
<specimenCount id="1DC9FD02E9368532FB26E9E60B0EFF3D" box="[1246,1284,188,212]" pageId="39" pageNumber="40" type="generic">2♂</specimenCount>
, 2♀; 
<collectionCode id="6DDEAE4EE9368532FABAE9E60B9FFF3D" box="[1346,1429,188,212]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="39" pageNumber="40">USNM</collectionCode>
) bear the same label data as the 
<typeStatus id="D4748829E9368532FE04E9850C68FF11" box="[508,610,223,248]" pageId="39" pageNumber="40" type="holotype">holotype</typeStatus>
. Other 
<typeStatus id="D4748829E9368532FD4EE9BA0D2BFF11" box="[694,801,224,248]" pageId="39" pageNumber="40" type="paratype">paratypes</typeStatus>
are as follows: 
<emphasis id="39BBEA99E9368532FC2FE9BA0AC9FF11" box="[983,1219,224,248]" italics="true" pageId="39" pageNumber="40">
<collectingCountry id="73D8761BE9368532FC2FE9BA0A78FF1E" box="[983,1138,224,247]" name="Argentina" pageId="39" pageNumber="40">ARGENTINA</collectingCountry>
. Jujuy:
</emphasis>
Cangrejillos (S La Quiaca; 
<geoCoordinate id="6EFB504CE9368532FF0DE85E0F45FEF4" box="[245,335,260,285]" direction="south" orientation="latitude" pageId="39" pageNumber="40" precision="925" value="-22.416666">22°25'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE9368532FEA3E85E0FCAFEF4" box="[347,448,260,285]" direction="west" orientation="longitude" pageId="39" pageNumber="40" precision="925" value="-65.566666">65°34'W</geoCoordinate>
; 
<quantity id="CC379B6EE9368532FE33E85E0C29FEF5" box="[459,547,260,285]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="39" pageNumber="40" unit="m" value="3500.0">3500 m</quantity>
), 
<date id="7F71104BE9368532FDCFE85E0CCFFEF4" box="[567,709,260,285]" pageId="39" pageNumber="40" value="1968-10-28">28 Oct 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E9368532FCADE85E0D70FEF5" box="[853,890,260,284]" pageId="39" pageNumber="40" type="generic">1♂</specimenCount>
, 
<collectionCode id="6DDEAE4EE9368532FC7EE85E0DCEFEF5" box="[902,964,260,284]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="39" pageNumber="40">CNC</collectionCode>
).
</paragraph>
<caption id="5FB06603E9368532FF6FEE390D85F85E" httpUri="https://zenodo.org/record/399425/files/figure.png" pageId="39" pageNumber="40" targetBox="[153,1436,345,1860]" targetPageId="39">
<paragraph id="0B70368BE9368532FF6FEE390D85F85E" blockId="39.[151,1436,1891,1975]" pageId="39" pageNumber="40">
<emphasis id="39BBEA99E9368532FF6FEE390F5FF890" bold="true" box="[151,341,1891,1914]" pageId="39" pageNumber="40">FIGURES 53–57.</emphasis>
Head, thorax, and structures of male terminalia of 
<taxonomicName id="CCCF4D08E9368532FC9EEE3E0A5BF893" box="[870,1105,1892,1914]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="mallonota" status="sp. nov.">
<emphasis id="39BBEA99E9368532FC9EEE3E0A5BF893" box="[870,1105,1892,1914]" italics="true" pageId="39" pageNumber="40">Neoephydra mallonota</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228857E2E9368532FBA0EE3E0AA1F890" box="[1112,1195,1892,1913]" pageId="39" pageNumber="40" rank="species">
<emphasis id="39BBEA99E9368532FBA0EE3E0AA1F890" bold="true" box="[1112,1195,1892,1913]" pageId="39" pageNumber="40">sp. nov.</emphasis>
</taxonomicNameLabel>
(53) head, lateroblique view; (54) mesonotum, dorsal view; (55) tergite 5, epandrium, cerci, surstyli, lateral view; (56) same, posterior view; (57) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view.
</paragraph>
</caption>
<paragraph id="0B70368BE939853DFF6FED290D19FB61" blockId="40.[151,787,1139,1161]" box="[151,787,1139,1161]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF6FED290EFCFB61" bold="true" box="[151,246,1139,1160]" pageId="40" pageNumber="41">MAP 10.</emphasis>
Distribution map for 
<taxonomicName id="CCCF4D08E939853DFE2BED290CB6FB60" box="[467,700,1139,1161]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mallonota" status="sp. nov.">
<emphasis id="39BBEA99E939853DFE2BED290CB6FB60" box="[467,700,1139,1161]" italics="true" pageId="40" pageNumber="41">Neoephydra mallonota</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228857E2E939853DFD3AED290D19FB61" box="[706,787,1139,1160]" pageId="40" pageNumber="41" rank="species">
<emphasis id="39BBEA99E939853DFD3AED290D19FB61" bold="true" box="[706,787,1139,1160]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="43D56500E939853DFF3FEDEC0EEBFA4F" pageId="40" pageNumber="41" type="description">
<paragraph id="0B70368BE939853DFF3FEDEC0B1BFAD3" blockId="40.[151,1436,1205,1662]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF3FEDEC0FD7FB27" box="[199,477,1205,1230]" italics="true" pageId="40" pageNumber="41">
<collectingCountry id="73D8761BE939853DFF3FEDEC0F3FFB24" box="[199,309,1206,1229]" name="Bolivia" pageId="40" pageNumber="41">BOLIVIA</collectingCountry>
. Cochabamba:
</emphasis>
<collectingCountry id="73D8761BE939853DFE1BEDED0C1DFB27" box="[483,535,1207,1230]" name="Japan" pageId="40" pageNumber="41">Japo</collectingCountry>
(
<quantity id="CC379B6EE939853DFDDEEDEC0C67FB27" box="[550,621,1205,1230]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="40" pageNumber="41" unit="km" value="18.0">18 km</quantity>
W; 
<geoCoordinate id="6EFB504CE939853DFD63EDEC0CFFFB27" box="[667,757,1205,1230]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="925" value="-17.583334">17°35'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE939853DFCF8EDEF0D70FB27" box="[768,890,1205,1230]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="92" value="-66.93667">66°56.2'W</geoCoordinate>
; 
<quantity id="CC379B6EE939853DFC7DEDEF0DD7FB27" box="[901,989,1205,1230]" metricMagnitude="3" metricUnit="m" metricValue="4.06" pageId="40" pageNumber="41" unit="m" value="4060.0">4060 m</quantity>
), 
<date id="7F71104BE939853DFC08EDEC0A89FB27" box="[1008,1155,1206,1230]" pageId="40" pageNumber="41" value="2001-03-23">23 Mar 2001</date>
, W. N. Mathis (
<specimenCount id="1DC9FD02E939853DFACAEDEF0B6FFB27" box="[1330,1381,1205,1230]" pageId="40" pageNumber="41" type="generic">15♂</specimenCount>
, 6♀; USNM); Lequepalca (
<quantity id="CC379B6EE939853DFE6CED800FC1FB1B" box="[404,459,1241,1266]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="km" value="1.0">1 km</quantity>
E; 1737.7'S, 
<geoCoordinate id="6EFB504CE939853DFD9AED830CC2FB1B" box="[610,712,1241,1266]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="925" value="-66.95">66°57'W</geoCoordinate>
; 
<quantity id="CC379B6EE939853DFD2BED800D21FB1B" box="[723,811,1242,1266]" metricMagnitude="3" metricUnit="m" metricValue="3.97" pageId="40" pageNumber="41" unit="m" value="3970.0">3970 m</quantity>
), 
<date id="7F71104BE939853DFCB8ED830DDEFB1B" box="[832,980,1241,1266]" pageId="40" pageNumber="41" value="2001-03-26">26 Mar 2001</date>
, A. Freidberg, W. N. Mathis (
<specimenCount id="1DC9FD02E939853DFADBED800B52FB1B" box="[1315,1368,1242,1266]" pageId="40" pageNumber="41" type="generic">22♂</specimenCount>
, 15♀; USNM). 
<emphasis id="39BBEA99E939853DFEF8EDA50F53FAFF" box="[256,345,1279,1302]" italics="true" pageId="40" pageNumber="41">La Paz:</emphasis>
El Alto (de La Paz; 16°27.'S, 
<geoCoordinate id="6EFB504CE939853DFD4BEDA70D24FAFF" box="[691,814,1277,1302]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="92" value="-68.096664">68°05.8'W</geoCoordinate>
; 
<quantity id="CC379B6EE939853DFCC2EDA70DB7FAFF" box="[826,957,1277,1302]" metricMagnitude="3" metricUnit="m" metricValue="2.3225" metricValueMax="4.6" metricValueMin="0.045" pageId="40" pageNumber="41" unit="m" value="2322.5" valueMax="4600.0" valueMin="45.0">45–4600 m</quantity>
), 
<date id="7F71104BE939853DFC2BEDA40A68FAFF" box="[979,1122,1277,1302]" pageId="40" pageNumber="41" value="1968-10-28">28 Oct 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E939853DFB0FEDA40B16FAFF" box="[1271,1308,1278,1302]" pageId="40" pageNumber="41" type="generic">3♂</specimenCount>
, 2♀; CNC, USNM); El Alto (
<quantity id="CC379B6EE939853DFE99EC780FA3FAD3" box="[353,425,1313,1338]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="40" pageNumber="41" unit="km" value="23.0">23 km</quantity>
S; 
<geoCoordinate id="6EFB504CE939853DFE36EC7B0C34FAD3" box="[462,574,1313,1338]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="92" value="-16.711666">16°42.7'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE939853DFDB1EC7B0CC8FAD3" box="[585,706,1313,1338]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="92" value="-68.18667">68°11.2'W</geoCoordinate>
; 
<quantity id="CC379B6EE939853DFD35EC7B0D2FFAD3" box="[717,805,1313,1338]" metricMagnitude="3" metricUnit="m" metricValue="3.86" pageId="40" pageNumber="41" unit="m" value="3860.0">3860 m</quantity>
), 
<date id="7F71104BE939853DFCC1EC780DC6FAD3" box="[825,972,1314,1338]" pageId="40" pageNumber="41" value="2001-03-21">21 Mar 2001</date>
, W. N. Mathis (1♀; USNM).
</paragraph>
<paragraph id="0B70368BE939853DFF3FEC1C0EEBFA4F" blockId="40.[151,1436,1205,1662]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF3FEC1C0FBCFAB7" box="[199,438,1350,1374]" italics="true" pageId="40" pageNumber="41">
<collectingCountry id="73D8761BE939853DFF3FEC1C0F16FAB4" box="[199,284,1350,1373]" name="Chile" pageId="40" pageNumber="41">CHILE</collectingCountry>
. Antofagasta:
</emphasis>
Mucar (
<geoCoordinate id="6EFB504CE939853DFDEDEC1C0C65FAB7" box="[533,623,1350,1374]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="925" value="-23.333334">23°20'S</geoCoordinate>
on 
<collectingCountry id="73D8761BE939853DFD60EC1F0D0DFAB7" box="[664,775,1349,1374]" name="Argentina" pageId="40" pageNumber="41">Argentine</collectingCountry>
border; 
<quantity id="CC379B6EE939853DFC9EEC1C0A0EFAB7" box="[870,1028,1350,1374]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="40" pageNumber="41" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000–4100 m</quantity>
), 
<date id="7F71104BE939853DFBE0EC1C0ADEFAB7" box="[1048,1236,1349,1374]" pageId="40" pageNumber="41" value="1965-12-12" valueMax="1965-12-16" valueMin="1965-12-12">12–16 Dec 1965</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E939853DFA9BEC1C0B9DFAB7" box="[1379,1431,1350,1374]" pageId="40" pageNumber="41" type="generic">30♂</specimenCount>
, 37♀; CNC); Tumbre (E of Atacama Salt Lake; 
<geoCoordinate id="6EFB504CE939853DFD49EC300D00FA6B" box="[689,778,1386,1410]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="925" value="-23.35">23°21'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE939853DFCEFEC330D76FA6B" box="[791,892,1385,1410]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="925" value="-67.8">67°48'W</geoCoordinate>
; 
<quantity id="CC379B6EE939853DFC70EC330A2DFA6B" box="[904,1063,1385,1410]" metricMagnitude="3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="40" pageNumber="41" unit="m" value="3700.0" valueMax="3800.0" valueMin="3600.0">3600–3800 m</quantity>
), 
<date id="7F71104BE939853DFBC4EC330AD4FA6B" box="[1084,1246,1385,1410]" pageId="40" pageNumber="41" value="1965-12-06" valueMax="1965-12-09" valueMin="1965-12-06">6–9 Dec 1965</date>
, L. E. Peña (1♀; CNC).
</paragraph>
</subSubSection>
<subSubSection id="43D56500E939853DFF3FECEB0A4AFA23" box="[199,1088,1457,1482]" pageId="40" pageNumber="41" type="materials_examined">
<paragraph id="0B70368BE939853DFF3FECEB0A4AFA23" blockId="40.[151,1436,1205,1662]" box="[199,1088,1457,1482]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF3FECEB0F6FFA23" bold="true" box="[199,357,1457,1482]" pageId="40" pageNumber="41">
<typeStatus id="D4748829E939853DFF3FECEB0F08FA23" box="[199,258,1457,1482]" pageId="40" pageNumber="41">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="73D8761BE939853DFE94ECEB0FD5FA23" box="[364,479,1457,1482]" name="Argentina" pageId="40" pageNumber="41">Argentina</collectingCountry>
. Jujuy: Cangrejillos (S La Quiaca; 
<geoCoordinate id="6EFB504CE939853DFC9FECEB0DCBFA23" box="[871,961,1457,1482]" direction="south" orientation="latitude" pageId="40" pageNumber="41" precision="925" value="-22.416666">22°25'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE939853DFC35ECEB0A3EFA23" box="[973,1076,1457,1482]" direction="west" orientation="longitude" pageId="40" pageNumber="41" precision="925" value="-65.566666">65°34'W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="43D56500E939853DFF3FEC8F0CCAF9F8" pageId="40" pageNumber="41" type="distribution">
<paragraph id="0B70368BE939853DFF3FEC8F0CCAF9F8" blockId="40.[151,1436,1205,1662]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF3FEC8F0F53FA07" bold="true" box="[199,345,1493,1518]" pageId="40" pageNumber="41">Distribution</emphasis>
(Map 10). 
<emphasis id="39BBEA99E939853DFE04EC8F0C80FA07" box="[508,650,1493,1518]" italics="true" pageId="40" pageNumber="41">Neotropical:</emphasis>
<collectingCountry id="73D8761BE939853DFD66EC8F0D07FA07" box="[670,781,1493,1518]" name="Argentina" pageId="40" pageNumber="41">Argentina</collectingCountry>
(Jujuy), 
<collectingCountry id="73D8761BE939853DFC72EC8F0DD7FA07" box="[906,989,1493,1518]" name="Bolivia" pageId="40" pageNumber="41">Bolivia</collectingCountry>
(Cochabamba, La Paz), and 
<collectingCountry id="73D8761BE939853DFAA7EC8F0B91FA07" box="[1375,1435,1493,1518]" name="Chile" pageId="40" pageNumber="41">Chile</collectingCountry>
(Antofagasta), between 17°–27°S and 65°–68°W.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E939853DFF3FEF470A45F9B0" pageId="40" pageNumber="41" type="etymology">
<paragraph id="0B70368BE939853DFF3FEF470A45F9B0" blockId="40.[151,1436,1205,1662]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF3FEF470F44F9DF" bold="true" box="[199,334,1565,1590]" pageId="40" pageNumber="41">Etymology.</emphasis>
The specific epithet, 
<emphasis id="39BBEA99E939853DFDB1EF470CCBF9DF" box="[585,705,1565,1590]" italics="true" pageId="40" pageNumber="41">
<taxonomicName id="CCCF4D08E939853DFDB1EF470CB7F9DF" box="[585,701,1565,1590]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="mallonota">mallonota</taxonomicName>
,
</emphasis>
is of Greek derivation and is a combination of 
<emphasis id="39BBEA99E939853DFB13EF470B38F9DF" box="[1259,1330,1565,1590]" italics="true" pageId="40" pageNumber="41">mallo,</emphasis>
meaning wool, and 
<emphasis id="39BBEA99E939853DFEF4EF190F4FF9B3" box="[268,325,1603,1626]" italics="true" pageId="40" pageNumber="41">nota,</emphasis>
meaning the back, in allusion to the abundant pilosity on the scutum.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E939853DFF3FEF3F0F74F8E4" pageId="40" pageNumber="41" type="discussion">
<paragraph id="0B70368BE939853DFF3FEF3F0AB0F997" blockId="40.[151,1436,1205,1662]" box="[199,1210,1637,1662]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF3FEF3F0F31F997" bold="true" box="[199,315,1637,1662]" pageId="40" pageNumber="41">Remarks.</emphasis>
Our comments concerning the identification of 
<taxonomicName id="CCCF4D08E939853DFCADEF3D0A08F997" box="[853,1026,1637,1662]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dasycephala">
<emphasis id="39BBEA99E939853DFCADEF3D0A08F997" box="[853,1026,1637,1662]" italics="true" pageId="40" pageNumber="41">N. dasycephala</emphasis>
</taxonomicName>
apply here also.
</paragraph>
<paragraph id="0B70368BE939853DFF6FEF880F74F8E4" blockId="40.[151,542,1745,1805]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF6FEF880FB7F902" bold="true" box="[151,445,1745,1772]" pageId="40" pageNumber="41">
11. 
<taxonomicName id="CCCF4D08E939853DFF39EF8B0FB7F902" box="[193,445,1745,1771]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="shewelli" status="sp. nov.">
<emphasis id="39BBEA99E939853DFF39EF8B0FB7F902" bold="true" box="[193,445,1745,1771]" italics="true" pageId="40" pageNumber="41">Neoephydra shewelli</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="228857E2E939853DFE3CEF880C14F902" box="[452,542,1746,1771]" pageId="40" pageNumber="41" rank="species">
<emphasis id="39BBEA99E939853DFE3CEF880C14F902" bold="true" box="[452,542,1746,1771]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
<figureCitation id="93F42A0EE939853DFF6FEFAE0F14F8E4" box="[151,286,1780,1805]" captionStart="FIGURES 58 – 60" captionStartId="41.[151,264,1185,1208]" captionTargetBox="[194,1390,209,1155]" captionTargetId="figure@41.[190,1396,193,1164]" captionTargetPageId="41" captionText="FIGURES 58 – 60. Structures of male terminalia of Neoephydra shewelli sp. nov. (58) tergite 5, epandrium, cerci, surstyli, lateral view; (59) same, posterior view; (60) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399426/files/figure.png" pageId="40" pageNumber="41">Figs. 58–60</figureCitation>
, Map 11
</paragraph>
</subSubSection>
<subSubSection id="43D56500E939853CFF6FEE660F85F9FB" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="diagnosis">
<paragraph id="0B70368BE939853DFF6FEE660FC9F80C" blockId="40.[151,1436,1852,2021]" pageId="40" pageNumber="41">
<emphasis id="39BBEA99E939853DFF6FEE660F1BF8BC" bold="true" box="[151,273,1852,1877]" pageId="40" pageNumber="41">Diagnosis.</emphasis>
This is the most distinctive species of the 
<taxonomicName id="CCCF4D08E939853DFCF9EE660D87F8BC" box="[769,909,1852,1877]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="dasycephala">
<emphasis id="39BBEA99E939853DFCF9EE660D87F8BC" box="[769,909,1852,1877]" italics="true" pageId="40" pageNumber="41">dasycephala</emphasis>
</taxonomicName>
group and specimens are easily distinguished from those of related congeners by the following characters: general coloration paler, especially the tibiae and tarsi; size smaller; structures of the male terminalia very distinctive. Medium-sized shore flies, body length 
<quantity id="CC379B6EE939853DFAD4EEDE0EC9F829" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="3.8" metricValueMin="3.5" pageId="40" pageNumber="41" unit="mm" value="3.65" valueMax="3.8" valueMin="3.5">3.50–3.80 mm</quantity>
(averaging 
<quantity id="CC379B6EE939853DFEB1EEF20FBBF828" box="[329,433,1960,1985]" metricMagnitude="-3" metricUnit="m" metricValue="3.66" pageId="40" pageNumber="41" unit="mm" value="3.66">3.66 mm</quantity>
); generally microtomentose, dull, pale greenish gray to slightly orange tan, abdomen less microtomentose, subshiny.
</paragraph>
<caption id="5FB06603E938853CFF6FEDFB0B5EFB3F" httpUri="https://zenodo.org/record/399426/files/figure.png" pageId="41" pageNumber="42" targetBox="[194,1390,209,1155]" targetPageId="41">
<paragraph id="0B70368BE938853CFF6FEDFB0B5EFB3F" blockId="41.[151,1436,1185,1238]" pageId="41" pageNumber="42">
<emphasis id="39BBEA99E938853CFF6FEDFB0F5DFB5F" bold="true" box="[151,343,1185,1208]" pageId="41" pageNumber="42">FIGURES 58–60.</emphasis>
Structures of male terminalia of 
<taxonomicName id="CCCF4D08E938853CFD4EEDFB0D84FB5E" box="[694,910,1185,1207]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="42" phylum="Arthropoda" rank="species" species="shewelli" status="sp. nov.">
<emphasis id="39BBEA99E938853CFD4EEDFB0D84FB5E" box="[694,910,1185,1207]" italics="true" pageId="41" pageNumber="42">Neoephydra shewelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228857E2E938853CFC6FEDF80DE1FB5E" box="[919,1003,1186,1207]" pageId="41" pageNumber="42" rank="species">
<emphasis id="39BBEA99E938853CFC6FEDF80DE1FB5E" bold="true" box="[919,1003,1186,1207]" pageId="41" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
(58) tergite 5, epandrium, cerci, surstyli, lateral view; (59) same, posterior view; (60) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view.
</paragraph>
</caption>
<paragraph id="0B70368BE938853CFF3FEDA70F85F9FB" blockId="41.[151,1437,1277,2022]" pageId="41" pageNumber="42">
<emphasis id="39BBEA99E938853CFF3FEDA70F07FAFF" box="[199,269,1277,1302]" italics="true" pageId="41" pageNumber="42">Head:</emphasis>
Head ratio 0.56–0.58; frontal ratio 0.49–0.53, mesofrons with metallic bronzish green luster; ocellar triangle and posterior portion of a parafrons microtomentose, concolorous, orangish tan, becoming gray to charcoal gray anteriorly; thin strip of parafrons adjacent to dorsal margin of eye tannish gray to white. Antenna microtomentose, mostly unicolorous, grayish brown to grayish black, sometimes becoming paler, grayer basally. Facial ratio 0.83–0.87; face, including antennal groove mostly dull, microtomentose, unicolorous, brown to pale yellowish tan but frequently becoming paler in color ventrally. Eye ratio 0.89–0.93; gena-to-eye ratio 0.57–0.60; gena very high, mostly grayish white to silvery anteriorly, becoming darker, tinged with pale greenish gray to tan coloration posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E938853FFF3FEF470C4DFF3C" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="0B70368BE938853CFF3FEF470B3EF8BF" blockId="41.[151,1437,1277,2022]" pageId="41" pageNumber="42">
<emphasis id="39BBEA99E938853CFF3FEF470F2BF9DF" box="[199,289,1565,1590]" italics="true" pageId="41" pageNumber="42">Thorax:</emphasis>
Generally microtomentose, brown to brownish gray; scutum mostly pale bronzish tan, more microtomentose and grayer anteriorly and laterally, subshiny and darker posteriorly, with very faint metallic vittae, 1 medial and 2 lateral pairs, vittae more evident posteriorly; dorsum of scutellum shiny anteromedially, with metallic bronze luster nearly concolorous with shiny mesofrons. Pleural areas more or less unicolorous but with dorsal portion, especially anepisternum nearly concolorous with scutum, ventrally paler and grayer. Wing mostly infumate, pale brown; length 
<quantity id="CC379B6EE938853CFE1DEF880C83F900" box="[485,649,1746,1770]" metricMagnitude="-3" metricUnit="m" metricValue="3.315" metricValueMax="3.49" metricValueMin="3.14" pageId="41" pageNumber="42" unit="mm" value="3.315" valueMax="3.49" valueMin="3.14">3.14–3.49 mm</quantity>
(averaging 
<quantity id="CC379B6EE938853CFCE8EF880D73F903" box="[784,889,1746,1770]" metricMagnitude="-3" metricUnit="m" metricValue="3.31" pageId="41" pageNumber="42" unit="mm" value="3.31">3.31 mm</quantity>
); costal vein ratio 0.61–0.64; M vein ratio 0.85– 0.88. Forecoxa pale brownish white to grayish white; femora microtomentose, very palely tinged with some green color but mostly gray; tibiae mostly concolorous, mostly pale orange but with some grayer and darker areas toward apices; tarsi with basal tarsomeres concolorous with apex of tibiae, other tarsomeres apically dark brown.
</paragraph>
<paragraph id="0B70368BE938853FFF3FEE3B0C4DFF3C" blockId="41.[151,1437,1277,2022]" lastBlockId="42.[151,1437,151,861]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<emphasis id="39BBEA99E938853CFF3FEE3B0F30F893" box="[199,314,1889,1914]" italics="true" pageId="41" pageNumber="42">Abdomen:</emphasis>
Visible segments mostly concolorous, subshiny to shiny dorsally with metallic green to bronzish green luster, more microtomentose, grayer toward lateral and posterior margins. Male terminalia (
<figureCitation id="93F42A0EE938853CFAFBEEDF0B9AF877" box="[1283,1424,1925,1950]" captionStart="FIGURES 58 – 60" captionStartId="41.[151,264,1185,1208]" captionTargetBox="[194,1390,209,1155]" captionTargetId="figure@41.[190,1396,193,1164]" captionTargetPageId="41" captionText="FIGURES 58 – 60. Structures of male terminalia of Neoephydra shewelli sp. nov. (58) tergite 5, epandrium, cerci, surstyli, lateral view; (59) same, posterior view; (60) internal male terminalia (fused hypandrium gonite, aedeagus), lateral view." httpUri="https://zenodo.org/record/399426/files/figure.png" pageId="41" pageNumber="42">Figs. 58–60</figureCitation>
): epandrium parallel sided; basal portion of fused surstyli wide, narrowed more or less abruptly; posterior surstylar processes more widely separated basally, becoming slightly divergent apically; 1 pair of lateral processes, each bearing rather stout setae apically; phallapodeme evenly rounded along outer margin, wide; aedeagus thick over most of length, medial margin sinuate.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E93B853FFF3FE9850EF3FDD5" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="0B70368BE93B853FFF3FE9850D34FE1D" blockId="42.[151,1437,151,861]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF3FE9850F7CFF11" bold="true" box="[199,374,223,248]" pageId="42" pageNumber="43">
<typeStatus id="D4748829E93B853FFF3FE9850F08FF11" box="[199,258,223,248]" pageId="42" pageNumber="43">Type</typeStatus>
material.
</emphasis>
The 
<typeStatus id="D4748829E93B853FFE4AE9850C1EFF11" box="[434,532,223,248]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
male is labeled “ 
<collectingCountry id="73D8761BE93B853FFD22E9BA0D2DFF11" box="[730,807,224,248]" name="Peru" pageId="42" pageNumber="43">PERU</collectingCountry>
. Cuzco: Quispicamchis [sic, Quispicanchi], Huarcapay, 
<quantity id="CC379B6EE93B853FFF6FE85E0EE2FEF5" box="[151,232,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="42" pageNumber="43" unit="m" value="2900.0">2900m</quantity>
, 
<date id="7F71104BE93B853FFF00E85E0F89FEF5" box="[248,387,260,284]" pageId="42" pageNumber="43" value="1988-09-01">1 Sep 1988</date>
, WNMathis/ 
<typeStatus id="D4748829E93B853FFDEFE85E0CA1FEF4" box="[535,683,260,285]" pageId="42" pageNumber="43" type="holotype">HOLOTYPE</typeStatus>
♂ 
<taxonomicName id="CCCF4D08E93B853FFD20E85E0A2FFEF4" authority="Mathis" authorityName="Mathis" box="[728,1061,260,285]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="shewelli">Neoephydra shewelli Mathis</taxonomicName>
<collectionCode id="6DDEAE4EE93B853FFBC9E85E0A88FEF5" box="[1073,1154,260,284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
[red].” The 
<typeStatus id="D4748829E93B853FFAE3E85E0B77FEF4" box="[1307,1405,260,285]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
is double mounted (minuten in a plastic elastomer block), is in excellent condition, and is deposited in the 
<collectionCode id="6DDEAE4EE93B853FFAD1E8720B77FEA9" box="[1321,1405,296,320]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
. A male 
<typeStatus id="D4748829E93B853FFF23E8170F36FE8C" box="[219,316,333,357]" pageId="42" pageNumber="43" type="paratype">paratype</typeStatus>
(
<collectionCode id="6DDEAE4EE93B853FFEA8E8160FAEFE8D" box="[336,420,332,356]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
) bears the same locality label data as the 
<typeStatus id="D4748829E93B853FFC57E8160A1FFE8C" box="[943,1045,332,357]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
. Other 
<typeStatus id="D4748829E93B853FFB8BE8170AD4FE8C" box="[1139,1246,333,357]" pageId="42" pageNumber="43" type="paratype">paratypes</typeStatus>
are as follows: 
<emphasis id="39BBEA99E93B853FFF6FE82A0F8EFE61" box="[151,388,368,392]" italics="true" pageId="42" pageNumber="43">
<collectingCountry id="73D8761BE93B853FFF6FE82A0F38FE6E" box="[151,306,368,391]" name="Argentina" pageId="42" pageNumber="43">ARGENTINA</collectingCountry>
. Jujuy:
</emphasis>
Cangrejillos (S La Quiaca; 
<geoCoordinate id="6EFB504CE93B853FFD30E8350D28FE61" box="[712,802,367,392]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-22.416666">22°25'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFCC8E8350D9FFE61" box="[816,917,367,392]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-65.566666">65°34'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFC5BE8350DF7FE61" box="[931,1021,367,392]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="42" pageNumber="43" unit="m" value="3500.0">3500 m</quantity>
), 
<date id="7F71104BE93B853FFBEBE82A0AAFFE61" box="[1043,1189,367,392]" pageId="42" pageNumber="43" value="1968-10-28">28 Oct 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E93B853FFAC7E82A0B69FE61" box="[1343,1379,368,392]" pageId="42" pageNumber="43" type="generic">1♂</specimenCount>
, 1♀; 
<collectionCode id="6DDEAE4EE93B853FFF6FE8CE0ED9FE45" box="[151,211,404,428]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="43">CNC</collectionCode>
, 
<collectionCode id="6DDEAE4EE93B853FFF27E8CE0F3FFE45" box="[223,309,404,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
); Cerrillos (
<geoCoordinate id="6EFB504CE93B853FFE43E8CE0C1CFE45" box="[443,534,404,428]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-22.316668">22°19'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFDDAE8CE0C8DFE44" box="[546,647,404,429]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-65.816666">65°49'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFD6BE8CE0CE6FE45" box="[659,748,404,429]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="42" pageNumber="43" unit="m" value="3600.0">3600 m</quantity>
), 
<date id="7F71104BE93B853FFCF9E8CE0D85FE44" box="[769,911,404,429]" pageId="42" pageNumber="43" value="1968-10-31">31 Oct 1968</date>
, L. E. Peña (1♀; 
<collectionCode id="6DDEAE4EE93B853FFBACE8CE0A98FE45" box="[1108,1170,404,428]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="43">CNC</collectionCode>
); Coyaguayma (
<quantity id="CC379B6EE93B853FFAB3E8CE0B8FFE44" box="[1355,1413,404,429]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="42" pageNumber="43" unit="km" value="7.0">7 km</quantity>
S 
<taxonomicName id="CCCF4D08E93B853FFF6FE8ED0F5AFE39" authority="Periquitas" authorityName="Periquitas" box="[151,336,439,464]" class="Magnoliopsida" family="Convolvulaceae" genus="Mina" higherTaxonomySource="CoL" kingdom="Plantae" order="Solanales" pageId="42" pageNumber="43" phylum="Tracheophyta" rank="genus">Mina Periquitas</taxonomicName>
; 
<geoCoordinate id="6EFB504CE93B853FFEA5E8ED0FC7FE39" box="[349,461,439,464]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="92" value="-22.778334">22°46.7'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFE23E8ED0C5EFE39" box="[475,596,439,464]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="92" value="-66.558334">66°33.5'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFD9BE8E20CB4FE39" box="[611,702,440,464]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="42" pageNumber="43" unit="m" value="4100.0">4100 m</quantity>
), 
<date id="7F71104BE93B853FFD2DE8E20D6BFE39" box="[725,865,439,464]" pageId="42" pageNumber="43" value="1968-11-04">4 Nov 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E93B853FFC04E8ED0A28FE39" box="[1020,1058,439,464]" pageId="42" pageNumber="43" type="generic">6♂</specimenCount>
; 
<collectionCode id="6DDEAE4EE93B853FFBC8E8E20A65FE39" box="[1072,1135,440,464]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="43">CNC</collectionCode>
); Santa Catalina (
<geoCoordinate id="6EFB504CE93B853FFAC5E8ED0B9DFE39" box="[1341,1431,439,464]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-22.083334">22°05'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFF6FE8860EF6FE1C" box="[151,252,476,501]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-66.3">66°18'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFEFFE8860F55FE1D" box="[263,351,476,500]" metricMagnitude="3" metricUnit="m" metricValue="3.7" pageId="42" pageNumber="43" unit="m" value="3700.0">3700 m</quantity>
), 
<date id="7F71104BE93B853FFE8BE8860C0BFE1C" box="[371,513,476,501]" pageId="42" pageNumber="43" value="1968-10-25">25 Oct 1968</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E93B853FFD68E8860CBCFE1D" box="[656,694,476,500]" pageId="42" pageNumber="43" type="generic">2♂</specimenCount>
, 1♀; 
<collectionCode id="6DDEAE4EE93B853FFD0CE8860D38FE1D" box="[756,818,476,500]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="42" pageNumber="43">CNC</collectionCode>
).
</paragraph>
<paragraph id="0B70368BE93B853FFF3FEB5A0EF3FDD5" blockId="42.[151,1437,151,861]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF3FEB5A0F85FDFE" box="[199,399,512,535]" italics="true" pageId="42" pageNumber="43">
<collectingCountry id="73D8761BE93B853FFF3FEB5A0F3FFDFE" box="[199,309,512,535]" name="Bolivia" pageId="42" pageNumber="43">BOLIVIA</collectingCountry>
. Oruro:
</emphasis>
Pazña (S of town; 
<geoCoordinate id="6EFB504CE93B853FFD8AE8A50CEBFDF1" box="[626,737,511,536]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="92" value="-18.603333">18°36.2'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFD08E8A50D60FDF1" box="[752,874,511,536]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="92" value="-66.91167">66°54.7'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFC80E8A50DD8FDF1" box="[888,978,511,536]" metricMagnitude="3" metricUnit="m" metricValue="3.75" pageId="42" pageNumber="43" unit="m" value="3750.0">3750 m</quantity>
), 
<date id="7F71104BE93B853FFC11EB5A0A8BFDF1" box="[1001,1153,512,536]" pageId="42" pageNumber="43" value="2001-03-22">22 Mar 2001</date>
, W. N. Mathis (
<specimenCount id="1DC9FD02E93B853FFAC5EB5A0B69FDF1" box="[1341,1379,512,536]" pageId="42" pageNumber="43" type="generic">2♂</specimenCount>
, 1♀; 
<collectionCode id="6DDEAE4EE93B853FFF6FEB7E0EE7FDD5" box="[151,237,548,572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="42" pageNumber="43">USNM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="43D56500E93B853FFF3FEB120F81FD6D" pageId="42" pageNumber="43" type="description">
<paragraph id="0B70368BE93B853FFF3FEB120F81FD6D" blockId="42.[151,1437,151,861]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF3FEB120FBDFD89" box="[199,439,584,608]" italics="true" pageId="42" pageNumber="43">
<collectingCountry id="73D8761BE93B853FFF3FEB120F16FDB6" box="[199,284,584,607]" name="Chile" pageId="42" pageNumber="43">CHILE</collectingCountry>
. Antofagasta:
</emphasis>
Mucar (on 
<collectingCountry id="73D8761BE93B853FFDC5EB1D0CA7FD89" box="[573,685,583,608]" name="Argentina" pageId="42" pageNumber="43">Argentina</collectingCountry>
border; 
<geoCoordinate id="6EFB504CE93B853FFCF6EB120D62FD89" box="[782,872,584,608]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-23.366667">23°22'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFC8DEB1D0DD0FD89" box="[885,986,583,608]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-67.11667">67°07'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFC1EEB120A8FFD89" box="[998,1157,584,608]" metricMagnitude="3" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="42" pageNumber="43" unit="m" value="4050.0" valueMax="4100.0" valueMin="4000.0">4000–4100 m</quantity>
), 
<date id="7F71104BE93B853FFB62EB1D0B52FD89" box="[1178,1368,583,608]" pageId="42" pageNumber="43" value="1965-12-12" valueMax="1965-12-16" valueMin="1965-12-12">12–16 Dec 1965</date>
, L. E. Peña (
<specimenCount id="1DC9FD02E93B853FFF25EB360F09FD6D" box="[221,259,620,644]" pageId="42" pageNumber="43" type="generic">2♂</specimenCount>
, 1♀; CNC).
</paragraph>
</subSubSection>
<subSubSection id="43D56500E93B853FFF3FEBD50A61FD41" box="[199,1131,655,680]" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="0B70368BE93B853FFF3FEBD50A61FD41" blockId="42.[151,1437,151,861]" box="[199,1131,655,680]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF3FEBD50F6FFD41" bold="true" box="[199,357,655,680]" pageId="42" pageNumber="43">
<typeStatus id="D4748829E93B853FFF3FEBD50F08FD41" box="[199,258,655,680]" pageId="42" pageNumber="43">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="73D8761BE93B853FFE94EBCA0FA9FD4E" box="[364,419,656,679]" name="Peru" pageId="42" pageNumber="43">Peru</collectingCountry>
. Cuzco: Quispicanchi, Huarcapay (
<geoCoordinate id="6EFB504CE93B853FFCD7EBCA0D83FD41" box="[815,905,656,680]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-13.633333">13°38'S</geoCoordinate>
, 
<geoCoordinate id="6EFB504CE93B853FFC6DEBCA0DF0FD41" box="[917,1018,656,680]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-71.666664">71°40'W</geoCoordinate>
; 
<quantity id="CC379B6EE93B853FFBFDEBCA0A54FD41" box="[1029,1118,656,680]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="42" pageNumber="43" unit="m" value="2900.0">2900 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="43D56500E93B853FFF3FEBEE0C88FD19" pageId="42" pageNumber="43" type="distribution">
<paragraph id="0B70368BE93B853FFF3FEBEE0C88FD19" blockId="42.[151,1437,151,861]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF3FEBEE0F53FD24" bold="true" box="[199,345,692,717]" pageId="42" pageNumber="43">Distribution</emphasis>
(Map 11). 
<emphasis id="39BBEA99E93B853FFE14EBEE0C70FD24" box="[492,634,692,717]" italics="true" pageId="42" pageNumber="43">Neotropical:</emphasis>
<collectingCountry id="73D8761BE93B853FFD71EBEE0CF3FD24" box="[649,761,692,717]" name="Argentina" pageId="42" pageNumber="43">Argentina</collectingCountry>
(Jujuy), 
<collectingCountry id="73D8761BE93B853FFC94EBEE0DB4FD24" box="[876,958,692,717]" name="Bolivia" pageId="42" pageNumber="43">Bolivia</collectingCountry>
(Oruro), 
<collectingCountry id="73D8761BE93B853FFBCFEBEE0A79FD24" box="[1079,1139,692,717]" name="Chile" pageId="42" pageNumber="43">Chile</collectingCountry>
(Antofagasta), and 
<collectingCountry id="73D8761BE93B853FFA91EBEF0B96FD25" box="[1385,1436,693,716]" name="Peru" pageId="42" pageNumber="43">Peru</collectingCountry>
(Cuzco), between 13°–24°S and 65°–72°W.
</paragraph>
</subSubSection>
<subSubSection id="43D56500E93B853FFF3FEBA60CF4F8BE" pageId="42" pageNumber="43" type="etymology">
<paragraph id="0B70368BE93B853FFF3FEBA60FC8FCB4" blockId="42.[151,1437,151,861]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF3FEBA60F44FCFC" bold="true" box="[199,334,764,789]" pageId="42" pageNumber="43">Etymology.</emphasis>
The specific epithet, 
<emphasis id="39BBEA99E93B853FFDB9EBA60CA9FCFC" box="[577,675,764,789]" italics="true" pageId="42" pageNumber="43">
<taxonomicName id="CCCF4D08E93B853FFDB9EBA60C95FCFC" box="[577,671,764,789]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="shewelli">shewelli</taxonomicName>
,
</emphasis>
is a Latin genitive patronym, honoring Guy E. Shewell (deceased), who graciously loaned the specimens of the 
<typeStatus id="D4748829E93B853FFD74EA7A0CB6FCD1" box="[652,700,800,824]" pageId="42" pageNumber="43">type</typeStatus>
series and who has contributed so much to our knowledge of the 
<taxonomicName id="CCCF4D08E93B853FFF6FEA1E0EE1FCB4" box="[151,235,836,861]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
of the New World.
</paragraph>
<paragraph id="0B70368BE93B853FFF6FEE180CF4F8BE" blockId="42.[151,766,1858,1880]" box="[151,766,1858,1880]" pageId="42" pageNumber="43">
<emphasis id="39BBEA99E93B853FFF6FEE180EFFF8BE" bold="true" box="[151,245,1858,1879]" pageId="42" pageNumber="43">MAP 11.</emphasis>
Distribution map for 
<taxonomicName id="CCCF4D08E93B853FFE2AEE180CADF8B1" box="[466,679,1858,1880]" class="Insecta" family="Ephydridae" genus="Neoephydra" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="shewelli" status="sp. nov.">
<emphasis id="39BBEA99E93B853FFE2AEE180CADF8B1" box="[466,679,1858,1880]" italics="true" pageId="42" pageNumber="43">Neoephydra shewelli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="228857E2E93B853FFD55EE180CF4F8BE" box="[685,766,1858,1879]" pageId="42" pageNumber="43" rank="species">
<emphasis id="39BBEA99E93B853FFD55EE180CF4F8BE" bold="true" box="[685,766,1858,1879]" pageId="42" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>